Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood Author Frolov, Andrey V. Author Montreuil, Olivier Author Akhmetova, Lilia A. text Zootaxa 2016 4207 1 1 93 journal article 37549 10.11646/zootaxa.4207.1.1 cef35771-ff95-4a41-9af6-196dea899a64 1175-5326 200444 F2875582-31E2-496F-AEEF-1D657DD86C33 Key to the Madagascar Orphninae genera (both sexes) 1. Protibiae with 2 outer teeth apically and 1 much smaller outer tooth basally ( Fig. 7 D)........... Pseudorphnus Benderitter - Protibiae with 3 outer teeth in apical half ( Figs. 10 H, 30C, 28F).................................................2 2. Body length smaller (usually less than 7 mm ); lateral margin of pronotum with 4 long setae: 2 near anterior margin, 1 in the middle, and 1 near posterior margin; some or all these setae may be abraded, but 4 chaetiferous punctures are always distinct; left and right mandibles may differ strongly in length in males, and frontoclypeus is somewhat asymmetrical in such cases; disc of pronotum smooth, without processes or excavations.................................. Madecorphnus Paulian - Body length larger (usually longer than 7 mm ); lateral margin of pronotum with numerous setae (at least more than 10) evenly distributed along the margin, setae may differ strongly in length; left and right mandibles of about same length, frontoclypeus symmetrical; disc of pronotum may be with more-or-less developed tubercles, ridges, and excavations in males........... 3 3. Eyes smaller (width of eye in dorsal view more than 7 times smaller than distance between eyes); propleurae with a longitudinal ridge bearing a row of setae parallel to lateral margin of pronotum; disc of pronotum with more-or-less developed tubercles, ridges, and excavations in males, and smooth in females; apical outer tooth of protibia in males directed laterally, at reasonable angle to internal margin of tibia ( Fig. 30 C)....................................... Triodontus Westwood - Eyes larger (width of eye in dorsal view less than 4 times smaller than distance between eyes); propleurae without a longitudinal ridge, only with a sparser row of setae; disc of pronotum with 2 feebly developed tubercles in middle or without tubercles; apical outer tooth of protibia almost parallel to internal margin of tibia ( Fig. 28 F)........ .. Renorphnus Frolov & Montreuil