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<document id="A4B0159E1B39D83770BD1A435B193887" ID-CLB-Dataset="4934" ID-DOI="10.11646/zootaxa.4985.1.2" ID-GBIF-Dataset="bd94b499-c7a6-418e-bb8f-3c5d59675260" ID-ISSN="1175-5326" ID-Zenodo-Dep="4929647" ID-ZooBank="5B23C14A-C7AB-4F22-B0A8-79E700F29FFF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623439671578" checkinUser="plazi" docAuthor="Ramos-Pastrana, Yardany &amp; Rafael, José Albertino" docDate="2021" docId="507C87C0CD7D1C5CEEEA3998FC62FBE4" docLanguage="en" docName="zootaxa.4985.1.2.pdf" docOrigin="Zootaxa 4985 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tomosvaryella similis" docType="treatment" docVersion="11" lastPageNumber="54" masterDocId="AC45FFB8CD6C1C4DEE7D390FFFCEFFBC" masterDocTitle="Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species" masterLastPageNumber="68" masterPageNumber="37" pageNumber="54" updateTime="1699012508356" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0AB25163A15B8C25670ACCB90293D770">Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species</mods:title>
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<mods:namePart id="25D48290A11DAD0A970C5733F24A8F18">Ramos-Pastrana, Yardany</mods:namePart>
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<mods:affiliation id="1FFB5506BFFBD003139EDD8F711F1281">Universidad de la Amazonia, Grupo de Investigación en Entomología Universidad de la Amazonia - GIEU-, Laboratório de Entomologia, Av, 11 5 - 69 Juan XXIII, Florencia, Caquetá, Colombia. ya. ramos @ udla. edu. co; https: // orcid. org / 0000 - 0002 - 3193 - 6659 Instituto Nacional de Pesquisas da Amazônia, INPA, Manaus, Amazonas, Brazil.</mods:affiliation>
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<mods:namePart id="F2ABEEBDD70CB325B8B2CF18B9AF47D3">Rafael, José Albertino</mods:namePart>
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<treatment id="507C87C0CD7D1C5CEEEA3998FC62FBE4" ID-DOI="http://doi.org/10.5281/zenodo.4929088" ID-GBIF-Taxon="184587559" ID-Zenodo-Dep="4929088" LSID="urn:lsid:plazi:treatment:507C87C0CD7D1C5CEEEA3998FC62FBE4" httpUri="http://treatment.plazi.org/id/507C87C0CD7D1C5CEEEA3998FC62FBE4" lastPageNumber="54" pageId="17" pageNumber="54">
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<taxonomicName id="1FD54D55CD7D1C5CEEEA3998FD9AFF0D" ID-CoL="7CH3H" authority="(Hough, 1899)" baseAuthorityName="Hough" baseAuthorityYear="1899" box="[151,596,151,178]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="EAA1EAC4CD7D1C5CEEEA3998FD9AFF0D" bold="true" box="[151,596,151,178]" pageId="17" pageNumber="54">
<emphasis id="EAA1EAC4CD7D1C5CEEEA3998FE55FF0E" bold="true" box="[151,411,151,178]" italics="true" pageId="17" pageNumber="54">Tomosvaryella similis</emphasis>
(
<bibRefCitation id="BC444B27CD7D1C5CEFD63998FD82FF0D" author="Hough, G." box="[427,588,151,177]" pageId="17" pageNumber="54" pagination="77 - 86" refId="ref22797" refString="Hough, G. (1899) Studies in Diptera Cyclorrhapha 1. The Pipunculidae of the United States. Proceeding of the Boston Society of Natural History, 29, 77 - 86." type="journal article" year="1899">Hough, 1899</bibRefCitation>
)
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<paragraph id="D86A36D6CD7D1C5CEEEA39B4FE82FF69" blockId="17.[151,596,151,213]" box="[151,332,187,213]" pageId="17" pageNumber="54">
<figureCitation id="40EE2A53CD7D1C5CEEEA39B4FED6FF69" box="[151,280,187,213]" captionStart="FIGURES 83–92" captionStartId="18.[151,264,1898,1922]" captionTargetBox="[226,1361,181,1874]" captionTargetId="figure-29@18.[226,1361,181,1874]" captionTargetPageId="18" captionText="FIGURES 83–92. Tomosvaryella similis (Hough, 1899) (IAvH–2583). Male. 83, Habitus, left lateral view; 84, Left hind trochanter, lateral view; 85, Tergites and sternites 6 and 7, ventral view; 86, Tip of abdomen with syntergosternite 8, ventral view; 87, Susrtyli, dorsal view; 88, Left surstylus, lateral view; 89, Right surstylus, lateral view; 90, Phallic guide and phallus, left lateral view; 91, Ejaculatory apodeme, lateral view; 92, Hypandrium and gonopods, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4930399" httpUri="https://zenodo.org/record/4930399/files/figure.png" pageId="17" pageNumber="54">Figs 83–92</figureCitation>
,
<figureCitation id="40EE2A53CD7D1C5CEF5F39B4FE82FF69" box="[290,332,187,213]" captionStart="FIGURE 133" captionStartId="27.[151,250,1909,1933]" captionTargetBox="[264,1323,1205,1885]" captionTargetId="figure-148@27.[264,1323,1205,1885]" captionTargetPageId="27" captionText="FIGURE 133. Geographical records of Tomosvaryella species in Colombia. T. lynchi, T. macarenensis sp. nov., T. similis, T. venezuelana, T. subvirescens and T. scopulata." figureDoi="http://doi.org/10.5281/zenodo.4930417" httpUri="https://zenodo.org/record/4930417/files/figure.png" pageId="17" pageNumber="54">133</figureCitation>
</paragraph>
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<subSubSection id="90CF655DCD7D1C5CEEEA39F5FC40FEEC" pageId="17" pageNumber="54" type="reference_group">
<paragraph id="D86A36D6CD7D1C5CEEEA39F5FE32FEAE" blockId="17.[151,1436,250,336]" box="[151,508,250,274]" pageId="17" pageNumber="54">
<treatmentCitationGroup id="F8C511F8CD7D1C5CEEEA39F5FE32FEAE" box="[151,508,250,274]" pageId="17" pageNumber="54">
<treatmentCitation id="597410C7CD7D1C5CEEEA39F5FE32FEAE" author="Hough, G." box="[151,508,250,274]" page="84" pageId="17" pageNumber="54" year="1899">
<taxonomicName id="1FD54D55CD7D1C5CEEEA39F5FE32FEAE" ID-CoL="88Y86" authority="Hough, 1899: 84" authorityName="Hough" authorityPageNumber="84" authorityYear="1899" box="[151,508,250,274]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="EAA1EAC4CD7D1C5CEEEA39F5FE9CFEAE" box="[151,338,250,274]" italics="true" pageId="17" pageNumber="54">Pipunculus similis</emphasis>
<bibRefCitation id="BC444B27CD7D1C5CEF2539F5FE32FEAE" author="Hough, G." box="[344,508,250,274]" pageId="17" pageNumber="54" pagination="77 - 86" refId="ref22797" refString="Hough, G. (1899) Studies in Diptera Cyclorrhapha 1. The Pipunculidae of the United States. Proceeding of the Boston Society of Natural History, 29, 77 - 86." type="journal article" year="1899">Hough, 1899:84</bibRefCitation>
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</paragraph>
<paragraph id="D86A36D6CD7D1C5CEEEA3816FC40FEEC" blockId="17.[151,1436,250,336]" pageId="17" pageNumber="54">
<treatmentCitationGroup id="F8C511F8CD7D1C5CEEEA3816FC40FEEC" pageId="17" pageNumber="54">
<treatmentCitation id="597410C7CD7D1C5CEEEA3816FDF8FE8D" author="Hardy, D. E." box="[151,566,281,305]" page="174" pageId="17" pageNumber="54" year="1943">
<taxonomicName id="1FD54D55CD7D1C5CEEEA3816FDF8FE8D" ID-CoL="7CH3H" authority="Hardy, 1943: 174" authorityName="Hardy" authorityPageNumber="174" authorityYear="1943" box="[151,566,281,305]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="54" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="EAA1EAC4CD7D1C5CEEEA3816FEB8FE8D" box="[151,374,281,305]" italics="true" pageId="17" pageNumber="54">Tomosvaryella similis</emphasis>
<bibRefCitation id="BC444B27CD7D1C5CEF023816FDF8FE8D" author="Hardy, D. E." box="[383,566,281,305]" pageId="17" pageNumber="54" pagination="3 - 23" refId="ref22147" refString="Hardy, D. E. (1943) A revision of Nearctic Dorilaidae (Pipunculidae). Kansas University Science Bulletin, 29, 3 - 23." type="journal article" year="1943">Hardy, 1943: 174</bibRefCitation>
</taxonomicName>
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, figs 98 a–h;
<treatmentCitation id="597410C7CD7D1C5CECBB3816FCA0FE8D" author="Aczel, M." box="[710,878,281,305]" page="22" pageId="17" pageNumber="54" year="1948">
<bibRefCitation id="BC444B27CD7D1C5CECBB3816FCA0FE8D" author="Aczel, M." box="[710,878,281,305]" pageId="17" pageNumber="54" pagination="5 - 168" refId="ref21292" refString="Aczel, M. (1948) Grundlagen einer Monographie der Dorilaiden. (Diptera). Dorilaiden Studien VI. Acta Zoologica Lilloana, 6, 5 - 168." type="journal article" year="1948">Aczél, 1948: 22</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="597410C7CD7D1C5CED063816FBF0FE8D" author="Hardy, D. E." box="[891,1086,281,305]" page="856" pageId="17" pageNumber="54" year="1965">
<bibRefCitation id="BC444B27CD7D1C5CED063816FBF0FE8D" author="Hardy, D. E." box="[891,1086,281,305]" pageId="17" pageNumber="54" refId="ref22525" refString="Hardy, D. E. (1965 c) A Catalogue of the Diptera of America North of Mexico. Agriculture Handbook. No. 276. United State Departament Agriculture, Washington, D. C., 1696 pp." type="book" year="1965">Hardy, 1965c: 856</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="597410C7CD7D1C5CEA343816FA56FE8D" author="Ale-Rocha, R. &amp; Rafael, J. A." box="[1097,1432,281,305]" page="420" pageId="17" pageNumber="54" year="1995">
<bibRefCitation id="BC444B27CD7D1C5CEA343816FA56FE8D" author="Ale-Rocha, R. &amp; Rafael, J. A." box="[1097,1432,281,305]" pageId="17" pageNumber="54" pagination="407 - 427" refId="ref21517" refString="Ale-Rocha, R. &amp; Rafael, J. A. (1995) Tomosvaryella Aczel do Mexico, America Central e Antilhas (Diptera, Pipunculidae). Revista Brasileira de Zoologia, 12, 407 - 427. https: // doi. org / 10.1590 / S 0101 - 81751995000200017" type="journal article" year="1995">Ale-Rocha &amp; Rafael, 1995: 420</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="597410C7CD7D1C5CEEBA3837FE31FEF3" ID-DOI="http://doi.org/10.5281/zenodo.6269534" ID-Zenodo-Dep="6269534" author="Rafael, J. A. &amp; Ale-Rocha, R." box="[199,511,311,336]" httpUri="http://treatment.plazi.org/id/C779878C9A4A2224FECECA8BCDCEF94A" page="16" pageId="17" pageNumber="54" year="2004">
<bibRefCitation id="BC444B27CD7D1C5CEEBA3837FE31FEF3" author="Rafael, J. A. &amp; Ale-Rocha, R." box="[199,511,311,336]" pageId="17" pageNumber="54" pagination="1 - 18" refId="ref23196" refString="Rafael, J. A. &amp; Ale-Rocha, R. (2004) Nicaraguan Pipunculidae (Diptera): new records and description of new species. Zootaxa, 529 (1), 1 - 18. https: // doi. org / 10.11646 / zootaxa. 529.1.1" type="journal article" year="2004">Rafael &amp; Ale-Rocha, 2004: 16</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="597410C7CD7D1C5CEC743837FCF3FEEC" author="Rodriguez, H. C. &amp; Rafael, J. A." box="[521,829,312,336]" page="33" pageId="17" pageNumber="54" year="2012">
<bibRefCitation id="BC444B27CD7D1C5CEC743837FCF3FEEC" author="Rodriguez, H. C. &amp; Rafael, J. A." box="[521,829,312,336]" pageId="17" pageNumber="54" refId="ref23353" refString="Rodriguez, H. C. &amp; Rafael, J. A. (2012) Pipunculidae (Diptera) of the Latin America and the Caribbean: A catalog of Species with Notes on Biology and Pipunculid-Host Associations. LAP Lambert Academic Publishing GmbH &amp; Co. KG, Saarbrucken, 48 pp." type="book" year="2012">Rodriguez &amp; Rafael, 2012: 33</bibRefCitation>
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, 35, 38.
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</paragraph>
</subSubSection>
<subSubSection id="90CF655DCD7D1C5CEEEA3877FDAAFD6E" pageId="17" pageNumber="54" type="diagnosis">
<paragraph id="D86A36D6CD7D1C5CEEEA3877FDAAFD6E" blockId="17.[151,1437,376,1113]" pageId="17" pageNumber="54">
<emphasis id="EAA1EAC4CD7D1C5CEEEA3877FEDCFE2E" bold="true" box="[151,274,376,402]" pageId="17" pageNumber="54">Diagnosis.</emphasis>
Male. Body shiny, robust (
<figureCitation id="40EE2A53CD7D1C5CEC433877FD5EFE2F" box="[574,656,376,403]" captionStart="FIGURES 83–92" captionStartId="18.[151,264,1898,1922]" captionTargetBox="[226,1361,181,1874]" captionTargetId="figure-29@18.[226,1361,181,1874]" captionTargetPageId="18" captionText="FIGURES 83–92. Tomosvaryella similis (Hough, 1899) (IAvH–2583). Male. 83, Habitus, left lateral view; 84, Left hind trochanter, lateral view; 85, Tergites and sternites 6 and 7, ventral view; 86, Tip of abdomen with syntergosternite 8, ventral view; 87, Susrtyli, dorsal view; 88, Left surstylus, lateral view; 89, Right surstylus, lateral view; 90, Phallic guide and phallus, left lateral view; 91, Ejaculatory apodeme, lateral view; 92, Hypandrium and gonopods, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4930399" httpUri="https://zenodo.org/record/4930399/files/figure.png" pageId="17" pageNumber="54">Fig. 83</figureCitation>
). Postcranium with lower three quarters grayish-pruinose, upper quarter brownish-pruinose (
<figureCitation id="40EE2A53CD7D1C5CEFDD3893FE3AFE0A" box="[416,500,412,438]" captionStart="FIGURES 83–92" captionStartId="18.[151,264,1898,1922]" captionTargetBox="[226,1361,181,1874]" captionTargetId="figure-29@18.[226,1361,181,1874]" captionTargetPageId="18" captionText="FIGURES 83–92. Tomosvaryella similis (Hough, 1899) (IAvH–2583). Male. 83, Habitus, left lateral view; 84, Left hind trochanter, lateral view; 85, Tergites and sternites 6 and 7, ventral view; 86, Tip of abdomen with syntergosternite 8, ventral view; 87, Susrtyli, dorsal view; 88, Left surstylus, lateral view; 89, Right surstylus, lateral view; 90, Phallic guide and phallus, left lateral view; 91, Ejaculatory apodeme, lateral view; 92, Hypandrium and gonopods, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4930399" httpUri="https://zenodo.org/record/4930399/files/figure.png" pageId="17" pageNumber="54">Fig. 83</figureCitation>
). Scutum mainly brownish-pruinose; thoracic setae long. Legs brownish-pruinose, except hind femur posteriorly and ventrally shiny. Hind trochanter with bulging, drop-shaped, protuberance, usually with a single ventral and a single dorsal seta (
<figureCitation id="40EE2A53CD7D1C5CEC0F38ECFD0FFE41" box="[626,705,483,509]" captionStart="FIGURES 83–92" captionStartId="18.[151,264,1898,1922]" captionTargetBox="[226,1361,181,1874]" captionTargetId="figure-29@18.[226,1361,181,1874]" captionTargetPageId="18" captionText="FIGURES 83–92. Tomosvaryella similis (Hough, 1899) (IAvH–2583). Male. 83, Habitus, left lateral view; 84, Left hind trochanter, lateral view; 85, Tergites and sternites 6 and 7, ventral view; 86, Tip of abdomen with syntergosternite 8, ventral view; 87, Susrtyli, dorsal view; 88, Left surstylus, lateral view; 89, Right surstylus, lateral view; 90, Phallic guide and phallus, left lateral view; 91, Ejaculatory apodeme, lateral view; 92, Hypandrium and gonopods, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4930399" httpUri="https://zenodo.org/record/4930399/files/figure.png" pageId="17" pageNumber="54">Fig. 84</figureCitation>
). Tergites and sternites 6 and 7 as in
<figureCitation id="40EE2A53CD7D1C5CEA4238ECFB42FE42" box="[1087,1164,483,510]" captionStart="FIGURES 83–92" captionStartId="18.[151,264,1898,1922]" captionTargetBox="[226,1361,181,1874]" captionTargetId="figure-29@18.[226,1361,181,1874]" captionTargetPageId="18" captionText="FIGURES 83–92. Tomosvaryella similis (Hough, 1899) (IAvH–2583). Male. 83, Habitus, left lateral view; 84, Left hind trochanter, lateral view; 85, Tergites and sternites 6 and 7, ventral view; 86, Tip of abdomen with syntergosternite 8, ventral view; 87, Susrtyli, dorsal view; 88, Left surstylus, lateral view; 89, Right surstylus, lateral view; 90, Phallic guide and phallus, left lateral view; 91, Ejaculatory apodeme, lateral view; 92, Hypandrium and gonopods, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4930399" httpUri="https://zenodo.org/record/4930399/files/figure.png" pageId="17" pageNumber="54">Fig. 85</figureCitation>
. Surstyli subsymmetrical, with external and internal margins somewhat straight, having slightly hooked-tips, right surstylus slightly shorter (
<figureCitation id="40EE2A53CD7D1C5CEEDD3B25FEEEFDF9" box="[160,288,554,581]" captionStart="FIGURES 83–92" captionStartId="18.[151,264,1898,1922]" captionTargetBox="[226,1361,181,1874]" captionTargetId="figure-29@18.[226,1361,181,1874]" captionTargetPageId="18" captionText="FIGURES 83–92. Tomosvaryella similis (Hough, 1899) (IAvH–2583). Male. 83, Habitus, left lateral view; 84, Left hind trochanter, lateral view; 85, Tergites and sternites 6 and 7, ventral view; 86, Tip of abdomen with syntergosternite 8, ventral view; 87, Susrtyli, dorsal view; 88, Left surstylus, lateral view; 89, Right surstylus, lateral view; 90, Phallic guide and phallus, left lateral view; 91, Ejaculatory apodeme, lateral view; 92, Hypandrium and gonopods, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4930399" httpUri="https://zenodo.org/record/4930399/files/figure.png" pageId="17" pageNumber="54">Fig. 86–87</figureCitation>
), both surstyli with tips slightly downward-directed when seen in lateral view (
<figureCitation id="40EE2A53CD7D1C5CEAD53B25FAE2FDF8" box="[1192,1324,554,580]" captionStart="FIGURES 83–92" captionStartId="18.[151,264,1898,1922]" captionTargetBox="[226,1361,181,1874]" captionTargetId="figure-29@18.[226,1361,181,1874]" captionTargetPageId="18" captionText="FIGURES 83–92. Tomosvaryella similis (Hough, 1899) (IAvH–2583). Male. 83, Habitus, left lateral view; 84, Left hind trochanter, lateral view; 85, Tergites and sternites 6 and 7, ventral view; 86, Tip of abdomen with syntergosternite 8, ventral view; 87, Susrtyli, dorsal view; 88, Left surstylus, lateral view; 89, Right surstylus, lateral view; 90, Phallic guide and phallus, left lateral view; 91, Ejaculatory apodeme, lateral view; 92, Hypandrium and gonopods, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4930399" httpUri="https://zenodo.org/record/4930399/files/figure.png" pageId="17" pageNumber="54">Figs 88–89</figureCitation>
). Apex of phallic guide short, with inconspicuous spine, angled at about 20 degrees (
<figureCitation id="40EE2A53CD7D1C5CEDA73B42FBE0FDDB" box="[986,1070,589,615]" captionStart="FIGURES 83–92" captionStartId="18.[151,264,1898,1922]" captionTargetBox="[226,1361,181,1874]" captionTargetId="figure-29@18.[226,1361,181,1874]" captionTargetPageId="18" captionText="FIGURES 83–92. Tomosvaryella similis (Hough, 1899) (IAvH–2583). Male. 83, Habitus, left lateral view; 84, Left hind trochanter, lateral view; 85, Tergites and sternites 6 and 7, ventral view; 86, Tip of abdomen with syntergosternite 8, ventral view; 87, Susrtyli, dorsal view; 88, Left surstylus, lateral view; 89, Right surstylus, lateral view; 90, Phallic guide and phallus, left lateral view; 91, Ejaculatory apodeme, lateral view; 92, Hypandrium and gonopods, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4930399" httpUri="https://zenodo.org/record/4930399/files/figure.png" pageId="17" pageNumber="54">Fig. 90</figureCitation>
). Phallus with ejaculatory ejaculary ducts elongated slender branches; one ejaculatory duct with three basal spines, the other two ducts without spines (
<figureCitation id="40EE2A53CD7D1C5CEE8C3B9BFE87FD12" box="[241,329,660,686]" captionStart="FIGURES 83–92" captionStartId="18.[151,264,1898,1922]" captionTargetBox="[226,1361,181,1874]" captionTargetId="figure-29@18.[226,1361,181,1874]" captionTargetPageId="18" captionText="FIGURES 83–92. Tomosvaryella similis (Hough, 1899) (IAvH–2583). Male. 83, Habitus, left lateral view; 84, Left hind trochanter, lateral view; 85, Tergites and sternites 6 and 7, ventral view; 86, Tip of abdomen with syntergosternite 8, ventral view; 87, Susrtyli, dorsal view; 88, Left surstylus, lateral view; 89, Right surstylus, lateral view; 90, Phallic guide and phallus, left lateral view; 91, Ejaculatory apodeme, lateral view; 92, Hypandrium and gonopods, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4930399" httpUri="https://zenodo.org/record/4930399/files/figure.png" pageId="17" pageNumber="54">Fig. 90</figureCitation>
). Ejaculatory apodeme sandal-shaped, darker near base (
<figureCitation id="40EE2A53CD7D1C5CED9D3B9BFBF6FD12" box="[992,1080,660,686]" captionStart="FIGURES 83–92" captionStartId="18.[151,264,1898,1922]" captionTargetBox="[226,1361,181,1874]" captionTargetId="figure-29@18.[226,1361,181,1874]" captionTargetPageId="18" captionText="FIGURES 83–92. Tomosvaryella similis (Hough, 1899) (IAvH–2583). Male. 83, Habitus, left lateral view; 84, Left hind trochanter, lateral view; 85, Tergites and sternites 6 and 7, ventral view; 86, Tip of abdomen with syntergosternite 8, ventral view; 87, Susrtyli, dorsal view; 88, Left surstylus, lateral view; 89, Right surstylus, lateral view; 90, Phallic guide and phallus, left lateral view; 91, Ejaculatory apodeme, lateral view; 92, Hypandrium and gonopods, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4930399" httpUri="https://zenodo.org/record/4930399/files/figure.png" pageId="17" pageNumber="54">Fig. 91</figureCitation>
). Gonopods asymmetrical, left gonopod slender than right one (
<figureCitation id="40EE2A53CD7D1C5CEC7E3BB7FD99FD6E" box="[515,599,696,722]" captionStart="FIGURES 83–92" captionStartId="18.[151,264,1898,1922]" captionTargetBox="[226,1361,181,1874]" captionTargetId="figure-29@18.[226,1361,181,1874]" captionTargetPageId="18" captionText="FIGURES 83–92. Tomosvaryella similis (Hough, 1899) (IAvH–2583). Male. 83, Habitus, left lateral view; 84, Left hind trochanter, lateral view; 85, Tergites and sternites 6 and 7, ventral view; 86, Tip of abdomen with syntergosternite 8, ventral view; 87, Susrtyli, dorsal view; 88, Left surstylus, lateral view; 89, Right surstylus, lateral view; 90, Phallic guide and phallus, left lateral view; 91, Ejaculatory apodeme, lateral view; 92, Hypandrium and gonopods, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4930399" httpUri="https://zenodo.org/record/4930399/files/figure.png" pageId="17" pageNumber="54">Fig. 92</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="90CF655DCD7D1C5CEEBA3BD4FD5DFCDC" pageId="17" pageNumber="54" type="materials_examined">
<paragraph id="D86A36D6CD7D1C5CEEBA3BD4FD5DFCDC" blockId="17.[151,1437,376,1113]" pageId="17" pageNumber="54">
<emphasis id="EAA1EAC4CD7D1C5CEEBA3BD4FE61FD49" bold="true" box="[199,431,731,757]" pageId="17" pageNumber="54">Material examined.</emphasis>
<materialsCitation id="68BD3C8BCD7D1C5CEFC83BD4FAE5FCA4" ID-GBIF-Occurrence="3335985387" collectingDate="2001-11-16" collectingDateMax="2001-11-30" collectingDateMin="2001-11-16" collectionCode="LEUA" collectorName="Malaise &amp; P. Reina" country="Colombia" elevation="2600" latitude="5.4166665" location="Iguaque" longLatPrecision="1305" longitude="-73.45" pageId="17" pageNumber="54" specimenCount="1" specimenCount-male="1" stateProvince="Boyaca">
<collectingCountry id="A0C27646CD7D1C5CEFC83BD4FD83FD49" box="[437,589,731,757]" name="Colombia" pageId="17" pageNumber="54">COLOMBIA</collectingCountry>
,
<emphasis id="EAA1EAC4CD7D1C5CEC2A3BD5FD60FD49" bold="true" box="[599,686,730,757]" pageId="17" pageNumber="54">
<collectingRegion id="1A11F834CD7D1C5CEC2A3BD5FD60FD49" box="[599,686,730,757]" country="Colombia" name="Boyaca" pageId="17" pageNumber="54">Boyacá</collectingRegion>
</emphasis>
, SFF[Santuario de Fauna y Flora]
<location id="DD0A600DCD7D1C5CEA563BD3FB46FD4A" LSID="urn:lsid:plazi:treatment:507C87C0CD7D1C5CEEEA3998FC62FBE4:DD0A600DCD7D1C5CEA563BD3FB46FD4A" box="[1067,1160,732,758]" country="Colombia" latitude="5.4166665" longLatPrecision="1305" longitude="-73.45" name="Iguaque" pageId="17" pageNumber="54" stateProvince="Boyaca">Iguaque</location>
,
<geoCoordinate id="BDE15011CD7D1C5CEAEF3BD4FB2DFD4A" box="[1170,1251,731,758]" degrees="5" direction="north" minutes="25" orientation="latitude" pageId="17" pageNumber="54" precision="925" value="5.4166665">5º25’N</geoCoordinate>
/
<geoCoordinate id="BDE15011CD7D1C5CEA8B3BD4FA94FD49" box="[1270,1370,731,757]" degrees="73" direction="west" minutes="27" orientation="longitude" pageId="17" pageNumber="54" precision="925" value="-73.45">73º27’W</geoCoordinate>
,
<quantity id="1F2D9B33CD7D1C5CEB193BD4FF61FCA5" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="17" pageNumber="54" unit="m" value="2600.0">
<elevation id="53F8D1E5CD7D1C5CEB193BD4FF61FCA5" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="17" pageNumber="54" unit="m" value="2600.0">2600 m</elevation>
</quantity>
[eters],
<collectorName id="75205300CD7D1C5CEF7E3BF1FE90FCA4" box="[259,350,766,792]" pageId="17" pageNumber="54">Malaise</collectorName>
,
<date id="AC6B1016CD7D1C5CEF173BF1FDC0FCA4" box="[362,526,766,792]" pageId="17" pageNumber="54" value="2001-11-16" valueMax="2001-11-30" valueMin="2001-11-16">
<collectingDate id="BC2FE9FECD7D1C5CEF173BF1FDC0FCA4" box="[362,526,766,792]" pageId="17" pageNumber="54" value="2001-11-16" valueMax="2001-11-30" valueMin="2001-11-16">16–30.xi.2001</collectingDate>
</date>
,
<collectorName id="75205300CD7D1C5CEC673BF0FDB8FCA4" box="[538,630,766,793]" pageId="17" pageNumber="54">P. Reina</collectorName>
leg, “M2583” (
<specimenCount id="CED3FD5FCD7D1C5CED543BF0FC83FCA5" box="[809,845,767,793]" pageId="17" pageNumber="54" type="male">1♂</specimenCount>
,
<collectionCode id="BEC4AE13CD7D1C5CED243BF0FC68FCA5" box="[857,934,767,793]" pageId="17" pageNumber="54">LEUA</collectionCode>
) (photographed specimen); “
<emphasis id="EAA1EAC4CD7D1C5CEA933BF1FAE5FCA4" box="[1262,1323,766,792]" italics="true" pageId="17" pageNumber="54">idem,</emphasis>
</materialsCitation>
<materialsCitation id="68BD3C8BCD7D1C5CEB4E3BF1FD5DFCDC" ID-GBIF-Occurrence="3336171303" collectorName="V. Villalba" country="Colombia" county="Parque Nacional Natural" elevation="140" latitude="5.35" location="Centro Administrativo" longLatPrecision="1305" longitude="-67.85" municipality="Tuparro" pageId="17" pageNumber="54" specimenCount="1" specimenCount-male="1" stateProvince="Vichada">
<emphasis id="EAA1EAC4CD7D1C5CEB4E3BF1FA58FCA4" bold="true" box="[1331,1430,766,792]" pageId="17" pageNumber="54">
<collectingRegion id="1A11F834CD7D1C5CEB4E3BF1FA58FCA4" box="[1331,1430,766,792]" country="Colombia" name="Vichada" pageId="17" pageNumber="54">Vichada</collectingRegion>
</emphasis>
, PNN[
<collectingCounty id="310B4E5ACD7D1C5CEEAA3A2CFE36FC80" box="[215,504,801,829]" pageId="17" pageNumber="54">Parque Nacional Natural</collectingCounty>
]
<collectingMunicipality id="380EACACCD7D1C5CEC753A2CFDAAFC81" box="[520,612,803,829]" pageId="17" pageNumber="54">Tuparro</collectingMunicipality>
,
<location id="DD0A600DCD7D1C5CEC0E3A2DFCBBFC80" LSID="urn:lsid:plazi:treatment:507C87C0CD7D1C5CEEEA3998FC62FBE4:DD0A600DCD7D1C5CEC0E3A2DFCBBFC80" box="[627,885,802,828]" country="Colombia" county="Parque Nacional Natural" latitude="5.35" longLatPrecision="1305" longitude="-67.85" municipality="Tuparro" name="Centro Administrativo" pageId="17" pageNumber="54" stateProvince="Vichada">Centro Administrativo</location>
,
<geoCoordinate id="BDE15011CD7D1C5CEDFE3A2DFC2CFC81" box="[899,994,802,829]" degrees="05" direction="north" minutes="21" orientation="latitude" pageId="17" pageNumber="54" precision="925" value="5.35">05º21’N</geoCoordinate>
/
<geoCoordinate id="BDE15011CD7D1C5CED823A2DFBAAFC81" box="[1023,1124,802,829]" degrees="67" direction="west" minutes="51" orientation="longitude" pageId="17" pageNumber="54" precision="925" value="-67.85">67º51’W</geoCoordinate>
,
<quantity id="1F2D9B33CD7D1C5CEA0F3A2DFB71FC80" box="[1138,1215,801,828]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="17" pageNumber="54" unit="m" value="140.0">
<elevation id="53F8D1E5CD7D1C5CEA0F3A2DFB71FC80" box="[1138,1215,801,828]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="17" pageNumber="54" unit="m" value="140.0">140 m</elevation>
</quantity>
[eters], Malaise 17, 12–29.Jun[vi].2000,
<collectorName id="75205300CD7D1C5CEFFD3A49FE39FCE3" box="[384,503,837,864]" pageId="17" pageNumber="54">V. Villalba</collectorName>
(
<specimenCount id="CED3FD5FCD7D1C5CEC753A49FDE2FCDC" box="[520,556,838,864]" pageId="17" pageNumber="54" type="male">1♂</specimenCount>
, IAvH)”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="90CF655DCD7D1C5CEEBA3A66FA9DFC1B" pageId="17" pageNumber="54" type="distribution">
<paragraph id="D86A36D6CD7D1C5CEEBA3A66FA9DFC1B" blockId="17.[151,1437,376,1113]" pageId="17" pageNumber="54">
<emphasis id="EAA1EAC4CD7D1C5CEEBA3A66FDCBFC3F" bold="true" box="[199,517,873,899]" pageId="17" pageNumber="54">Geographical distribution.</emphasis>
<collectingCountry id="A0C27646CD7D1C5CEC713A66FDB8FC3F" box="[524,630,873,899]" name="Bahamas" pageId="17" pageNumber="54">Bahamas</collectingCountry>
;
<collectingCountry id="A0C27646CD7D1C5CECFC3A66FD0EFC3F" box="[641,704,873,899]" name="United States of America" pageId="17" pageNumber="54">USA</collectingCountry>
.;
<collectingCountry id="A0C27646CD7D1C5CECB23A66FCE6FC3F" box="[719,808,873,899]" name="Mexico" pageId="17" pageNumber="54">Mexico</collectingCountry>
;
<collectingCountry id="A0C27646CD7D1C5CED4E3A66FC68FC3F" box="[819,934,873,899]" name="Nicaragua" pageId="17" pageNumber="54">Nicaragua</collectingCountry>
(
<bibRefCitation id="BC444B27CD7D1C5CEDC93A66FB12FC3F" author="Rodriguez, H. C. &amp; Rafael, J. A." box="[948,1244,873,899]" pageId="17" pageNumber="54" refId="ref23353" refString="Rodriguez, H. C. &amp; Rafael, J. A. (2012) Pipunculidae (Diptera) of the Latin America and the Caribbean: A catalog of Species with Notes on Biology and Pipunculid-Host Associations. LAP Lambert Academic Publishing GmbH &amp; Co. KG, Saarbrucken, 48 pp." type="book" year="2012">Rodriguez &amp; Rafael 2012</bibRefCitation>
);
<collectingCountry id="A0C27646CD7D1C5CEA923A66FA93FC3F" box="[1263,1373,873,899]" name="Colombia" pageId="17" pageNumber="54">Colombia</collectingCountry>
(new record),
<collectingRegion id="1A11F834CD7D1C5CEE8B3A82FE83FC1B" box="[246,333,909,935]" country="Colombia" name="Boyaca" pageId="17" pageNumber="54">Boyacá</collectingRegion>
, Santuario de Fauna y Flora Iguaque;
<collectingRegion id="1A11F834CD7D1C5CEC8A3A83FC98FC1A" box="[759,854,908,934]" country="Colombia" name="Vichada" pageId="17" pageNumber="54">Vichada</collectingRegion>
, Parque Nacional Natural Tuparro (
<figureCitation id="40EE2A53CD7D1C5CEA993A83FA88FC1B" box="[1252,1350,908,935]" captionStart="FIGURE 133" captionStartId="27.[151,250,1909,1933]" captionTargetBox="[264,1323,1205,1885]" captionTargetId="figure-148@27.[264,1323,1205,1885]" captionTargetPageId="27" captionText="FIGURE 133. Geographical records of Tomosvaryella species in Colombia. T. lynchi, T. macarenensis sp. nov., T. similis, T. venezuelana, T. subvirescens and T. scopulata." figureDoi="http://doi.org/10.5281/zenodo.4930417" httpUri="https://zenodo.org/record/4930417/files/figure.png" pageId="17" pageNumber="54">Fig. 133</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="90CF655DCD7D1C5CEEBA3ABFFBB6FBAD" pageId="17" pageNumber="54" type="biology_ecology">
<paragraph id="D86A36D6CD7D1C5CEEBA3ABFFBB6FBAD" blockId="17.[151,1437,376,1113]" pageId="17" pageNumber="54">
<emphasis id="EAA1EAC4CD7D1C5CEEBA3ABFFEE4FC76" bold="true" box="[199,298,944,970]" pageId="17" pageNumber="54">Habitat.</emphasis>
The specimens were collected with Malaise traps at ground level in areas of tropical dry forest, in the Northeast region in the Santuario de Fauna y Flora Iguaque, and in gallery forest with low elevation of the canopy from the ground in Parque Nacional Natural Tuparro in the Orinoco region of
<collectingCountry id="A0C27646CD7D1C5CEA7F3AF8FBBAFBAD" box="[1026,1140,1015,1041]" name="Colombia" pageId="17" pageNumber="54">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="90CF655DCD7D1C5CEEBA3D15FC62FBE4" pageId="17" pageNumber="54" type="discussion">
<paragraph id="D86A36D6CD7D1C5CEEBA3D15FC62FBE4" blockId="17.[151,1437,376,1113]" pageId="17" pageNumber="54">
<emphasis id="EAA1EAC4CD7D1C5CEEBA3D15FE81FB88" bold="true" box="[199,335,1050,1076]" pageId="17" pageNumber="54">Comments.</emphasis>
The studied specimens are similar to the
<typeStatus id="076E8874CD7D1C5CED5A3D15FC47FB88" box="[807,905,1050,1076]" pageId="17" pageNumber="54" type="holotype">holotype</typeStatus>
illustrated by
<bibRefCitation id="BC444B27CD7D1C5CEA563D15FAA9FB89" author="Ale-Rocha, R. &amp; Rafael, J. A." box="[1067,1383,1050,1077]" pageId="17" pageNumber="54" pagination="407 - 427" refId="ref21517" refString="Ale-Rocha, R. &amp; Rafael, J. A. (1995) Tomosvaryella Aczel do Mexico, America Central e Antilhas (Diptera, Pipunculidae). Revista Brasileira de Zoologia, 12, 407 - 427. https: // doi. org / 10.1590 / S 0101 - 81751995000200017" type="journal article" year="1995">Ale-Rocha &amp; Rafael (1995)</bibRefCitation>
, differing only the length, which exceeds in 0.58 mm that of the
<typeStatus id="076E8874CD7D1C5CED3F3D31FC66FBE4" box="[834,936,1086,1112]" pageId="17" pageNumber="54" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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