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<document id="3AEC82F1D5FFD6D8370CFB605FEA51AE" ID-CLB-Dataset="51238" ID-DOI="10.5281/zenodo.156768" ID-GBIF-Dataset="0ae8bbb8-3209-479e-bd4e-efa5e38555e4" ID-ISSN="1175­5326" ID-Zenodo-Dep="156768" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459639206332" checkinUser="plazi" docAuthor="Wappler, Torsten" docDate="2003" docId="D33887DEFF963B0AFEF101C9E74AD344" docLanguage="en" docName="zt00374.pdf" docOrigin="Zootaxa 374" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Oblongomorpha lutetia Wappler, 2003, n. sp." docType="treatment" docVersion="10" lastPageNumber="15" masterDocId="2F01FFA6FF9D3B04FFF9072DE524D40A" masterDocTitle="New fossil lace bugs (Heteroptera: Tingidae) from the Middle Eocene of the Grube Messel (Germany), with a catalog of fossil lace bugs" masterLastPageNumber="26" masterPageNumber="1" pageNumber="12" updateTime="1698205971885" updateUser="ExternalLinkService">
<mods:mods id="FBB3276B64912850CE169EFC67E405A1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8FCB86747EC5AC7C42531E9FA134FE14">
<mods:title id="D09230CD98CA09E842D5DDC7F1AAC836">New fossil lace bugs (Heteroptera: Tingidae) from the Middle Eocene of the Grube Messel (Germany), with a catalog of fossil lace bugs</mods:title>
</mods:titleInfo>
<mods:name id="3408DA8F5CE1E0C5914FC5DCE690AC0C" type="personal">
<mods:role id="9106755BF32746A469389DECCD05CCBD">
<mods:roleTerm id="B84C4F0481E97F353B2062DD354FEC3C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="52694BB4E5211C65D3ECD642E06100DF">Wappler, Torsten</mods:namePart>
</mods:name>
<mods:typeOfResource id="9BE962EA070BDA7F84F74E46F62B59A9">text</mods:typeOfResource>
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<mods:title id="058EFFA41B7B5C06CD3650965BA49DEC">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8154D0DCA47B3CC4B4A4D1F702777B64">
<mods:date id="673D8FEF2A238958EAA0057371743A86">2003</mods:date>
<mods:detail id="3E8AD1245205DA9AA60E0AF714DCCC2C" type="volume">
<mods:number id="6EA621BFC5F866BC24B4DEC6896222C3">374</mods:number>
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<mods:classification id="28B829AAEC862AC5F2D08B11300467CB">journal article</mods:classification>
<mods:identifier id="A4F0962624096CB98644A663E2007F3A" type="CLB-Dataset">51238</mods:identifier>
<mods:identifier id="2BEB4AC65BC17E23D4B7C85CEAD6C56F" type="DOI">10.5281/zenodo.156768</mods:identifier>
<mods:identifier id="116B102D90F7BE6A3177C8E6B9E405F3" type="GBIF-Dataset">0ae8bbb8-3209-479e-bd4e-efa5e38555e4</mods:identifier>
<mods:identifier id="B98577DFB1CAF24FF255F16BF7EE7E34" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="CC905621C78AA25CA6DE16B6BF06BFA1" type="Zenodo-Dep">156768</mods:identifier>
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<treatment id="D33887DEFF963B0AFEF101C9E74AD344" ID-DOI="http://doi.org/10.5281/zenodo.6273739" ID-GBIF-Taxon="119332626" ID-Zenodo-Dep="6273739" LSID="urn:lsid:zoobank.org:act:CA015707-CC21-4CB8-AD59-276AFCF278EB" httpUri="http://treatment.plazi.org/id/D33887DEFF963B0AFEF101C9E74AD344" lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12">
<subSubSection id="138B6543FF963B0FFEF101C9E635D2F4" box="[264,785,1764,1790]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="5B2E36C8FF963B0FFEF101C9E635D2F4" blockId="11.[264,785,1764,1790]" box="[264,785,1764,1790]" pageId="11" pageNumber="12">
<heading id="006681A4FF963B0FFEF101C9E635D2F4" bold="true" box="[264,785,1764,1790]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<emphasis id="69E5EADAFF963B0FFEF101C9E635D2F4" bold="true" box="[264,785,1764,1790]" pageId="11" pageNumber="12">
<taxonomicName id="9C914D4BFF963B0FFEF101C9E707D2F4" box="[264,547,1764,1790]" class="Insecta" family="Tingidae" genus="Oblongomorpha" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="lutetia" status="sp. nov.">
<emphasis id="69E5EADAFF963B0FFEF101C9E707D2F4" bold="true" box="[264,547,1764,1790]" italics="true" pageId="11" pageNumber="12">Oblongomorpha lutetia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="72D657A1FF963B0FFDD301C9E748D2F4" box="[554,620,1764,1790]" pageId="11" pageNumber="12" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="C3AA2A4DFF963B0FFD8201C9E7F8D2F4" box="[635,732,1764,1790]" captionStart="FIGURE 11" captionStartId="12.[264,367,1810,1834]" captionTargetBox="[360,1234,313,1795]" captionTargetId="figure@12.[336,1250,294,1806]" captionTargetPageId="12" captionText="FIGURE 11. Oblongomorpha lutetia gen. et sp. n., reconstruction of the holotype, (macropterous form), compression fossil; coll. Forschungsinstitut Senckenberg (SMF), Frankfurt am Main (Germany), specimen SMF MeI 10508; scale bar represents 1 mm." httpUri="https://zenodo.org/record/156773/files/figure.png" pageId="11" pageNumber="12">Figs. 11</figureCitation>
–16)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="138B6543FF963B0FFEF10003E74ED37B" pageId="11" pageNumber="12" type="etymology">
<paragraph id="5B2E36C8FF963B0FFEF10003E74ED37B" blockId="11.[264,1320,1838,1905]" pageId="11" pageNumber="12">
<emphasis id="69E5EADAFF963B0FFEF10003E4D5D342" bold="true" box="[264,497,1838,1864]" pageId="11" pageNumber="12">Derivatio nominis:</emphasis>
The specific epithet is a reference to the Lutetian, the age of the fossil deposit in which it was found.
</paragraph>
</subSubSection>
<caption id="0FEE6640FF913B08FEF1003FE685D360" httpUri="https://zenodo.org/record/156773/files/figure.png" pageId="12" pageNumber="13" targetBox="[360,1234,313,1795]" targetPageId="12">
<paragraph id="5B2E36C8FF913B08FEF1003FE685D360" blockId="12.[264,1322,1810,1898]" pageId="12" pageNumber="13">
<emphasis id="69E5EADAFF913B08FEF1003FE4B3D320" bold="true" box="[264,407,1810,1834]" pageId="12" pageNumber="13">FIGURE 11.</emphasis>
<taxonomicName id="9C914D4BFF913B08FE66003EE7BED320" box="[415,666,1811,1834]" class="Insecta" family="Tingidae" genus="Oblongomorpha" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="lutetia">
<emphasis id="69E5EADAFF913B08FE66003EE7BED320" box="[415,666,1811,1834]" italics="true" pageId="12" pageNumber="13">Oblongomorpha lutetia</emphasis>
</taxonomicName>
<emphasis id="69E5EADAFF913B08FD5B003FE615D320" bold="true" box="[674,817,1810,1834]" pageId="12" pageNumber="13">gen. et sp. n.</emphasis>
, reconstruction of the holotype, (macropterous form), compression fossil; coll. Forschungsinstitut Senckenberg (SMF), Frankfurt am Main (Germany), specimen SMF MeI 10508; scale bar represents 1 mm.
</paragraph>
</caption>
<subSubSection id="138B6543FF903B09FEC10636E650D5DC" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="5B2E36C8FF903B09FEC10636E412D58C" blockId="13.[264,1325,283,470]" pageId="13" pageNumber="14">
<emphasis id="69E5EADAFF903B09FEC10636E497D53F" bold="true" box="[312,435,283,309]" pageId="13" pageNumber="14">
<typeStatus id="842A886AFF903B09FEC10636E48AD53F" box="[312,430,283,309]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
:
</emphasis>
A complete individual, dorsoventrally compressed, coll. Forschungsinstitut Senckenberg (
<collectionCode id="3D80AE0DFF903B09FE490669E4D4D554" LSID="urn:lsid:biocol.org:col:34838" box="[432,496,324,350]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="13" pageNumber="14">SMF</collectionCode>
), Frankfurt am Main (
<collectingCountry id="23867658FF903B09FD0C0669E64DD554" box="[757,873,324,350]" name="Germany" pageId="13" pageNumber="14">Germany</collectingCountry>
), specimen
<collectionCode id="3D80AE0DFF903B09FC0D0669E10BD554" LSID="urn:lsid:biocol.org:col:34838" box="[1012,1071,324,350]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="13" pageNumber="14">SMF</collectionCode>
MeI 10508 (
<figureCitation id="C3AA2A4DFF903B09FB330669E038D554" box="[1226,1308,324,350]" captionStart="FIGURE 11" captionStartId="12.[264,367,1810,1834]" captionTargetBox="[360,1234,313,1795]" captionTargetId="figure@12.[336,1250,294,1806]" captionTargetPageId="12" captionText="FIGURE 11. Oblongomorpha lutetia gen. et sp. n., reconstruction of the holotype, (macropterous form), compression fossil; coll. Forschungsinstitut Senckenberg (SMF), Frankfurt am Main (Germany), specimen SMF MeI 10508; scale bar represents 1 mm." httpUri="https://zenodo.org/record/156773/files/figure.png" pageId="13" pageNumber="14">Fig. 11</figureCitation>
– 12).
</paragraph>
<paragraph id="5B2E36C8FF903B09FEC106BEE650D5DC" blockId="13.[264,1325,283,470]" pageId="13" pageNumber="14">
<emphasis id="69E5EADAFF903B09FEC106BEE49AD5A7" bold="true" box="[312,446,403,429]" pageId="13" pageNumber="14">
<typeStatus id="842A886AFF903B09FEC106BEE49ED5A7" box="[312,442,403,429]" pageId="13" pageNumber="14" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectionCode id="3D80AE0DFF903B09FE3306B9E721D5A4" LSID="urn:lsid:biocol.org:col:34838" box="[458,517,404,430]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="13" pageNumber="14">SMF</collectionCode>
MeI 6210 (dorsoventrally compressed),
<collectionCode id="3D80AE0DFF903B09FC0106B9E117D5A4" LSID="urn:lsid:biocol.org:col:34838" box="[1016,1075,404,430]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="13" pageNumber="14">SMF</collectionCode>
MeI 6616 (laterally compressed),
<collectionCode id="3D80AE0DFF903B09FE520691E4C2D5DC" LSID="urn:lsid:biocol.org:col:34838" box="[427,486,444,470]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="13" pageNumber="14">SMF</collectionCode>
MeI 8766 (laterally compressed).
</paragraph>
</subSubSection>
<subSubSection id="138B6543FF903B0AFEF101B7E1A8D5F4" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="description">
<paragraph id="5B2E36C8FF903B09FEF101B7E77AD331" blockId="13.[264,1323,1690,1851]" pageId="13" pageNumber="14">
<emphasis id="69E5EADAFF903B09FEF101B7E482D2B8" bold="true" box="[264,422,1690,1714]" pageId="13" pageNumber="14">
Figure
<date id="2F2F1008FF903B09FEA501B7E482D2B8" box="[348,422,1690,1714]" pageId="13" pageNumber="14" value="1912-12-16">12–16.</date>
</emphasis>
12, Dorsal habitus of
<typeStatus id="842A886AFF903B09FD5801B7E7DAD2BB" box="[673,766,1690,1713]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
of
<taxonomicName id="9C914D4BFF903B09FCD101B7E102D2BB" box="[808,1062,1690,1713]" class="Insecta" family="Tingidae" genus="Oblongomorpha" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lutetia">
<emphasis id="69E5EADAFF903B09FCD101B7E102D2BB" box="[808,1062,1690,1713]" italics="true" pageId="13" pageNumber="14">Oblongomorpha lutetia</emphasis>
</taxonomicName>
(SMF MeI 10508); 13, Lateral habitus of
<typeStatus id="842A886AFF903B09FE300191E701D2D9" box="[457,549,1724,1747]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
(SMF MeI 8766); 14, Detail of head and basal antennae structures (SMF MeI 6210) (black arrow indicates elongate antennal segment I); 15, Lateral habitus of
<typeStatus id="842A886AFF903B09FB3601F2E00FD2FC" box="[1231,1323,1759,1782]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
(SMF MeI 6616); 16, Tarsal structure of
<taxonomicName id="9C914D4BFF903B09FD3D002FE60DD313" box="[708,809,1794,1817]" class="Insecta" family="Tingidae" genus="Oblongomorpha" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lutetia">
<emphasis id="69E5EADAFF903B09FD3D002FE60DD313" box="[708,809,1794,1817]" italics="true" pageId="13" pageNumber="14">
<collectingCountry id="23867658FF903B09FD3D002FE7FED313" box="[708,730,1794,1817]" name="Iceland" pageId="13" pageNumber="14">O</collectingCountry>
. lutetia
</emphasis>
</taxonomicName>
detailing bristle­like setae on the lower surface (white arrow) (SMF MeI 6616).
</paragraph>
<paragraph id="5B2E36C8FF933B0AFEC10636E1A8D5F4" blockId="14.[264,1324,283,1870]" pageId="14" pageNumber="15">
<emphasis id="69E5EADAFF933B0AFEC10636E761D53F" bold="true" box="[312,581,283,309]" pageId="14" pageNumber="15">Dimensions (in mm):</emphasis>
Body length 5.03, width 1.25; length of head (from apex of clypeus to posterior margin of pronotum) 0.59, width 0.66; interocular width 0.13; antennomeres 1–4: 0.59, 0.26, 2.36, 1.66; length of pronotum 1.83, width 0.96; procoxa 0.19 (SMF MeI 6616); length of fore leg (SMF MeI 6616): trochanter 0.16, femur 1.13, tibia 1.43; tarsus 0.23; length of middle leg (SMF MeI 6616): femur 1.23, tibia 1.13, tarsus 0.26; length of hind leg (SMF MeI 6616): femur 1.13, tibia 1.23, tarsus 0.26.
</paragraph>
</subSubSection>
<subSubSection id="138B6543FF933B0AFEC10526E4BDD694" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="5B2E36C8FF933B0AFEC10526E132D62C" blockId="14.[264,1324,283,1870]" box="[312,1046,523,550]" pageId="14" pageNumber="15">
<emphasis id="69E5EADAFF933B0AFEC10526E4C5D62F" bold="true" box="[312,481,523,549]" pageId="14" pageNumber="15">
<typeStatus id="842A886AFF933B0AFEC10526E451D62F" box="[312,373,523,549]" pageId="14" pageNumber="15">Type</typeStatus>
locality:
</emphasis>
Grube Messel near Darmstadt, grid square E89.
</paragraph>
<paragraph id="5B2E36C8FF933B0AFEC1051EE4BDD694" blockId="14.[264,1324,283,1870]" pageId="14" pageNumber="15">
<emphasis id="69E5EADAFF933B0AFEC1051EE65ED647" bold="true" box="[312,890,563,589]" pageId="14" pageNumber="15">Stratigraphical and geographical distribution:</emphasis>
Known only from the lower Middle Eocene (Lower Lutetian,
<collectionCode id="3D80AE0DFF933B0AFDB80571E7B0D67C" box="[577,660,604,630]" pageId="14" pageNumber="15">ELMA</collectionCode>
Geiseltalian, MP11), Messel Formation; 1.25–
<quantity id="9C699B2DFF933B0AFB280571E00DD67C" box="[1233,1321,604,630]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="15" unit="m" value="0.1">0.10 m</quantity>
under alpha.
</paragraph>
</subSubSection>
<subSubSection id="138B6543FF933B0AFEC10581E632D6CC" box="[312,790,684,710]" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="5B2E36C8FF933B0AFEC10581E632D6CC" blockId="14.[264,1324,283,1870]" box="[312,790,684,710]" pageId="14" pageNumber="15">
<emphasis id="69E5EADAFF933B0AFEC10581E49ED6CC" bold="true" box="[312,442,684,710]" pageId="14" pageNumber="15">Diagnosis:</emphasis>
<collectingCountry id="23867658FF933B0AFE380581E4C5D6CC" box="[449,481,684,710]" name="American Samoa" pageId="14" pageNumber="15">As</collectingCountry>
for the genus (see above).
</paragraph>
</subSubSection>
<subSubSection id="138B6543FF933B0AFEC105F9E74AD344" pageId="14" pageNumber="15" type="description">
<paragraph id="5B2E36C8FF933B0AFEC105F9E6F6D71C" blockId="14.[264,1324,283,1870]" pageId="14" pageNumber="15">
<emphasis id="69E5EADAFF933B0AFEC105F9E4F6D6E4" bold="true" box="[312,466,724,750]" pageId="14" pageNumber="15">Description:</emphasis>
Body surface areolate. Body oblong and slender, 4 times longer than wide, dorsal side of the body, legs, and antennae pale brown.
</paragraph>
<paragraph id="5B2E36C8FF933B0AFEC10409E470D00C" blockId="14.[264,1324,283,1870]" pageId="14" pageNumber="15">Head very short, protruding little in front of the antennal insertion, sharply declivent. Head without cephalic spines or tubercles. Vertex between eyes slightly elevated. Eyes globular, strongly projecting laterally of the head. Antennae long and thin, first antennal segment 3 to 4 times as long as second, 3rd prolonged, 4th clubbed and covered with dense setae. Labium short and not reaching the middle of the metasternum (e.g. SMF MeI 6210, 8766).</paragraph>
<paragraph id="5B2E36C8FF933B0AFEC10339E17DD114" blockId="14.[264,1324,283,1870]" pageId="14" pageNumber="15">Pronotum 1.9 times longer as wide, areolate, with a ring­like vesicular at the anterior margin, broad (tectiform) in longitudinal direction, bearing two transverse rows of cells. Paranota absent. Triangular areolate posterior pronotal process is well developed, extremely elongated backwards along hemelytra covering whole clavi and approximately half of discoidal area. Pronotal disc perceptible convex (cyst absent!), with 3 longitudinal carinae. Median carina extending from calli to posterior margin. Lateral carinae parallel to median but only half as long. Anterior pronotal margin slightly concave.</paragraph>
<paragraph id="5B2E36C8FF933B0AFEC10201E4CED2DC" blockId="14.[264,1324,283,1870]" pageId="14" pageNumber="15">Hemelytra (macropterous form) very long and extended much beyond abdominal apex, with distinctly carinated veins, separating the hemelytra in costal, subcostal, discoidal, and sutural areas. Outline of hemelytra distinctly sinuate about midlength, apical rounded. Costal area very narrow, for most of its length with two rows of small cells. Subcostal area also very narrow, for most of its length with two rows of small cells, in the apical third only with a single row of nearly rectangular cells. Discoidal area triangular in shape, occupying nearly 0.54 of hemelytron length, in the widest part with 4–5 rows of small cells. Clavi entirely concealed by the posterior pronotal process. Sutural area (membrane) in broadest place with 11–12 of angled cells, size of cells slightly increasing from the anterior margin to the apex of hemelytra. Tips of sutural areas of both hemelytra fully overlapping at rest.</paragraph>
<paragraph id="5B2E36C8FF933B0AFEC101C9E74AD344" blockId="14.[264,1324,283,1870]" pageId="14" pageNumber="15">Legs moderately long. Tarsi 2­segmented, with second segment definitely broader and longer than segment I. Upper surface of segment II slightly convex, lower surface flat and covered with bristle­like hairs.</paragraph>
</subSubSection>
</treatment>
</document>
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