<documentid="9FE460AA3BD15A2F61890866AEA73254"ID-DOI="10.5070/P9401057774"ID-ISSN="0031-0298"ID-Zenodo-Dep="10913295"ID-ZooBank="11600574-2B0E-4C13-BC08-A3A5EF9EE562"IM.bibliography_approvedBy="carolina"IM.illustrations_approvedBy="carolina"IM.materialsCitations_approvedBy="carolina"IM.metadata_approvedBy="felipe"IM.tables_requiresApprovalFor="operationResults"IM.taxonomicNames_approvedBy="carolina"IM.treatmentCitations_approvedBy="felipe"IM.treatments_approvedBy="carolina"checkinTime="1712167020321"checkinUser="felipe"docAuthor="Groves, Lindsey & Squires, Richard L."docDate="2023"docId="921FD94CFFC2FFB9FEDDFF78FCD8FB53"docLanguage="en"docName="PaleoBios.40.10.1-52.pdf"docOrigin="PaleoBios 40 (10)"docSource="http://dx.doi.org/10.5070/p9401057774"docStyle="DocumentStyle:76F55B8832C4952BA7DD61B33E1F0E81.3:PaleoBios.2023-.journal_article"docStyleId="76F55B8832C4952BA7DD61B33E1F0E81"docStyleName="PaleoBios.2023-.journal_article"docStyleVersion="3"docTitle="Bernaya sp."docType="treatment"docVersion="7"lastPageNumber="16"masterDocId="6E26A134FFD3FFA8FFA2FFBBFFCAFFE5"masterDocTitle="Revision of northeast Pacific Paleogene cypraeoidean gastropods, including recognition of three new species: implications for paleobiogeographic distribution and faunal turnover"masterLastPageNumber="52"masterPageNumber="1"pageNumber="16"updateTime="1727793805776"updateUser="ExternalLinkService"zenodo-license-document="CC-BY-NC-SA-4.0">
<mods:titleid="EE3EADF4A874DBC8505D9DDBA46FCD48">Revision of northeast Pacific Paleogene cypraeoidean gastropods, including recognition of three new species: implications for paleobiogeographic distribution and faunal turnover</mods:title>
<figureCitationid="828D74DFFFC2FFB9FE2EFF5DFDCEFEE4"box="[396,516,230,257]"captionStart="Figure 5"captionStartId="15.[113,200,200,226]"captionTargetId="figure-12@14.[141,1485,197,1966]"captionTargetPageId="14"captionText="Figure 5 (previous page) A–C. Bernaya grovesi Squires and Demetrion, 1992, Bateque Formation (Eocene, “Ca- pay Stage”), Mesa La Salina, Baja California Sur, México. Basal (A), dorsal (B), and right (labral) (C) views, all 2.6x, length 17 mm, width 12 mm, height 7.7 mm, of holotype IGM 5172, LACMIP Locality 401220a (ex CSUN Locality 1220b). D–F. Bernaya kaylinae n. sp., Llajas Formation, (Eocene, “Domengine Stage”), Las Llajas Canyon, north side Simi Valley, Ventura County, southern California. Basal (D), dorsal (E), and right (labral) side (F) views, 3.8x, length 25.6 mm, width 17.3 mm, height 12.9 mm, of holotype, LACMIP 40374.80, LACMIP Type 14938 (LACMIP Locality 40374). G–I. Bernaya squiresi Groves, 2011, unnamed Eocene strata (“Capay Stage”), southwest end of Discovery Bay, Jefferson County, Washington. Basal (G), dorsal (H), and right (labral) side (I) views, 0.97x, length 52.7 mm, width 40 mm, height 26.3 mm, of holotype LACMIP 22341.1, LACMIP Type 13644, LACMIP Locality 22341. J–L. Bernaya sp., unnamed Paleocene strata (Thanetian Stage), near Lower Lake, Lake County, northern California. Basal (J), dorsal (K), and right (labral) side (L) views, 1.7x, length 33.8 mm, width 23 mm, height 18.3 mm, hypotype LACMIP 7047.72, LACMIP Type 14923 (LACMIP Locality 7047). M–O. Protocypraea? simiensis (Nelson, 1925), Santa Susana Formation (Thanetian Stage), Simi Hills, Ventura County, Paleocene, southern Cali- fornia. Basal (M), dorsal (N), and right (labral) side (O) views, 1.6x, length 32.3 mm, width 24.6 mm, height 17.3 mm, of holotype UCMP 30498, UCMP Locality 3818. P.Protocypraea? sp. 1, Oyster Bay Formation, Paleocene, Appian Way, Central Island District, Vancouver Island, British Columbia. Basal view, 1.8x, length 48.1 mm, width 31.4 mm, height 25.3 mm, of CDM 997.91.3, informal locality AP3."figureDoi="http://doi.org/10.5281/zenodo.13869594"httpUri="https://zenodo.org/record/13869594/files/figure.png"pageId="17"pageNumber="16">Figs. 5J–L</figureCitation>
<figureCitationid="828D74DFFFC2FFB9FFDBFEEBFF3DFE8F"box="[121,247,335,362]"captionStart="Figure 5"captionStartId="15.[113,200,200,226]"captionTargetId="figure-12@14.[141,1485,197,1966]"captionTargetPageId="14"captionText="Figure 5 (previous page) A–C. Bernaya grovesi Squires and Demetrion, 1992, Bateque Formation (Eocene, “Ca- pay Stage”), Mesa La Salina, Baja California Sur, México. Basal (A), dorsal (B), and right (labral) (C) views, all 2.6x, length 17 mm, width 12 mm, height 7.7 mm, of holotype IGM 5172, LACMIP Locality 401220a (ex CSUN Locality 1220b). D–F. Bernaya kaylinae n. sp., Llajas Formation, (Eocene, “Domengine Stage”), Las Llajas Canyon, north side Simi Valley, Ventura County, southern California. Basal (D), dorsal (E), and right (labral) side (F) views, 3.8x, length 25.6 mm, width 17.3 mm, height 12.9 mm, of holotype, LACMIP 40374.80, LACMIP Type 14938 (LACMIP Locality 40374). G–I. Bernaya squiresi Groves, 2011, unnamed Eocene strata (“Capay Stage”), southwest end of Discovery Bay, Jefferson County, Washington. Basal (G), dorsal (H), and right (labral) side (I) views, 0.97x, length 52.7 mm, width 40 mm, height 26.3 mm, of holotype LACMIP 22341.1, LACMIP Type 13644, LACMIP Locality 22341. J–L. Bernaya sp., unnamed Paleocene strata (Thanetian Stage), near Lower Lake, Lake County, northern California. Basal (J), dorsal (K), and right (labral) side (L) views, 1.7x, length 33.8 mm, width 23 mm, height 18.3 mm, hypotype LACMIP 7047.72, LACMIP Type 14923 (LACMIP Locality 7047). M–O. Protocypraea? simiensis (Nelson, 1925), Santa Susana Formation (Thanetian Stage), Simi Hills, Ventura County, Paleocene, southern Cali- fornia. Basal (M), dorsal (N), and right (labral) side (O) views, 1.6x, length 32.3 mm, width 24.6 mm, height 17.3 mm, of holotype UCMP 30498, UCMP Locality 3818. P.Protocypraea? sp. 1, Oyster Bay Formation, Paleocene, Appian Way, Central Island District, Vancouver Island, British Columbia. Basal view, 1.8x, length 48.1 mm, width 31.4 mm, height 25.3 mm, of CDM 997.91.3, informal locality AP3."figureDoi="http://doi.org/10.5281/zenodo.13869594"httpUri="https://zenodo.org/record/13869594/files/figure.png"pageId="17"pageNumber="16">Figs. 5J–L</figureCitation>
<bibRefCitationid="7E2715ABFFC2FFB9FE88FDBBFE7FFDFF"author="Gabb, W. M."box="[298,437,512,538]"pageId="17"pageNumber="16"pagination="55 - 217"refId="ref37370"refString="Gabb, W. M. 1864. Description of the Cretaceous fos- sils. Geological Survey of California, Palaeontology 1 (4): 55 - 217, pls. 9 - 32."type="journal article"year="1864">Gabb, 1864</bibRefCitation>
</taxonomicName>
, in the Herndon Creek, Lower Lake, Lake County, northern
<collectingRegionid="D872A6B8FFC2FFB9FE7BFD98FD99FDD8"box="[473,595,547,573]"country="United States of America"name="California"pageId="17"pageNumber="16">California</collectingRegion>
(
<bibRefCitationid="7E2715ABFFC2FFB9FDC7FD98FCC5FDD8"author="Stanton, T. W."box="[613,783,547,573]"pageId="17"pageNumber="16"pagination="63 - 67"refId="ref43868"refString="Stanton, T. W. 1896. The faunal relations of the Eocene and Upper Cretaceous on the Pacific Coast. U. S. Geological Survey 17 th Annual Report, pt. 1, 1005 - 1060, pls. 63 - 67."type="book chapter"year="1896">Stanton 1896</bibRefCitation>
).
<bibRefCitationid="7E2715ABFFC2FFB9FFD2FDFDFF33FD85"author="Saul, L. R."box="[112,249,582,608]"pageId="17"pageNumber="16"pagination="1 - 7"refId="ref41604"refString="Saul, L. R. 1983. Turritella zonation across the Cretaceous- Tertiary boundary, California. University of California Publications, Geological Sciences 125: x + 1 - 164, figs. 1 - 27, pls. 1 - 7."type="journal article"year="1983">Saul (1983</bibRefCitation>
: p. 102) confirmed
<bibRefCitationid="7E2715ABFFC2FFB9FE57FDFDFD02FD84"author="Stanton, T. W."box="[501,712,582,609]"pageId="17"pageNumber="16"pagination="63 - 67"refId="ref43868"refString="Stanton, T. W. 1896. The faunal relations of the Eocene and Upper Cretaceous on the Pacific Coast. U. S. Geological Survey 17 th Annual Report, pt. 1, 1005 - 1060, pls. 63 - 67."type="book chapter"year="1896">Stanton’s (1896)</bibRefCitation>
report of this turritellid and assigned it to the Thanetian Stage.
from the Crescent Formation (middle lower Eocene “Capay Stage”). Both specimens have similar shell shape, a small exposed spire, numerous strong teeth on the apertural lips, and are poorly preserved in the critically important anterior end. Three additional specimens of the Lower Lake taxon (
the Lower Lake area, are likely the same genus but their preservation is too poor for confirmation. A single specimen from the Santa Susana Formation, Simi Valley, Ventura County, southern Cali- fornia (