treatments-xml/data/54/22/87/542287ABFFBA41202CD644B54FFEFC89.xml

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<mods:title id="8718E0A43F95D3DBC898E866D9C3CB5C">A revision of the Carebara pygmaea (Emery, 1887) species group (Hymenoptera: Formicidae) in Thailand</mods:title>
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<mods:namePart id="11C535E72B35B097BC24CAEE214E4C76">Wimolsuthikul, Chaipat</mods:namePart>
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<mods:namePart id="BF386B4BF475EF2CBB766D49A31BC6C7">Yamane, Seiki</mods:namePart>
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<mods:affiliation id="5FFA6AFC779A9450E29D52BE97EA8CA4">Khlong &amp; Department of Medical Entomology, Faculty of Tropical Medicine, Mahidol University, Bangkok 10400, Thailand.</mods:affiliation>
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<mods:namePart id="5ABB97FE7DA21F82D023197BA0E03BFD">Jaitrong, Weeyawat</mods:namePart>
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<taxonomicName id="1B8B4D3EFFBA412F2CD644B54EB3FF2C" ID-CoL="CLG6K" authority="Matsuura &amp; Hosoishi, 2024" authorityName="Matsuura &amp; Hosoishi" authorityYear="2024" box="[151,755,151,178]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="85" phylum="Arthropoda" rank="species" species="bokorensis">
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<emphasis id="EEFFEAAFFFBA412F2CD644B54DD3FF2F" bold="true" box="[151,403,151,178]" italics="true" pageId="6" pageNumber="85">Carebara bokorensis</emphasis>
<bibRefCitation id="B81A4B4CFFBA412F2DDB44B54EB3FF2C" author="Matsuura, K. &amp; Hosoishi, S. &amp; Sokh, H." box="[410,755,151,177]" pageId="6" pageNumber="85" pagination="1 - 9" refId="ref12643" refString="Matsuura, K., Hosoishi, S. &amp; Sokh, H. (2024) A remarkable new species from the pygmaea species group of the genus Carebara Westwood, 1840 (Hymenoptera: Formicidae: Myrmicinae) from Cambodia. Far Eastern Entomologist, 496, 1 - 9. https: // doi. org / 10.25221 / fee. 496.1" type="journal article" year="2024">Matsuura &amp; Hosoishi, 2024</bibRefCitation>
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<paragraph id="DC3436BDFFBA412F2CD644994D52FF4B" blockId="6.[151,755,151,214]" box="[151,274,187,214]" pageId="6" pageNumber="85">
(
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)
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<taxonomicName id="1B8B4D3EFFBA412F2CD644D84E0FFE8F" ID-CoL="CLG6K" authority="Matsuura &amp; Hosoishi" authorityName="Matsuura &amp; Hosoishi" authorityYear="2024" box="[151,591,250,274]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="85" phylum="Arthropoda" rank="species" species="bokorensis">
<emphasis id="EEFFEAAFFFBA412F2CD644D84D2CFE8F" box="[151,364,250,274]" italics="true" pageId="6" pageNumber="85">Carebara bokorensis</emphasis>
Matsuura &amp; Hosoishi
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, in 
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<bibRefCitation id="B81A4B4CFFBA412F2E3244D94F20FE8F" author="Matsuura, K. &amp; Hosoishi, S. &amp; Sokh, H." box="[627,864,250,274]" pageId="6" pageNumber="85" pagination="1 - 9" refId="ref12643" refString="Matsuura, K., Hosoishi, S. &amp; Sokh, H. (2024) A remarkable new species from the pygmaea species group of the genus Carebara Westwood, 1840 (Hymenoptera: Formicidae: Myrmicinae) from Cambodia. Far Eastern Entomologist, 496, 1 - 9. https: // doi. org / 10.25221 / fee. 496.1" type="journal article" year="2024">
Matsuura 
<emphasis id="EEFFEAAFFFBA412F2E9844D94F46FE8F" box="[729,774,250,274]" italics="true" pageId="6" pageNumber="85">et al</emphasis>
. 2024: 3
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, figs. 1–2.
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<paragraph id="DC3436BDFFBA412F2CD645194854FE78" blockId="6.[151,1436,315,486]" pageId="6" pageNumber="85">
<emphasis id="EEFFEAAFFFBA412F2CD645194D0AFEC8" bold="true" box="[151,330,315,341]" pageId="6" pageNumber="85">Type materials</emphasis>
: 
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<typeStatus id="0330881FFFBA412F2D1845194D81FEC8" box="[345,449,315,341]" pageId="6" pageNumber="85" type="holotype">Holotype</typeStatus>
major worker (
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, collection code KUECCRB001, images in 
<bibRefCitation id="B81A4B4CFFBA412F28F3451E49DCFEC8" author="Matsuura, K. &amp; Hosoishi, S. &amp; Sokh, H." box="[1202,1436,315,342]" pageId="6" pageNumber="85" pagination="1 - 9" refId="ref12643" refString="Matsuura, K., Hosoishi, S. &amp; Sokh, H. (2024) A remarkable new species from the pygmaea species group of the genus Carebara Westwood, 1840 (Hymenoptera: Formicidae: Myrmicinae) from Cambodia. Far Eastern Entomologist, 496, 1 - 9. https: // doi. org / 10.25221 / fee. 496.1" type="journal article" year="2024">
Matsuura 
<emphasis id="EEFFEAAFFFBA412F2965451E4916FEC8" box="[1316,1366,315,341]" italics="true" pageId="6" pageNumber="85">et al</emphasis>
. 2024
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were examined), 
<specimenCount id="CA8DFD34FFBA412F2D24457D4DA0FEE7" box="[357,480,351,378]" count="2" pageId="6" pageNumber="85" type="generic" typeStatus="paratype">2 paratype</specimenCount>
major workers (
<collectionCode id="BA9AAE78FFBA412F2EEA457D4EBCFEE4" box="[683,764,351,377]" country="Japan" httpUri="http://grbio.org/cool/txbq-e9hf" name="Kyushu University Entomology Collection" pageId="6" pageNumber="85">KUEC</collectionCode>
, collection code KUECCRB002, KUECCRB003), and 
<specimenCount id="CA8DFD34FFBA412F29CF45424CB7FE03" count="7" pageId="6" pageNumber="85" type="generic" typeStatus="paratype">7 paratype</specimenCount>
minor workers (
<collectionCode id="BA9AAE78FFBA412F2DEC45A14DBEFE00" box="[429,510,387,413]" country="Japan" httpUri="http://grbio.org/cool/txbq-e9hf" name="Kyushu University Entomology Collection" pageId="6" pageNumber="85">KUEC</collectionCode>
, collection code KUECCRB004, KUECCRB005, KUECCRB006, KUECCRB007, KUECCRB008, KUECCRB009, KUECCRB010), 
<collectingCountry id="A49C762DFFBA412F2E9845854F0FFE5C" box="[729,847,423,449]" name="Cambodia" pageId="6" pageNumber="85">Cambodia</collectingCountry>
, 
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, 
<quantity id="1B739B58FFBA412F2814458548D4FE5C" box="[1109,1172,423,449]" metricMagnitude="2" metricUnit="m" metricValue="6.63" pageId="6" pageNumber="85" unit="m" value="663.0">
<elevation id="57A6D18EFFBA412F2814458548D4FE5C" box="[1109,1172,423,449]" metricMagnitude="2" metricUnit="m" metricValue="6.63" pageId="6" pageNumber="85" unit="m" value="663.0">663m</elevation>
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alt., 
<collectingRegion id="1E4FF85FFFBA412F288E458A49D8FE5C" box="[1231,1432,423,450]" country="Cambodia" name="Kampot" pageId="6" pageNumber="85">Kampot Province</collectingRegion>
, 
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, 
<geoCoordinate id="B9BF507AFFBA412F2CBC45E94D22FE78" box="[253,354,459,485]" degrees="104" direction="east" minutes="06" orientation="longitude" pageId="6" pageNumber="85" precision="925" value="104.1">104°06E</geoCoordinate>
, 
<date id="A835107DFFBA412F2D2C45E94DB7FE78" box="[365,503,459,485]" pageId="6" pageNumber="85" value="2011-12-19">
<collectingDate id="B871E995FFBA412F2D2C45E94DB7FE78" box="[365,503,459,485]" pageId="6" pageNumber="85" value="2011-12-19">19.XII.2011</collectingDate>
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, 
<collectorName id="717E536BFFBA412F2E4245E94EC4FE78" box="[515,644,459,485]" pageId="6" pageNumber="85">S. Hosoishi</collectorName>
leg. SH11-Cam-55, decayed wood.
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</paragraph>
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<emphasis id="EEFFEAAFFFBA412F2CD6439A4D52F84D" bold="true" box="[151,274,1976,2000]" pageId="6" pageNumber="85">FIGURE 3.</emphasis>
<taxonomicName id="1B8B4D3EFFBA412F2D59439A4DADF84D" authorityName="Matsuura &amp; Hosoishi" authorityYear="2024" box="[280,493,1976,2000]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="85" phylum="Arthropoda" rank="species" species="bokorensis">
<emphasis id="EEFFEAAFFFBA412F2D59439A4DADF84D" box="[280,493,1976,2000]" italics="true" pageId="6" pageNumber="85">Carebara bokorensis</emphasis>
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, non-type major worker (THNHM-I-00028081). A, Body in profile view; B, head in full-face view; C, body in dorsal view.
</paragraph>
</caption>
<caption id="88F46635FFBB412E2CD642484DD0F93B" ID-DOI="http://doi.org/10.5281/zenodo.12742651" ID-Zenodo-Dep="12742651" httpUri="https://zenodo.org/record/12742651/files/figure.png" pageId="7" pageNumber="86" startId="7.[151,250,1642,1666]" targetBox="[151,1436,181,1618]" targetPageId="7" targetType="figure">
<paragraph id="DC3436BDFFBB412E2CD642484DD0F93B" blockId="7.[151,1436,1642,1702]" pageId="7" pageNumber="86">
<emphasis id="EEFFEAAFFFBB412E2CD642484D54F91F" bold="true" box="[151,276,1642,1666]" pageId="7" pageNumber="86">FIGURE 4.</emphasis>
<taxonomicName id="1B8B4D3EFFBB412E2D5A42484DB1F91F" authorityName="Matsuura &amp; Hosoishi" authorityYear="2024" box="[283,497,1642,1666]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="86" phylum="Arthropoda" rank="species" species="bokorensis">
<emphasis id="EEFFEAAFFFBB412E2D5A42484DB1F91F" box="[283,497,1642,1666]" italics="true" pageId="7" pageNumber="86">Carebara bokorensis</emphasis>
</taxonomicName>
, ergatoid queen (THNHM-I-00028220). A, Body in profile view; B, head in full-face view; 2C, body in dorsal view.
</paragraph>
</caption>
<paragraph id="DC3436BDFFBB41212C8642DA4871FF64" blockId="7.[151,1437,1784,2027]" lastBlockId="8.[151,1437,150,501]" lastPageId="8" lastPageNumber="87" pageId="7" pageNumber="86">
<emphasis id="EEFFEAAFFFBB412E2C8642DA4EE7F88F" bold="true" box="[199,679,1784,1810]" pageId="7" pageNumber="86">Non-type material examined. Western</emphasis>
<materialsCitation id="6CE33CE0FFBB412E2EF442DA4FEBF85B" ID-GBIF-Occurrence="4906507342" collectingDate="2020-07-28" collectionCode="THNHM, SKYC" collectorName="W. Jaitrong &amp; Eastern" country="Thailand" county="Suan Phueng District" elevation="1005" latitude="13.53884" location="Hill Evergreen Forest" longLatPrecision="1" longitude="99.20996" municipality="Khao Lam" pageId="7" pageNumber="86" specimenCode="THNHM-I-00028081, THNHM-I-00028201, THNHM-I-00028202, THNHM-I-00028203, THNHM-I-00028216, THNHM-I-00028220, THNHM-I-00028215, THNHM-I-00028204, THNHM-I-00028213, THNHM-I-00028214" specimenCount="1" stateProvince="Ratchaburi">
<emphasis id="EEFFEAAFFFBB412E2EF442DA4F6CF88F" bold="true" box="[693,812,1784,1810]" pageId="7" pageNumber="86">
<collectingCountry id="A49C762DFFBB412E2EF442DA4F66F88F" box="[693,806,1784,1810]" name="Thailand" pageId="7" pageNumber="86">Thailand</collectingCountry>
:
</emphasis>
4 major worker (
<specimenCode id="8C2D9EC6FFBB412E284E42DA4946F88F" box="[1039,1286,1784,1810]" collectionCode="THNHM-I-" pageId="7" pageNumber="86">THNHM-I-00028081</specimenCode>
, 
<specimenCode id="8C2D9EC6FFBB412E295B42DB4D47F8AB" collectionCode="THNHM-I-" pageId="7" pageNumber="86">THNHM-I-00028201</specimenCode>
to 
<specimenCode id="8C2D9EC6FFBB412E2D78433E4E6FF8AB" box="[313,559,1820,1846]" collectionCode="THNHM-I-" pageId="7" pageNumber="86">THNHM-I-00028202</specimenCode>
, 
<collectionCode id="BA9AAE78FFBB412E2E00433F4EEBF8AA" box="[577,683,1821,1847]" pageId="7" pageNumber="86">THNHM</collectionCode>
; 
<specimenCode id="8C2D9EC6FFBB412E2EFC433E4FF2F8AB" box="[701,946,1820,1846]" collectionCode="THNHM-I-" pageId="7" pageNumber="86">THNHM-I-00028203</specimenCode>
, 
<collectionCode id="BA9AAE78FFBB412E2F84433E4854F8AB" box="[965,1044,1820,1846]" pageId="7" pageNumber="86">SKYC</collectionCode>
), 6 ergatoid queens (
<specimenCode id="8C2D9EC6FFBB412E2959433E4D47F8C7" collectionCode="THNHM-I-" pageId="7" pageNumber="86">THNHM-I-00028216</specimenCode>
to 
<specimenCode id="8C2D9EC6FFBB412E2D7343624E67F8C7" box="[306,551,1856,1882]" collectionCode="THNHM-I-" pageId="7" pageNumber="86">THNHM-I-00028220</specimenCode>
, 
<collectionCode id="BA9AAE78FFBB412E2E7443634EDFF8C6" box="[565,671,1857,1883]" pageId="7" pageNumber="86">THNHM</collectionCode>
; 
<specimenCode id="8C2D9EC6FFBB412E2EEF43624FE3F8C6" box="[686,931,1856,1883]" collectionCode="THNHM-I-" pageId="7" pageNumber="86">THNHM-I-00028215</specimenCode>
, 
<collectionCode id="BA9AAE78FFBB412E2FF343624841F8C7" box="[946,1025,1856,1882]" pageId="7" pageNumber="86">SKYC</collectionCode>
), and 12 minor workers (THNHM-I-00028082, 
<specimenCode id="8C2D9EC6FFBB412E2D6843464E5BF8E3" box="[297,539,1892,1918]" collectionCode="THNHM-I-" pageId="7" pageNumber="86">THNHM-I-00028204</specimenCode>
to 
<specimenCode id="8C2D9EC6FFBB412E2E0243464F78F8E3" box="[579,824,1892,1918]" collectionCode="THNHM-I-" pageId="7" pageNumber="86">THNHM-I-00028213</specimenCode>
, 
<collectionCode id="BA9AAE78FFBB412E2F0543474FEEF8E2" box="[836,942,1893,1919]" pageId="7" pageNumber="86">THNHM</collectionCode>
; 
<specimenCode id="8C2D9EC6FFBB412E2FFA434648F1F8E3" box="[955,1201,1892,1918]" collectionCode="THNHM-I-" pageId="7" pageNumber="86">THNHM-I-00028214</specimenCode>
, 
<collectionCode id="BA9AAE78FFBB412E28FF4346494DF8E3" box="[1214,1293,1892,1918]" pageId="7" pageNumber="86">SKYC</collectionCode>
), 
<collectingRegion id="1E4FF85FFFBB412E296043464CBDF83F" country="Thailand" name="Ratchaburi" pageId="7" pageNumber="86">Ratchaburi Province</collectingRegion>
, 
<collectingCounty id="35554E31FFBB412E2D4C43AA4E47F83F" box="[269,519,1928,1954]" pageId="7" pageNumber="86">Suan Phueng District</collectingCounty>
, 
<collectingMunicipality id="3C50ACC7FFBB412E2E5643AA4ED5F83E" box="[535,661,1928,1955]" pageId="7" pageNumber="86">Khao Lam</collectingMunicipality>
, 
<location id="D9546066FFBB412E2EE443AA4FE3F83E" LSID="urn:lsid:plazi:treatment:542287ABFFBA41202CD644B54FFEFC89:D9546066FFBB412E2EE443AA4FE3F83E" box="[677,931,1928,1955]" country="Thailand" county="Suan Phueng District" latitude="13.53884" longLatPrecision="1" longitude="99.20996" municipality="Khao Lam" name="Hill Evergreen Forest" pageId="7" pageNumber="86" stateProvince="Ratchaburi">Hill Evergreen Forest</location>
, 
<geoCoordinate id="B9BF507AFFBB412E2FF343AA4809F83E" box="[946,1097,1928,1955]" degrees="13.538840" direction="north" orientation="latitude" pageId="7" pageNumber="86" precision="1" value="13.53884">13.538840ºN</geoCoordinate>
, 
<geoCoordinate id="B9BF507AFFBB412E281843AA48ACF83F" box="[1113,1260,1928,1954]" degrees="99.209962" direction="east" orientation="longitude" pageId="7" pageNumber="86" precision="1" value="99.20996">99.209962ºE</geoCoordinate>
, 
<elevation id="57A6D18EFFBB412E28BD43AA49D7F83F" box="[1276,1431,1928,1955]" metricMagnitude="3" metricUnit="m" metricValue="1.005" pageId="7" pageNumber="86" unit="m" value="1005.0">
<quantity id="1B739B58FFBB412E28BD43AA4916F83E" box="[1276,1366,1928,1955]" metricMagnitude="3" metricUnit="m" metricValue="1.005" pageId="7" pageNumber="86" unit="m" value="1005.0">1005 m</quantity>
a.s.l.
</elevation>
, 
<location id="D9546066FFBB412E2CD6438E4CABF85B" LSID="urn:lsid:plazi:treatment:542287ABFFBA41202CD644B54FFEFC89:D9546066FFBB412E2CD6438E4CABF85B" box="[151,235,1964,1990]" country="Thailand" county="Suan Phueng District" latitude="13.53884" longLatPrecision="1" longitude="99.20996" municipality="Khao Lam" name="Rotting" pageId="7" pageNumber="86" stateProvince="Ratchaburi">Rotting</location>
log, 
<date id="A835107DFFBB412E2D6B438E4DF6F85B" box="[298,438,1964,1990]" pageId="7" pageNumber="86" value="2020-07-28">
<collectingDate id="B871E995FFBB412E2D6B438E4DF6F85B" box="[298,438,1964,1990]" pageId="7" pageNumber="86" value="2020-07-28">28.VII.2020</collectingDate>
</date>
, 
<collectorName id="717E536BFFBB412E2D85438F4E07F85B" box="[452,583,1964,1991]" pageId="7" pageNumber="86">W. Jaitrong</collectorName>
leg., TH20-WJT-082. 
<collectorName id="717E536BFFBB412E2F0F438E4FEBF85B" box="[846,939,1964,1990]" pageId="7" pageNumber="86">
<emphasis id="EEFFEAAFFFBB412E2F0F438E4FEBF85B" bold="true" box="[846,939,1964,1990]" pageId="7" pageNumber="86">Eastern</emphasis>
</collectorName>
</materialsCitation>
<materialsCitation id="6CE33CE0FFBB41212FF4438E4DB8FF2C" ID-GBIF-Occurrence="4906507315" collectingDate="2008-01-19" collectionCode="THNHM" collectorName="W. Jaitrong" country="Thailand" county="Soi Dao District" elevation="285" lastPageId="8" lastPageNumber="87" latitude="13.106065" location="Khao Soi Dao" longLatPrecision="1" longitude="102.19612" municipality="Sai Khao Subdistrict" pageId="7" pageNumber="86" specimenCount="1" stateProvince="Chanthaburi">
<emphasis id="EEFFEAAFFFBB412E2FF4438E486BF85B" bold="true" box="[949,1067,1964,1990]" pageId="7" pageNumber="86">
<collectingCountry id="A49C762DFFBB412E2FF4438E4866F85B" box="[949,1062,1964,1990]" name="Thailand" pageId="7" pageNumber="86">Thailand</collectingCountry>
:
</emphasis>
<collectingRegion id="1E4FF85FFFBB412E2874438E496FF85B" box="[1077,1327,1964,1990]" country="Thailand" name="Chanthaburi" pageId="7" pageNumber="86">Chanthaburi Province</collectingRegion>
, 
<collectingCounty id="35554E31FFBB412E297F438E4CAEF877" pageId="7" pageNumber="86">Soi Dao District</collectingCounty>
, 
<collectingMunicipality id="3C50ACC7FFBB412E2CB943F24DA1F877" box="[248,481,2000,2026]" pageId="7" pageNumber="86">Sai Khao Subdistrict</collectingMunicipality>
, near 
<location id="D9546066FFBB412E2E6043F24E81F876" LSID="urn:lsid:plazi:treatment:542287ABFFBA41202CD644B54FFEFC89:D9546066FFBB412E2E6043F24E81F876" box="[545,705,2000,2027]" country="Thailand" county="Soi Dao District" latitude="13.106065" longLatPrecision="1" longitude="102.19612" municipality="Sai Khao Subdistrict" name="Khao Soi Dao" pageId="7" pageNumber="86" stateProvince="Chanthaburi">Khao Soi Dao</location>
, 
<geoCoordinate id="B9BF507AFFBB412E2E8A43F24F22F876" box="[715,866,2000,2027]" degrees="13.106065" direction="north" orientation="latitude" pageId="7" pageNumber="86" precision="1" value="13.106065">13.106065ºN</geoCoordinate>
, 
<geoCoordinate id="B9BF507AFFBB412E2F2D43F2484DF877" box="[876,1037,2000,2026]" degrees="102.196120" direction="east" orientation="longitude" pageId="7" pageNumber="86" precision="1" value="102.19612">102.196120ºE</geoCoordinate>
, 
<elevation id="57A6D18EFFBB412E285943F248DAF877" box="[1048,1178,2000,2027]" metricMagnitude="2" metricUnit="m" metricValue="2.85" pageId="7" pageNumber="86" unit="m" value="285.0">
<quantity id="1B739B58FFBB412E285943F2481EF876" box="[1048,1118,2000,2027]" metricMagnitude="2" metricUnit="m" metricValue="2.85" pageId="7" pageNumber="86" unit="m" value="285.0">285 m</quantity>
a.s.l.
</elevation>
, 
<date id="A835107DFFBB412E28E443F24953F877" box="[1189,1299,2000,2026]" pageId="7" pageNumber="86" value="2008-01-19">
<collectingDate id="B871E995FFBB412E28E443F24953F877" box="[1189,1299,2000,2026]" pageId="7" pageNumber="86" value="2008-01-19">19.I.2008</collectingDate>
</date>
, 
<collectorName id="717E536BFFBB412E295C43F349DCF877" box="[1309,1436,2000,2027]" pageId="7" pageNumber="86">W. Jaitrong</collectorName>
leg., TH08-WJT-91 (
<collectionCode id="BA9AAE78FFB441212DC244B54DB0FF2C" box="[387,496,151,177]" pageId="8" pageNumber="87">THNHM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="6CE33CE0FFB441212E4444B54F29FF48" ID-GBIF-Occurrence="4906507331" collectingDate="2005-08-27" collectionCode="THNHM" collectorName="W. Jaitrong" country="Thailand" elevation="153" latitude="13.371229" location="Thatakeab District" longLatPrecision="1" longitude="101.8176" pageId="8" pageNumber="87" specimenCount="1" stateProvince="Chachoengsao">
<collectingRegion id="1E4FF85FFFB441212E4444B54F56FF2C" box="[517,790,151,177]" country="Thailand" name="Chachoengsao" pageId="8" pageNumber="87">Chachoengsao Province</collectingRegion>
, 
<location id="D9546066FFB441212F6244B54FB7FF2C" LSID="urn:lsid:plazi:treatment:542287ABFFBA41202CD644B54FFEFC89:D9546066FFB441212F6244B54FB7FF2C" box="[803,1015,151,177]" country="Thailand" latitude="13.371229" longLatPrecision="1" longitude="101.8176" name="Thatakeab District" pageId="8" pageNumber="87" stateProvince="Chachoengsao">Thatakeab District</location>
, 
<geoCoordinate id="B9BF507AFFB44121284244B548DAFF2C" box="[1027,1178,151,177]" degrees="13.371229" direction="north" orientation="latitude" pageId="8" pageNumber="87" precision="1" value="13.371229">13.371229ºN</geoCoordinate>
, 
<geoCoordinate id="B9BF507AFFB4412128E744B44908FF2C" box="[1190,1352,150,177]" degrees="101.817599" direction="east" orientation="longitude" pageId="8" pageNumber="87" precision="1" value="101.8176">101.817599ºE</geoCoordinate>
, 
<elevation id="57A6D18EFFB44121291544B44C8DFF48" metricMagnitude="2" metricUnit="m" metricValue="1.53" pageId="8" pageNumber="87" unit="m" value="153.0">
<quantity id="1B739B58FFB44121291544B449DCFF2C" box="[1364,1436,150,177]" metricMagnitude="2" metricUnit="m" metricValue="1.53" pageId="8" pageNumber="87" unit="m" value="153.0">153 m</quantity>
a.s.l.
</elevation>
, 
<date id="A835107DFFB441212C9844984D2EFF48" box="[217,366,186,213]" pageId="8" pageNumber="87" value="2005-08-27">
<collectingDate id="B871E995FFB441212C9844984D2EFF48" box="[217,366,186,213]" pageId="8" pageNumber="87" value="2005-08-27">27.VIII.2005</collectingDate>
</date>
, 
<collectorName id="717E536BFFB441212D3844994DB8FF48" box="[377,504,187,213]" pageId="8" pageNumber="87">W. Jaitrong</collectorName>
leg., WJT270805-01 (
<collectionCode id="BA9AAE78FFB441212EB544994F21FF48" box="[756,865,187,213]" pageId="8" pageNumber="87">THNHM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="6CE33CE0FFB441212F354499486DFF64" ID-GBIF-Occurrence="4906507307" collectingDate="2003-08-21" collectionCode="SKYC" collectorName="Khao Ang Reu Nai WS &amp; Sk. Yamane" country="Thailand" location="Lumchangwat" pageId="8" pageNumber="87" specimenCount="1" stateProvince="Chachoengsao">
<collectingRegion id="1E4FF85FFFB441212F35449948C4FF48" box="[884,1156,187,213]" country="Thailand" name="Chachoengsao" pageId="8" pageNumber="87">Chachoengsao Province</collectingRegion>
, 
<collectorName id="717E536BFFB4412128D1449949D8FF48" box="[1168,1432,187,213]" pageId="8" pageNumber="87">Khao Ang Reu Nai WS</collectorName>
, 
<location id="D9546066FFB441212CD644FD4D77FF64" LSID="urn:lsid:plazi:treatment:542287ABFFBA41202CD644B54FFEFC89:D9546066FFB441212CD644FD4D77FF64" box="[151,311,223,249]" country="Thailand" name="Lumchangwat" pageId="8" pageNumber="87" stateProvince="Chachoengsao">Lumchangwat</location>
stn., 
<date id="A835107DFFB441212D3544FD4E49FF64" box="[372,521,223,249]" pageId="8" pageNumber="87" value="2003-08-21">
<collectingDate id="B871E995FFB441212D3544FD4E49FF64" box="[372,521,223,249]" pageId="8" pageNumber="87" value="2003-08-21">21.VIII.2003</collectingDate>
</date>
, 
<collectorName id="717E536BFFB441212E5544FD4ED9FF64" box="[532,665,223,249]" pageId="8" pageNumber="87">Sk. Yamane</collectorName>
leg., TH03-SKY-27 &amp; -44 (
<collectionCode id="BA9AAE78FFB441212F9544FD4865FF64" box="[980,1061,223,249]" pageId="8" pageNumber="87">SKYC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="94916536FFB441212C8645214CA5FE68" pageId="8" pageNumber="87" type="description">
<paragraph id="DC3436BDFFB441212C8645214CA5FE68" blockId="8.[151,1437,150,501]" pageId="8" pageNumber="87">
<emphasis id="EEFFEAAFFFB441212C8645214EDEFE80" bold="true" box="[199,670,258,285]" pageId="8" pageNumber="87">Measurements (in mm). Major worker</emphasis>
(n = 4): TL 3.7–3.99, HL 0.92–0.97, HW 0.84–0.96, SL 0.4–0.46, ED 0.06–0.07, ML 0.83–0.86, MW 0.5–0.53, DPW 0.23–0.26, PL 0.36–0.4, 
<collectingCountry id="A49C762DFFB44121284E45054873FEDC" box="[1039,1075,295,321]" name="Philippines" pageId="8" pageNumber="87">PH</collectingCountry>
0.26–0.3, CI 91–93, SI 47–52, PI1 133–138, PI2 64–67. 
<emphasis id="EEFFEAAFFFB441212DFC45684E29FEF8" bold="true" box="[445,617,330,357]" pageId="8" pageNumber="87">Minor worker</emphasis>
(n = 5): TL 1.52–1.62, HL 0.33–0.36, HW 0.3–0.33, SL 0.23–0.26, ED 0.01–0.02, ML 0.33–0.36, MW 0.20–0.21, DPW 0.07–0.1, PL 0.17–0.2, 
<collectingCountry id="A49C762DFFB441212F9E454D4843FE14" box="[991,1027,367,393]" name="Philippines" pageId="8" pageNumber="87">PH</collectingCountry>
0.1–0.13, CI 90–91, SI 89–90, PI1 150–167, PI2 40–60. 
<emphasis id="EEFFEAAFFFB441212DC745B04E7EFE30" bold="true" box="[390,574,402,429]" pageId="8" pageNumber="87">Ergatoid queen</emphasis>
(n = 5): TL 3.76–4.03, HL 0.99–1.02, HW 0.89–0.92, SL 0.43–0.46, ED 0.07, ML 0.86–0.89, MW 0.53–0.56, DPW 0.26–0.3, PL 0.4–0.43, 
<collectingCountry id="A49C762DFFB441212F0C45954F31FE4C" box="[845,881,439,465]" name="Philippines" pageId="8" pageNumber="87">PH</collectingCountry>
0.3–0.33, CI 90–91, SI 46–52, PI1 130–133, PI2 61–69.
</paragraph>
</subSubSection>
<caption id="88F46635FFB441212CD642E24D8EF961" ID-DOI="http://doi.org/10.5281/zenodo.12742653" ID-Zenodo-Dep="12742653" httpUri="https://zenodo.org/record/12742653/files/figure.png" pageId="8" pageNumber="87" startId="8.[151,250,1728,1752]" targetBox="[206,1381,525,1705]" targetPageId="8" targetType="figure">
<paragraph id="DC3436BDFFB441212CD642E24D8EF961" blockId="8.[151,1436,1728,1788]" pageId="8" pageNumber="87">
<emphasis id="EEFFEAAFFFB441212CD642E24D52F945" bold="true" box="[151,274,1728,1752]" pageId="8" pageNumber="87">FIGURE 5.</emphasis>
<taxonomicName id="1B8B4D3EFFB441212D5942E24DACF945" authorityName="Matsuura &amp; Hosoishi" authorityYear="2024" box="[280,492,1728,1752]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="87" phylum="Arthropoda" rank="species" species="bokorensis">
<emphasis id="EEFFEAAFFFB441212D5942E24DACF945" box="[280,492,1728,1752]" italics="true" pageId="8" pageNumber="87">Carebara bokorensis</emphasis>
</taxonomicName>
, non-type minor worker (THNHM-I-00028204). A, Body in profile view; B, head in full-face view; 2C, body in dorsal view.
</paragraph>
</caption>
<subSubSection id="94916536FFB441202C86436C496EFE81" lastPageId="9" lastPageNumber="88" pageId="8" pageNumber="87" type="diagnosis">
<paragraph id="DC3436BDFFB441212C86436C4EDFF865" blockId="8.[151,1437,1870,2041]" pageId="8" pageNumber="87">
<emphasis id="EEFFEAAFFFB441212C86436C4DD6F8F5" bold="true" box="[199,406,1870,1896]" pageId="8" pageNumber="87">Diagnosis. Major</emphasis>
(
<figureCitation id="44B02A38FFB441212DE6436C4DADF8F5" box="[423,493,1870,1896]" captionStart="FIGURE 3" captionStartId="6.[151,250,1976,2000]" captionTargetBox="[157,1429,509,1946]" captionTargetId="figure-104@6.[157,1429,509,1946]" captionTargetPageId="6" captionText="FIGURE 3. Carebara bokorensis, non-type major worker (THNHM-I-00028081).A, Body in profile view; B, head in full-face view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742649" httpUri="https://zenodo.org/record/12742649/files/figure.png" pageId="8" pageNumber="87">Fig. 3</figureCitation>
): head, promesonotum, and gastral tergite I with dense and long erect hairs mixed with decumbent hairs; with head in full-face view, posterior one-third with dense fovea, lacking transverse striation, while anterior two-thirds with longitudinal striation; eye small, with ca. 13–15 ommatidia; gastral tergite I punctate. 
<emphasis id="EEFFEAAFFFB441212CD643984CA3F849" bold="true" box="[151,227,1978,2004]" pageId="8" pageNumber="87">Minor</emphasis>
(
<figureCitation id="44B02A38FFB441212CB643984D7EF848" box="[247,318,1978,2005]" captionStart="FIGURE 5" captionStartId="8.[151,250,1728,1752]" captionTargetBox="[206,1381,525,1705]" captionTargetId="figure-159@8.[206,1381,525,1705]" captionTargetPageId="8" captionText="FIGURE 5. Carebara bokorensis, non-type minor worker (THNHM-I-00028204).A, Body in profile view; B, head in full-face view; 2C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742653" httpUri="https://zenodo.org/record/12742653/files/figure.png" pageId="8" pageNumber="87">Fig. 5</figureCitation>
): propodeal dorsum entirely punctate; petiole and postpetiole with decumbent hairs; propodeal junction roundly convex; body reddish brown.
</paragraph>
<paragraph id="DC3436BDFFB541202C8644B4496EFE81" blockId="9.[151,1436,150,789]" pageId="9" pageNumber="88">
<emphasis id="EEFFEAAFFFB541202C8644B44DC1FF2D" bold="true" box="[199,385,150,176]" pageId="9" pageNumber="88">Ergatoid queen</emphasis>
(
<figureCitation id="44B02A38FFB541202DD044B44D97FF2D" box="[401,471,150,176]" captionStart="FIGURE 4" captionStartId="7.[151,250,1642,1666]" captionTargetBox="[151,1436,181,1618]" captionTargetId="figure-17@7.[151,1436,181,1618]" captionTargetPageId="7" captionText="FIGURE 4. Carebara bokorensis, ergatoid queen (THNHM-I-00028220). A, Body in profile view; B, head in full-face view; 2C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12742651" httpUri="https://zenodo.org/record/12742651/files/figure.png" pageId="9" pageNumber="88">Fig. 4</figureCitation>
). Similar to the major worker in structure, sculpture, coloration and pilosity, with the following conditions that should be noted: body slightly larger in average (HW 
<quantity id="1B739B58FFB541202853449848E0FF49" box="[1042,1184,186,212]" metricMagnitude="-2" metricUnit="m" metricValue="2.2987" metricValueMax="2.3368" metricValueMin="2.2606" pageId="9" pageNumber="88" unit="in" value="0.905" valueMax="0.92" valueMin="0.89">0.89–0.92 in</quantity>
ergatoid queen; 
<quantity id="1B739B58FFB54120291C44984CBEFF64" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="9.6" metricValueMin="8.4" pageId="9" pageNumber="88" unit="mm" value="0.8999999999999999" valueMax="0.96" valueMin="0.84">0.84– 0.96 mm</quantity>
in major worker); ocelli present (absent in major worker), located on vertex; distance between anterior ocellus and lateral ocelli slightly shorter than distance between lateral ocelli; eye larger than in the major.
</paragraph>
</subSubSection>
<subSubSection id="94916536FFB541202C86450448B8FEDD" box="[199,1272,294,321]" pageId="9" pageNumber="88" type="biology_ecology">
<paragraph id="DC3436BDFFB541202C86450448B8FEDD" blockId="9.[151,1436,150,789]" box="[199,1272,294,321]" pageId="9" pageNumber="88">
<emphasis id="EEFFEAAFFFB541202C8645044D6AFEDD" bold="true" box="[199,298,294,320]" pageId="9" pageNumber="88">Habitat.</emphasis>
This species nests in rotting wood in lowland dry evergreen forests (
<quantity id="1B739B58FFB54120286D450448EDFEDC" box="[1068,1197,294,321]" metricMagnitude="2" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="9" pageNumber="88" unit="m" value="350.0" valueMax="500.0" valueMin="200.0">200–500 m</quantity>
a.s.l.).
</paragraph>
</subSubSection>
<subSubSection id="94916536FFB541202C86456848F2FEF9" box="[199,1202,330,356]" pageId="9" pageNumber="88" type="distribution">
<paragraph id="DC3436BDFFB541202C86456848F2FEF9" blockId="9.[151,1436,150,789]" box="[199,1202,330,356]" pageId="9" pageNumber="88">
<emphasis id="EEFFEAAFFFB541202C8645684D21FEF9" bold="true" box="[199,353,330,356]" pageId="9" pageNumber="88">Distribution.</emphasis>
<collectingCountry id="A49C762DFFB541202D2645684D8BFEF9" box="[359,459,330,356]" name="Thailand" pageId="9" pageNumber="88">Thailand</collectingCountry>
(
<collectingRegion id="1E4FF85FFFB541202D9845684E28FEF9" box="[473,616,330,356]" country="Thailand" name="Chanthaburi" pageId="9" pageNumber="88">Chanthaburi</collectingRegion>
, 
<collectingRegion id="1E4FF85FFFB541202E3545684F5AFEF9" box="[628,794,330,356]" country="Thailand" name="Chachoengsao" pageId="9" pageNumber="88">Chachoengsao</collectingRegion>
, and 
<collectingRegion id="1E4FF85FFFB541202F1545684F8FFEF9" box="[852,975,330,356]" country="Thailand" name="Ratchaburi" pageId="9" pageNumber="88">Ratchaburi</collectingRegion>
Provinces, 
<figureCitation id="44B02A38FFB541202812456848E5FEF9" box="[1107,1189,330,356]" captionStart="FIGURE 13" captionStartId="19.[151,250,1999,2023]" captionTargetBox="[213,1378,244,1975]" captionTargetId="figure-45@19.[199,1389,237,1975]" captionTargetPageId="19" captionText="FIGURE 13. Distribution of Carebara pygmea species group in Thailand." figureDoi="http://doi.org/10.5281/zenodo.12742675" httpUri="https://zenodo.org/record/12742675/files/figure.png" pageId="9" pageNumber="88">Fig. 13</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="94916536FFB541202C86454C4FFEFC89" pageId="9" pageNumber="88" type="discussion">
<paragraph id="DC3436BDFFB541202C86454C4DAEFD85" blockId="9.[151,1436,150,789]" pageId="9" pageNumber="88">
<emphasis id="EEFFEAAFFFB541202C86454C4D7BFE15" bold="true" box="[199,315,366,392]" pageId="9" pageNumber="88">Remarks.</emphasis>
The major worker of this species can be easily distinguished from other species of the group in having dense erect and decumbent hairs on the head, promesonotum and gaster; posterior one third of head with dense punctures (with transverse striation in other species); and dense punctures on gastral tergite I (more or less smooth and shiny in other species). This species is similar to 
<taxonomicName id="1B8B4D3EFFB541202F7645F94FD7FE69" authorityName="Wimolsuthikul &amp; Yamane &amp; Samung &amp; Jaitrong" authorityYear="2024" box="[823,919,475,501]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="88" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="EEFFEAAFFFB541202F7645F94F0AFE68" box="[823,842,475,501]" italics="true" pageId="9" pageNumber="88">C</emphasis>
. 
<emphasis id="EEFFEAAFFFB541202F1745F94FD7FE69" box="[854,919,475,500]" italics="true" pageId="9" pageNumber="88">parva</emphasis>
</taxonomicName>
and 
<taxonomicName id="1B8B4D3EFFB541202F8D45F94879FE69" authorityName="Jaitrong, Pitaktunsakul &amp; Jantarit" authorityYear="2021" box="[972,1081,474,501]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="88" phylum="Arthropoda" rank="species" species="panhai">
<emphasis id="EEFFEAAFFFB541202F8D45F94879FE69" box="[972,1081,474,501]" italics="true" pageId="9" pageNumber="88">C. panhai</emphasis>
</taxonomicName>
. See “Remarks” under 
<taxonomicName id="1B8B4D3EFFB54120297A45F949DCFE69" authorityName="Wimolsuthikul &amp; Yamane &amp; Samung &amp; Jaitrong" authorityYear="2024" box="[1339,1436,475,501]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="88" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="EEFFEAAFFFB54120297A45F9490EFE68" box="[1339,1358,475,501]" italics="true" pageId="9" pageNumber="88">C</emphasis>
. 
<emphasis id="EEFFEAAFFFB54120291A45F949DCFE69" box="[1371,1436,475,500]" italics="true" pageId="9" pageNumber="88">parva</emphasis>
</taxonomicName>
and 
<taxonomicName id="1B8B4D3EFFB541202C8645DD4D75FD85" authorityName="Jaitrong, Pitaktunsakul &amp; Jantarit" authorityYear="2021" box="[199,309,510,537]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="88" phylum="Arthropoda" rank="species" species="panhai">
<emphasis id="EEFFEAAFFFB541202C8645DD4D75FD85" box="[199,309,510,537]" italics="true" pageId="9" pageNumber="88">C. panhai</emphasis>
</taxonomicName>
for comparison.
</paragraph>
<paragraph id="DC3436BDFFB541202C8646004FFEFC89" blockId="9.[151,1436,150,789]" pageId="9" pageNumber="88">
<taxonomicName id="1B8B4D3EFFB541202C8646004DF4FDA1" authorityName="Matsuura &amp; Hosoishi" authorityYear="2024" box="[199,436,546,572]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="88" phylum="Arthropoda" rank="species" species="bokorensis">
<emphasis id="EEFFEAAFFFB541202C8646004DF4FDA1" box="[199,436,546,572]" italics="true" pageId="9" pageNumber="88">Carebara bokorensis</emphasis>
</taxonomicName>
is also similar to 
<taxonomicName id="1B8B4D3EFFB541202EC446004FA1FDA1" authority="(Xu, 2003)" baseAuthorityName="Xu" baseAuthorityYear="2003" box="[645,993,546,572]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="88" phylum="Arthropoda" rank="species" species="altinoda">
<emphasis id="EEFFEAAFFFB541202EC446004F17FDA1" box="[645,855,546,572]" italics="true" pageId="9" pageNumber="88">Carebara altinoda</emphasis>
(Xu, 2003)
</taxonomicName>
. However, 
<taxonomicName id="1B8B4D3EFFB541202822460148BEFDA1" authorityName="Matsuura &amp; Hosoishi" authorityYear="2024" box="[1123,1278,546,573]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="88" phylum="Arthropoda" rank="species" species="bokorensis">
<emphasis id="EEFFEAAFFFB541202822460148BEFDA1" box="[1123,1278,546,573]" italics="true" pageId="9" pageNumber="88">C. bokorensis</emphasis>
</taxonomicName>
can be easily separated from 
<taxonomicName id="1B8B4D3EFFB541202D0A46654D8CFDFD" baseAuthorityName="Xu" baseAuthorityYear="2003" box="[331,460,582,609]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="88" phylum="Arthropoda" rank="species" species="altinoda">
<emphasis id="EEFFEAAFFFB541202D0A46654D8CFDFD" box="[331,460,582,609]" italics="true" pageId="9" pageNumber="88">C. altinoda</emphasis>
</taxonomicName>
by 1) in major worker, Dorsa of head, mesosoma, and gaster with dense erect hairs mixed with dense decumbent hairs (with dense decumbent on head and with sparse erect hairs on mesosoma and gaster in 
<taxonomicName id="1B8B4D3EFFB541202D4046AD4DC1FD35" baseAuthorityName="Xu" baseAuthorityYear="2003" box="[257,385,654,681]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="88" phylum="Arthropoda" rank="species" species="altinoda">
<emphasis id="EEFFEAAFFFB541202D4046AD4DC1FD35" box="[257,385,654,681]" italics="true" pageId="9" pageNumber="88">C. altinoda</emphasis>
</taxonomicName>
); 2) in major worker, without a pair of minute horns on vertex (with a pair of minute horns in 
<taxonomicName id="1B8B4D3EFFB541202CF646914D77FD51" baseAuthorityName="Xu" baseAuthorityYear="2003" box="[183,311,690,717]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="88" phylum="Arthropoda" rank="species" species="altinoda">
<emphasis id="EEFFEAAFFFB541202CF646914D77FD51" box="[183,311,690,717]" italics="true" pageId="9" pageNumber="88">C. altinoda</emphasis>
</taxonomicName>
); 3) in minor worker, propodeum in profile roundly convex (bluntly angulates in 
<taxonomicName id="1B8B4D3EFFB5412028AA4691492BFD51" baseAuthorityName="Xu" baseAuthorityYear="2003" box="[1259,1387,690,717]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="88" phylum="Arthropoda" rank="species" species="altinoda">
<emphasis id="EEFFEAAFFFB5412028AA4691492BFD51" box="[1259,1387,690,717]" italics="true" pageId="9" pageNumber="88">C. altinoda</emphasis>
</taxonomicName>
); in minor worker, distance between propodeal spiracle and metanotal groove ca. 4 times as long as distance between propodeal spiracle and propodeal junction (ca. 8.5 times in 
<taxonomicName id="1B8B4D3EFFB541202F7146D94FEEFC89" baseAuthorityName="Xu" baseAuthorityYear="2003" box="[816,942,762,789]" class="Insecta" family="Formicidae" genus="Carebara" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="88" phylum="Arthropoda" rank="species" species="altinoda">
<emphasis id="EEFFEAAFFFB541202F7146D94FEEFC89" box="[816,942,762,789]" italics="true" pageId="9" pageNumber="88">C. altinoda</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>