168 lines
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168 lines
18 KiB
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<document id="8844E8770A2191992F6B52000BB13677" ID-DOI="10.5252/geodiversitas2024v46a15" ID-ISSN="1638-9395" ID-Zenodo-Dep="13985760" ID-ZooBank="urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1729758589633" checkinUser="plazi" docAuthor="Merle, Didier, Pacaud, Jean-Michel, Ledon, Daniel & Goret, Bernard" docDate="2024" docId="038E87DAFFFBE41BFC1EFBC2FAA5F89F" docLanguage="en" docName="geodiversitas.2024.46.15.pdf" docOrigin="Geodiversitas 46 (15)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="(Pterymarchia) Houart 1995" docType="treatment" docVersion="3" lastPageNumber="522" masterDocId="FFB7FFA2FFE6E406FFB8FFD6FF81FFB3" masterDocTitle="New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe" masterLastPageNumber="551" masterPageNumber="495" pageNumber="522" updateTime="1729856536171" updateUser="valdenar" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:titleInfo id="C38372B35139DE8C522CC28CC9F24855">
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<mods:title id="92C98DEFA2132AA05745B1D967B73246">New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe</mods:title>
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<mods:namePart id="11DE77AC5062B69D53534C2175A34F9A">Merle, Didier</mods:namePart>
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<mods:namePart id="F310FCB4CE4127841911E8D55928B151">Pacaud, Jean-Michel</mods:namePart>
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</mods:name>
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<mods:namePart id="8CA3DAE06B4773218CE9343916AB74CE">Ledon, Daniel</mods:namePart>
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<mods:affiliation id="78FB3D7993F445BF5D75FF403B2CC881">Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr</mods:affiliation>
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<mods:nameIdentifier id="5D6CFE32D5D171ED196CE33FE44D0858" type="email">didier.merle@mnhn.fr</mods:nameIdentifier>
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<mods:namePart id="A822BC8294FC0E2C48B9A88AEEBABA52">Goret, Bernard</mods:namePart>
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<mods:affiliation id="EFEECE0D64BEF79808DAE7F3CFF4288A">Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com</mods:affiliation>
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<mods:nameIdentifier id="40E5D370C153C3834E8CCEAA5B4182C3" type="email">bernardgoret@hotmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="09744AF32D37D36B66DE0EA18E232DD5">text</mods:typeOfResource>
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<mods:title id="594C5B59C12AA38A1B4CD5908571D957">Geodiversitas</mods:title>
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<mods:part id="91F80651F2DE2438869EEAF84591D214">
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<mods:date id="2CC5FE656FC4EC644C6FAD187429E2F5">2024</mods:date>
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<mods:number id="5552ECB253BD35AF6D23605DA10C750F">2024-10-24</mods:number>
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<mods:number id="C8B1AE904A10A0650A82BA82F6AA0F1B">46</mods:number>
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<mods:number id="6BE6B6F22223C21369DAAB31C065A612">15</mods:number>
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<mods:start id="01A57790D7641C392AE6E39851D30A18">495</mods:start>
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<mods:url id="BDA036BE11290DFB4ADA6B546C4F5BC5">https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf</mods:url>
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<mods:classification id="D8704517A07F52616ED337CAFFFB1BF5">journal article</mods:classification>
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<mods:identifier id="7DD4B8D8C0855059BC7B68FE9A45106D" type="DOI">10.5252/geodiversitas2024v46a15</mods:identifier>
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<mods:identifier id="E4BEFCC8C8E1B893D6082A7AA431597B" type="ISSN">1638-9395</mods:identifier>
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<mods:identifier id="62E2FFC2909372CB0D48E7C09605A2D0" type="Zenodo-Dep">13985760</mods:identifier>
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<subSubSection id="C33D6547FFFBE41BFC1EFBC2FAB7FB9C" box="[934,1334,1044,1071]" pageId="29" pageNumber="522" type="nomenclature">
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<paragraph id="8B9836CCFFFBE41BFC1EFBC2FAB7FB9C" blockId="29.[934,1334,1044,1071]" box="[934,1334,1044,1071]" pageId="29" pageNumber="522">
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<heading id="D0D081A0FFFBE41BFC1EFBC2FAB7FB9C" box="[934,1334,1044,1071]" centered="true" fontSize="11" level="2" pageId="29" pageNumber="522" reason="2">
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Subgenus
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<taxonomicName id="4C274D4FFFFBE41BFBABFBC2FAB7FB9C" authority="Houart, 1995" authorityName="Houart" authorityYear="1995" box="[1043,1334,1044,1071]" kingdom="Animalia" pageId="29" pageNumber="522" rank="subGenus" subGenus="Pterymarchia">
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<emphasis id="B953EADEFFFBE41BFBABFBC2FB1DFB9D" box="[1043,1180,1044,1070]" italics="true" pageId="29" pageNumber="522">Pterymarchia</emphasis>
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<bibRefCitation id="EFB64B3DFFFBE41BFB1AFBC3FAB7FB9C" author="HOUART R." box="[1186,1334,1044,1071]" pageId="29" pageNumber="522" pagination="127 - 136" refId="ref44916" refString="HOUART R. 1995. - Pterymarchia n. gen. and Vaughtia n. gen., two new muricid genera (Gastropoda: Muricidae: Muricinae and Ocenebrinae). Apex 10 (4): 127 - 136. https: // www. biodiversitylibrary. org / page / 42257087" type="journal article" year="1995">Houart, 1995</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33D6547FFFBE41BFC95FB98FBADFB33" pageId="29" pageNumber="522" type="materials_examined">
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<paragraph id="8B9836CCFFFBE41BFC95FB98FBADFB33" blockId="29.[813,1456,1101,1152]" pageId="29" pageNumber="522">
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<typeStatus id="549C886EFFFBE41BFC95FB98FCDCFBD0" box="[813,861,1102,1125]" pageId="29" pageNumber="522">TYPE</typeStatus>
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</smallCapsWord>
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<smallCapsWord id="8D7EA010FFFBE41BFCDDFB85FC2AFBD7" baselines="1120" box="[869,939,1107,1124]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="species" pageId="29" pageNumber="522">SPECIES</smallCapsWord>
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. —
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<taxonomicName id="4C274D4FFFFBE41BFC62FB98FB67FBD6" authority="Born, 1778" authorityName="Born" authorityYear="1778" box="[986,1254,1101,1125]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="29" pageNumber="522" phylum="Mollusca" rank="species" species="tripterus">
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<emphasis id="B953EADEFFFBE41BFC62FB98FBEAFBD6" box="[986,1131,1102,1125]" italics="true" pageId="29" pageNumber="522">Murex tripterus</emphasis>
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Born, 1778
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</taxonomicName>
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, by original designation. Recent: Indo-Pacific.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33D6547FFFBE41BFC95FB62FAA5F89F" pageId="29" pageNumber="522" type="discussion">
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<paragraph id="8B9836CCFFFBE41BFC95FB62FBF6FB7F" blockId="29.[810,1457,1204,1836]" box="[813,1143,1204,1230]" pageId="29" pageNumber="522">
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<smallCapsWord id="8D7EA010FFFBE41BFC0AFB6FFC52FB7F" baselines="1223" box="[946,979,1209,1228]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="on" pageId="29" pageNumber="522">ON</smallCapsWord>
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<smallCapsWord id="8D7EA010FFFBE41BFC61FB6CFB85FB7E" baselines="1224" box="[985,1028,1210,1229]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="the" pageId="29" pageNumber="522">THE</smallCapsWord>
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<smallCapsWord id="8D7EA010FFFBE41BFBB3FB6FFBF6FB7F" baselines="1223" box="[1035,1143,1209,1228]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="subgenus" pageId="29" pageNumber="522">SUBGENUS</smallCapsWord>
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</paragraph>
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<paragraph id="8B9836CCFFFBE41BFC95FB02FAA5F89F" blockId="29.[810,1457,1204,1836]" pageId="29" pageNumber="522">
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According toHouart (1995),
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<taxonomicName id="4C274D4FFFFBE41BFBE0FB02FB5DFB5D" authorityName="Houart" authorityYear="1995" box="[1112,1244,1236,1262]" class="Gastropoda" family="Muricidae" genus="Pterymarchia" kingdom="Animalia" order="Neogastropoda" pageId="29" pageNumber="522" phylum="Mollusca" rank="genus">
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<emphasis id="B953EADEFFFBE41BFBE0FB02FB5DFB5D" box="[1112,1244,1236,1262]" italics="true" pageId="29" pageNumber="522">Pterymarchia</emphasis>
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</taxonomicName>
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can be distinguished from
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<taxonomicName id="4C274D4FFFFBE41BFCD0FB22FC55FABE" authorityName="Swainson" authorityYear="1833" box="[872,980,1268,1293]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="29" pageNumber="522" phylum="Mollusca" rank="genus">
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<emphasis id="B953EADEFFFBE41BFCD0FB22FC55FABE" box="[872,980,1268,1293]" italics="true" pageId="29" pageNumber="522">Pterynotus</emphasis>
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</taxonomicName>
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(
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<emphasis id="B953EADEFFFBE41BFC5BFB22FB80FABE" box="[995,1025,1268,1293]" italics="true" pageId="29" pageNumber="522">s.s.</emphasis>
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) by having more numerous axial varices on the early teleoconch whorls (9-11 vs 6-8), later appearance in ontogeny of the trivaricate sculpture, when present, sometimes more varices on the last whorls (3-5 vs 3), and more strongly developed denticles.
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<bibRefCitation id="EFB64B3DFFFBE41BFB78FAA2FADDFA3D" author="HOUART R." box="[1216,1372,1395,1422]" pageId="29" pageNumber="522" pagination="127 - 136" refId="ref44916" refString="HOUART R. 1995. - Pterymarchia n. gen. and Vaughtia n. gen., two new muricid genera (Gastropoda: Muricidae: Muricinae and Ocenebrinae). Apex 10 (4): 127 - 136. https: // www. biodiversitylibrary. org / page / 42257087" type="journal article" year="1995">Houart (1995</bibRefCitation>
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,
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<bibRefCitation id="EFB64B3DFFFBE41BFAD4FAA5FA26FA3E" author="HOUART R." box="[1388,1447,1395,1421]" pageId="29" pageNumber="522" pagination="37 - 66" refId="ref45005" refString="HOUART R. 2018. - Historique et classification des especes actuelles de Muricidae (Neogastropoda, Muricoidea). Novapex 19 (2): 37 - 66." type="journal article" year="2018">2018</bibRefCitation>
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) proposed full generic status for
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<taxonomicName id="4C274D4FFFFBE41BFB33FA45FA92FA1E" authorityName="Houart" authorityYear="1995" box="[1163,1299,1427,1453]" class="Gastropoda" family="Muricidae" genus="Pterymarchia" kingdom="Animalia" order="Neogastropoda" pageId="29" pageNumber="522" phylum="Mollusca" rank="genus">
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<emphasis id="B953EADEFFFBE41BFB33FA45FA92FA1E" box="[1163,1299,1427,1453]" italics="true" pageId="29" pageNumber="522">Pterymarchia</emphasis>
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</taxonomicName>
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. However, we prefer to maintain it at subgeneric rank because: 1) based on molecular data,
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<taxonomicName id="4C274D4FFFFBE41BFC59FA02FBCFFA5E" authorityName="Swainson" authorityYear="1833" box="[993,1102,1492,1517]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="29" pageNumber="522" phylum="Mollusca" rank="genus">
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<emphasis id="B953EADEFFFBE41BFC59FA02FBCFFA5E" box="[993,1102,1492,1517]" italics="true" pageId="29" pageNumber="522">Pterynotus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C274D4FFFFBE41BFB31FA05FA94FA5E" authorityName="Houart" authorityYear="1995" box="[1161,1301,1491,1517]" class="Gastropoda" family="Muricidae" genus="Pterymarchia" kingdom="Animalia" order="Neogastropoda" pageId="29" pageNumber="522" phylum="Mollusca" rank="genus">
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<emphasis id="B953EADEFFFBE41BFB31FA05FA94FA5E" box="[1161,1301,1491,1517]" italics="true" pageId="29" pageNumber="522">Pterymarchia</emphasis>
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</taxonomicName>
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are very close (
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<bibRefCitation id="EFB64B3DFFFBE41BFC8EFA25FC71F9BE" author="BARCO A. & CLAREMONT M. & REID D. G. & HOUART R. & BOUCHET P. & WILLIAMS S. T. & CRUAUD C. & COULOUX A. & OLIVERIO M." box="[822,1008,1523,1549]" pageId="29" pageNumber="522" pagination="1025 - 1039" refId="ref42748" refString="BARCO A., CLAREMONT M., REID D. G., HOUART R., BOUCHET P., WILLIAMS S. T., CRUAUD C., COULOUX A. & OLIVERIO M. 2010. - A molecular phylogenetic framework for the Muricidae, a diverse family of carnivorous gastropods. Molecular Phylogenetics and Evolution 55 (3): 1025 - 1039. https: // doi. org / 10.1016 / j. ympev. 2010.03.008" type="journal article" year="2010">
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Barco
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<emphasis id="B953EADEFFFBE41BFCC3FA25FC31F9BE" box="[891,944,1523,1549]" italics="true" pageId="29" pageNumber="522">et al.</emphasis>
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2010
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</bibRefCitation>
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); and 2) based on shell characters,
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<taxonomicName id="4C274D4FFFFBE41BFACAFA25FC03F99F" authorityName="Houart" authorityYear="1995" class="Gastropoda" family="Muricidae" genus="Pterymarchia" kingdom="Animalia" order="Neogastropoda" pageId="29" pageNumber="522" phylum="Mollusca" rank="genus">
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<emphasis id="B953EADEFFFBE41BFACAFA25FC03F99F" italics="true" pageId="29" pageNumber="522">Pterymarchia</emphasis>
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</taxonomicName>
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shares with
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<taxonomicName id="4C274D4FFFFBE41BFBB2F9C5FBF7F99F" authorityName="Swainson" authorityYear="1833" box="[1034,1142,1555,1580]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="29" pageNumber="522" phylum="Mollusca" rank="genus">
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|
<emphasis id="B953EADEFFFBE41BFBB2F9C5FBF7F99F" box="[1034,1142,1555,1580]" italics="true" pageId="29" pageNumber="522">Pterynotus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
a scabrous surface appearing early in ontogeny, generally fine to moderately fine primary cords (from IP to ABP), and a series of internal denticles from ID to D6. Furthermore, several other characters are occasionally shared by both genera, such as the trivaricate sculpture, columellar tubercles [e.g.,
|
|||
|
|
<emphasis id="B953EADEFFFBE41BFBFFF965FBDDF97F" box="[1095,1116,1715,1740]" italics="true" pageId="29" pageNumber="522">P.</emphasis>
|
|||
|
|
(
|
|||
|
|
<emphasis id="B953EADEFFFBE41BFBD1F965FB07F97F" box="[1129,1158,1715,1740]" italics="true" pageId="29" pageNumber="522">s.s.</emphasis>
|
|||
|
|
)
|
|||
|
|
<emphasis id="B953EADEFFFBE41BFB2DF965FB7FF97F" box="[1173,1278,1715,1740]" italics="true" pageId="29" pageNumber="522">patagiatus</emphasis>
|
|||
|
|
(Hedley, 1912),
|
|||
|
|
<emphasis id="B953EADEFFFBE41BFC94F905FCBEF95F" box="[812,831,1747,1772]" italics="true" pageId="29" pageNumber="522">P.</emphasis>
|
|||
|
|
(
|
|||
|
|
<emphasis id="B953EADEFFFBE41BFCE8F905FCEFF95F" box="[848,878,1747,1772]" italics="true" pageId="29" pageNumber="522">s.s.</emphasis>
|
|||
|
|
)
|
|||
|
|
<emphasis id="B953EADEFFFBE41BFCC7F905FB94F95F" box="[895,1045,1747,1772]" italics="true" pageId="29" pageNumber="522">praepatagiatus</emphasis>
|
|||
|
|
<bibRefCitation id="EFB64B3DFFFBE41BFBA6F904FB27F95F" author="VOKES E. H." box="[1054,1190,1746,1772]" pageId="29" pageNumber="522" pagination="1 - 108" refId="ref48996" refString="VOKES E. H. 1992. - Cenozoic Muricidae of the Western Atlantic region, Part IX - Pterynotus, Poirieria, Aspella, Dermomurex, Calotrophon, Acantolabia, and Attiliosa; additions and corrections. Tulane Studies in Geology and Paleontology 25 (1 - 3): 1 - 108." type="journal article" year="1992">Vokes, 1992</bibRefCitation>
|
|||
|
|
and
|
|||
|
|
<emphasis id="B953EADEFFFBE41BFB5BF905FB7BF95F" box="[1251,1274,1747,1772]" italics="true" pageId="29" pageNumber="522">P.</emphasis>
|
|||
|
|
(
|
|||
|
|
<emphasis id="B953EADEFFFBE41BFABCF905FAA3F95F" box="[1284,1314,1747,1772]" italics="true" pageId="29" pageNumber="522">s.s.</emphasis>
|
|||
|
|
)
|
|||
|
|
<emphasis id="B953EADEFFFBE41BFA8BF904FA2EF95F" box="[1331,1455,1746,1772]" italics="true" pageId="29" pageNumber="522">granuliferus</emphasis>
|
|||
|
|
(Grateloup, 1833)], and even developed denticles in some
|
|||
|
|
<emphasis id="B953EADEFFFBE41BFC96F8C4FCC0F898" box="[814,833,1810,1835]" italics="true" pageId="29" pageNumber="522">P.</emphasis>
|
|||
|
|
(
|
|||
|
|
<emphasis id="B953EADEFFFBE41BFCF6F8C4FCEAF898" box="[846,875,1810,1835]" italics="true" pageId="29" pageNumber="522">s.s.</emphasis>
|
|||
|
|
)
|
|||
|
|
<emphasis id="B953EADEFFFBE41BFCC3F8C4FC79F89F" box="[891,1016,1810,1836]" italics="true" pageId="29" pageNumber="522">albobruneus</emphasis>
|
|||
|
|
Bertsch & D’Attilio, 1980.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|