<documentID-CLB-Dataset="24202"ID-DOI="http://dx.doi.org/10.3897/zse.96.48312"ID-GBIF-Dataset="7de3ea60-ba35-4831-ae1d-048dc362f8b0"ID-Pensoft-Pub="1860-0743-1-25"ID-Pensoft-UUID="4EDA9B5B777551B7BBB0152DD188E5EF"ID-ZooBank="F2C8585A1268443693348B64AE20F6EE"ModsDocID="1860-0743-1-25"checkinTime="1580744411276"checkinUser="pensoft"docAuthor="Sands, Arthur F, Gloeer, Peter, Guerlek, Mustafa E, Albrecht, Christian & Neubauer, Thomas A"docDate="2020"docId="A185AFE24AA551278669452B0925B575"docLanguage="en"docName="ZoosystEvol 96(1): 25-66"docOrigin="Zoosystematics and Evolution 96 (1)"docSource="http://dx.doi.org/10.3897/zse.96.48312"docTitle="Theodoxus jordani"docType="treatment"docVersion="8"id="4EDA9B5B777551B7BBB0152DD188E5EF"lastPageNumber="25"masterDocId="4EDA9B5B777551B7BBB0152DD188E5EF"masterDocTitle="A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species"masterLastPageNumber="66"masterPageNumber="25"pageNumber="25"updateTime="1732731517510"updateUser="ExternalLinkService">
<mods:titleid="6A015BA6BAD0330C0A608538EC423AB1">A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species</mods:title>
<mods:namePartid="999DE2D7C85933DFA10345064C46B8FD">Neubauer, Thomas A</mods:namePart>
<mods:affiliationid="ED13E11AD419EA7334F7251BE4236DFE">Justus Liebig University Giessen, Giessen, Germany & Naturalis Biodiversity Centre, Leiden, Netherlands</mods:affiliation>
<taxonomicNameid="D232ABCA9C386999886BFB574D325A7E"ID-CoL="7TCK2"LSID="A185AFE2-4AA5-5127-8669-452B0925B575"authority="(G. B. Sowerby I, 1836)"baseAuthorityName="G. B. Sowerby I"baseAuthorityYear="1836"class="Gastropoda"family="Neritidae"genus="Theodoxus"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Theodoxus jordani"order="Cycloneritida"pageId="0"pageNumber="25"phylum="Mollusca"rank="species"species="jordani">Theodoxus jordani (G.B. Sowerby I, 1836)</taxonomicName>
<figureCitationid="2E464572D12E2985AB328661BCE3A2B4"captionStart="Figure 15"captionStartId="F15"captionText="Figure 15. Theodoxus jordani (G. B. Sowerby I, 1836). A - D. Specimen from the Avakas Gorge, Peyia, Cyprus (UGSB 18321) incorporated into the phylogeny (Fig. 2); E - H. Specimen collected in the Ceyhan River, Eski Misis, Adana, Turkey (UGSB 20777); I - L. Specimen from the Orontes River, Antakya, Hatay, Turkey (UGSB 24175) used in the phylogeny (Fig. 2); M - O. Specimen (T. euphraticus - morphotype) from the Seyed Hosein Park spring, near the Bishapur, Fars, Iran (UGSB 22233); P - R. Topotype of N. jordani from the River Jordan, Israel (UGSB 24177) incorporated into the phylogeny (Fig. 2); S, T. Specimen (T. euphraticus - morphotype) from the Karun River, Ahvaz, Iran (UGSB 21681); U. Specimen (T. niloticus - morphotype) collected in the Nile River, Egypt (UGSB 24178) used in the phylogeny (Fig. 2); V. Specimen (T. euphraticus - morphotype) from the Karun River, Ahvaz, Iran (UGSB 21682). Scale bars: 1 mm."figureDoi="10.3897/zse.96.48312.figure15"httpUri="https://binary.pensoft.net/fig/376464"pageId="0"pageNumber="25">Figures 15A-V</figureCitation>
<figureCitationid="4A4F5877C88FB52C85CA30163737E5D0"captionStart="Figure 16"captionStartId="F16"captionText="Figure 16. Theodoxus jordani (G. B. Sowerby I, 1836). A, B. Syntype of N. euphratica from the Euphrates River at Samawah, Iraq (ZMZ 528916); C - E. Syntype of N. michonii from Syria (MHNG-MOLL- 111706); F - H. Syntype of N. karasuna collected from the Karasu River, south-eastern Turkey (ZMZ 528937); I, J; K, L. Two syntypes of N. ponsoti from Lake Muzayrīb, Syria (MNHN-IM- 2000 - 32754); M. Syntype (labelled as paratype) of N. ponsoti from Lake Muzayrīb (MT 820, RBINS); N. Syntype of N. jordani var. turris collected in the Sea of Galilee (ZMZ 528930); O, P. Syntype of N. bellardii from the Litani River, Beqaa valley, Lebanon (ZMZ 528918). Scale bars: 1 mm."figureDoi="10.3897/zse.96.48312.figure16"httpUri="https://binary.pensoft.net/fig/376465"pageId="0"pageNumber="25">, 16A-P</figureCitation>
<figureCitationid="144D0E39BCB6CFDCB0DAD45EF7E92A4D"captionStart="Figure 17"captionStartId="F17"captionText="Figure 17. Theodoxus jordani (G. B. Sowerby I, 1836). A - C. Syntype of N. meridionalis var. mesopotamica from near Diyarbakir, Turkey (ZMZ 528914); D, E. Specimen (T. mesopotamicus - morphotype) collected in Sanliurfa, Turkey (UGSB 23426) and used in the phylogeny (Fig. 2); F - H; K - M. Two syntypes of N. cinctella collected at the source of River Chabur near Ras al-Ayn, Syria (ZMB / Moll 21732); I, J. Topotype of N. cinctella from the same locality (UGSB 18762). Scale bars: 1 mm."figureDoi="10.3897/zse.96.48312.figure17"httpUri="https://binary.pensoft.net/fig/376466"pageId="0"pageNumber="25">, 17A-J</figureCitation>
<bibRefCitationid="508B436DB51A82C630C0ED880BFB7779"author="Bourguignat, JR"journalOrPublisher="Baudry, Paris"pageId="0"pageNumber="25"publicationUrl="http://gallica.bnf.fr/ark:/12148/bpt6k99038p/f1.item"refId="B31"refString="Bourguignat, JR, 1853. Catalogue raisonne des mollusques terrestres et fluviatiles recueillis par M. F. de Saulcy pendant son voyage en Orient. Gide et J. Baudry, Paris, http://gallica.bnf.fr/ark:/12148/bpt6k99038p/f1.item"title="Catalogue raisonne des mollusques terrestres et fluviatiles recueillis par M. F. de Saulcy pendant son voyage en Orient. Gide et J."url="http://gallica.bnf.fr/ark:/12148/bpt6k99038p/f1.item"year="1853">Bourguignat 1853</bibRefCitation>
<bibRefCitationid="EF779AA644E6120B5FD40D00F219C69B"author="Martens, Ev"journalOrPublisher="Bauer & Raspe, Nuernberg"pageId="0"pageNumber="25"publicationUrl="https://biodiversitylibrary.org/page/52484536"refId="B99"refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer & Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536"title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)."url="https://biodiversitylibrary.org/page/52484536"year="1879">Martens 1879</bibRefCitation>
<bibRefCitationid="642007A58510FF9AB05BFA05D66FF965"author="Kobelt, W"journalOrPublisher="Aquila"pageId="0"pageNumber="25"publicationUrl="https://biodiversitylibrary.org/page/16299729"refId="B83"refString="Kobelt, W, 1899. Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1-11. [pls 211-240] https://biodiversitylibrary.org/page/16299729"title="Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1 - 11. [pls 211 - 240]"url="https://biodiversitylibrary.org/page/16299729"year="1899">Kobelt 1899</bibRefCitation>
<bibRefCitationid="7360D310A027210972F06F412A693759"author="Bourguignat, JR"journalOrPublisher="Bailliere, Lutetia"pageId="0"pageNumber="25"publicationUrl="https://gallica.bnf.fr/ark:/12148/bpt6k99037b.image"refId="B30"refString="Bourguignat, JR, 1852. Testacea novissima quae Cl. de Saulcy in itinere per Orientem annis 1850 et 1851, collegit. J.-B. Bailliere, Lutetia, https://gallica.bnf.fr/ark:/12148/bpt6k99037b.image"title="Testacea novissima quae Cl. de Saulcy in itinere per Orientem annis 1850 et 1851, collegit. J. - B."url="https://gallica.bnf.fr/ark:/12148/bpt6k99037b.image"year="1852">Bourguignat 1852</bibRefCitation>
<bibRefCitationid="E8BC32AF5A68BA32799F13504F642AC9"author="Bourguignat, JR"journalOrPublisher="Baudry, Paris"pageId="0"pageNumber="25"publicationUrl="http://gallica.bnf.fr/ark:/12148/bpt6k99038p/f1.item"refId="B31"refString="Bourguignat, JR, 1853. Catalogue raisonne des mollusques terrestres et fluviatiles recueillis par M. F. de Saulcy pendant son voyage en Orient. Gide et J. Baudry, Paris, http://gallica.bnf.fr/ark:/12148/bpt6k99038p/f1.item"title="Catalogue raisonne des mollusques terrestres et fluviatiles recueillis par M. F. de Saulcy pendant son voyage en Orient. Gide et J."url="http://gallica.bnf.fr/ark:/12148/bpt6k99038p/f1.item"year="1853">Bourguignat 1853</bibRefCitation>
<bibRefCitationid="8A8A43F354E7392428CEDD0222927E56"author="Mousson, A"journalOrPublisher="Zurcher et Furrer, Zuerich"pageId="0"pageNumber="25"publicationUrl="https://biodiversitylibrary.org/page/47588377"refId="B109"refString="Mousson, A, 1854. Coquilles terrestres et fluviatiles recueillies par M. le prof Bellardi dans un voyage en Orient. Zurcher et Furrer, Zuerich, https://biodiversitylibrary.org/page/47588377"title="Coquilles terrestres et fluviatiles recueillies par M. le prof Bellardi dans un voyage en Orient."url="https://biodiversitylibrary.org/page/47588377"year="1854">Mousson 1854</bibRefCitation>
<bibRefCitationid="ABFBDB29705E5B249B1E80485DEAE815"author="Reeve, LA"journalOrPublisher="The Annals and Magazine of Natural History"pageId="0"pageNumber="25"publicationUrl="https://biodiversitylibrary.org/page/9141496"refId="B134"refString="Reeve, LA, 1856. Monograph of the genus Neritina. In: Reeve LA (Ed.) Conchologia iconica: or, Illustrations of the shells of molluscous animals. Vol. 4. Privately published, London. [1-37 pls. and unpaginated index] https://biodiversitylibrary.org/page/9141496"title="Monograph of the genus Neritina. In: Reeve LA (Ed.) Conchologia iconica: or, Illustrations of the shells of molluscous animals. Vol. 4. Privately published, London. [1 - 37 pls. and unpaginated index]"url="https://biodiversitylibrary.org/page/9141496"year="1856">Reeve 1856</bibRefCitation>
<bibRefCitationid="E62119CA1D9E7C7819141104925B2D28"author="Reeve, LA"journalOrPublisher="The Annals and Magazine of Natural History"pageId="0"pageNumber="25"publicationUrl="https://biodiversitylibrary.org/page/9141496"refId="B134"refString="Reeve, LA, 1856. Monograph of the genus Neritina. In: Reeve LA (Ed.) Conchologia iconica: or, Illustrations of the shells of molluscous animals. Vol. 4. Privately published, London. [1-37 pls. and unpaginated index] https://biodiversitylibrary.org/page/9141496"title="Monograph of the genus Neritina. In: Reeve LA (Ed.) Conchologia iconica: or, Illustrations of the shells of molluscous animals. Vol. 4. Privately published, London. [1 - 37 pls. and unpaginated index]"url="https://biodiversitylibrary.org/page/9141496"year="1856">Reeve 1856</bibRefCitation>
<bibRefCitationid="FA045FFF7CC12E025174139F29E2266A"author="Martens, Ev"journalOrPublisher="Bauer & Raspe, Nuernberg"pageId="0"pageNumber="25"publicationUrl="https://biodiversitylibrary.org/page/52484536"refId="B99"refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer & Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536"title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)."url="https://biodiversitylibrary.org/page/52484536"year="1879">Martens 1879</bibRefCitation>
<bibRefCitationid="7D68949D5C79EEE78E9032B0B61CCB1B"author="Kobelt, W"journalOrPublisher="Aquila"pageId="0"pageNumber="25"publicationUrl="https://biodiversitylibrary.org/page/16299729"refId="B83"refString="Kobelt, W, 1899. Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1-11. [pls 211-240] https://biodiversitylibrary.org/page/16299729"title="Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1 - 11. [pls 211 - 240]"url="https://biodiversitylibrary.org/page/16299729"year="1899">Kobelt 1899</bibRefCitation>
<bibRefCitationid="ACCB54CF64D5D24B871D3CD2D53A8342"author="Mousson, A"journalOrPublisher="Zurcher et Furrer, Zuerich"pageId="0"pageNumber="25"publicationUrl="https://biodiversitylibrary.org/page/8742661"refId="B110"refString="Mousson, A, 1861. Coquilles terrestres et fluviatiles recueillies par M. le Prof. J. R. Roth dans son dernier voyage en Orient. Vierteljahrsschrift der Naturforschenden Gesellschaft in Zuerich 6: 1-34, 124-156. https://biodiversitylibrary.org/page/8742661"title="Coquilles terrestres et fluviatiles recueillies par M. le Prof. J. R. Roth dans son dernier voyage en Orient. Vierteljahrsschrift der Naturforschenden Gesellschaft in Zuerich 6: 1 - 34, 124 - 156."url="https://biodiversitylibrary.org/page/8742661"year="1861">Mousson 1861</bibRefCitation>
<bibRefCitationid="212B81EA7C0BCE64316E7BE0CAB34176"author="Kobelt, W"journalOrPublisher="Aquila"pageId="0"pageNumber="25"publicationUrl="https://biodiversitylibrary.org/page/16299729"refId="B83"refString="Kobelt, W, 1899. Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1-11. [pls 211-240] https://biodiversitylibrary.org/page/16299729"title="Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1 - 11. [pls 211 - 240]"url="https://biodiversitylibrary.org/page/16299729"year="1899">Kobelt 1899</bibRefCitation>
<bibRefCitationid="1CFBFDF67C2E2217DF3910F6FD3BE81E"author="Mousson, A"journalOrPublisher="Journal de Conchyliologie"pageId="0"pageNumber="25"pagination="5 - 60"publicationUrl="https://biodiversitylibrary.org/page/15661760"refId="B111"refString="Mousson, A, 1874. Coquilles terrestres et fluviatiles recueillies par M. le Dr Alex. Schlaefli en Orient. Journal de Conchyliologie 22 (1): 5 - 60, https://biodiversitylibrary.org/page/15661760"title="Coquilles terrestres et fluviatiles recueillies par M. le Dr Alex. Schlaefli en Orient."url="https://biodiversitylibrary.org/page/15661760"volume="22"year="1874">Mousson 1874</bibRefCitation>
<bibRefCitationid="8E72DCB8A8AB6CED43C1876CDC4F50E9"author="Blanckenhorn, M"journalOrPublisher="Palaeontographica"pageId="0"pageNumber="25"pagination="71 - 144"publicationUrl="https://biodiversitylibrary.org/page/33130542"refId="B25"refString="Blanckenhorn, M, 1897. Zur Kenntnis der Suesswasserablagerungen und Mollusken Syriens. Palaeontographica 44: 71 - 144, https://biodiversitylibrary.org/page/33130542"title="Zur Kenntnis der Suesswasserablagerungen und Mollusken Syriens."url="https://biodiversitylibrary.org/page/33130542"volume="44"year="1897">Blanckenhorn 1897</bibRefCitation>
<bibRefCitationid="103808F8169DE1A80092F9B912A2AA96"author="Mousson, A"journalOrPublisher="Journal de Conchyliologie"pageId="0"pageNumber="25"pagination="5 - 60"publicationUrl="https://biodiversitylibrary.org/page/15661760"refId="B111"refString="Mousson, A, 1874. Coquilles terrestres et fluviatiles recueillies par M. le Dr Alex. Schlaefli en Orient. Journal de Conchyliologie 22 (1): 5 - 60, https://biodiversitylibrary.org/page/15661760"title="Coquilles terrestres et fluviatiles recueillies par M. le Dr Alex. Schlaefli en Orient."url="https://biodiversitylibrary.org/page/15661760"volume="22"year="1874">Mousson 1874</bibRefCitation>
<bibRefCitationid="041D8B78CDA08B6BF7F536B626D57A5E"author="Mousson, A"journalOrPublisher="Journal de Conchyliologie"pageId="0"pageNumber="25"pagination="5 - 60"publicationUrl="https://biodiversitylibrary.org/page/15661760"refId="B111"refString="Mousson, A, 1874. Coquilles terrestres et fluviatiles recueillies par M. le Dr Alex. Schlaefli en Orient. Journal de Conchyliologie 22 (1): 5 - 60, https://biodiversitylibrary.org/page/15661760"title="Coquilles terrestres et fluviatiles recueillies par M. le Dr Alex. Schlaefli en Orient."url="https://biodiversitylibrary.org/page/15661760"volume="22"year="1874">Mousson 1874</bibRefCitation>
<bibRefCitationid="8BCF0FDA476D398F7E5CCBBFC2301CCE"author="Kobelt, W"journalOrPublisher="Aquila"pageId="0"pageNumber="25"publicationUrl="https://biodiversitylibrary.org/page/16299729"refId="B83"refString="Kobelt, W, 1899. Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1-11. [pls 211-240] https://biodiversitylibrary.org/page/16299729"title="Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1 - 11. [pls 211 - 240]"url="https://biodiversitylibrary.org/page/16299729"year="1899">Kobelt 1899</bibRefCitation>
<bibRefCitationid="8D9B88B8428067B9670347C348AFA82C"author="Martens, Ev"journalOrPublisher="Novitates Conchologicae, Supplement"pageId="0"pageNumber="25"pagination="1 - 127"publicationUrl="https://biodiversitylibrary.org/page/12980992"refId="B98"refString="Martens, Ev, 1874. Ueber vorderasiatische Conchylien nach den Sammlungen des Prof. Hausknecht. Novitates Conchologicae, Supplement 5: 1 - 127, https://biodiversitylibrary.org/page/12980992"title="Ueber vorderasiatische Conchylien nach den Sammlungen des Prof. Hausknecht."url="https://biodiversitylibrary.org/page/12980992"volume="5"year="1874">Martens 1874</bibRefCitation>
<bibRefCitationid="FFE18DC6AD0B7E69A52677984037CF73"author="Martens, Ev"journalOrPublisher="Novitates Conchologicae, Supplement"pageId="0"pageNumber="25"pagination="1 - 127"publicationUrl="https://biodiversitylibrary.org/page/12980992"refId="B98"refString="Martens, Ev, 1874. Ueber vorderasiatische Conchylien nach den Sammlungen des Prof. Hausknecht. Novitates Conchologicae, Supplement 5: 1 - 127, https://biodiversitylibrary.org/page/12980992"title="Ueber vorderasiatische Conchylien nach den Sammlungen des Prof. Hausknecht."url="https://biodiversitylibrary.org/page/12980992"volume="5"year="1874">Martens 1874</bibRefCitation>
<bibRefCitationid="33BE073DADDEDFD27F3B32923D0D694E"author="Martens, Ev"journalOrPublisher="Bauer & Raspe, Nuernberg"pageId="0"pageNumber="25"publicationUrl="https://biodiversitylibrary.org/page/52484536"refId="B99"refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer & Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536"title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)."url="https://biodiversitylibrary.org/page/52484536"year="1879">Martens 1879</bibRefCitation>
<bibRefCitationid="3C08A7B4B0E395D603D98D0F8C605FE0"author="Kobelt, W"journalOrPublisher="Aquila"pageId="0"pageNumber="25"publicationUrl="https://biodiversitylibrary.org/page/16299729"refId="B83"refString="Kobelt, W, 1899. Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1-11. [pls 211-240] https://biodiversitylibrary.org/page/16299729"title="Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1 - 11. [pls 211 - 240]"url="https://biodiversitylibrary.org/page/16299729"year="1899">Kobelt 1899</bibRefCitation>
<bibRefCitationid="70A281DF15EE1FE5A704C2DA7C201014"author="Martens, Ev"journalOrPublisher="Bauer & Raspe, Nuernberg"pageId="0"pageNumber="25"publicationUrl="https://biodiversitylibrary.org/page/52484536"refId="B99"refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer & Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536"title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)."url="https://biodiversitylibrary.org/page/52484536"year="1879">Martens 1879</bibRefCitation>
<bibRefCitationid="4F6692A22E5D7A56456531F08CEB9F53"author="Martens, Ev"journalOrPublisher="Bauer & Raspe, Nuernberg"pageId="0"pageNumber="25"publicationUrl="https://biodiversitylibrary.org/page/52484536"refId="B99"refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer & Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536"title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)."url="https://biodiversitylibrary.org/page/52484536"year="1879">Martens 1879</bibRefCitation>
<bibRefCitationid="6121034C22DD883DD026BCE56AFE5EFA"author="Kobelt, W"journalOrPublisher="Aquila"pageId="0"pageNumber="25"publicationUrl="https://biodiversitylibrary.org/page/16299729"refId="B83"refString="Kobelt, W, 1899. Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1-11. [pls 211-240] https://biodiversitylibrary.org/page/16299729"title="Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1 - 11. [pls 211 - 240]"url="https://biodiversitylibrary.org/page/16299729"year="1899">Kobelt 1899</bibRefCitation>
<bibRefCitationid="5ADBEBDC0D9FDB7ECA8125CF69C0D447"author="Dautzenberg, P"journalOrPublisher="Revue biologique du nord de la France"pageId="0"pageNumber="25"pagination="329 - 353"publicationUrl="https://biodiversitylibrary.org/page/10877573"refId="B42"refString="Dautzenberg, P, 1894. Liste des mollusques terrestres et fluviatiles recueillis par M. Th. Barrois en Palestine et en Syrie. Revue biologique du nord de la France 6: 329 - 353, https://biodiversitylibrary.org/page/10877573"title="Liste des mollusques terrestres et fluviatiles recueillis par M. Th. Barrois en Palestine et en Syrie."url="https://biodiversitylibrary.org/page/10877573"volume="6"year="1894">Dautzenberg 1894</bibRefCitation>
<bibRefCitationid="398C29F9B2525DF308C876563082BC71"author="Blanckenhorn, M"journalOrPublisher="Palaeontographica"pageId="0"pageNumber="25"pagination="71 - 144"publicationUrl="https://biodiversitylibrary.org/page/33130542"refId="B25"refString="Blanckenhorn, M, 1897. Zur Kenntnis der Suesswasserablagerungen und Mollusken Syriens. Palaeontographica 44: 71 - 144, https://biodiversitylibrary.org/page/33130542"title="Zur Kenntnis der Suesswasserablagerungen und Mollusken Syriens."url="https://biodiversitylibrary.org/page/33130542"volume="44"year="1897">Blanckenhorn 1897</bibRefCitation>
<bibRefCitationid="37DF231F42EAFBAC8CD0F275D08C9B3A"author="Germain, L"journalOrPublisher="Bailliere et fils, Paris"pageId="0"pageNumber="25"publicationUrl="https://biodiversitylibrary.org/page/13187112"refId="B59"refString="Germain, L, 1921. Mollusques terrestres et fluviatiles de Syrie. Tome premier - Introduction et Gasteropodes. J.-B. Bailliere et fils, Paris, https://biodiversitylibrary.org/page/13187112"title="Mollusques terrestres et fluviatiles de Syrie. Tome premier - Introduction et Gasteropodes. J. - B."url="https://biodiversitylibrary.org/page/13187112"year="1921">Germain 1921</bibRefCitation>
<bibRefCitationid="0945816939C595BFB55287EF31E6B96C"author="Germain, L"journalOrPublisher="Bailliere et fils, Paris"pageId="0"pageNumber="25"publicationUrl="https://biodiversitylibrary.org/page/13187112"refId="B59"refString="Germain, L, 1921. Mollusques terrestres et fluviatiles de Syrie. Tome premier - Introduction et Gasteropodes. J.-B. Bailliere et fils, Paris, https://biodiversitylibrary.org/page/13187112"title="Mollusques terrestres et fluviatiles de Syrie. Tome premier - Introduction et Gasteropodes. J. - B."url="https://biodiversitylibrary.org/page/13187112"year="1921">Germain 1921</bibRefCitation>
<bibRefCitationid="6631CAB2418301EEF9EEC29F56E806D1"author="Pallary, P"journalOrPublisher="Bulletin du Museum National d'Histoire Naturelle, deuxieme serie"pageId="0"pageNumber="25"pagination="286 - 290"publicationUrl="http://gallica.bnf.fr/ark:/12148/bpt6k5532113m/f44.image.r"refId="B117"refString="Pallary, P, 1930. Mollusques aquatiques nouveaux du Levant. Bulletin du Museum National d'Histoire Naturelle, deuxieme serie 2 (3): 286 - 290, http://gallica.bnf.fr/ark:/12148/bpt6k5532113m/f44.image.r"title="Mollusques aquatiques nouveaux du Levant."url="http://gallica.bnf.fr/ark:/12148/bpt6k5532113m/f44.image.r"volume="2"year="1930">Pallary 1930</bibRefCitation>
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<taxonomicNameid="8FD34A91943B7B463D546841BCAFAEC5"class="Gastropoda"family="Neritidae"genus="Theodoxus"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Theodoxus jordani var. bicarinatus"order="Cycloneritida"pageId="0"pageNumber="25"phylum="Mollusca"rank="variety"species="jordani"variety="bicarinatus">Theodoxus (Neritaea) jordani var. bicarinatus</taxonomicName>
<bibRefCitationid="D9D56DD7E42C02CE6CD4CA70E3E5FB89"author="Pallary, P"journalOrPublisher="Memoires de l'Institut d'Egypte"pageId="0"pageNumber="25"pagination="1 - 141"refId="B118"refString="Pallary, P, 1939. Deuxieme addition a la faune malacologique de la Syrie. Memoires de l'Institut d'Egypte 39: 1 - 141"title="Deuxieme addition a la faune malacologique de la Syrie."volume="39"year="1939">Pallary 1939</bibRefCitation>
<bibRefCitationid="DE3A69A0857C82AF244857437CFEBEBF"author="Pallary, P"journalOrPublisher="Memoires de l'Institut d'Egypte"pageId="0"pageNumber="25"pagination="1 - 141"refId="B118"refString="Pallary, P, 1939. Deuxieme addition a la faune malacologique de la Syrie. Memoires de l'Institut d'Egypte 39: 1 - 141"title="Deuxieme addition a la faune malacologique de la Syrie."volume="39"year="1939">Pallary 1939</bibRefCitation>
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<bibRefCitationid="715FB8532867E7821C2ABA7FD569E817"author="Pallary, P"journalOrPublisher="Memoires de l'Institut d'Egypte"pageId="0"pageNumber="25"pagination="1 - 141"refId="B118"refString="Pallary, P, 1939. Deuxieme addition a la faune malacologique de la Syrie. Memoires de l'Institut d'Egypte 39: 1 - 141"title="Deuxieme addition a la faune malacologique de la Syrie."volume="39"year="1939">Pallary 1939</bibRefCitation>
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<bibRefCitationid="91DB18C97171337FED41B7BA953C8E25"author="Pallary, P"journalOrPublisher="Memoires de l'Institut d'Egypte"pageId="0"pageNumber="25"pagination="1 - 141"refId="B118"refString="Pallary, P, 1939. Deuxieme addition a la faune malacologique de la Syrie. Memoires de l'Institut d'Egypte 39: 1 - 141"title="Deuxieme addition a la faune malacologique de la Syrie."volume="39"year="1939">Pallary 1939</bibRefCitation>
<bibRefCitationid="183CCA78153EF02B5877018DC1029577"author="Pallary, P"journalOrPublisher="Memoires de l'Institut d'Egypte"pageId="0"pageNumber="25"pagination="1 - 141"refId="B118"refString="Pallary, P, 1939. Deuxieme addition a la faune malacologique de la Syrie. Memoires de l'Institut d'Egypte 39: 1 - 141"title="Deuxieme addition a la faune malacologique de la Syrie."volume="39"year="1939">Pallary 1939</bibRefCitation>
<bibRefCitationid="D13AFBEA23B67C8BF7E9B42D51568B9F"author="Tchernov, E"journalOrPublisher="Malacologia"pageId="0"pageNumber="25"pagination="147 - 184"publicationUrl="https://biodiversitylibrary.org/page/13122806"refId="B154"refString="Tchernov, E, 1975. The molluscs of the Sea of Galilee. Malacologia 15 (1): 147 - 184, https://biodiversitylibrary.org/page/13122806"title="The molluscs of the Sea of Galilee."url="https://biodiversitylibrary.org/page/13122806"volume="15"year="1975">Tchernov 1975</bibRefCitation>
<bibRefCitationid="B5D9E48EDF7436E587C5931717A55B70"DOI="https://doi.org/10.1080/09397140.1993.10637638"author="Schuett, H"journalOrPublisher="Zoology in the Middle East"pageId="0"pageNumber="25"pagination="69 - 112"refId="B143"refString="Schuett, H, Ortal, R, 1993. A preliminary correlation between the Plio-Pleistocene malacofaunas of the Jordan Valley (Israel) and the Orontes Valley (Syria). Zoology in the Middle East 8 (1): 69 - 112, DOI: https://doi.org/10.1080/09397140.1993.10637638"title="A preliminary correlation between the Plio-Pleistocene malacofaunas of the Jordan Valley (Israel) and the Orontes Valley (Syria)."url="https://doi.org/10.1080/09397140.1993.10637638"volume="8"year="1993">
<bibRefCitationid="482EBF550E703285FC761ABBC3A8DA99"author="Bandel, K"journalOrPublisher="Mitteilungen aus dem geologisch-palaeontologischen Institut der Universitaet Hamburg"pageId="0"pageNumber="25"pagination="65 - 164"refId="B20"refString="Bandel, K, 2001. The history of Theodoxus and Neritina connected with description and systematic evaluation of related Neritimorpha (Gastropoda). Mitteilungen aus dem geologisch-palaeontologischen Institut der Universitaet Hamburg 85: 65 - 164"title="The history of Theodoxus and Neritina connected with description and systematic evaluation of related Neritimorpha (Gastropoda)."volume="85"year="2001">Bandel 2001</bibRefCitation>
<bibRefCitationid="80EA83FB94AD1F757F5B727CEFAE096D"author="Amr, ZS"journalOrPublisher="Denisia"pageId="0"pageNumber="25"pagination="221 - 227"publicationUrl="http://www.zobodat.at/stable/pdf/DENISIA_0014_0221-0227.pdf"refId="B10"refString="Amr, ZS, Abu Baker, M, 2004. Freshwater snails of Jordan. Denisia 14: 221 - 227, http://www.zobodat.at/stable/pdf/DENISIA_0014_0221-0227.pdf"title="Freshwater snails of Jordan."url="http://www.zobodat.at/stable/pdf/DENISIA_0014_0221-0227.pdf"volume="14"year="2004">Amr and Abu Baker 2004</bibRefCitation>
<bibRefCitationid="65DD180068812544E07D0FEBF8D0B417"DOI="https://doi.org/10.1080/09397140.1989.10637575"author="Schuett, H"journalOrPublisher="Zoology in the Middle East"pageId="0"pageNumber="25"pagination="55 - 58"refId="B144"refString="Schuett, H, Sesen, R, 1989a. The freshwater molluscs of Ceylanpinar. Zoology in the Middle East 3 (1): 55 - 58, DOI: https://doi.org/10.1080/09397140.1989.10637575"title="The freshwater molluscs of Ceylanpinar."url="https://doi.org/10.1080/09397140.1989.10637575"volume="3"year="1989 a">
<bibRefCitationid="0A19F63D1685C0D921990E49E88A2160"DOI="https://doi.org/10.1080/09397140.1993.10637638"author="Schuett, H"journalOrPublisher="Zoology in the Middle East"pageId="0"pageNumber="25"pagination="69 - 112"refId="B143"refString="Schuett, H, Ortal, R, 1993. A preliminary correlation between the Plio-Pleistocene malacofaunas of the Jordan Valley (Israel) and the Orontes Valley (Syria). Zoology in the Middle East 8 (1): 69 - 112, DOI: https://doi.org/10.1080/09397140.1993.10637638"title="A preliminary correlation between the Plio-Pleistocene malacofaunas of the Jordan Valley (Israel) and the Orontes Valley (Syria)."url="https://doi.org/10.1080/09397140.1993.10637638"volume="8"year="1993">
<bibRefCitationid="FD848A5E5304348DF5B1A5B32C0ABCB3"DOI="https://doi.org/10.1080/09397140.1993.10637638"author="Schuett, H"journalOrPublisher="Zoology in the Middle East"pageId="0"pageNumber="25"pagination="69 - 112"refId="B143"refString="Schuett, H, Ortal, R, 1993. A preliminary correlation between the Plio-Pleistocene malacofaunas of the Jordan Valley (Israel) and the Orontes Valley (Syria). Zoology in the Middle East 8 (1): 69 - 112, DOI: https://doi.org/10.1080/09397140.1993.10637638"title="A preliminary correlation between the Plio-Pleistocene malacofaunas of the Jordan Valley (Israel) and the Orontes Valley (Syria)."url="https://doi.org/10.1080/09397140.1993.10637638"volume="8"year="1993">
<bibRefCitationid="4F57352214CD1AC559558214F1DECEF8"author="Brown, DS"journalOrPublisher="Taylor & Francis, London"pageId="0"pageNumber="25"refId="B33"refString="Brown, DS, 1994. Freshwater snails of Africa and their medical importance. Taylor & Francis, London"title="Freshwater snails of Africa and their medical importance."year="1994">Brown 1994</bibRefCitation>
<bibRefCitationid="31EF802254BCF7133A3D8728A1AB3B4B"author="Gloeer, P"journalOrPublisher="Peter Gloeer, Hetlingen, Germany"pageId="0"pageNumber="25"refId="B62"refString="Gloeer, P, 2019. The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails. Peter Gloeer, Hetlingen, Germany"title="The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails."year="2019">
<bibRefCitationid="B9B047DFCAB994980FD990C0611D0B95"author="Amr, ZS"journalOrPublisher="Denisia"pageId="0"pageNumber="25"pagination="221 - 227"publicationUrl="http://www.zobodat.at/stable/pdf/DENISIA_0014_0221-0227.pdf"refId="B10"refString="Amr, ZS, Abu Baker, M, 2004. Freshwater snails of Jordan. Denisia 14: 221 - 227, http://www.zobodat.at/stable/pdf/DENISIA_0014_0221-0227.pdf"title="Freshwater snails of Jordan."url="http://www.zobodat.at/stable/pdf/DENISIA_0014_0221-0227.pdf"volume="14"year="2004">Amr and Abu Baker 2004</bibRefCitation>
<bibRefCitationid="2EE9ED14AFE03450D63F96B82A3B66B5"author="Handal, EN"journalOrPublisher="Jordan Journal of Natural History"pageId="0"pageNumber="25"pagination="23 - 29"refId="B70"refString="Handal, EN, Amr, Z, Qumsiyeh, MB, 2015. Some records of freshwater snail from the Occupied Palestinian territories. Jordan Journal of Natural History 1 (2): 23 - 29"title="Some records of freshwater snail from the Occupied Palestinian territories."volume="1"year="2015">Handal et al. 2015</bibRefCitation>
<bibRefCitationid="36313A69CB4D46BBEEB2F2BF413CC1EE"DOI="https://doi.org/10.3897/zookeys.219.3406"author="Gloeer, P"journalOrPublisher="ZooKeys"pageId="0"pageNumber="25"pagination="11 - 61"refId="B63"refString="Gloeer, P, Pesic, V, 2012. The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species. ZooKeys 219: 11 - 61, DOI: https://doi.org/10.3897/zookeys.219.3406"title="The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species."url="https://doi.org/10.3897/zookeys.219.3406"volume="219"year="2012">
<bibRefCitationid="D93D6715ABFC76002261C8F2030942E5"DOI="https://doi.org/10.3897/zookeys.219.3406"author="Gloeer, P"journalOrPublisher="ZooKeys"pageId="0"pageNumber="25"pagination="11 - 61"refId="B63"refString="Gloeer, P, Pesic, V, 2012. The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species. ZooKeys 219: 11 - 61, DOI: https://doi.org/10.3897/zookeys.219.3406"title="The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species."url="https://doi.org/10.3897/zookeys.219.3406"volume="219"year="2012">
<bibRefCitationid="7C23CB3C4B775577686646DF8AC02735"author="Mansoorian, AB"journalOrPublisher="Privately published, Tehran"pageId="0"pageNumber="25"refId="B96"refString="Mansoorian, AB, Samaee, A, 2012. Gastropoda of the freshwaters of Iran. Privately published, Tehran"title="Gastropoda of the freshwaters of Iran."year="2012">Mansoorian and Samaee 2012</bibRefCitation>
<bibRefCitationid="7C7E7C980B008DB3404F449DA69D3857"DOI="https://doi.org/10.3897/zookeys.219.3406"author="Gloeer, P"journalOrPublisher="ZooKeys"pageId="0"pageNumber="25"pagination="11 - 61"refId="B63"refString="Gloeer, P, Pesic, V, 2012. The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species. ZooKeys 219: 11 - 61, DOI: https://doi.org/10.3897/zookeys.219.3406"title="The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species."url="https://doi.org/10.3897/zookeys.219.3406"volume="219"year="2012">
<bibRefCitationid="07F85523E7F55938B8E20DFAD8784C5C"author="Alhejoj, I"journalOrPublisher="Freiberger Forschungshefte: Palaeontologie, Stratigraphie, Fazies C"pageId="0"pageNumber="25"pagination="141 - 173"refId="B7"refString="Alhejoj, I, Bandel, K, 2013. Mollusks of the Pleistocene Al-Qarn Formation of the Jordan Rift-Valley in Jordan. Freiberger Forschungshefte: Palaeontologie, Stratigraphie, Fazies C 545 (21): 141 - 173"title="Mollusks of the Pleistocene Al-Qarn Formation of the Jordan Rift-Valley in Jordan."volume="545"year="2013">Alhejoj and Bandel 2013</bibRefCitation>
<bibRefCitationid="41E850CA3369D3F7BF3ACA63950D126B"author="Mienis, HK"journalOrPublisher="Ellipsaria"pageId="0"pageNumber="25"pagination="37 - 38"refId="B102"refString="Mienis, HK, Rittner, O, 2017. Some information concerning Theodoxus (Neritaea) octagonus Eichhorst, 2016, an endemic subterranean species from Israel. Ellipsaria 19 (2): 37 - 38"title="Some information concerning Theodoxus (Neritaea) octagonus Eichhorst, 2016, an endemic subterranean species from Israel."volume="19"year="2017">Mienis and Rittner 2017</bibRefCitation>
<bibRefCitationid="EC37AFDEFCDFDC0A8E5C2EF88E7993B8"author="Gloeer, P"journalOrPublisher="Peter Gloeer, Hetlingen, Germany"pageId="0"pageNumber="25"refId="B62"refString="Gloeer, P, 2019. The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails. Peter Gloeer, Hetlingen, Germany"title="The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails."year="2019">
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<bibRefCitationid="334B7CD3D7949F7C16A606CEA70D5281"author="Martens, Ev"journalOrPublisher="Bauer & Raspe, Nuernberg"pageId="0"pageNumber="25"publicationUrl="https://biodiversitylibrary.org/page/52484536"refId="B99"refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer & Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536"title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)."url="https://biodiversitylibrary.org/page/52484536"year="1879">Martens 1879</bibRefCitation>
<bibRefCitationid="AD6359D4AA457252353AE1E222FC37BD"DOI="https://doi.org/10.1007/978-3-319-51856-5_17"author="Alhejoj, I"journalOrPublisher="Freiberger Forschungshefte: Palaeontologie, Stratigraphie, Fazies C"pageId="0"pageNumber="25"refId="B8"refString="Alhejoj, I, Bandel, K, Salameh, E, 2017. Aquatic mollusks: occurrences, identification and their use as bioindicators of environmental conditions (salinity, trace elements and pollution parameters) in Jordan. In Abdalla O, Kacimov A, Chen M, Al-Maktoumi A, Al-Hosni T and Clark I (Eds) Water resources in arid areas: the way forward. Springer, Cham, 295-318. https://doi.org/10.1007/978-3-319-51856-5_17"title="Aquatic mollusks: occurrences, identification and their use as bioindicators of environmental conditions (salinity, trace elements and pollution parameters) in Jordan. In Abdalla O, Kacimov A, Chen M, Al-Maktoumi A, Al-Hosni T and Clark I (Eds) Water resources in arid areas: the way forward. Springer, Cham, 295 - 318."url="https://doi.org/10.1007/978-3-319-51856-5_17"year="2017">Alhejoj et al. 2017</bibRefCitation>
<bibRefCitationid="967CABBEE899021BD4657D9F2A9C21B0"DOI="https://doi.org/10.1111/jbi.13590"author="Sands, AF"journalOrPublisher="Journal of Biogeography"pageId="0"pageNumber="25"pagination="1274 - 1286"refId="B138"refString="Sands, AF, Sereda, SV, Stelbrink, B, Neubauer, TA, Lazarev, S, Wilke, T, Albrecht, C, 2019a. Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography 46 (6): 1274 - 1286, DOI: https://doi.org/10.1111/jbi.13590"title="Contributions of biogeographical functions to species accumulation may change over time in refugial regions."url="https://doi.org/10.1111/jbi.13590"volume="46"year="2019 a">Sands et al. (2019a)</bibRefCitation>
<bibRefCitationid="EF0F7BFAB2B061BD9D9422F15BDE9928"author="Brown, DS"journalOrPublisher="Taylor & Francis, London"pageId="0"pageNumber="25"refId="B33"refString="Brown, DS, 1994. Freshwater snails of Africa and their medical importance. Taylor & Francis, London"title="Freshwater snails of Africa and their medical importance."year="1994">Brown 1994</bibRefCitation>
<bibRefCitationid="4B57086F469850E46237EF9EF2A82345"author="Gloeer, P"journalOrPublisher="Peter Gloeer, Hetlingen, Germany"pageId="0"pageNumber="25"refId="B62"refString="Gloeer, P, 2019. The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails. Peter Gloeer, Hetlingen, Germany"title="The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails."year="2019">
<taxonomicNameid="24F3DFEEDD70CC943D20B6E955F90430"lsidName="N. anatolica var. mesopotamica"pageId="0"pageNumber="25"rank="variety"species="anatolica"variety="mesopotamica">N. anatolica var. mesopotamica</taxonomicName>
<bibRefCitationid="9CAF99DE4D6EE9199904F2722CABD7A8"DOI="https://doi.org/10.1080/09397140.1989.10637575"author="Schuett, H"journalOrPublisher="Zoology in the Middle East"pageId="0"pageNumber="25"pagination="55 - 58"refId="B144"refString="Schuett, H, Sesen, R, 1989a. The freshwater molluscs of Ceylanpinar. Zoology in the Middle East 3 (1): 55 - 58, DOI: https://doi.org/10.1080/09397140.1989.10637575"title="The freshwater molluscs of Ceylanpinar."url="https://doi.org/10.1080/09397140.1989.10637575"volume="3"year="1989 a">
<figureCitationid="7E0EC07867A00E03B32F4DBE5E00C6DA"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the " Systematic checklist " for more information). Small green (PP ≥ 0.95) and red (PP <0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)."figureDoi="10.3897/zse.96.48312.figure2"httpUri="https://binary.pensoft.net/fig/376451"pageId="0"pageNumber="25">2</figureCitation>
<bibRefCitationid="65B1129CDA9C3A45F1BAD6E35C0A8090"author="Roth, G"editor="Krupp, F"journalOrPublisher="Dr. ludwig Reichert Verlag, Wiesbaden, Germany"pageId="0"pageNumber="25"pagination="72 - 79"refId="B136"refString="Roth, G, 1987. Data on the distribution and faunal history of the genus Theodoxus in the Middle East (Gastropoda: Neritidae). In: Krupp, F, Schneider, W, Kinzelbach, R, Eds., Proceedings of the Symposium on the Fauna and Zoogeography of the Middle East. Dr. ludwig Reichert Verlag, Wiesbaden, Germany: 72 - 79"title="Data on the distribution and faunal history of the genus Theodoxus in the Middle East (Gastropoda: Neritidae)."volumeTitle="Proceedings of the Symposium on the Fauna and Zoogeography of the Middle East."year="1987">Roth (1987)</bibRefCitation>
<figureCitationid="8E83C0407E8E78C3EAA0AD121CBF0E98"captionStart="Figure 15"captionStartId="F15"captionText="Figure 15. Theodoxus jordani (G. B. Sowerby I, 1836). A - D. Specimen from the Avakas Gorge, Peyia, Cyprus (UGSB 18321) incorporated into the phylogeny (Fig. 2); E - H. Specimen collected in the Ceyhan River, Eski Misis, Adana, Turkey (UGSB 20777); I - L. Specimen from the Orontes River, Antakya, Hatay, Turkey (UGSB 24175) used in the phylogeny (Fig. 2); M - O. Specimen (T. euphraticus - morphotype) from the Seyed Hosein Park spring, near the Bishapur, Fars, Iran (UGSB 22233); P - R. Topotype of N. jordani from the River Jordan, Israel (UGSB 24177) incorporated into the phylogeny (Fig. 2); S, T. Specimen (T. euphraticus - morphotype) from the Karun River, Ahvaz, Iran (UGSB 21681); U. Specimen (T. niloticus - morphotype) collected in the Nile River, Egypt (UGSB 24178) used in the phylogeny (Fig. 2); V. Specimen (T. euphraticus - morphotype) from the Karun River, Ahvaz, Iran (UGSB 21682). Scale bars: 1 mm."figureDoi="10.3897/zse.96.48312.figure15"httpUri="https://binary.pensoft.net/fig/376464"pageId="0"pageNumber="25">15</figureCitation>
<figureCitationid="ACE1FFE094C77A4DCA119026ADEA516D"captionStart="Figure 17"captionStartId="F17"captionText="Figure 17. Theodoxus jordani (G. B. Sowerby I, 1836). A - C. Syntype of N. meridionalis var. mesopotamica from near Diyarbakir, Turkey (ZMZ 528914); D, E. Specimen (T. mesopotamicus - morphotype) collected in Sanliurfa, Turkey (UGSB 23426) and used in the phylogeny (Fig. 2); F - H; K - M. Two syntypes of N. cinctella collected at the source of River Chabur near Ras al-Ayn, Syria (ZMB / Moll 21732); I, J. Topotype of N. cinctella from the same locality (UGSB 18762). Scale bars: 1 mm."figureDoi="10.3897/zse.96.48312.figure17"httpUri="https://binary.pensoft.net/fig/376466"pageId="0"pageNumber="25">17</figureCitation>
<figureCitationid="E49874CBDAA69A036288E56FAD8EE207"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the " Systematic checklist " for more information). Small green (PP ≥ 0.95) and red (PP <0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)."figureDoi="10.3897/zse.96.48312.figure2"httpUri="https://binary.pensoft.net/fig/376451"pageId="0"pageNumber="25">2</figureCitation>
<figureCitationid="AEDC5D76BD3F308F5476384B5E3AC42A"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the " Systematic checklist " for more information). Small green (PP ≥ 0.95) and red (PP <0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)."figureDoi="10.3897/zse.96.48312.figure2"httpUri="https://binary.pensoft.net/fig/376451"pageId="0"pageNumber="25">2</figureCitation>
<figureCitationid="528CBBDF3C55833D7A5D2FDBB1BBFE48"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the " Systematic checklist " for more information). Small green (PP ≥ 0.95) and red (PP <0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)."figureDoi="10.3897/zse.96.48312.figure2"httpUri="https://binary.pensoft.net/fig/376451"pageId="0"pageNumber="25">2</figureCitation>
<figureCitationid="68BAE4B00C51D5FAA9F0F87FEE83EEF3"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the " Systematic checklist " for more information). Small green (PP ≥ 0.95) and red (PP <0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)."figureDoi="10.3897/zse.96.48312.figure2"httpUri="https://binary.pensoft.net/fig/376451"pageId="0"pageNumber="25">2</figureCitation>
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<taxonomicNameid="7AF33AF34614CF5C92788AB12B6FED2D"lsidName="N. jordani var. turris"pageId="0"pageNumber="25"rank="variety"species="jordani"variety="turris">N. jordani var. turris</taxonomicName>
<taxonomicNameid="7121FB8F9E3C68FBF9F2F3E2B13B235B"lsidName="N. meridionalis var. mesopotamica"pageId="0"pageNumber="25"rank="variety"species="meridionalis"variety="mesopotamica">N. meridionalis var. mesopotamica</taxonomicName>
<figureCitationid="15ABACF74CE02610ACEC122B58697C89"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the " Systematic checklist " for more information). Small green (PP ≥ 0.95) and red (PP <0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)."figureDoi="10.3897/zse.96.48312.figure2"httpUri="https://binary.pensoft.net/fig/376451"pageId="0"pageNumber="25">2</figureCitation>
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besides overlapping ranges; the operculum structures of the two species are near identical with otherwise large pseudo-apophyses and both occasionally show keeling of the shell. Moreover, key characters used to distinguish
, such as the pronounced aperture and colouration of the shells, may be mitigated when intraspecific phenotypic plasticity of both species is considered (
<bibRefCitationid="ED5DE27915DB4423E6D37A8D34F340DD"DOI="https://doi.org/10.1007/BF00346406"author="Heller, J"journalOrPublisher="Oecologia"pageId="0"pageNumber="25"pagination="98 - 104"refId="B71"refString="Heller, J, 1979. Visual versus non-visual selection of shell colour in an Israeli freshwater snail. Oecologia 44 (1): 98 - 104, DOI: https://doi.org/10.1007/BF00346406"title="Visual versus non-visual selection of shell colour in an Israeli freshwater snail."url="https://doi.org/10.1007/BF00346406"volume="44"year="1979">Heller 1979</bibRefCitation>
<bibRefCitationid="E0EAC7414231C3C86E22E627EBB94570"author="Bandel, K"journalOrPublisher="Mitteilungen aus dem geologisch-palaeontologischen Institut der Universitaet Hamburg"pageId="0"pageNumber="25"pagination="65 - 164"refId="B20"refString="Bandel, K, 2001. The history of Theodoxus and Neritina connected with description and systematic evaluation of related Neritimorpha (Gastropoda). Mitteilungen aus dem geologisch-palaeontologischen Institut der Universitaet Hamburg 85: 65 - 164"title="The history of Theodoxus and Neritina connected with description and systematic evaluation of related Neritimorpha (Gastropoda)."volume="85"year="2001">Bandel 2001</bibRefCitation>
<bibRefCitationid="B91194778334946DD6483B9437E7BE4C"author="Amr, Z"journalOrPublisher="Jordan Journal of Natural History"pageId="0"pageNumber="25"pagination="83 - 115"refId="B11"refString="Amr, Z, Nasarat, H, Neubert, E, 2014. Notes on the current and past freshwater snail fauna of Jordan. Jordan Journal of Natural History 1: 83 - 115"title="Notes on the current and past freshwater snail fauna of Jordan."volume="1"year="2014">Amr et al. 2014</bibRefCitation>
<bibRefCitationid="DC2980FB3B789EB8BCA80578D1E4AFD4"author="Mienis, HK"journalOrPublisher="Ellipsaria"pageId="0"pageNumber="25"pagination="37 - 38"refId="B102"refString="Mienis, HK, Rittner, O, 2017. Some information concerning Theodoxus (Neritaea) octagonus Eichhorst, 2016, an endemic subterranean species from Israel. Ellipsaria 19 (2): 37 - 38"title="Some information concerning Theodoxus (Neritaea) octagonus Eichhorst, 2016, an endemic subterranean species from Israel."volume="19"year="2017">Mienis and Rittner 2017</bibRefCitation>
<figureCitationid="F1399CD2CA045E4FE5506D3C7DC74DAD"captionStart="Figure 15"captionStartId="F15"captionText="Figure 15. Theodoxus jordani (G. B. Sowerby I, 1836). A - D. Specimen from the Avakas Gorge, Peyia, Cyprus (UGSB 18321) incorporated into the phylogeny (Fig. 2); E - H. Specimen collected in the Ceyhan River, Eski Misis, Adana, Turkey (UGSB 20777); I - L. Specimen from the Orontes River, Antakya, Hatay, Turkey (UGSB 24175) used in the phylogeny (Fig. 2); M - O. Specimen (T. euphraticus - morphotype) from the Seyed Hosein Park spring, near the Bishapur, Fars, Iran (UGSB 22233); P - R. Topotype of N. jordani from the River Jordan, Israel (UGSB 24177) incorporated into the phylogeny (Fig. 2); S, T. Specimen (T. euphraticus - morphotype) from the Karun River, Ahvaz, Iran (UGSB 21681); U. Specimen (T. niloticus - morphotype) collected in the Nile River, Egypt (UGSB 24178) used in the phylogeny (Fig. 2); V. Specimen (T. euphraticus - morphotype) from the Karun River, Ahvaz, Iran (UGSB 21682). Scale bars: 1 mm."figureDoi="10.3897/zse.96.48312.figure15"httpUri="https://binary.pensoft.net/fig/376464"pageId="0"pageNumber="25">15</figureCitation>
<figureCitationid="687503BCD526787B82B0B967D4729563"captionStart="Figure 17"captionStartId="F17"captionText="Figure 17. Theodoxus jordani (G. B. Sowerby I, 1836). A - C. Syntype of N. meridionalis var. mesopotamica from near Diyarbakir, Turkey (ZMZ 528914); D, E. Specimen (T. mesopotamicus - morphotype) collected in Sanliurfa, Turkey (UGSB 23426) and used in the phylogeny (Fig. 2); F - H; K - M. Two syntypes of N. cinctella collected at the source of River Chabur near Ras al-Ayn, Syria (ZMB / Moll 21732); I, J. Topotype of N. cinctella from the same locality (UGSB 18762). Scale bars: 1 mm."figureDoi="10.3897/zse.96.48312.figure17"httpUri="https://binary.pensoft.net/fig/376466"pageId="0"pageNumber="25">17</figureCitation>
<taxonomicNameid="9D0E8959FE5B48527524CB99E0219FB5"lsidName="N. jordani var. turris"pageId="0"pageNumber="25"rank="variety"species="jordani"variety="turris">N. jordani var. turris</taxonomicName>
<taxonomicNameid="80964604980BCB25BEF0A07AA7E078F3"lsidName="T. jordani var. unicarinatus"pageId="0"pageNumber="25"rank="variety"species="jordani"variety="unicarinatus">T. jordani var. unicarinatus</taxonomicName>
<taxonomicNameid="4E831DE0774CDA0D2F1E49A9C9ED47DA"lsidName="T. jordani var. bicarinatus"pageId="0"pageNumber="25"rank="variety"species="jordani"variety="bicarinatus">T. jordani var. bicarinatus</taxonomicName>
<bibRefCitationid="2D197558D946386DA3297E8105937971"DOI="https://doi.org/10.1111/jbi.13590"author="Sands, AF"journalOrPublisher="Journal of Biogeography"pageId="0"pageNumber="25"pagination="1274 - 1286"refId="B138"refString="Sands, AF, Sereda, SV, Stelbrink, B, Neubauer, TA, Lazarev, S, Wilke, T, Albrecht, C, 2019a. Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography 46 (6): 1274 - 1286, DOI: https://doi.org/10.1111/jbi.13590"title="Contributions of biogeographical functions to species accumulation may change over time in refugial regions."url="https://doi.org/10.1111/jbi.13590"volume="46"year="2019 a">Sands et al. (2019a)</bibRefCitation>
<bibRefCitationid="452C54985FF1A97CE17F8E9BA54D8105"DOI="https://doi.org/10.1111/jbi.13590"author="Sands, AF"journalOrPublisher="Journal of Biogeography"pageId="0"pageNumber="25"pagination="1274 - 1286"refId="B138"refString="Sands, AF, Sereda, SV, Stelbrink, B, Neubauer, TA, Lazarev, S, Wilke, T, Albrecht, C, 2019a. Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography 46 (6): 1274 - 1286, DOI: https://doi.org/10.1111/jbi.13590"title="Contributions of biogeographical functions to species accumulation may change over time in refugial regions."url="https://doi.org/10.1111/jbi.13590"volume="46"year="2019 a">Sands et al. (2019a)</bibRefCitation>
<figureCitationid="9117C853AEDC1C241080A0FEB55BFF3E"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the " Systematic checklist " for more information). Small green (PP ≥ 0.95) and red (PP <0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)."figureDoi="10.3897/zse.96.48312.figure2"httpUri="https://binary.pensoft.net/fig/376451"pageId="0"pageNumber="25">2</figureCitation>
<bibRefCitationid="B8FDA848B7B5EE7BD904DAA011167EC0"DOI="https://doi.org/10.1111/jbi.13590"author="Sands, AF"journalOrPublisher="Journal of Biogeography"pageId="0"pageNumber="25"pagination="1274 - 1286"refId="B138"refString="Sands, AF, Sereda, SV, Stelbrink, B, Neubauer, TA, Lazarev, S, Wilke, T, Albrecht, C, 2019a. Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography 46 (6): 1274 - 1286, DOI: https://doi.org/10.1111/jbi.13590"title="Contributions of biogeographical functions to species accumulation may change over time in refugial regions."url="https://doi.org/10.1111/jbi.13590"volume="46"year="2019 a">Sands et al. 2019a</bibRefCitation>
<figureCitationid="36507A314B6F762D48F626127B3BF18B"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the " Systematic checklist " for more information). Small green (PP ≥ 0.95) and red (PP <0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)."figureDoi="10.3897/zse.96.48312.figure2"httpUri="https://binary.pensoft.net/fig/376451"pageId="0"pageNumber="25">2</figureCitation>
<bibRefCitationid="1435E7CB4A9BD42FA54AB8F8B03EEB16"author="Martens, Ev"journalOrPublisher="Novitates Conchologicae, Supplement"pageId="0"pageNumber="25"pagination="1 - 127"publicationUrl="https://biodiversitylibrary.org/page/12980992"refId="B98"refString="Martens, Ev, 1874. Ueber vorderasiatische Conchylien nach den Sammlungen des Prof. Hausknecht. Novitates Conchologicae, Supplement 5: 1 - 127, https://biodiversitylibrary.org/page/12980992"title="Ueber vorderasiatische Conchylien nach den Sammlungen des Prof. Hausknecht."url="https://biodiversitylibrary.org/page/12980992"volume="5"year="1874">Martens (1874)</bibRefCitation>
<taxonomicNameid="1E914935DD93AB95509D0F54571ECA63"class="Gastropoda"family="Neritidae"genus="Neritina"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Neritina anatolica var. mesopotamica"order="Cycloneritida"pageId="0"pageNumber="25"phylum="Mollusca"rank="variety"species="anatolica"variety="mesopotamica">Neritina anatolica var. mesopotamica</taxonomicName>
<taxonomicNameid="548C6D1F13B1DD62E0925EE6A7B20EB2"lsidName="N. meridionalis var. mesopotamica"pageId="0"pageNumber="25"rank="variety"species="meridionalis"variety="mesopotamica">N. meridionalis var. mesopotamica</taxonomicName>
<bibRefCitationid="2EEEC4D8AED83B51C0637CCA57A0FC3F"author="Martens, Ev"journalOrPublisher="Novitates Conchologicae, Supplement"pageId="0"pageNumber="25"pagination="1 - 127"publicationUrl="https://biodiversitylibrary.org/page/12980992"refId="B98"refString="Martens, Ev, 1874. Ueber vorderasiatische Conchylien nach den Sammlungen des Prof. Hausknecht. Novitates Conchologicae, Supplement 5: 1 - 127, https://biodiversitylibrary.org/page/12980992"title="Ueber vorderasiatische Conchylien nach den Sammlungen des Prof. Hausknecht."url="https://biodiversitylibrary.org/page/12980992"volume="5"year="1874">Martens 1874</bibRefCitation>
<bibRefCitationid="4BC63BE5E576AFC6F7F492EFFF84A876"author="Martens, Ev"journalOrPublisher="Bauer & Raspe, Nuernberg"pageId="0"pageNumber="25"publicationUrl="https://biodiversitylibrary.org/page/52484536"refId="B99"refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer & Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536"title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)."url="https://biodiversitylibrary.org/page/52484536"year="1879">Martens 1879</bibRefCitation>
<bibRefCitationid="995B7113EA2160A2A045F9144C3789A9"DOI="https://doi.org/10.1080/09397140.1989.10637575"author="Schuett, H"journalOrPublisher="Zoology in the Middle East"pageId="0"pageNumber="25"pagination="55 - 58"refId="B144"refString="Schuett, H, Sesen, R, 1989a. The freshwater molluscs of Ceylanpinar. Zoology in the Middle East 3 (1): 55 - 58, DOI: https://doi.org/10.1080/09397140.1989.10637575"title="The freshwater molluscs of Ceylanpinar."url="https://doi.org/10.1080/09397140.1989.10637575"volume="3"year="1989 a">
<taxonomicNameid="AFF02086757212F4B018F83A80517476"lsidName="N. anatolica var. mesopotamica"pageId="0"pageNumber="25"rank="variety"species="anatolica"variety="mesopotamica">N. anatolica var. mesopotamica</taxonomicName>
<taxonomicNameid="4681E19EDE213E3011F6F9F13A37F699"lsidName="N. meridionalis var. mesopotamica"pageId="0"pageNumber="25"rank="variety"species="meridionalis"variety="mesopotamica">N. meridionalis var. mesopotamica</taxonomicName>
is common throughout southern Anatolia, the Levant, Mesopotamia, and parts of the Middle East, extending to at least southern Iran, although it is probably not found eastward of the Zagros Mountains (
<bibRefCitationid="2E5C0F9A7CECBCEB36371F77DE16F4A9"author="Roth, G"editor="Krupp, F"journalOrPublisher="Dr. ludwig Reichert Verlag, Wiesbaden, Germany"pageId="0"pageNumber="25"pagination="72 - 79"refId="B136"refString="Roth, G, 1987. Data on the distribution and faunal history of the genus Theodoxus in the Middle East (Gastropoda: Neritidae). In: Krupp, F, Schneider, W, Kinzelbach, R, Eds., Proceedings of the Symposium on the Fauna and Zoogeography of the Middle East. Dr. ludwig Reichert Verlag, Wiesbaden, Germany: 72 - 79"title="Data on the distribution and faunal history of the genus Theodoxus in the Middle East (Gastropoda: Neritidae)."volumeTitle="Proceedings of the Symposium on the Fauna and Zoogeography of the Middle East."year="1987">Roth 1987</bibRefCitation>
<bibRefCitationid="F5EC08DC67A4368C878A93373C8DA463"author="Bandel, K"journalOrPublisher="Mitteilungen aus dem geologisch-palaeontologischen Institut der Universitaet Hamburg"pageId="0"pageNumber="25"pagination="65 - 164"refId="B20"refString="Bandel, K, 2001. The history of Theodoxus and Neritina connected with description and systematic evaluation of related Neritimorpha (Gastropoda). Mitteilungen aus dem geologisch-palaeontologischen Institut der Universitaet Hamburg 85: 65 - 164"title="The history of Theodoxus and Neritina connected with description and systematic evaluation of related Neritimorpha (Gastropoda)."volume="85"year="2001">Bandel 2001</bibRefCitation>
<bibRefCitationid="C7BB02C421BBA92A34758D958F495516"DOI="https://doi.org/10.1111/jbi.13590"author="Sands, AF"journalOrPublisher="Journal of Biogeography"pageId="0"pageNumber="25"pagination="1274 - 1286"refId="B138"refString="Sands, AF, Sereda, SV, Stelbrink, B, Neubauer, TA, Lazarev, S, Wilke, T, Albrecht, C, 2019a. Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography 46 (6): 1274 - 1286, DOI: https://doi.org/10.1111/jbi.13590"title="Contributions of biogeographical functions to species accumulation may change over time in refugial regions."url="https://doi.org/10.1111/jbi.13590"volume="46"year="2019 a">Sands et al. 2019a</bibRefCitation>
<figureCitationid="F95C5666EC6FD07F51514FF9009D0507"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Occurrence map for Theodoxus spp. in Asia. All points conform to specimens used in the phylogenetic analyses (Suppl. material 1: Table S 1), except for T. gloeri for which no molecular data are available and only the type and paratype localities are indicated. Species are partitioned across four identical maps (A - D) to limit overlap of points as far as possible. Barring T. gloeri, closer related species are grouped together (Fig. 2)."figureDoi="10.3897/zse.96.48312.figure3"httpUri="https://binary.pensoft.net/fig/376452"pageId="0"pageNumber="25">3A</figureCitation>
<bibRefCitationid="A51BE9B26EE1A6D189638E0FFA053CAB"DOI="https://doi.org/10.1111/jbi.13590"author="Sands, AF"journalOrPublisher="Journal of Biogeography"pageId="0"pageNumber="25"pagination="1274 - 1286"refId="B138"refString="Sands, AF, Sereda, SV, Stelbrink, B, Neubauer, TA, Lazarev, S, Wilke, T, Albrecht, C, 2019a. Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography 46 (6): 1274 - 1286, DOI: https://doi.org/10.1111/jbi.13590"title="Contributions of biogeographical functions to species accumulation may change over time in refugial regions."url="https://doi.org/10.1111/jbi.13590"volume="46"year="2019 a">Sands et al. 2019a</bibRefCitation>
Mousson, 1874 which inhabits the lower reaches of the Euphrates, Karun, and Tigris rivers, as well as other brackish water systems around the Persian Gulf (
<bibRefCitationid="068876E73B0C0B5377BDE7CE53BBEC94"DOI="https://doi.org/10.3897/zookeys.219.3406"author="Gloeer, P"journalOrPublisher="ZooKeys"pageId="0"pageNumber="25"pagination="11 - 61"refId="B63"refString="Gloeer, P, Pesic, V, 2012. The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species. ZooKeys 219: 11 - 61, DOI: https://doi.org/10.3897/zookeys.219.3406"title="The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species."url="https://doi.org/10.3897/zookeys.219.3406"volume="219"year="2012">
