treatments-xml/data/83/3F/87/833F87A34300FFFFFF43FF79FBC9FE49.xml

<document id="B3DAB7E5504114AF21B45AC680EC2D10" ID-DOI="10.11646/zootaxa.5555.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="14508494" ID-ZooBank="A27CB74D-EB39-4475-A059-D649C2DBEFB6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1734446566316" checkinUser="plazi" docAuthor="Pinzón-M, Conchita &amp; Pinto-Da-Rocha, Ricardo" docDate="2024" docId="833F87A34300FFFFFF43FF79FBC9FE49" docLanguage="en" docName="zootaxa.5555.1.3.pdf" docOrigin="Zootaxa 5555 (1)" docSource="https://doi.org/10.11646/zootaxa.5555.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhaucus marmoratus Pinzón-M &amp; Pinto-Da-Rocha 2024, New Combination" docType="treatment" docVersion="3" lastPageNumber="48" masterDocId="7F06FFDB430CFFF2FFD4FFEEFFF9FFC0" masterDocTitle="Two new species of Rhaucus Simon, 1879 reveal putative synapomorphies related to the microstructural morphology of male genitalia (Opiliones: Cosmetidae)" masterLastPageNumber="56" masterPageNumber="35" pageNumber="47" updateTime="1734699100454" updateUser="guilherme" zenodo-license-document="CLOSED">
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<mods:title id="57CB0A6A92F40D7ABFFDC5C6DCDAEB79">Two new species of Rhaucus Simon, 1879 reveal putative synapomorphies related to the microstructural morphology of male genitalia (Opiliones: Cosmetidae)</mods:title>
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<mods:namePart id="E40906A95AE0851F288DD6FFDEF8AC49">Pinzón-M, Conchita</mods:namePart>
<mods:affiliation id="BFF20F6107E29BD9B3F9489FCB656576">Departamento de Zoologia, Instituto de Biociências, Universidade de São Paulo, Caixa Postal: 11.461, 05422 - 970, São Paulo, SP, Brazil</mods:affiliation>
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<mods:namePart id="284E26A203A0A40822B6142DFCD37106">Pinto-Da-Rocha, Ricardo</mods:namePart>
<mods:affiliation id="2FAF1EE548237445272378F43AAAB136">Departamento de Zoologia, Instituto de Biociências, Universidade de São Paulo, Caixa Postal: 11.461, 05422 - 970, São Paulo, SP, Brazil</mods:affiliation>
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<mods:date id="9AFB332BD7179CBCEAE145FB76E9BF5C">2024</mods:date>
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<mods:url id="72ABBC6B1BA9C7B20A582228F94B3C27">https://doi.org/10.11646/zootaxa.5555.1.3</mods:url>
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<treatment id="833F87A34300FFFFFF43FF79FBC9FE49" ID-DOI="http://doi.org/10.5281/zenodo.14508613" ID-Zenodo-Dep="14508613" LSID="urn:lsid:plazi:treatment:833F87A34300FFFFFF43FF79FBC9FE49" httpUri="http://treatment.plazi.org/id/833F87A34300FFFFFF43FF79FBC9FE49" lastPageId="13" lastPageNumber="48" pageId="12" pageNumber="47">
<subSubSection id="438C653E4300FFFEFF43FF79FCB2FF71" box="[151,843,151,178]" pageId="12" pageNumber="47" type="nomenclature">
<paragraph id="0B2936B54300FFFEFF43FF79FCB2FF71" blockId="12.[151,843,151,213]" box="[151,843,151,178]" pageId="12" pageNumber="47">
<heading id="506181D94300FFFEFF43FF79FCB2FF71" bold="true" box="[151,843,151,178]" fontSize="11" level="1" pageId="12" pageNumber="47" reason="1">
<emphasis id="39E2EAA74300FFFEFF43FF79FCB2FF71" bold="true" box="[151,843,151,178]" pageId="12" pageNumber="47">
<taxonomicName id="CC964D364300FFFEFF43FF79FD99FF71" authority="(Roewer, 1912)" authorityName="Pinzón-M &amp; Pinto-Da-Rocha" authorityYear="2024" baseAuthorityName="Roewer" baseAuthorityYear="1912" box="[151,608,151,178]" class="Arachnida" family="Cosmetidae" genus="Rhaucus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="marmoratus" status="comb. nov.">
<emphasis id="39E2EAA74300FFFEFF43FF79FE67FF71" bold="true" box="[151,414,151,178]" italics="true" pageId="12" pageNumber="47">Rhaucus marmoratus</emphasis>
(Roewer, 1912)
</taxonomicName>
<taxonomicNameLabel id="22D157DC4300FFFEFDB3FF79FCB2FF71" box="[615,843,151,177]" pageId="12" pageNumber="47" rank="species">New Combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="438C653E4300FFFEFF43FF55FBBBFEF1" pageId="12" pageNumber="47" type="description">
<paragraph id="0B2936B54300FFFEFF43FF55FEE3FF15" blockId="12.[151,843,151,213]" box="[151,282,187,213]" pageId="12" pageNumber="47">
(
<figureCitation id="93AD2A304300FFFEFF4BFF55FF1EFF15" box="[159,231,187,213]" captionStart="FIGURE 8" captionStartId="12.[152,255,1937,1961]" captionTargetBox="[206,1380,899,1911]" captionTargetId="figure-333@12.[206,1380,899,1911]" captionTargetPageId="12" captionText="FIGURE 8. Rhaucus marmoratus comb. nov. (ICN-Ar-1659) A. Male. Habitus, dorsal view; B. Female. Habitus, dorsal view; C–F. Penis: C. Dorsal view; D. Ventral view, arrow showing the areolated MS B.; E–F. Lateral view. Scales: 0.05 mm. Micrographs C–F. Courtesy of B. N. Damrom." figureDoi="http://doi.org/10.5281/zenodo.14508516" httpUri="https://zenodo.org/record/14508516/files/figure.png" pageId="12" pageNumber="47">Figs 8</figureCitation>
, 
<figureCitation id="93AD2A304300FFFEFF20FF55FEE8FF15" box="[244,273,187,213]" captionStart="FIGURE 12" captionStartId="16.[152,255,1648,1672]" captionTargetBox="[170,1416,179,1622]" captionTargetId="figure-25@16.[170,1416,179,1622]" captionTargetPageId="16" captionText="FIGURE 12. Distribution map of Rhaucus spp." figureDoi="http://doi.org/10.5281/zenodo.14508526" httpUri="https://zenodo.org/record/14508526/files/figure.png" pageId="12" pageNumber="47">12</figureCitation>
)
</paragraph>
<paragraph id="0B2936B54300FFFEFF43FF14FBBBFEF1" blockId="12.[151,1436,250,305]" pageId="12" pageNumber="47">
<taxonomicName id="CC964D364300FFFEFF43FF14FD70FED2" authority="Roewer, 1912 c: 142" authorityName="Roewer" authorityPageNumber="142" authorityYear="1912" box="[151,649,250,274]" class="Arachnida" family="Cosmetidae" genus="Pararhaucus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="marmoratus">
<emphasis id="39E2EAA74300FFFEFF43FF14FE53FED2" bold="true" box="[151,426,250,274]" italics="true" pageId="12" pageNumber="47">Pararhaucus marmoratus</emphasis>
Roewer, 1912c: 142
</taxonomicName>
, pl. 7, 
<figureCitation id="93AD2A304300FFFEFD0DFF14FCC7FED2" box="[729,830,250,274]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="5.[152,255,1644,1668]" captionStartId-1="6.[152,255,1909,1933]" captionTargetBox-0="[151,1436,179,1619]" captionTargetBox-1="[186,1393,187,1874]" captionTargetId-0="figure-24@5.[151,1436,179,1619]" captionTargetId-1="figure-25@6.[186,1401,179,1882]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 2. Rhaucus artifex sp.nov. A–B: Male holotype (minor male) (MUSENUV-Ar-2618). A. Habitus, dorsal view. B. Habitus, lateral view. C. Male Paratype (major male) (MNRJ 1594). Dorsal view. D. Female paratype (MUSENUV-Ar 2619). Habitus, dorsal view. Foto C courtesy of Daniela Ahumada, scale bar 5mm." captionText-1="FIGURE 3. Rhaucus artifex sp. nov. A–J: Male holotype (MUSENUV-Ar-2618).A. Habitus, dorsal view; B. Right chelicera, dorsal view; C. Right hand of chelicera, anterior view; D. Right leg, Pa, dorsal view; E. Left leg I, tarsomeres, mesal view; F–H. Left leg IV: F. Ti, ventral view; G. Fe, ventral view; H. Fe, dorsal view; I–J. Left pedipalp: I. Tro, Fe, Pa, Ti, Ta, Dorsal view; J. Same, Mesal view. Scales: 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14508498" figureDoi-1="http://doi.org/10.5281/zenodo.14508500" httpUri-0="https://zenodo.org/record/14508498/files/figure.png" httpUri-1="https://zenodo.org/record/14508500/files/figure.png" pageId="12" pageNumber="47">Figs. 2–3</figureCitation>
; Flórez &amp; Sánchez, 1995: 368. 
<taxonomicName id="CC964D364300FFFEFB47FF14FA6CFED2" baseAuthorityName="Roewer" baseAuthorityYear="1912" box="[1171,1429,250,274]" class="Arachnida" family="Cosmetidae" genus="Meterginus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="marmoratus">
<emphasis id="39E2EAA74300FFFEFB47FF14FA6CFED2" bold="true" box="[1171,1429,250,274]" italics="true" pageId="12" pageNumber="47">Meterginus marmoratus</emphasis>
</taxonomicName>
: Roewer, 1912b: 106; 1923: 383, Fig. 467; Mello-Leitão, 1932: 87; Kury, 2003: 72, 280.
</paragraph>
</subSubSection>
<subSubSection id="438C653E4300FFFEFF43FEB5FBF8FE22" pageId="12" pageNumber="47" type="materials_examined">
<paragraph id="0B2936B54300FFFEFF43FEB5FCB2FE59" blockId="12.[151,1437,347,878]" pageId="12" pageNumber="47">
<emphasis id="39E2EAA74300FFFEFF43FEB5FEB4FEB5" bold="true" box="[151,333,347,373]" pageId="12" pageNumber="47">Type Material:</emphasis>
<materialsCitation id="BBFE3CE84300FFFEFE82FEB5FEF5FE59" collectionCode="MNHN" country="Colombia" location="Between Tambo and Boca del Monte" municipality="Between Tambo" pageId="12" pageNumber="47" specimenCount="4" specimenCount-male="4" stateProvince="Cundinamarca" typeStatus="syntype">
<collectingCountry id="738176254300FFFEFE82FEB5FE3EFEB5" box="[342,455,347,373]" name="Colombia" pageId="12" pageNumber="47">Colombia</collectingCountry>
, 
<collectingRegion id="C952F8574300FFFEFE01FEB5FD82FEB5" box="[469,635,347,373]" country="Colombia" name="Cundinamarca" pageId="12" pageNumber="47">Cundinamarca</collectingRegion>
, 
<location id="0E49606E4300FFFEFD5CFEB2FBCAFEB6" LSID="urn:lsid:plazi:treatment:833F87A34300FFFFFF43FF79FBC9FE49:0E49606E4300FFFEFD5CFEB2FBCAFEB6" box="[648,1075,347,374]" country="Colombia" municipality="Between Tambo" name="Between Tambo and Boca del Monte" pageId="12" pageNumber="47" stateProvince="Cundinamarca">Between Tambo and Boca del Monte</location>
(
<specimenCount id="1D90FD3C4300FFFEFB90FEB5FB67FEB5" box="[1092,1182,347,373]" count="4" pageId="12" pageNumber="47" type="male">4 males</specimenCount>
<typeStatus id="D42D88174300FFFEFB73FEB2FAF0FEB6" box="[1191,1289,348,374]" pageId="12" pageNumber="47" type="syntype">syntypes</typeStatus>
<collectionCode id="6D87AE704300FFFEFAC7FEB2FA92FEB6" box="[1299,1387,348,374]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="12" pageNumber="47">MNHN</collectionCode>
, not examined;
</materialsCitation>
<materialsCitation id="BBFE3CE84300FFFEFEC7FE91FCBEFE59" box="[275,839,383,410]" collectionCode="SMF" country="Colombia" location="Boca del Monte" municipality="Between Tambo" pageId="12" pageNumber="47" specimenCode="SMF 285-32" specimenCount="1" specimenCount-male="1" stateProvince="Cundinamarca" typeStatus="syntype">
<specimenCount id="1D90FD3C4300FFFEFEC7FE91FEA7FE59" box="[275,350,383,409]" count="1" pageId="12" pageNumber="47" type="male">1 male</specimenCount>
<typeStatus id="D42D88174300FFFEFEB1FE6EFE45FE5A" box="[357,444,384,410]" pageId="12" pageNumber="47" type="syntype">syntype</typeStatus>
<specimenCode id="5B309ECE4300FFFEFE17FE91FDABFE5A" box="[451,594,383,410]" pageId="12" pageNumber="47">SMF 285-32</specimenCode>
examined by photos)
</materialsCitation>
.
</paragraph>
<paragraph id="0B2936B54300FFFEFF13FE4DFBF8FE22" blockId="12.[151,1437,347,878]" pageId="12" pageNumber="47">
<emphasis id="39E2EAA74300FFFEFF13FE4DFDFDFE7D" bold="true" box="[199,516,419,445]" pageId="12" pageNumber="47">Other Examined Material:</emphasis>
<materialsCitation id="BBFE3CE84300FFFEFDDDFE4DFC04FE22" collectingDate="2016-10-15" collectorName="Grupo Biologia" country="Colombia" latitude="4.6015553" location="Parque Nacional Natural Chicaque" longLatPrecision="15" longitude="-94.31278" pageId="12" pageNumber="47" specimenCode="ICN-Ar-1659" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Cundinamarca">
<collectingCountry id="738176254300FFFEFDDDFE4DFD83FE7D" box="[521,634,419,445]" name="Colombia" pageId="12" pageNumber="47">Colombia</collectingCountry>
, 
<collectingRegion id="C952F8574300FFFEFD50FE4DFCD5FE7D" box="[644,812,419,445]" country="Colombia" name="Cundinamarca" pageId="12" pageNumber="47">Cundinamarca</collectingRegion>
: 
<specimenCount id="1D90FD3C4300FFFEFCE1FE4DFC70FE7D" box="[821,905,419,445]" count="3" pageId="12" pageNumber="47" type="male">3 males</specimenCount>
and 
<specimenCount id="1D90FD3C4300FFFEFC69FE4DFBDEFE7D" box="[957,1063,419,445]" count="3" pageId="12" pageNumber="47" type="female">3 females</specimenCount>
(
<specimenCode id="5B309ECE4300FFFEFBE1FE4DFB2AFE7E" box="[1077,1235,419,446]" pageId="12" pageNumber="47">ICN-Ar-1659</specimenCode>
), 
<location id="0E49606E4300FFFEFB30FE4AFEA6FE21" LSID="urn:lsid:plazi:treatment:833F87A34300FFFFFF43FF79FBC9FE49:0E49606E4300FFFEFB30FE4AFEA6FE21" country="Colombia" latitude="4.6015553" longLatPrecision="15" longitude="-94.31278" name="Parque Nacional Natural Chicaque" pageId="12" pageNumber="47" stateProvince="Cundinamarca">Parque Nacional Natural Chicaque</location>
, 
<geoCoordinate id="6EA250724300FFFEFEBEFE29FE0AFE22" box="[362,499,455,482]" degrees="4" direction="north" minutes="36" orientation="latitude" pageId="12" pageNumber="47" precision="1" seconds="5.6" value="4.6015553">4°36’5.6’’N</geoCoordinate>
, 
<geoCoordinate id="6EA250724300FFFEFE2AFE29FD6BFE21" box="[510,658,455,481]" degrees="94" direction="west" minutes="18" orientation="longitude" pageId="12" pageNumber="47" precision="15" seconds="46" value="-94.31278">94°18’46’’W</geoCoordinate>
. 
<collectorName id="A66353634300FFFEFD49FE29FCB5FE21" box="[669,844,455,481]" pageId="12" pageNumber="47">Grupo Biología</collectorName>
leg. 
<date id="7F2810754300FFFEFC57FE29FC04FE22" box="[899,1021,455,482]" pageId="12" pageNumber="47" value="2016-10-15">
<collectingDate id="6F6CE99D4300FFFEFC57FE29FC04FE22" box="[899,1021,455,482]" pageId="12" pageNumber="47" value="2016-10-15">15.X.2016</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="438C653E4300FFFEFF13FE05FA86FCAD" pageId="12" pageNumber="47" type="diagnosis">
<paragraph id="0B2936B54300FFFEFF13FE05FA86FCAD" blockId="12.[151,1437,347,878]" pageId="12" pageNumber="47">
<emphasis id="39E2EAA74300FFFEFF13FE05FEBDFDC5" bold="true" box="[199,324,491,517]" pageId="12" pageNumber="47">Diagnosis:</emphasis>
Ozopore dorsally located, with the ID mostly projected and an elongated slit of the ozopore extending towards the lateral margin of the DS (with a rounded ozopore in 
<taxonomicName id="CC964D364300FFFEFC2DFDFEFB9AFDE9" authorityName="Pinzón-M &amp; Pinto-Da-Rocha" authorityYear="2024" box="[1017,1123,527,553]" class="Arachnida" family="Cosmetidae" genus="Rhaucus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="artifex" status="sp. nov.">
<emphasis id="39E2EAA74300FFFEFC2DFDFEFB9AFDE9" box="[1017,1123,527,553]" italics="true" pageId="12" pageNumber="47">R. artifex</emphasis>
</taxonomicName>
<emphasis id="39E2EAA74300FFFEFBB8FDE1FB3EFDE9" bold="true" box="[1132,1223,527,553]" pageId="12" pageNumber="47">
<taxonomicNameLabel id="22D157DC4300FFFEFBB8FDE1FB3EFDE9" box="[1132,1223,527,553]" pageId="12" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="CC964D364300FFFEFB03FDFEFA6CFDE9" baseAuthorityName="Simon" baseAuthorityYear="1879" box="[1239,1429,527,553]" class="Arachnida" family="Cosmetidae" genus="Rhaucus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="papilionaceus">
<emphasis id="39E2EAA74300FFFEFB03FDFEFA6CFDE9" box="[1239,1429,527,553]" italics="true" pageId="12" pageNumber="47">R. papilionaceus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CC964D364300FFFEFF43FDDAFE92FD8D" authorityName="Simon" authorityYear="1879" box="[151,363,563,589]" class="Arachnida" family="Cosmetidae" genus="Rhaucus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="quinquelineatus">
<emphasis id="39E2EAA74300FFFEFF43FDDAFE92FD8D" box="[151,363,563,589]" italics="true" pageId="12" pageNumber="47">R. quinquelineatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CC964D364300FFFEFEAEFDDAFE0DFD8D" baseAuthorityName="Simon" baseAuthorityYear="1879" box="[378,500,564,589]" class="Arachnida" family="Cosmetidae" genus="Rhaucus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="serripes">
<emphasis id="39E2EAA74300FFFEFEAEFDDAFE0DFD8D" box="[378,500,564,589]" italics="true" pageId="12" pageNumber="47">R. serripes</emphasis>
</taxonomicName>
, and 
<taxonomicName id="CC964D364300FFFEFDE0FDDAFD33FD8D" authorityName="Simon" authorityYear="1879" box="[564,714,563,589]" class="Arachnida" family="Cosmetidae" genus="Rhaucus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="vulneratus">
<emphasis id="39E2EAA74300FFFEFDE0FDDAFD33FD8D" box="[564,714,563,589]" italics="true" pageId="12" pageNumber="47">R. vulneratus</emphasis>
</taxonomicName>
; in 
<taxonomicName id="CC964D364300FFFEFD2DFDDAFC8CFD8D" authorityName="Pinzón-M &amp; Pinto-Da-Rocha" authorityYear="2024" baseAuthorityName="Roewer" baseAuthorityYear="1912" box="[761,885,564,589]" class="Arachnida" family="Cosmetidae" genus="Rhaucus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="serratus" status="comb. nov.">
<emphasis id="39E2EAA74300FFFEFD2DFDDAFC8CFD8D" box="[761,885,564,589]" italics="true" pageId="12" pageNumber="47">R. serratus</emphasis>
</taxonomicName>
<emphasis id="39E2EAA74300FFFEFCA9FDDDFC06FD8D" bold="true" box="[893,1023,563,589]" pageId="12" pageNumber="47">
<taxonomicNameLabel id="22D157DC4300FFFEFCA9FDDDFC06FD8D" box="[893,1023,563,589]" pageId="12" pageNumber="47" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, the integumentary dome covers the ozopore and is well projected dorsally; in 
<taxonomicName id="CC964D364300FFFEFDA1FDB6FD08FDB1" authorityName="Pinzón-M &amp; Pinto-Da-Rocha" authorityYear="2024" box="[629,753,600,625]" class="Arachnida" family="Cosmetidae" genus="Rhaucus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="gaiterus" status="sp. nov.">
<emphasis id="39E2EAA74300FFFEFDA1FDB6FD08FDB1" box="[629,753,600,625]" italics="true" pageId="12" pageNumber="47">R. gaiterus</emphasis>
</taxonomicName>
<emphasis id="39E2EAA74300FFFEFD2DFDB9FCADFDB1" bold="true" box="[761,852,599,625]" pageId="12" pageNumber="47">
<taxonomicNameLabel id="22D157DC4300FFFEFD2DFDB9FCADFDB1" box="[761,852,599,625]" pageId="12" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, there is no dorsal fold). The ocularium has a well-marked medial depression (not well-marked in 
<taxonomicName id="CC964D364300FFFEFD66FD92FCE5FD55" authorityName="Pinzón-M &amp; Pinto-Da-Rocha" authorityYear="2024" box="[690,796,635,661]" class="Arachnida" family="Cosmetidae" genus="Rhaucus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="artifex" status="sp. nov.">
<emphasis id="39E2EAA74300FFFEFD66FD92FCE5FD55" box="[690,796,635,661]" italics="true" pageId="12" pageNumber="47">R. artifex</emphasis>
</taxonomicName>
<emphasis id="39E2EAA74300FFFEFCF0FD95FC86FD55" bold="true" box="[804,895,635,661]" pageId="12" pageNumber="47">
<taxonomicNameLabel id="22D157DC4300FFFEFCF0FD95FC86FD55" box="[804,895,635,661]" pageId="12" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="CC964D364300FFFEFC5BFD92FC0FFD55" authorityName="Garcia &amp; Kury" authorityYear="2017" box="[911,1014,635,661]" class="Arachnida" family="Cosmetidae" genus="Rhaucus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="florezi">
<emphasis id="39E2EAA74300FFFEFC5BFD92FC0FFD55" box="[911,1014,635,661]" italics="true" pageId="12" pageNumber="47">R. florezi</emphasis>
</taxonomicName>
, 
<taxonomicName id="CC964D364300FFFEFBD1FD92FB7BFD55" authorityName="Pinzón-M &amp; Pinto-Da-Rocha" authorityYear="2024" box="[1029,1154,636,661]" class="Arachnida" family="Cosmetidae" genus="Rhaucus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="gaiterus" status="sp. nov.">
<emphasis id="39E2EAA74300FFFEFBD1FD92FB7BFD55" box="[1029,1154,636,661]" italics="true" pageId="12" pageNumber="47">R. gaiterus</emphasis>
</taxonomicName>
<emphasis id="39E2EAA74300FFFEFB5EFD95FB1CFD55" bold="true" box="[1162,1253,635,661]" pageId="12" pageNumber="47">
<taxonomicNameLabel id="22D157DC4300FFFEFB5EFD95FB1CFD55" box="[1162,1253,635,661]" pageId="12" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="CC964D364300FFFEFB21FD92FA8DFD55" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1941" box="[1269,1396,635,661]" class="Arachnida" family="Cosmetidae" genus="Rhaucus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="39E2EAA74300FFFEFB21FD92FA8DFD55" box="[1269,1396,635,661]" italics="true" pageId="12" pageNumber="47">R. robustus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CC964D364300FFFEFA50FD92FEB3FD79" authorityName="Simon" authorityYear="1879" class="Arachnida" family="Cosmetidae" genus="Rhaucus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="quinquelineatus">
<emphasis id="39E2EAA74300FFFEFA50FD92FEB3FD79" italics="true" pageId="12" pageNumber="47">R. quinquelineatus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="CC964D364300FFFEFE5EFD4EFDFEFD79" authorityName="Pinzón-M &amp; Pinto-Da-Rocha" authorityYear="2024" baseAuthorityName="Roewer" baseAuthorityYear="1912" box="[394,519,672,697]" class="Arachnida" family="Cosmetidae" genus="Rhaucus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="serratus" status="comb. nov.">
<emphasis id="39E2EAA74300FFFEFE5EFD4EFDFEFD79" box="[394,519,672,697]" italics="true" pageId="12" pageNumber="47">R. serratus</emphasis>
</taxonomicName>
<emphasis id="39E2EAA74300FFFEFDDBFD71FD68FD79" bold="true" box="[527,657,671,697]" pageId="12" pageNumber="47">
<taxonomicNameLabel id="22D157DC4300FFFEFDDBFD71FD68FD79" box="[527,657,671,697]" pageId="12" pageNumber="47" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
). Area III of the DS has a pair of prominent mammiliform tubercles close to each other (spiniform tubercles in 
<taxonomicName id="CC964D364300FFFEFDA0FD2AFD25FD1D" authorityName="Pinzón-M &amp; Pinto-Da-Rocha" authorityYear="2024" box="[628,732,707,733]" class="Arachnida" family="Cosmetidae" genus="Rhaucus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="artifex" status="sp. nov.">
<emphasis id="39E2EAA74300FFFEFDA0FD2AFD25FD1D" box="[628,732,707,733]" italics="true" pageId="12" pageNumber="47">R. artifex</emphasis>
</taxonomicName>
<emphasis id="39E2EAA74300FFFEFD37FD2DFCC5FD1D" bold="true" box="[739,828,707,733]" pageId="12" pageNumber="47">
<taxonomicNameLabel id="22D157DC4300FFFEFD37FD2DFCC5FD1D" box="[739,828,707,733]" pageId="12" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="CC964D364300FFFEFCA6FD2AFC14FD1D" authorityName="Pinzón-M &amp; Pinto-Da-Rocha" authorityYear="2024" box="[882,1005,708,733]" class="Arachnida" family="Cosmetidae" genus="Rhaucus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="gaiterus" status="sp. nov.">
<emphasis id="39E2EAA74300FFFEFCA6FD2AFC14FD1D" box="[882,1005,708,733]" italics="true" pageId="12" pageNumber="47">R. gaiterus</emphasis>
</taxonomicName>
<emphasis id="39E2EAA74300FFFEFC20FD2DFBB4FD1D" bold="true" box="[1012,1101,707,733]" pageId="12" pageNumber="47">
<taxonomicNameLabel id="22D157DC4300FFFEFC20FD2DFBB4FD1D" box="[1012,1101,707,733]" pageId="12" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; mammiliform and smaller in 
<taxonomicName id="CC964D364300FFFEFF43FD06FF05FCC1" authorityName="Garcia &amp; Kury" authorityYear="2017" box="[151,252,743,769]" class="Arachnida" family="Cosmetidae" genus="Rhaucus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="florezi">
<emphasis id="39E2EAA74300FFFEFF43FD06FF05FCC1" box="[151,252,743,769]" italics="true" pageId="12" pageNumber="47">R. florezi</emphasis>
</taxonomicName>
and 
<taxonomicName id="CC964D364300FFFEFEE5FD06FE3CFCC1" authorityName="Simon" authorityYear="1879" box="[305,453,743,769]" class="Arachnida" family="Cosmetidae" genus="Rhaucus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="vulneratus">
<emphasis id="39E2EAA74300FFFEFEE5FD06FE3CFCC1" box="[305,453,743,769]" italics="true" pageId="12" pageNumber="47">R. vulneratus</emphasis>
</taxonomicName>
). Cx IV has 
<emphasis id="39E2EAA74300FFFEFD85FD09FD15FCC1" box="[593,748,743,769]" italics="true" pageId="12" pageNumber="47">clavi inguines</emphasis>
formed by a mound with few granules (strongly granulated in 
<taxonomicName id="CC964D364300FFFEFF43FCE2FF05FCE5" authorityName="Garcia &amp; Kury" authorityYear="2017" box="[151,252,779,805]" class="Arachnida" family="Cosmetidae" genus="Rhaucus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="florezi">
<emphasis id="39E2EAA74300FFFEFF43FCE2FF05FCE5" box="[151,252,779,805]" italics="true" pageId="12" pageNumber="47">R. florezi</emphasis>
</taxonomicName>
, 
<taxonomicName id="CC964D364300FFFEFEDDFCE2FE23FCE5" authorityName="Simon" authorityYear="1879" box="[265,474,779,805]" class="Arachnida" family="Cosmetidae" genus="Rhaucus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="quinquelineatus">
<emphasis id="39E2EAA74300FFFEFEDDFCE2FE23FCE5" box="[265,474,779,805]" italics="true" pageId="12" pageNumber="47">R. quinquelineatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CC964D364300FFFEFE33FCE2FDA6FCE5" baseAuthorityName="Simon" baseAuthorityYear="1879" box="[487,607,780,805]" class="Arachnida" family="Cosmetidae" genus="Rhaucus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="serripes">
<emphasis id="39E2EAA74300FFFEFE33FCE2FDA6FCE5" box="[487,607,780,805]" italics="true" pageId="12" pageNumber="47">R. serripes</emphasis>
</taxonomicName>
, and 
<taxonomicName id="CC964D364300FFFEFD4EFCE2FCD7FCE5" authorityName="Simon" authorityYear="1879" box="[666,814,779,805]" class="Arachnida" family="Cosmetidae" genus="Rhaucus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="vulneratus">
<emphasis id="39E2EAA74300FFFEFD4EFCE2FCD7FCE5" box="[666,814,779,805]" italics="true" pageId="12" pageNumber="47">R. vulneratus</emphasis>
</taxonomicName>
). The dorsal apical apophysis on Cx IV is multicapitate, forming a keel slightly flattened laterally (not laterally flattened in the remaining species). The color pattern consists of two lateral longitudinal patches extending from the prosoma to the posterior areas of the DS (see 
<figureCitation id="93AD2A304300FFFEFB2CFCBDFA8BFCAD" box="[1272,1394,851,877]" captionStart="FIGURE 8" captionStartId="12.[152,255,1937,1961]" captionTargetBox="[206,1380,899,1911]" captionTargetId="figure-333@12.[206,1380,899,1911]" captionTargetPageId="12" captionText="FIGURE 8. Rhaucus marmoratus comb. nov. (ICN-Ar-1659) A. Male. Habitus, dorsal view; B. Female. Habitus, dorsal view; C–F. Penis: C. Dorsal view; D. Ventral view, arrow showing the areolated MS B.; E–F. Lateral view. Scales: 0.05 mm. Micrographs C–F. Courtesy of B. N. Damrom." figureDoi="http://doi.org/10.5281/zenodo.14508516" httpUri="https://zenodo.org/record/14508516/files/figure.png" pageId="12" pageNumber="47">Fig. 8A–B</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="5FE9663D4300FFFEFF4CF87FFD89F830" ID-DOI="http://doi.org/10.5281/zenodo.14508516" ID-Zenodo-Dep="14508516" httpUri="https://zenodo.org/record/14508516/files/figure.png" pageId="12" pageNumber="47" startId="12.[152,255,1937,1961]" targetBox="[206,1380,899,1911]" targetPageId="12" targetType="figure">
<paragraph id="0B2936B54300FFFEFF4CF87FFD89F830" blockId="12.[151,1437,1937,2033]" pageId="12" pageNumber="47">
<emphasis id="39E2EAA74300FFFEFF4CF87FFEE5F869" bold="true" box="[152,284,1937,1961]" pageId="12" pageNumber="47">FIGURE 8.</emphasis>
<taxonomicName id="CC964D364300FFFEFEF2F87FFDFAF869" authorityName="Pinzón-M &amp; Pinto-Da-Rocha" authorityYear="2024" baseAuthorityName="Roewer" baseAuthorityYear="1912" box="[294,515,1937,1961]" class="Arachnida" family="Cosmetidae" genus="Rhaucus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="marmoratus" status="comb. nov.">
<emphasis id="39E2EAA74300FFFEFEF2F87FFDFAF869" box="[294,515,1937,1961]" italics="true" pageId="12" pageNumber="47">Rhaucus marmoratus</emphasis>
</taxonomicName>
<emphasis id="39E2EAA74300FFFEFDD9F87FFD7CF869" bold="true" box="[525,645,1937,1961]" pageId="12" pageNumber="47">
<taxonomicNameLabel id="22D157DC4300FFFEFDD9F87FFD7CF869" box="[525,645,1937,1961]" pageId="12" pageNumber="47" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
(ICN-Ar-1659) A. Male. Habitus, dorsal view; B. Female. Habitus, dorsal view; C–F. Penis: C. Dorsal view; D. Ventral view, arrow showing the areolated MS B.; E–F. Lateral view. Scales: 0.05 mm. Micrographs C–F. Courtesy of B. N. Damrom.
</paragraph>
</caption>
<subSubSection id="438C653E4301FFFFFF13FF78FBC9FE49" pageId="13" pageNumber="48" type="description">
<paragraph id="0B2936B54301FFFFFF13FF78FBC9FE49" blockId="13.[151,1437,150,393]" pageId="13" pageNumber="48">
<emphasis id="39E2EAA74301FFFFFF13FF78FE5AFF70" bold="true" box="[199,419,150,176]" pageId="13" pageNumber="48">Penis Description:</emphasis>
(
<figureCitation id="93AD2A304301FFFFFE67FF79FDD1FF71" box="[435,552,151,177]" captionStart="FIGURE 8" captionStartId="12.[152,255,1937,1961]" captionTargetBox="[206,1380,899,1911]" captionTargetId="figure-333@12.[206,1380,899,1911]" captionTargetPageId="12" captionText="FIGURE 8. Rhaucus marmoratus comb. nov. (ICN-Ar-1659) A. Male. Habitus, dorsal view; B. Female. Habitus, dorsal view; C–F. Penis: C. Dorsal view; D. Ventral view, arrow showing the areolated MS B.; E–F. Lateral view. Scales: 0.05 mm. Micrographs C–F. Courtesy of B. N. Damrom." figureDoi="http://doi.org/10.5281/zenodo.14508516" httpUri="https://zenodo.org/record/14508516/files/figure.png" pageId="13" pageNumber="48">Fig. 8C–F</figureCitation>
) VP is short and squared. The glans has a dorsal process that is elongated and cylindrical, with wattle extending ventrally and bearing stylar barbs. Two pairs of MS C: C1 is mostly more curved than C2, with both being long and flattened. Two pairs of MS D: D1 is cylindrical and straight, closer to the pair C, while D2 is dorsally attached, short, and located in the middle portion of the VP. Two pairs of MS A: dorsally attached, straight, cylindrical, and bifurcated at the top. Two pairs of MS E are volcano-shaped, and a pair of areolate MS B is located at the base of the VP. The lateral fields of the VP are covered by microsetae 
<typeStatus id="D42D88174301FFFFFB70FEA5FB2DFEA5" box="[1188,1236,331,357]" pageId="13" pageNumber="48">type</typeStatus>
4, and the medial portion has some microsetae T2. The lateral corners of the VP are curved dorsally.
</paragraph>
</subSubSection>
</treatment>
</document>