132 lines
14 KiB
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132 lines
14 KiB
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<document id="E509A25A08425BC64881FC6279A4C5A0" ID="10.11646/zootaxa.4207.1.1" ID-CLB-Dataset="37549" ID-DOI="10.11646/zootaxa.4207.1.1" ID-GBIF-Dataset="cef35771-ff95-4a41-9af6-196dea899a64" ID-ISSN="1175-5326" ID-Zenodo-Dep="200444" ID-ZooBank="F2875582-31E2-496F-AEEF-1D657DD86C33" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1481615616696" checkinUser="plazi" docAuthor="Frolov, Andrey V., Montreuil, Olivier & Akhmetova, Lilia A." docDate="2016" docId="5B4FBD00FFDA6161FF02618EFA16FA13" docLanguage="en" docName="zootaxa.4207.1.1.pdf" docOrigin="Zootaxa 4207 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Orphninae" docType="key" docVersion="16" lastPageNumber="12" masterDocId="A776C578FFD1616AFF95625CFF8AFFF5" masterDocTitle="Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood" masterLastPageNumber="93" masterPageNumber="1" pageNumber="12" updateTime="1698433759527" updateUser="ExternalLinkService">
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<mods:mods id="2C9EF355A1FB556895F569C4E3377096" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="42702B3D61D339851E913F9A00DF5779">
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<mods:title id="4BA751DDFD63E9F9F3CB22E2EE95E90F">Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood</mods:title>
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</mods:titleInfo>
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<mods:name id="B24C2DA16FB3829C4979E8813DB84886" type="personal">
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<mods:roleTerm id="F965C9441E7DBE7DADB1D4642C51D4DA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7F5E9DEA1FEDE504FE78915CEB0DC9F9">Frolov, Andrey V.</mods:namePart>
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</mods:name>
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<mods:name id="DAF3CA7BCC34D0CAED16135E75314434" type="personal">
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</mods:role>
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<mods:namePart id="256440A1A86497BEEDEDC131602F9155">Montreuil, Olivier</mods:namePart>
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</mods:name>
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<mods:name id="6146A55AD17B37E0E0781234F5F65BEA" type="personal">
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<mods:roleTerm id="749ADA852AB45D24A0D2342C9858CB14">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3778A182642E6327F848FB314AB6208E">Akhmetova, Lilia A.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="5ECF57B9B4DC313B86329B4EDBD303CC">text</mods:typeOfResource>
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<mods:title id="FE33C42898E618662D0989C943F38C8C">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="894AEF252883051C9853B3B8B5DADFA4">
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<mods:date id="D5755843CBED50C3458B50B4FCBB8B28">2016</mods:date>
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<mods:detail id="32AB37821AE15D32AC6E663D2D57992A" type="volume">
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<mods:number id="32BD8D0BCA5E13A5BFEB406C65F2CC2E">4207</mods:number>
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</mods:detail>
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<mods:detail id="189441441FBFA6C33B83F9E20CC60696" type="issue">
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<mods:number id="26993AFC37413D020F7850A19E608FDC">1</mods:number>
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<mods:start id="E1813352A9A1CBE424AEFFF8E8972BC2">1</mods:start>
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<mods:end id="8DA10CE7DC3EA58A030B8D4FC25FABDD">93</mods:end>
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</mods:extent>
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<mods:classification id="96817309053FFA48EC821101DF004A28">journal article</mods:classification>
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<mods:identifier id="1F60E89A4936E3F659E78D1F67512A6F" type="CLB-Dataset">37549</mods:identifier>
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<mods:identifier id="8DDB5F97ACF157DC7BF0B827F7C30580" type="DOI">10.11646/zootaxa.4207.1.1</mods:identifier>
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<mods:identifier id="70B6E25B3F9B448880D35F9019970CF5" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="68C6CB65D0D25EB930F3E3D716922937" type="Zenodo-Dep">200444</mods:identifier>
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<mods:identifier id="16CB77B7D0D167D1D0FB65F442763F03" type="ZooBank">F2875582-31E2-496F-AEEF-1D657DD86C33</mods:identifier>
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</mods:mods>
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<treatment id="5B4FBD00FFDA6161FF02618EFA16FA13" ID-DOI="http://doi.org/10.5281/zenodo.6057628" ID-GBIF-Taxon="125107259" ID-Zenodo-Dep="6057628" LSID="urn:lsid:plazi:treatment:5B4FBD00FFDA6161FF02618EFA16FA13" httpUri="http://treatment.plazi.org/id/5B4FBD00FFDA6161FF02618EFA16FA13" lastPageNumber="12" pageId="11" pageNumber="12">
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<subSubSection id="9BFC5F9DFFDA6161FF02618EFCCAFC19" box="[151,832,978,1004]" pageId="11" pageNumber="12" type="nomenclature">
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<paragraph id="D3590C16FFDA6161FF02618EFCCAFC19" blockId="11.[151,832,978,1004]" box="[151,832,978,1004]" pageId="11" pageNumber="12">
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<heading id="8811BB7AFFDA6161FF02618EFCCAFC19" bold="true" box="[151,832,978,1004]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
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<emphasis id="E192D004FFDA6161FF02618EFCCAFC19" bold="true" box="[151,832,978,1004]" pageId="11" pageNumber="12">
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Key to the Madagascar
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<taxonomicName id="14E67795FFDA6161FE54618EFDC0FC19" ID-CoL="8RYRW" box="[449,586,978,1004]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subFamily" subFamily="Orphninae">Orphninae</taxonomicName>
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genera (both sexes)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="9BFC5F9DFFDA6161FF026645FA16FA13" pageId="11" pageNumber="12" type="key">
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<keyStep id="681214B3FFDA6161FF026645FA16FBBE" pageId="11" pageNumber="12">
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<paragraph id="D3590C16FFDA6161FF026645FA16FBDB" blockId="11.[151,1436,1049,1510]" box="[151,1436,1049,1070]" pageId="11" pageNumber="12">
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<keyLead id="6817AF23FFDA6161FF026645FA16FBDB" box="[151,1436,1049,1070]" pageId="11" pageNumber="12">
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1. Protibiae with 2 outer teeth apically and 1 much smaller outer tooth basally (
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<figureCitation id="4BDD1093FFDA6161FC2A6645FC73FBDB" box="[959,1017,1049,1070]" captionStart="FIGURE 7" captionStartId="17.[151,250,1807,1829]" captionTargetBox="[220,1364,197,1782]" captionTargetId="figure@17.[206,1380,193,1796]" captionTargetPageId="17" captionText="FIGURE 7. Pseudorphnus hiboni. Habitus in dorsal view (A—male, F—female), head and pronotum in dorsolateral view (B), aedeagus in lateral view and parameres in dorsal view (C), protibia (D, male left, female right), base of elytra, SEM (E), labels of holotype (G), distributional records (H)." httpUri="https://zenodo.org/record/200451/files/figure.png" pageId="11" pageNumber="12">Fig. 7</figureCitation>
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D)...........
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<taxonomicName id="14E67795FFDA6161FB346645FA16FBDB" authority="Benderitter" authorityName="Benderitter" box="[1185,1436,1049,1070]" class="Insecta" family="Orphnidae" genus="Pseudorphnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="E192D004FFDA6161FB346645FAA0FBDB" box="[1185,1322,1049,1070]" italics="true" pageId="11" pageNumber="12">Pseudorphnus</emphasis>
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Benderitter
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="D3590C16FFDA6161FF02666AFA16FBBE" blockId="11.[151,1436,1049,1510]" box="[151,1436,1078,1099]" pageId="11" pageNumber="12">
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<keyLead id="6817AF23FFDA6161FF02666AFA16FBBE" box="[151,1436,1078,1099]" pageId="11" pageNumber="12">
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- Protibiae with 3 outer teeth in apical half (
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<figureCitation id="4BDD1093FFDA6161FDE3666AFD4EFBBE" box="[630,708,1078,1099]" captionStart="FIGURE 10" captionStartId="26.[151,250,1841,1863]" captionTargetBox="[203,1414,199,1811]" captionTargetId="figure@26.[170,1416,193,1820]" captionTargetPageId="26" captionText="FIGURE 10. Madecorphnus montreuili. Habitus in dorsal view (A—male, C—female), aedeagus in lateral view and parameres in dorsal view (B), labels of holotype (D), internal sac of aedeagus (E—photograph, F—schematic representation of armature), distributional records (G), protibia (H, male left, female right)." httpUri="https://zenodo.org/record/200454/files/figure.png" pageId="11" pageNumber="12">Figs. 10</figureCitation>
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H, 30C, 28F).................................................2
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="681214B3FFDA6161FF026608FA16FAEC" pageId="11" pageNumber="12">
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<paragraph id="D3590C16FFDA6161FF026608FA16FB34" blockId="11.[151,1436,1049,1510]" pageId="11" pageNumber="12">
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<keyLead id="6817AF23FFDA6161FF026608FA16FB34" pageId="11" pageNumber="12">
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2. Body length smaller (usually less than
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<quantity id="141EA1F3FFDA6161FDC96608FD13FB9C" box="[604,665,1108,1129]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="12" unit="mm" value="7.0">7 mm</quantity>
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); lateral margin of pronotum with 4 long setae: 2 near anterior margin,
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<quantity id="141EA1F3FFDA6161FAC46608FAFDFB9C" box="[1361,1399,1108,1129]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="12" unit="in" value="1.0">1 in</quantity>
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the middle, and 1 near posterior margin; some or all these setae may be abraded, but 4 chaetiferous punctures are always distinct; left and right mandibles may differ strongly in length in males, and frontoclypeus is somewhat asymmetrical in such cases; disc of pronotum smooth, without processes or excavations..................................
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<taxonomicName id="14E67795FFDA6161FB2B66F0FA16FB34" authority="Paulian" authorityName="Paulian" box="[1214,1436,1196,1217]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="E192D004FFDA6161FB2B66F0FAC4FB34" box="[1214,1358,1196,1217]" italics="true" pageId="11" pageNumber="12">Madecorphnus</emphasis>
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Paulian
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="D3590C16FFDA6161FF026695FA16FAEC" blockId="11.[151,1436,1049,1510]" pageId="11" pageNumber="12">
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<keyLead id="6817AF23FFDA6161FF026695FA16FAEC" pageId="11" pageNumber="12">
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- Body length larger (usually longer than
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<quantity id="141EA1F3FFDA6161FDF56695FD10FB2B" box="[608,666,1225,1246]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="12" unit="mm" value="7.0">7 mm</quantity>
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); lateral margin of pronotum with numerous setae (at least more than 10) evenly distributed along the margin, setae may differ strongly in length; left and right mandibles of about same length, frontoclypeus symmetrical; disc of pronotum may be with more-or-less developed tubercles, ridges, and excavations in males........... 3
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="681214B3FFDA6161FF02677DFA16FA13" pageId="11" pageNumber="12">
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<paragraph id="D3590C16FFDA6161FF02677DFA10FA7B" blockId="11.[151,1436,1049,1510]" pageId="11" pageNumber="12">
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<keyLead id="6817AF23FFDA6161FF02677DFA10FA7B" pageId="11" pageNumber="12">
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3. Eyes smaller (width of eye in dorsal view more than 7 times smaller than distance between eyes); propleurae with a longitudinal ridge bearing a row of setae parallel to lateral margin of pronotum; disc of pronotum with more-or-less developed tubercles, ridges, and excavations in males, and smooth in females; apical outer tooth of protibia in males directed laterally, at reasonable angle to internal margin of tibia (
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<figureCitation id="4BDD1093FFDA6161FD1F6725FD5AFA7B" box="[650,720,1401,1422]" captionStart="FIGURE 30" captionStartId="60.[151,250,932,954]" captionTargetBox="[193,1371,217,907]" captionTargetId="figure@60.[193,1393,193,910]" captionTargetPageId="60" captionText="FIGURE 30. Triodontus hova. Abdomen of male (A), syntype of O. nigrita, habitus in dorsal view (B), protibia (C, male left, female right)." httpUri="https://zenodo.org/record/200473/files/figure.png" pageId="11" pageNumber="12">Fig. 30</figureCitation>
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C).......................................
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<taxonomicName id="14E67795FFDA6161FB5C6725FA10FA7B" authority="Westwood" authorityName="Westwood" box="[1225,1434,1401,1422]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="E192D004FFDA6161FB5C6725FABBFA7B" box="[1225,1329,1401,1422]" italics="true" pageId="11" pageNumber="12">Triodontus</emphasis>
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Westwood
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="D3590C16FFDA6161FF0267CAFA16FA13" blockId="11.[151,1436,1049,1510]" pageId="11" pageNumber="12">
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<keyLead id="6817AF23FFDA6161FF0267CAFA16FA13" pageId="11" pageNumber="12">
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- Eyes larger (width of eye in dorsal view less than 4 times smaller than distance between eyes); propleurae without a longitudinal ridge, only with a sparser row of setae; disc of pronotum with 2 feebly developed tubercles in middle or without tubercles; apical outer tooth of protibia almost parallel to internal margin of tibia (
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<figureCitation id="4BDD1093FFDA6161FC1A678DFC5EFA13" box="[911,980,1489,1510]" captionStart="FIGURE 28" captionStartId="55.[151,250,1729,1751]" captionTargetBox="[192,1393,194,1698]" captionTargetId="figure@55.[191,1396,193,1708]" captionTargetPageId="55" captionText="FIGURE 28. Renorphnus clementi. Habitus in dorsal view (A—male, B—female), aedeagus in lateral view and parameres in dorsal view (C), internal sac of aedeagus (D), distributional records (E), protibia (F, male left, female right)." httpUri="https://zenodo.org/record/272644/files/figure.png" pageId="11" pageNumber="12">Fig. 28</figureCitation>
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F)........ ..
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<taxonomicName id="14E67795FFDA6161FBF1678DFA16FA13" authority="Frolov & Montreuil" authorityName="Frolov & Montreuil" box="[1124,1436,1489,1510]" class="Insecta" family="Orphnidae" genus="Renorphnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="E192D004FFDA6161FBF1678DFB52FA13" box="[1124,1240,1489,1510]" italics="true" pageId="11" pageNumber="12">Renorphnus</emphasis>
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Frolov & Montreuil
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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</subSubSection>
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</treatment>
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</document>
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