<documentID-DOI="http://dx.doi.org/10.3897/zookeys.1163.103650"ID-GBIF-Dataset="0f20ea37-1077-4923-afbf-7699ea7e73f4"ID-PMC="PMC10220496"ID-Pensoft-Pub="1313-2970-1163-47"ID-Pensoft-UUID="9480EC110E425078B0719BE5F4CDE4F9"ID-PubMed="37250364"ID-ZooBank="2DB90CB6E7464887B71BA5D7D7092CEF"ModsDocID="1313-2970-1163-47"checkinTime="1684532138151"checkinUser="pensoft"docAuthor="Tumpeesuwan, Sakboworn, Tanmuangpak, Kitti & Tumpeesuwan, Chanidaporn"docDate="2023"docId="2D76BE333DC0553E8CB91F12CE5A4AF2"docLanguage="en"docName="ZooKeys 1163: 47-59"docOrigin="ZooKeys 1163"docPubDate="2023-05-19"docSource="http://dx.doi.org/10.3897/zookeys.1163.103650"docTitle="Cryptosemelus niger S. Tumpeesuwan & C. Tumpeesuwan 2023, sp. nov."docType="treatment"docUuid="FA7C7967-4A62-4A6E-992C-4BE2E6DC9B36"docUuidSource="ZooBank"docVersion="5"id="9480EC110E425078B0719BE5F4CDE4F9"lastPageNumber="47"masterDocId="9480EC110E425078B0719BE5F4CDE4F9"masterDocTitle="New species of the dancing semislug Cryptosemelus Collinge, 1902 (Eupulmonata, Ariophantidae) from Loei Province, northeastern Thailand with a key to genera of mainland Southeast Asian semislugs and a key to species of the genus"masterLastPageNumber="59"masterPageNumber="47"pageNumber="47"updateTime="1732567316748"updateUser="ExternalLinkService">
<mods:titleid="0241C265CA625FEE847C6EEC2CC75B3C">New species of the dancing semislug Cryptosemelus Collinge, 1902 (Eupulmonata, Ariophantidae) from Loei Province, northeastern Thailand with a key to genera of mainland Southeast Asian semislugs and a key to species of the genus</mods:title>
<mods:affiliationid="A191381E9BA5EEC8661B445A0A674839">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Maha Sarakham 44150, Thailand</mods:affiliation>
<mods:affiliationid="41968FF56B590A3E1F4CEA7FDE5102BB">Palaeontological Research and Education Centre, Mahasarakham University, Kantharawichai District, Maha Sarakham 44150, Thailand</mods:affiliation>
<mods:affiliationid="49C92A76DCCE82DA20A2363A48ECD548">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Maha Sarakham 44150, Thailand</mods:affiliation>
<taxonomicNameid="9A8F2DD9AD1D9AC1532D58790AFBD50E"ID-CoL="BRHMV"LSID="https://zoobank.org/FA7C7967-4A62-4A6E-992C-4BE2E6DC9B36"authority="S. Tumpeesuwan & C. Tumpeesuwan"authorityName="S. Tumpeesuwan & C. Tumpeesuwan"authorityYear="2023"class="Gastropoda"family="Ariophantidae"genus="Cryptosemelus"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Cryptosemelus niger"order="Stylommatophora"pageId="0"pageNumber="47"phylum="Mollusca"rank="species"species="niger"status="sp. nov.">Cryptosemelus niger S. Tumpeesuwan & C. Tumpeesuwan</taxonomicName>
<figureCitationid="8A35E8ACDC8684CBA05D6690ABF79813"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. Map of type locality of Cryptosemelus niger S. Tumpeesuwan & C. Tumpeesuwan, sp. nov."figureDoi="10.3897/zookeys.1163.103650.figure1"httpUri="https://binary.pensoft.net/fig/852462"pageId="0"pageNumber="47">Figs 1</figureCitation>
<figureCitationid="BAF31DC4CD0408A6ED9EB4DA2C0BD4DB"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Cryptosemelus niger S. Tumpeesuwan & C. Tumpeesuwan, sp. nov. Photograph by Kitti Tanmuangpak A shell morphology of holotype NHMSU- 00054 B living snail."figureDoi="10.3897/zookeys.1163.103650.figure2"httpUri="https://binary.pensoft.net/fig/852463"pageId="0"pageNumber="47">, 2</figureCitation>
<figureCitationid="C94FAEDBCAE61DFCBFBAF6076AC8F267"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Cryptosemelus niger S. Tumpeesuwan & C. Tumpeesuwan, sp. nov. Living snail in natural habitats. Photograph by Chanidaporn Tumpeesuwan 12 December 2018 A C. niger crawling on soil in limestone crevice (white circle) B close-up view in living position C, D semislug was moved to green leaf for taking photo C left side represents left shell lobe (lsl) D right side represents right shell lobe (rsl), left shell lobes (lsl), and left dorsal lobe (ldl)."figureDoi="10.3897/zookeys.1163.103650.figure3"httpUri="https://binary.pensoft.net/fig/852464"pageId="0"pageNumber="47">, 3</figureCitation>
<figureCitationid="3833E7BA890D6598BD2DBD8A0C23AAF2"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Genital system of Cryptosemelus niger S. Tumpeesuwan & C. Tumpeesuwan, sp. nov. (paratype NHMSU- 00055) A photograph and B drawing by Kitti Tanmuangpak."figureDoi="10.3897/zookeys.1163.103650.figure4"httpUri="https://binary.pensoft.net/fig/852465"pageId="0"pageNumber="47">, 4</figureCitation>
<taxonomicNameid="59C9253C51EB8D09D50C47F4A102701B"authority="Tumpeesuwan & Tanmuangpak & Tumpeesuwan, 2023"authorityName="Tumpeesuwan & Tanmuangpak & Tumpeesuwan"authorityYear="2023"class="Gastropoda"family="Ariophantidae"genus="Cryptaustenia"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Cryptaustenia"order="Stylommatophora"pageId="0"pageNumber="47"phylum="Mollusca"rank="genus"status="sp. nov.">Cryptaustenia</taxonomicName>
sp.
<bibRefCitationid="161BB90627728A1AD5B1FA9547B5EDBB"author="Tanmuangpak, K"journalOrPublisher="Journal of Science and Technology MSU"pageId="0"pageNumber="47"refId="B21"refString="Tanmuangpak, K, 2016. Species diversity, abundance, distribution and habitat relationship of land snail in Phu Pha Lom limestone area, Loei Province. Unpublished PhD Thesis, Mahasarakham University, Maha Sarakham, Thailand."title="Species diversity, abundance, distribution and habitat relationship of land snail in Phu Pha Lom limestone area, Loei Province. Unpublished PhD Thesis, Mahasarakham University, Maha Sarakham, Thailand."year="2016">Tanmuangpak 2016</bibRefCitation>
<taxonomicNameid="45ABCE0E036BDE45DF84DAD7CDF38FF8"authority="Tumpeesuwan & Tanmuangpak & Tumpeesuwan, 2023"authorityName="Tumpeesuwan & Tanmuangpak & Tumpeesuwan"authorityYear="2023"class="Gastropoda"family="Ariophantidae"genus="Cryptaustenia"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Cryptaustenia"order="Stylommatophora"pageId="0"pageNumber="47"phylum="Mollusca"rank="genus"status="sp. nov.">Cryptaustenia</taxonomicName>
sp.
<bibRefCitationid="DBA27088D72A318C50CA458F5B72A460"author="Chimsaeng, N"journalOrPublisher="Taylor and Francis, London"pageId="0"pageNumber="47"refId="B3"refString="Chimsaeng, N, 2019. Reproductive system anatomy and radula moprphology of some land snail genera Dugella, Cryptaustenia and Aenigmatoconcha. Bachelor degree Science's project. Mahasarakham University, Maha Sarakham, Thailand. [In Thai with English abstract]"title="Reproductive system anatomy and radula moprphology of some land snail genera Dugella, Cryptaustenia and Aenigmatoconcha. Bachelor degree Science's project. Mahasarakham University, Maha Sarakham, Thailand. [In Thai with English abstract]"year="2019">Chimsaeng 2019</bibRefCitation>
<figureCitationid="1752BFC4F7AADC9CD36387219F5F9556"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Cryptosemelus niger S. Tumpeesuwan & C. Tumpeesuwan, sp. nov. Living snail in natural habitats. Photograph by Chanidaporn Tumpeesuwan 12 December 2018 A C. niger crawling on soil in limestone crevice (white circle) B close-up view in living position C, D semislug was moved to green leaf for taking photo C left side represents left shell lobe (lsl) D right side represents right shell lobe (rsl), left shell lobes (lsl), and left dorsal lobe (ldl)."figureDoi="10.3897/zookeys.1163.103650.figure3"httpUri="https://binary.pensoft.net/fig/852464"pageId="0"pageNumber="47">3</figureCitation>
Animal with blackish body, shell lobes with blackish reticulated skin (Figs
<figureCitationid="766E211CA1A65ACB17AEEACC8A3D2AED"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Cryptosemelus niger S. Tumpeesuwan & C. Tumpeesuwan, sp. nov. Photograph by Kitti Tanmuangpak A shell morphology of holotype NHMSU- 00054 B living snail."figureDoi="10.3897/zookeys.1163.103650.figure2"httpUri="https://binary.pensoft.net/fig/852463"pageId="0"pageNumber="47">2</figureCitation>
,
<figureCitationid="76B4E739F8ED0C881E38F5127B70E5ED"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Cryptosemelus niger S. Tumpeesuwan & C. Tumpeesuwan, sp. nov. Living snail in natural habitats. Photograph by Chanidaporn Tumpeesuwan 12 December 2018 A C. niger crawling on soil in limestone crevice (white circle) B close-up view in living position C, D semislug was moved to green leaf for taking photo C left side represents left shell lobe (lsl) D right side represents right shell lobe (rsl), left shell lobes (lsl), and left dorsal lobe (ldl)."figureDoi="10.3897/zookeys.1163.103650.figure3"httpUri="https://binary.pensoft.net/fig/852464"pageId="0"pageNumber="47">3</figureCitation>
). Genitalia with very short cylindrical vagina, smooth elongated cylindrical epiphallus, without penial caecum (Fig.
<figureCitationid="71240A5496C061DBD9858A0B889B3337"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Genital system of Cryptosemelus niger S. Tumpeesuwan & C. Tumpeesuwan, sp. nov. (paratype NHMSU- 00055) A photograph and B drawing by Kitti Tanmuangpak."figureDoi="10.3897/zookeys.1163.103650.figure4"httpUri="https://binary.pensoft.net/fig/852465"pageId="0"pageNumber="47">4</figureCitation>
). Radula with bicuspid lateral teeth (Fig.
<figureCitationid="D4F8DDCFF5D6509DD711139C2076CE50"captionStart="Figure 5"captionStartId="F5"captionText="Figure 5. Radula morphology of Cryptosemelus niger S. Tumpeesuwan & C. Tumpeesuwan, sp. nov. (paratype NHMSU- 00055) A radula plate, teeth rows arranged in wide V-shape B close-up view of middle part of radula C close-up view of right side of radula D close-up view of right side of radula showing marginal teeth. Central teeth indicated by ' C'; lateral teeth indicated by ' L'; marginal teeth indicated by ' M'."figureDoi="10.3897/zookeys.1163.103650.figure5"httpUri="https://binary.pensoft.net/fig/852466"pageId="0"pageNumber="47">5B, C</figureCitation>
<taxonomicNameid="8E9ED72F09DBB3CC2E4BCB7F768C9C18"authorityName="S. Tumpeesuwan & C. Tumpeesuwan"authorityYear="2023"class="Gastropoda"family="Ariophantidae"genus="Cryptosemelus"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Cryptosemelus niger"order="Stylommatophora"pageId="0"pageNumber="47"phylum="Mollusca"rank="species"species="niger">
<taxonomicNameid="2166F1F8171622BD65DCEACD00C7D75A"authorityName="S. Tumpeesuwan & C. Tumpeesuwan"authorityYear="2023"class="Gastropoda"family="Ariophantidae"genus="Cryptosemelus"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Cryptosemelus niger"order="Stylommatophora"pageId="0"pageNumber="47"phylum="Mollusca"rank="species"species="niger">
<figureCitationid="C540CCD84D7A57683B138F5D6735694F"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Cryptosemelus niger S. Tumpeesuwan & C. Tumpeesuwan, sp. nov. Photograph by Kitti Tanmuangpak A shell morphology of holotype NHMSU- 00054 B living snail."figureDoi="10.3897/zookeys.1163.103650.figure2"httpUri="https://binary.pensoft.net/fig/852463"pageId="0"pageNumber="47">2A</figureCitation>
<figureCitationid="7BEA5DFAA1CD2E30462A5F9E39E5F6A2"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Genital system of Cryptosemelus niger S. Tumpeesuwan & C. Tumpeesuwan, sp. nov. (paratype NHMSU- 00055) A photograph and B drawing by Kitti Tanmuangpak."figureDoi="10.3897/zookeys.1163.103650.figure4"httpUri="https://binary.pensoft.net/fig/852465"pageId="0"pageNumber="47">4</figureCitation>
). Atrium (at) short. Penis rather long and cylindrical, with thin penial sheath covering entire penis. Penial retractor muscle (prm) present, short, thin and attached at junction of e1 and e2. Epiphallus (e1+e2) length is slightly equal to penis length, surface smooth, e1 cylindrical and gradually smaller in diameter, e2 cylindrical and larger than e1. Flagellum absent. Vas deferens long. Vagina is shorter than penis, cylindrical. Gametolytic duct (gd) thickened at base, gametolytic sac (gs) swollen gland at distal end. Free oviduct (fo) is shorter than vagina. Uterus and prostate gland long and stout.
<figureCitationid="F5245E315817FFBF67A604EE9694AAC1"captionStart="Figure 5"captionStartId="F5"captionText="Figure 5. Radula morphology of Cryptosemelus niger S. Tumpeesuwan & C. Tumpeesuwan, sp. nov. (paratype NHMSU- 00055) A radula plate, teeth rows arranged in wide V-shape B close-up view of middle part of radula C close-up view of right side of radula D close-up view of right side of radula showing marginal teeth. Central teeth indicated by ' C'; lateral teeth indicated by ' L'; marginal teeth indicated by ' M'."figureDoi="10.3897/zookeys.1163.103650.figure5"httpUri="https://binary.pensoft.net/fig/852466"pageId="0"pageNumber="47">5</figureCitation>
). Teeth arranged in a wide V-shape with half row formula: 1-7-70+teeth. Central teeth symmetric tricuspid. Lateral teeth and marginal teeth gradually changing from broad to narrow bicuspid.
<figureCitationid="7F941FC239DC72DF1B4FE67030D3E4D4"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Cryptosemelus niger S. Tumpeesuwan & C. Tumpeesuwan, sp. nov. Photograph by Kitti Tanmuangpak A shell morphology of holotype NHMSU- 00054 B living snail."figureDoi="10.3897/zookeys.1163.103650.figure2"httpUri="https://binary.pensoft.net/fig/852463"pageId="0"pageNumber="47">2B</figureCitation>
,
<figureCitationid="9B7C8DE5A1498468C57E1E03DFC61D35"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Cryptosemelus niger S. Tumpeesuwan & C. Tumpeesuwan, sp. nov. Living snail in natural habitats. Photograph by Chanidaporn Tumpeesuwan 12 December 2018 A C. niger crawling on soil in limestone crevice (white circle) B close-up view in living position C, D semislug was moved to green leaf for taking photo C left side represents left shell lobe (lsl) D right side represents right shell lobe (rsl), left shell lobes (lsl), and left dorsal lobe (ldl)."figureDoi="10.3897/zookeys.1163.103650.figure3"httpUri="https://binary.pensoft.net/fig/852464"pageId="0"pageNumber="47">3</figureCitation>
). Living semislug with reticulated skin, blackish to dark body marked by conspicuous grooves running downward. Four mantle extensions well developed and same color body. Shell lobes do not cover entire shell. Caudal horn absent.
<taxonomicNameid="45A5974F7F7A1E890F61F60F1A9498A6"authorityName="S. Tumpeesuwan & C. Tumpeesuwan"authorityYear="2023"class="Gastropoda"family="Ariophantidae"genus="Cryptosemelus"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Cryptosemelus niger"order="Stylommatophora"pageId="0"pageNumber="47"phylum="Mollusca"rank="species"species="niger">
sp. nov. is a species restricted to the dry evergreen forest that covers the Phu Pha Lom limestone area, Loei Province, Thailand. We searched after rain and found the specimens normally hiding on the ground and in the leaf litter. When the snails were disturbed, they escaped by quickly flipping and waging their tail. Information on its natural predators is unknown, but the carnivorous land snails,
<taxonomicNameid="98FAFA591FCC6DEA7002A43E3A17FA8F"authorityName="J. E. Gray"authorityYear="1860"class="Gastropoda"family="Streptaxidae"kingdom="Animalia"lsidName=""order="Stylommatophora"pageId="0"pageNumber="47"phylum="Mollusca"rank="family">Streptaxidae</taxonomicName>
), and
<taxonomicNameid="739EC7FC19D2E1A983F3BCBCF7BA31E9"authorityName="Tanmuangpak & S. Tumpeesuwan"authorityYear="2015"class="Gastropoda"family="Diapheridae"genus="Sinoennea"kingdom="Animalia"lsidName="Sinoennea loeiensis"order="Stylommatophora"pageId="0"pageNumber="47"phylum="Mollusca"rank="species"species="loeiensis">
<taxonomicNameid="257246F98DBB06068E92ED19534DC93A"authorityName="Panha & Naggs"authorityYear="2010"class="Gastropoda"family="Diapheridae"kingdom="Animalia"lsidName=""order="Stylommatophora"pageId="0"pageNumber="47"phylum="Mollusca"rank="family">Diapheridae</taxonomicName>
) were found in the same locality (
<bibRefCitationid="F33D48CF86DD74241BB4CF6B31EA136B"author="Tanmuangpak, K"journalOrPublisher="Raffles Bulletin of Zoology"pageId="0"pageNumber="47"refId="B23"refString="Tanmuangpak, K, Tumpeesuwan, C, Tumpeesuwan, S, 2012. Diversity of Land Snails in Phu Pha Lom Forest Park at Muang District, Loei Province. Journal of Science and Technology Mahasarakham University 8(Special Issue): 15-26. [In Thai with English abstract]"title="Diversity of Land Snails in Phu Pha Lom Forest Park at Muang District, Loei Province. Journal of Science and Technology Mahasarakham University 8 (Special Issue): 15 - 26. [In Thai with English abstract]"year="2012">Tanmuangpak et al. 2012</bibRefCitation>
;
<bibRefCitationid="191E25CA0A017CF001BE68E5AB94B2FA"author="Tanmuangpak, K"journalOrPublisher="The Raffles Bulletin of Zoology"pageId="0"pageNumber="47"pagination="293 - 300"refId="B24"refString="Tanmuangpak, K, Dumrongrojwattana, P, Tumpeesuwan, C, Tumpeesuwan, S, 2015. Sinoennea loeiensis, a new species of diapherid microsnail (Pulmonata: Streptaxoidea: Diapheridae) from Phu Pha Lom Limestone Hill, Loei Province, Northeastern Thailand. The Raffles Bulletin of Zoology 63: 293 - 300"title="Sinoennea loeiensis, a new species of diapherid microsnail (Pulmonata: Streptaxoidea: Diapheridae) from Phu Pha Lom Limestone Hill, Loei Province, Northeastern Thailand."volume="63"year="2015">Tanmuangpak et al. 2015</bibRefCitation>
;
<bibRefCitationid="1B7D415D39824FB6EC653E82B1D2FEE7"author="Tanmuangpak, K"journalOrPublisher="Journal of Science and Technology MSU"pageId="0"pageNumber="47"refId="B21"refString="Tanmuangpak, K, 2016. Species diversity, abundance, distribution and habitat relationship of land snail in Phu Pha Lom limestone area, Loei Province. Unpublished PhD Thesis, Mahasarakham University, Maha Sarakham, Thailand."title="Species diversity, abundance, distribution and habitat relationship of land snail in Phu Pha Lom limestone area, Loei Province. Unpublished PhD Thesis, Mahasarakham University, Maha Sarakham, Thailand."year="2016">Tanmuangpak 2016</bibRefCitation>
<taxonomicNameid="807BE7EE8A0213D3A8336C709651351B"authorityName="S. Tumpeesuwan & C. Tumpeesuwan"authorityYear="2023"class="Gastropoda"family="Ariophantidae"genus="Cryptosemelus"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Cryptosemelus niger"order="Stylommatophora"pageId="0"pageNumber="47"phylum="Mollusca"rank="species"species="niger">
also lack a penial caecum but differs in the undulated surface patch on the proximal part of e2. The epiphallus and penis are cylindrical without a caecum, diverticulum, and granulate surface. The radula of the new species differs from all
in having bicuspid lateral teeth, whereas other species have tricuspid lateral teeth (Table
<tableCitationid="E1B6E01E2DFED4D94A7321A4457B5771"captionStart="Table 2"captionStartId="T2"captionText="Table 2. Comparison of shell, genital system and radula of Cryptosemelus spp. (data for C. gracilis, C. betarmon and C. tigrinus based on Pholyotha et al. 2021 a)."httpUri="http://table.plazi.org/id/BFB7330926E7374A7B241CC8CCC1D013"pageId="0"pageNumber="47"tableUuid="BFB7330926E7374A7B241CC8CCC1D013">2</tableCitation>
, Figs
<figureCitationid="517C694A17A8B46852E4103CA37CE2EF"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Cryptosemelus niger S. Tumpeesuwan & C. Tumpeesuwan, sp. nov. Photograph by Kitti Tanmuangpak A shell morphology of holotype NHMSU- 00054 B living snail."figureDoi="10.3897/zookeys.1163.103650.figure2"httpUri="https://binary.pensoft.net/fig/852463"pageId="0"pageNumber="47">2A, B</figureCitation>
,
<figureCitationid="014E1E46739BC469BDE784A32025763E"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Cryptosemelus niger S. Tumpeesuwan & C. Tumpeesuwan, sp. nov. Living snail in natural habitats. Photograph by Chanidaporn Tumpeesuwan 12 December 2018 A C. niger crawling on soil in limestone crevice (white circle) B close-up view in living position C, D semislug was moved to green leaf for taking photo C left side represents left shell lobe (lsl) D right side represents right shell lobe (rsl), left shell lobes (lsl), and left dorsal lobe (ldl)."figureDoi="10.3897/zookeys.1163.103650.figure3"httpUri="https://binary.pensoft.net/fig/852464"pageId="0"pageNumber="47">3A-D</figureCitation>
,
<figureCitationid="864FDC95353C8A40D7C7C391B355AD10"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Genital system of Cryptosemelus niger S. Tumpeesuwan & C. Tumpeesuwan, sp. nov. (paratype NHMSU- 00055) A photograph and B drawing by Kitti Tanmuangpak."figureDoi="10.3897/zookeys.1163.103650.figure4"httpUri="https://binary.pensoft.net/fig/852465"pageId="0"pageNumber="47">4</figureCitation>
,
<figureCitationid="D317F331E6C24AAE91FB2EE73FCA5E34"captionStart="Figure 5"captionStartId="F5"captionText="Figure 5. Radula morphology of Cryptosemelus niger S. Tumpeesuwan & C. Tumpeesuwan, sp. nov. (paratype NHMSU- 00055) A radula plate, teeth rows arranged in wide V-shape B close-up view of middle part of radula C close-up view of right side of radula D close-up view of right side of radula showing marginal teeth. Central teeth indicated by ' C'; lateral teeth indicated by ' L'; marginal teeth indicated by ' M'."figureDoi="10.3897/zookeys.1163.103650.figure5"httpUri="https://binary.pensoft.net/fig/852466"pageId="0"pageNumber="47">5B-D</figureCitation>
<bibRefCitationid="B29172C39A6AA765BC586B4438D74D32"DOI="https://doi.org/10.3897/zookeys.1076.75576"author="Pholyotha, A"journalOrPublisher="ZooKeys"pageId="0"pageNumber="47"pagination="43 - 65"refId="B14"refString="Pholyotha, A, Sucharit, C, Panha, S, 2021a. Rediscovering the dancing semislug genus Cryptosemelus Colling, 1902 (Eupulmonata, Ariophantidae) from Thailand with description of two new species. ZooKeys 1076: 43 - 65, DOI: https://doi.org/10.3897/zookeys.1076.75576"title="Rediscovering the dancing semislug genus Cryptosemelus Colling, 1902 (Eupulmonata, Ariophantidae) from Thailand with description of two new species."url="https://doi.org/10.3897/zookeys.1076.75576"volume="1076"year="2021 a">Pholyotha et al. 2021a</bibRefCitation>
<tdid="5F50DDE62DE67C08D0C7E1715ADA93D9"colspan="1"pageId="0"pageNumber="47"rowspan="1">Cylindrical and gradually tapering to distal end, smooth surface</td>
<tdid="C25B2311B17631DE5B67B56E74892B4E"colspan="1"pageId="0"pageNumber="47"rowspan="1">Cylindrical with prominently granulated surface</td>
<tdid="94BF988CF8009CAE6B5E7BCCC5419DAD"colspan="1"pageId="0"pageNumber="47"rowspan="1">Long cylindrical with smooth surface</td>
