230 lines
39 KiB
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230 lines
39 KiB
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<document ID-DOI="10.1111/zoj.12321" ID-GBIF-Dataset="da7659b3-437f-4db6-80d2-69fa35aa5b16" ID-ISSN="0024-4082" ID-PMC="PMC7165668" ID-PubMed="32336790" ID-Zenodo-Dep="5357812" approvalRequired="8" approvalRequired_for_textStreams="8" checkinTime="1630221662267" checkinUser="marcus" docAuthor="Gale, Andrew Scott" docDate="2016" docId="03D63555CC320D7A5324FD25BA79602A" docLanguage="en" docName="zoj12321.pdf" docOrigin="Zoological Journal of the Linnean Society 176 (2)" docSource="http://dx.doi.org/10.1111/zoj.12321" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.2:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="2" docTitle="Catherinum anglicum Gale 2016, SP. NOV." docType="treatment" docVersion="6" lastPageNumber="301" masterDocId="FFEF4D2DCC110D595225FFD9BF4B650B" masterDocTitle="Phylogeny of the deep-sea cirripede family Scalpellidae (Crustacea, Thoracica) based on shell capitular plate morphology" masterLastPageNumber="304" masterPageNumber="266" pageNumber="301" updateTime="1668248615449" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Phylogeny of the deep-sea cirripede family Scalpellidae (Crustacea, Thoracica) based on shell capitular plate morphology</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gale, Andrew Scott</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zoological Journal of the Linnean Society</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2016</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2016-01-25</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>176</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>266</mods:start>
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<mods:end>304</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.1111/zoj.12321</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1111/zoj.12321</mods:identifier>
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<mods:identifier type="GBIF-Dataset">da7659b3-437f-4db6-80d2-69fa35aa5b16</mods:identifier>
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<mods:identifier type="ISSN">0024-4082</mods:identifier>
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<mods:identifier type="PMC">PMC7165668</mods:identifier>
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<mods:identifier type="PubMed">32336790</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5357812</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7314181" ID-GBIF-Taxon="204903474" ID-Zenodo-Dep="7314181" LSID="urn:lsid:plazi:treatment:03D63555CC320D7A5324FD25BA79602A" httpUri="http://treatment.plazi.org/id/03D63555CC320D7A5324FD25BA79602A" lastPageNumber="301" pageId="35" pageNumber="301">
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<subSubSection box="[257,641,764,785]" pageId="35" pageNumber="301" type="nomenclature">
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<paragraph blockId="35.[257,641,764,818]" box="[257,641,764,785]" pageId="35" pageNumber="301">
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<heading box="[257,641,764,785]" centered="true" fontSize="9" level="2" pageId="35" pageNumber="301" reason="4">
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<taxonomicName authority="Gale, 2016" authorityName="Gale" authorityYear="2016" box="[257,543,764,785]" class="Maxillopoda" family="Scalpellidae" genus="Catherinum" kingdom="Animalia" order="Pedunculata" pageId="35" pageNumber="301" phylum="Arthropoda" rank="species" species="anglicum" status="sp. nov.">CATHERINUM ANGLICUM</taxonomicName>
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<emphasis bold="true" box="[550,641,764,785]" pageId="35" pageNumber="301">
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<taxonomicNameLabel box="[550,641,764,785]" pageId="35" pageNumber="301" rank="species">SP. NOV.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="35" pageNumber="301" type="description">
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<paragraph blockId="35.[257,641,764,818]" box="[296,602,796,818]" pageId="35" pageNumber="301">
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<figureCitation box="[296,602,796,818]" captionStart="Figure 19" captionStartId="29.[142,218,1650,1669]" captionTargetBox="[243,1320,193,1620]" captionTargetId="figure-0@29.[243,1320,193,1620]" captionTargetPageId="29" captionText="Figure 19. Fossil and recent Catherinum. C–K, N–Q, T, V, Catherinum anglicum sp. nov., Upper Campanian, Belemnitella mucronata zone, Keswick, Norwich, UK; (O) is the holotype, the other specimens are paratypes. C, D, F, G, carinolatera in external (C, D, F) and internal (G, H) aspects (C, NHMUK IC 1203; D, 1204; F–G, 1206; H, 1208). E, transverse section of carina (NHMUK IC 1205); I, T, dorsal view of carinae (NHMUK IC 1209, 1216); P, lateral view of carina (NHMUK IC 1214); V, internal view of base of carina (NHMUK IC 1217); J, K, O, internal views of scuta, to show large pit for dwarf males (NHMUK IC 1210, 1211, 1213); N, external view of scutum (NHMUK IC 1212); Q, external view of tergum (NHMUK IC 1215). A, B, L, M, R, S, U, Catherinum hispidum (G.O. Sars, 1890). Recent, Norwegian Sea. A, B, carinolatus in external and internal views. L, M, external and internal views of scutum. R, external view of tergum. S, U, dorsal and internal views of carina. Scale bars: S, U, V, 1 mm; A–D, F, G, K–O, Q, R, T, 0.5 mm; E, H, P, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5357846" httpUri="https://zenodo.org/record/5357846/files/figure.png" pageId="35" pageNumber="301">FIGURE 19C–K, N–Q, T, V</figureCitation>
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</paragraph>
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<paragraph blockId="35.[142,757,835,1040]" box="[142,254,835,856]" pageId="35" pageNumber="301">
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<emphasis box="[142,254,835,856]" italics="true" pageId="35" pageNumber="301">Diagnosis</emphasis>
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</paragraph>
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<paragraph blockId="35.[142,757,835,1040]" pageId="35" pageNumber="301">
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<taxonomicName authorityName="ZEVINA" authorityYear="1978" box="[142,284,865,886]" class="Maxillopoda" family="Scalpellidae" genus="Catherinum" kingdom="Animalia" order="Pedunculata" pageId="35" pageNumber="301" phylum="Arthropoda" rank="genus">
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<emphasis box="[142,284,865,886]" italics="true" pageId="35" pageNumber="301">Catherinum</emphasis>
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</taxonomicName>
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in which the occludent margin of the tergum is straight, and the external surface adjacent to this carries three ribs that diverge from the umbo. The upper part of the upper latus margin of the scutum is embayed. The carinolatus has a very broad upper latus margin.
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</paragraph>
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<paragraph blockId="35.[142,757,1081,1164]" box="[142,361,1081,1103]" pageId="35" pageNumber="301">
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<emphasis box="[142,361,1081,1103]" italics="true" pageId="35" pageNumber="301">Derivation of name</emphasis>
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</paragraph>
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<paragraph blockId="35.[142,757,1081,1164]" pageId="35" pageNumber="301">
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<materialsCitation ID-GBIF-Occurrence="3964107302" location="After East Anglia" municipality="East Anglia" pageId="35" pageNumber="301" specimenCount="1">
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<location LSID="urn:lsid:plazi:treatment:03D63555CC320D7A5324FD25BA79602A:8EA0D298CC320D7A52ABFB81BE2F6165" box="[142,356,1112,1134]" municipality="East Anglia" name="After East Anglia" pageId="35" pageNumber="301">
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After
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<collectingMunicipality box="[214,356,1112,1134]" pageId="35" pageNumber="301">East Anglia</collectingMunicipality>
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</location>
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, in which the
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<typeStatus box="[538,588,1112,1133]" pageId="35" pageNumber="301">type</typeStatus>
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material was collected
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</materialsCitation>
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.
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</paragraph>
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<paragraph blockId="35.[142,757,1205,1411]" box="[142,240,1205,1226]" pageId="35" pageNumber="301">
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<emphasis box="[142,240,1205,1226]" italics="true" pageId="35" pageNumber="301">Material</emphasis>
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</paragraph>
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<paragraph blockId="35.[142,757,1205,1411]" pageId="35" pageNumber="301">
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<materialsCitation ID-GBIF-Occurrence="3964107301" country="United Kingdom" county="The" location="Fifty" municipality="Norwich" pageId="35" pageNumber="301" specimenCount="1" stateProvince="Norfolk" typeStatus="holotype">
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<location LSID="urn:lsid:plazi:treatment:03D63555CC320D7A5324FD25BA79602A:8EA0D298CC320D7A52ABFB0DBF8361E1" box="[142,200,1236,1258]" country="United Kingdom" county="The" municipality="Norwich" name="Fifty" pageId="35" pageNumber="301" stateProvince="Norfolk">Fifty</location>
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valves, including carinae, scuta, terga, and carinolatera, from the
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<location LSID="urn:lsid:plazi:treatment:03D63555CC320D7A5324FD25BA79602A:8EA0D298CC320D7A53A8FB2ABD176003" box="[397,604,1266,1288]" country="United Kingdom" county="The" municipality="Norwich" name="upper Campanian" pageId="35" pageNumber="301" stateProvince="Norfolk">upper Campanian</location>
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,
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<taxonomicName baseAuthorityName="Schlotheim" baseAuthorityYear="1813" class="Cephalopoda" family="Dimitobelidae" genus="Belemnitella" higherTaxonomySource="CoL" kingdom="Animalia" order="Belemnitida" pageId="35" pageNumber="301" phylum="Mollusca" rank="species" species="mucronata">
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<emphasis italics="true" pageId="35" pageNumber="301">Belemnitella mucronata</emphasis>
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</taxonomicName>
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zone chalk of
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<location LSID="urn:lsid:plazi:treatment:03D63555CC320D7A5324FD25BA79602A:8EA0D298CC320D7A5386FAC8BEB4602C" box="[419,511,1297,1319]" country="United Kingdom" county="The" municipality="Norwich" name="Keswick" pageId="35" pageNumber="301" stateProvince="Norfolk">Keswick</location>
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and
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<location LSID="urn:lsid:plazi:treatment:03D63555CC320D7A5324FD25BA79602A:8EA0D298CC320D7A5012FAC8BE5C604E" country="United Kingdom" county="The" municipality="Norwich" name="Cringleford Newfound Farm" pageId="35" pageNumber="301" stateProvince="Norfolk">Cringleford Newfound Farm</location>
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pits,
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<collectingMunicipality box="[348,447,1327,1349]" pageId="35" pageNumber="301">Norwich</collectingMunicipality>
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,
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<collectingRegion box="[461,550,1327,1349]" country="United Kingdom" name="Norfolk" pageId="35" pageNumber="301">Norfolk</collectingRegion>
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,
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<collectingCountry box="[564,607,1328,1349]" name="United Kingdom" pageId="35" pageNumber="301">UK</collectingCountry>
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.
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<collectingCounty box="[619,663,1327,1349]" pageId="35" pageNumber="301">The</collectingCounty>
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scutum illustrated is the
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<typeStatus box="[340,436,1358,1380]" pageId="35" pageNumber="301">holotype</typeStatus>
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(
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<figureCitation box="[451,582,1358,1380]" captionStart="Figure 19" captionStartId="29.[142,218,1650,1669]" captionTargetBox="[243,1320,193,1620]" captionTargetId="figure-0@29.[243,1320,193,1620]" captionTargetPageId="29" captionText="Figure 19. Fossil and recent Catherinum. C–K, N–Q, T, V, Catherinum anglicum sp. nov., Upper Campanian, Belemnitella mucronata zone, Keswick, Norwich, UK; (O) is the holotype, the other specimens are paratypes. C, D, F, G, carinolatera in external (C, D, F) and internal (G, H) aspects (C, NHMUK IC 1203; D, 1204; F–G, 1206; H, 1208). E, transverse section of carina (NHMUK IC 1205); I, T, dorsal view of carinae (NHMUK IC 1209, 1216); P, lateral view of carina (NHMUK IC 1214); V, internal view of base of carina (NHMUK IC 1217); J, K, O, internal views of scuta, to show large pit for dwarf males (NHMUK IC 1210, 1211, 1213); N, external view of scutum (NHMUK IC 1212); Q, external view of tergum (NHMUK IC 1215). A, B, L, M, R, S, U, Catherinum hispidum (G.O. Sars, 1890). Recent, Norwegian Sea. A, B, carinolatus in external and internal views. L, M, external and internal views of scutum. R, external view of tergum. S, U, dorsal and internal views of carina. Scale bars: S, U, V, 1 mm; A–D, F, G, K–O, Q, R, T, 0.5 mm; E, H, P, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5357846" httpUri="https://zenodo.org/record/5357846/files/figure.png" pageId="35" pageNumber="301">Figure 19O</figureCitation>
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)
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</materialsCitation>
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, and the other illustrated valves are
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<typeStatus box="[400,516,1389,1410]" pageId="35" pageNumber="301">paratypes</typeStatus>
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.
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</paragraph>
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<paragraph blockId="35.[142,757,1452,1902]" box="[142,271,1452,1473]" pageId="35" pageNumber="301">
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<emphasis box="[142,271,1452,1473]" italics="true" pageId="35" pageNumber="301">Description</emphasis>
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</paragraph>
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<paragraph blockId="35.[142,757,1452,1902]" lastBlockId="35.[804,1420,197,372]" pageId="35" pageNumber="301">
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The carina (
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<figureCitation box="[291,521,1482,1504]" captionStart="Figure 19" captionStartId="29.[142,218,1650,1669]" captionTargetBox="[243,1320,193,1620]" captionTargetId="figure-0@29.[243,1320,193,1620]" captionTargetPageId="29" captionText="Figure 19. Fossil and recent Catherinum. C–K, N–Q, T, V, Catherinum anglicum sp. nov., Upper Campanian, Belemnitella mucronata zone, Keswick, Norwich, UK; (O) is the holotype, the other specimens are paratypes. C, D, F, G, carinolatera in external (C, D, F) and internal (G, H) aspects (C, NHMUK IC 1203; D, 1204; F–G, 1206; H, 1208). E, transverse section of carina (NHMUK IC 1205); I, T, dorsal view of carinae (NHMUK IC 1209, 1216); P, lateral view of carina (NHMUK IC 1214); V, internal view of base of carina (NHMUK IC 1217); J, K, O, internal views of scuta, to show large pit for dwarf males (NHMUK IC 1210, 1211, 1213); N, external view of scutum (NHMUK IC 1212); Q, external view of tergum (NHMUK IC 1215). A, B, L, M, R, S, U, Catherinum hispidum (G.O. Sars, 1890). Recent, Norwegian Sea. A, B, carinolatus in external and internal views. L, M, external and internal views of scutum. R, external view of tergum. S, U, dorsal and internal views of carina. Scale bars: S, U, V, 1 mm; A–D, F, G, K–O, Q, R, T, 0.5 mm; E, H, P, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5357846" httpUri="https://zenodo.org/record/5357846/files/figure.png" pageId="35" pageNumber="301">Fig. 19E, I, P, T, V</figureCitation>
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) is gently incurved towards the tergum, and the parietes are of even width; intraparietes are not developed. The tectal ridges are robust and broad (
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<figureCitation box="[355,485,1574,1596]" captionStart="Figure 19" captionStartId="29.[142,218,1650,1669]" captionTargetBox="[243,1320,193,1620]" captionTargetId="figure-0@29.[243,1320,193,1620]" captionTargetPageId="29" captionText="Figure 19. Fossil and recent Catherinum. C–K, N–Q, T, V, Catherinum anglicum sp. nov., Upper Campanian, Belemnitella mucronata zone, Keswick, Norwich, UK; (O) is the holotype, the other specimens are paratypes. C, D, F, G, carinolatera in external (C, D, F) and internal (G, H) aspects (C, NHMUK IC 1203; D, 1204; F–G, 1206; H, 1208). E, transverse section of carina (NHMUK IC 1205); I, T, dorsal view of carinae (NHMUK IC 1209, 1216); P, lateral view of carina (NHMUK IC 1214); V, internal view of base of carina (NHMUK IC 1217); J, K, O, internal views of scuta, to show large pit for dwarf males (NHMUK IC 1210, 1211, 1213); N, external view of scutum (NHMUK IC 1212); Q, external view of tergum (NHMUK IC 1215). A, B, L, M, R, S, U, Catherinum hispidum (G.O. Sars, 1890). Recent, Norwegian Sea. A, B, carinolatus in external and internal views. L, M, external and internal views of scutum. R, external view of tergum. S, U, dorsal and internal views of carina. Scale bars: S, U, V, 1 mm; A–D, F, G, K–O, Q, R, T, 0.5 mm; E, H, P, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5357846" httpUri="https://zenodo.org/record/5357846/files/figure.png" pageId="35" pageNumber="301">Fig. 19E, T</figureCitation>
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), and the tectal surface between these is slightly convex. The tergum (
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<figureCitation box="[655,749,1604,1626]" captionStart="Figure 19" captionStartId="29.[142,218,1650,1669]" captionTargetBox="[243,1320,193,1620]" captionTargetId="figure-0@29.[243,1320,193,1620]" captionTargetPageId="29" captionText="Figure 19. Fossil and recent Catherinum. C–K, N–Q, T, V, Catherinum anglicum sp. nov., Upper Campanian, Belemnitella mucronata zone, Keswick, Norwich, UK; (O) is the holotype, the other specimens are paratypes. C, D, F, G, carinolatera in external (C, D, F) and internal (G, H) aspects (C, NHMUK IC 1203; D, 1204; F–G, 1206; H, 1208). E, transverse section of carina (NHMUK IC 1205); I, T, dorsal view of carinae (NHMUK IC 1209, 1216); P, lateral view of carina (NHMUK IC 1214); V, internal view of base of carina (NHMUK IC 1217); J, K, O, internal views of scuta, to show large pit for dwarf males (NHMUK IC 1210, 1211, 1213); N, external view of scutum (NHMUK IC 1212); Q, external view of tergum (NHMUK IC 1215). A, B, L, M, R, S, U, Catherinum hispidum (G.O. Sars, 1890). Recent, Norwegian Sea. A, B, carinolatus in external and internal views. L, M, external and internal views of scutum. R, external view of tergum. S, U, dorsal and internal views of carina. Scale bars: S, U, V, 1 mm; A–D, F, G, K–O, Q, R, T, 0.5 mm; E, H, P, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5357846" httpUri="https://zenodo.org/record/5357846/files/figure.png" pageId="35" pageNumber="301">Fig. 19Q</figureCitation>
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) is elongated and triangular, and the carinal margin is slightly convex. Three radial ridges run from the umbo, subparallel with the occludent margin. The scuta (
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<figureCitation box="[150,336,1727,1749]" captionStart="Figure 19" captionStartId="29.[142,218,1650,1669]" captionTargetBox="[243,1320,193,1620]" captionTargetId="figure-0@29.[243,1320,193,1620]" captionTargetPageId="29" captionText="Figure 19. Fossil and recent Catherinum. C–K, N–Q, T, V, Catherinum anglicum sp. nov., Upper Campanian, Belemnitella mucronata zone, Keswick, Norwich, UK; (O) is the holotype, the other specimens are paratypes. C, D, F, G, carinolatera in external (C, D, F) and internal (G, H) aspects (C, NHMUK IC 1203; D, 1204; F–G, 1206; H, 1208). E, transverse section of carina (NHMUK IC 1205); I, T, dorsal view of carinae (NHMUK IC 1209, 1216); P, lateral view of carina (NHMUK IC 1214); V, internal view of base of carina (NHMUK IC 1217); J, K, O, internal views of scuta, to show large pit for dwarf males (NHMUK IC 1210, 1211, 1213); N, external view of scutum (NHMUK IC 1212); Q, external view of tergum (NHMUK IC 1215). A, B, L, M, R, S, U, Catherinum hispidum (G.O. Sars, 1890). Recent, Norwegian Sea. A, B, carinolatus in external and internal views. L, M, external and internal views of scutum. R, external view of tergum. S, U, dorsal and internal views of carina. Scale bars: S, U, V, 1 mm; A–D, F, G, K–O, Q, R, T, 0.5 mm; E, H, P, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5357846" httpUri="https://zenodo.org/record/5357846/files/figure.png" pageId="35" pageNumber="301">Fig. 19J, K, N, O</figureCitation>
|
|||
|
|
) are concavo–convex, trapezoidal, and the umbo is pointed and incurved; the upper latus margin has a distinctive concavity just beneath the tergal–upper latus angle. A narrow groove runs parallel with the occludent margin. On the interior of the valve, a deep, oval dwarf male receptacle lies between the apex and the scutal adductor scar. The carinolatus (
|
|||
|
|
<figureCitation box="[812,1014,227,249]" captionStart="Figure 19" captionStartId="29.[142,218,1650,1669]" captionTargetBox="[243,1320,193,1620]" captionTargetId="figure-0@29.[243,1320,193,1620]" captionTargetPageId="29" captionText="Figure 19. Fossil and recent Catherinum. C–K, N–Q, T, V, Catherinum anglicum sp. nov., Upper Campanian, Belemnitella mucronata zone, Keswick, Norwich, UK; (O) is the holotype, the other specimens are paratypes. C, D, F, G, carinolatera in external (C, D, F) and internal (G, H) aspects (C, NHMUK IC 1203; D, 1204; F–G, 1206; H, 1208). E, transverse section of carina (NHMUK IC 1205); I, T, dorsal view of carinae (NHMUK IC 1209, 1216); P, lateral view of carina (NHMUK IC 1214); V, internal view of base of carina (NHMUK IC 1217); J, K, O, internal views of scuta, to show large pit for dwarf males (NHMUK IC 1210, 1211, 1213); N, external view of scutum (NHMUK IC 1212); Q, external view of tergum (NHMUK IC 1215). A, B, L, M, R, S, U, Catherinum hispidum (G.O. Sars, 1890). Recent, Norwegian Sea. A, B, carinolatus in external and internal views. L, M, external and internal views of scutum. R, external view of tergum. S, U, dorsal and internal views of carina. Scale bars: S, U, V, 1 mm; A–D, F, G, K–O, Q, R, T, 0.5 mm; E, H, P, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5357846" httpUri="https://zenodo.org/record/5357846/files/figure.png" pageId="35" pageNumber="301">Fig. 19C, D, F–H</figureCitation>
|
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) has a low, incurved umbo, and a tall carinal margin. The long, straight upper latus and inframedian latus margins are set at right angles. The plate is very typical of
|
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|
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<taxonomicName authorityName="ZEVINA" authorityYear="1978" box="[1058,1192,319,340]" class="Maxillopoda" family="Scalpellidae" genus="Catherinum" kingdom="Animalia" order="Pedunculata" pageId="35" pageNumber="301" phylum="Arthropoda" rank="genus">
|
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|
|
<emphasis box="[1058,1192,319,340]" italics="true" pageId="35" pageNumber="301">Catherinum</emphasis>
|
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|
|
</taxonomicName>
|
|||
|
|
of the
|
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|
<taxonomicName authorityName="G. O. Sars" authorityYear="1890" box="[1270,1419,319,340]" class="Maxillopoda" family="Scalpellidae" genus="Scalpellum" kingdom="Animalia" order="Pedunculata" pageId="35" pageNumber="301" phylum="Arthropoda" rank="species" species="hispidum">
|
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|
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<emphasis box="[1270,1419,319,340]" italics="true" pageId="35" pageNumber="301">Ca. hispidum</emphasis>
|
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</taxonomicName>
|
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|
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(
|
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<figureCitation box="[812,943,350,372]" captionStart="Figure 19" captionStartId="29.[142,218,1650,1669]" captionTargetBox="[243,1320,193,1620]" captionTargetId="figure-0@29.[243,1320,193,1620]" captionTargetPageId="29" captionText="Figure 19. Fossil and recent Catherinum. C–K, N–Q, T, V, Catherinum anglicum sp. nov., Upper Campanian, Belemnitella mucronata zone, Keswick, Norwich, UK; (O) is the holotype, the other specimens are paratypes. C, D, F, G, carinolatera in external (C, D, F) and internal (G, H) aspects (C, NHMUK IC 1203; D, 1204; F–G, 1206; H, 1208). E, transverse section of carina (NHMUK IC 1205); I, T, dorsal view of carinae (NHMUK IC 1209, 1216); P, lateral view of carina (NHMUK IC 1214); V, internal view of base of carina (NHMUK IC 1217); J, K, O, internal views of scuta, to show large pit for dwarf males (NHMUK IC 1210, 1211, 1213); N, external view of scutum (NHMUK IC 1212); Q, external view of tergum (NHMUK IC 1215). A, B, L, M, R, S, U, Catherinum hispidum (G.O. Sars, 1890). Recent, Norwegian Sea. A, B, carinolatus in external and internal views. L, M, external and internal views of scutum. R, external view of tergum. S, U, dorsal and internal views of carina. Scale bars: S, U, V, 1 mm; A–D, F, G, K–O, Q, R, T, 0.5 mm; E, H, P, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5357846" httpUri="https://zenodo.org/record/5357846/files/figure.png" pageId="35" pageNumber="301">Fig. 19A, B</figureCitation>
|
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) and
|
|||
|
|
<taxonomicName authorityName="G. O. Sars" authorityYear="1877" box="[1014,1178,350,371]" class="Maxillopoda" family="Scalpellidae" genus="Scalpellum" kingdom="Animalia" order="Pedunculata" pageId="35" pageNumber="301" phylum="Arthropoda" rank="species" species="striolatum">
|
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|
|
<emphasis box="[1014,1178,350,371]" italics="true" pageId="35" pageNumber="301">Ca. striolatum</emphasis>
|
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|
|
</taxonomicName>
|
|||
|
|
(
|
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|
|
<figureCitation box="[1196,1311,350,372]" captionStart="Figure 7" captionStartId="11.[142,221,1682,1701]" captionTargetBox="[231,1332,193,1652]" captionTargetId="figure-0@11.[231,1332,193,1652]" captionTargetPageId="11" captionText="Figure 7. Evolution of lateral plate morphology, dorsal side: A–E, Weltnerium stroemii (M. Sars, 1859); F–J, Weltnerium nymphocola (Hoek, 1883); K–O, Catherinum novazelandiae (Hoek, 1883); P–T, Catherinum striolatum (G.O. Sars, 1877); U–Y, Amigdoscalpellum rigidum (Aurivillius, 1898). External aspect of carinolatus (A, F, K, P, U); interior of carinolatus (B, G, L, Q, V); external view of inframedian latus (C, H, M, R, W); dorsal (carinal) view of basal capitulum (D, I, N, S, X); enlarged lateral view of basal capitulum (E, J, O, T, Y). Note the changes in the nature of the articulation between the carinolatera of a pair (interpenetrant, X; opposed double articulation, D, I, N, S) and those in the shape of the inframedian latus (triangular, W; waisted hourglass, M, R; pentagonal, C, H). See Table 1 for details of localities. Scale bars: A–C, F–H, M, P-R, U–V, 0.5 mm; D, E, I, J, N, O, S, T, X, Y, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5357828" httpUri="https://zenodo.org/record/5357828/files/figure.png" pageId="35" pageNumber="301">Fig. 7P, Q</figureCitation>
|
|||
|
|
)
|
|||
|
|
<typeStatus box="[1328,1381,350,371]" pageId="35" pageNumber="301">type</typeStatus>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph blockId="35.[804,1420,434,1313]" box="[804,905,434,455]" pageId="35" pageNumber="301">
|
|||
|
|
<emphasis box="[804,905,434,455]" italics="true" pageId="35" pageNumber="301">Remarks</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph blockId="35.[804,1420,434,1313]" pageId="35" pageNumber="301">
|
|||
|
|
<taxonomicName authority="Gale, 2016" authorityName="Gale" authorityYear="2016" box="[804,1052,465,486]" class="Maxillopoda" family="Scalpellidae" genus="Catherinum" kingdom="Animalia" order="Pedunculata" pageId="35" pageNumber="301" phylum="Arthropoda" rank="species" species="anglicum" status="sp. nov.">
|
|||
|
|
<emphasis box="[804,1052,465,486]" italics="true" pageId="35" pageNumber="301">Catherinum anglicum</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel box="[1058,1141,465,486]" pageId="35" pageNumber="301" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
is perhaps closest to the extant
|
|||
|
|
<taxonomicName authorityName="G. O. Sars" authorityYear="1890" box="[891,1046,495,516]" class="Maxillopoda" family="Scalpellidae" genus="Scalpellum" kingdom="Animalia" order="Pedunculata" pageId="35" pageNumber="301" phylum="Arthropoda" rank="species" species="hispidum">
|
|||
|
|
<emphasis box="[891,1046,495,516]" italics="true" pageId="35" pageNumber="301">Ca. hispidum</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
from the Norwegian Sea. The carinae are very similar indeed, both carrying very broad, robust tectal ridges (
|
|||
|
|
<figureCitation box="[1133,1291,556,578]" captionStart="Figure 19" captionStartId="29.[142,218,1650,1669]" captionTargetBox="[243,1320,193,1620]" captionTargetId="figure-0@29.[243,1320,193,1620]" captionTargetPageId="29" captionText="Figure 19. Fossil and recent Catherinum. C–K, N–Q, T, V, Catherinum anglicum sp. nov., Upper Campanian, Belemnitella mucronata zone, Keswick, Norwich, UK; (O) is the holotype, the other specimens are paratypes. C, D, F, G, carinolatera in external (C, D, F) and internal (G, H) aspects (C, NHMUK IC 1203; D, 1204; F–G, 1206; H, 1208). E, transverse section of carina (NHMUK IC 1205); I, T, dorsal view of carinae (NHMUK IC 1209, 1216); P, lateral view of carina (NHMUK IC 1214); V, internal view of base of carina (NHMUK IC 1217); J, K, O, internal views of scuta, to show large pit for dwarf males (NHMUK IC 1210, 1211, 1213); N, external view of scutum (NHMUK IC 1212); Q, external view of tergum (NHMUK IC 1215). A, B, L, M, R, S, U, Catherinum hispidum (G.O. Sars, 1890). Recent, Norwegian Sea. A, B, carinolatus in external and internal views. L, M, external and internal views of scutum. R, external view of tergum. S, U, dorsal and internal views of carina. Scale bars: S, U, V, 1 mm; A–D, F, G, K–O, Q, R, T, 0.5 mm; E, H, P, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5357846" httpUri="https://zenodo.org/record/5357846/files/figure.png" pageId="35" pageNumber="301">Fig. 19I, S, T</figureCitation>
|
|||
|
|
), and with shallow short grooves for articulation with the carinolatus in the interior (
|
|||
|
|
<figureCitation box="[1139,1278,617,639]" captionStart="Figure 19" captionStartId="29.[142,218,1650,1669]" captionTargetBox="[243,1320,193,1620]" captionTargetId="figure-0@29.[243,1320,193,1620]" captionTargetPageId="29" captionText="Figure 19. Fossil and recent Catherinum. C–K, N–Q, T, V, Catherinum anglicum sp. nov., Upper Campanian, Belemnitella mucronata zone, Keswick, Norwich, UK; (O) is the holotype, the other specimens are paratypes. C, D, F, G, carinolatera in external (C, D, F) and internal (G, H) aspects (C, NHMUK IC 1203; D, 1204; F–G, 1206; H, 1208). E, transverse section of carina (NHMUK IC 1205); I, T, dorsal view of carinae (NHMUK IC 1209, 1216); P, lateral view of carina (NHMUK IC 1214); V, internal view of base of carina (NHMUK IC 1217); J, K, O, internal views of scuta, to show large pit for dwarf males (NHMUK IC 1210, 1211, 1213); N, external view of scutum (NHMUK IC 1212); Q, external view of tergum (NHMUK IC 1215). A, B, L, M, R, S, U, Catherinum hispidum (G.O. Sars, 1890). Recent, Norwegian Sea. A, B, carinolatus in external and internal views. L, M, external and internal views of scutum. R, external view of tergum. S, U, dorsal and internal views of carina. Scale bars: S, U, V, 1 mm; A–D, F, G, K–O, Q, R, T, 0.5 mm; E, H, P, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5357846" httpUri="https://zenodo.org/record/5357846/files/figure.png" pageId="35" pageNumber="301">Fig. 19U, V</figureCitation>
|
|||
|
|
). The terga are similar in shape, but the scutal margin is convex in
|
|||
|
|
<taxonomicName authorityName="G. O. Sars" authorityYear="1890" box="[835,988,679,700]" class="Maxillopoda" family="Scalpellidae" genus="Scalpellum" kingdom="Animalia" order="Pedunculata" pageId="35" pageNumber="301" phylum="Arthropoda" rank="species" species="hispidum">
|
|||
|
|
<emphasis box="[835,988,679,700]" italics="true" pageId="35" pageNumber="301">Ca. hispidum</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and straight in
|
|||
|
|
<taxonomicName authority="Gale, 2016" authorityName="Gale" authorityYear="2016" box="[1177,1329,679,700]" class="Maxillopoda" family="Scalpellidae" genus="Catherinum" kingdom="Animalia" order="Pedunculata" pageId="35" pageNumber="301" phylum="Arthropoda" rank="species" species="anglicum" status="sp. nov.">
|
|||
|
|
<emphasis box="[1177,1329,679,700]" italics="true" pageId="35" pageNumber="301">Ca. anglicum</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel box="[1336,1420,679,700]" pageId="35" pageNumber="301" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
Three divergent ribs run from the umbo, adjacent to the occludent margin in
|
|||
|
|
<taxonomicName authority="Gale, 2016" authorityName="Gale" authorityYear="2016" box="[1073,1218,740,761]" class="Maxillopoda" family="Scalpellidae" genus="Catherinum" kingdom="Animalia" order="Pedunculata" pageId="35" pageNumber="301" phylum="Arthropoda" rank="species" species="anglicum" status="sp. nov.">
|
|||
|
|
<emphasis box="[1073,1218,740,761]" italics="true" pageId="35" pageNumber="301">Ca. anglicum</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel box="[1224,1307,740,761]" pageId="35" pageNumber="301" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
, but these are absent in the living species (
|
|||
|
|
<figureCitation box="[1163,1291,770,792]" captionStart="Figure 19" captionStartId="29.[142,218,1650,1669]" captionTargetBox="[243,1320,193,1620]" captionTargetId="figure-0@29.[243,1320,193,1620]" captionTargetPageId="29" captionText="Figure 19. Fossil and recent Catherinum. C–K, N–Q, T, V, Catherinum anglicum sp. nov., Upper Campanian, Belemnitella mucronata zone, Keswick, Norwich, UK; (O) is the holotype, the other specimens are paratypes. C, D, F, G, carinolatera in external (C, D, F) and internal (G, H) aspects (C, NHMUK IC 1203; D, 1204; F–G, 1206; H, 1208). E, transverse section of carina (NHMUK IC 1205); I, T, dorsal view of carinae (NHMUK IC 1209, 1216); P, lateral view of carina (NHMUK IC 1214); V, internal view of base of carina (NHMUK IC 1217); J, K, O, internal views of scuta, to show large pit for dwarf males (NHMUK IC 1210, 1211, 1213); N, external view of scutum (NHMUK IC 1212); Q, external view of tergum (NHMUK IC 1215). A, B, L, M, R, S, U, Catherinum hispidum (G.O. Sars, 1890). Recent, Norwegian Sea. A, B, carinolatus in external and internal views. L, M, external and internal views of scutum. R, external view of tergum. S, U, dorsal and internal views of carina. Scale bars: S, U, V, 1 mm; A–D, F, G, K–O, Q, R, T, 0.5 mm; E, H, P, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5357846" httpUri="https://zenodo.org/record/5357846/files/figure.png" pageId="35" pageNumber="301">Fig. 19Q, R</figureCitation>
|
|||
|
|
). The scuta both possess an incurved umbo, and a large, deep pit for dwarf males (
|
|||
|
|
<figureCitation box="[999,1125,832,854]" captionStart="Figure 19" captionStartId="29.[142,218,1650,1669]" captionTargetBox="[243,1320,193,1620]" captionTargetId="figure-0@29.[243,1320,193,1620]" captionTargetPageId="29" captionText="Figure 19. Fossil and recent Catherinum. C–K, N–Q, T, V, Catherinum anglicum sp. nov., Upper Campanian, Belemnitella mucronata zone, Keswick, Norwich, UK; (O) is the holotype, the other specimens are paratypes. C, D, F, G, carinolatera in external (C, D, F) and internal (G, H) aspects (C, NHMUK IC 1203; D, 1204; F–G, 1206; H, 1208). E, transverse section of carina (NHMUK IC 1205); I, T, dorsal view of carinae (NHMUK IC 1209, 1216); P, lateral view of carina (NHMUK IC 1214); V, internal view of base of carina (NHMUK IC 1217); J, K, O, internal views of scuta, to show large pit for dwarf males (NHMUK IC 1210, 1211, 1213); N, external view of scutum (NHMUK IC 1212); Q, external view of tergum (NHMUK IC 1215). A, B, L, M, R, S, U, Catherinum hispidum (G.O. Sars, 1890). Recent, Norwegian Sea. A, B, carinolatus in external and internal views. L, M, external and internal views of scutum. R, external view of tergum. S, U, dorsal and internal views of carina. Scale bars: S, U, V, 1 mm; A–D, F, G, K–O, Q, R, T, 0.5 mm; E, H, P, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5357846" httpUri="https://zenodo.org/record/5357846/files/figure.png" pageId="35" pageNumber="301">Fig. 19J–O</figureCitation>
|
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|
|
), which is directed slightly more transversely in
|
|||
|
|
<taxonomicName authorityName="G. O. Sars" authorityYear="1890" box="[1071,1220,862,884]" class="Maxillopoda" family="Scalpellidae" genus="Scalpellum" kingdom="Animalia" order="Pedunculata" pageId="35" pageNumber="301" phylum="Arthropoda" rank="species" species="hispidum">
|
|||
|
|
<emphasis box="[1071,1220,862,884]" italics="true" pageId="35" pageNumber="301">Ca. hispidum</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. The carinolatera are similar in shape (
|
|||
|
|
<figureCitation box="[1048,1243,893,915]" captionStart="Figure 19" captionStartId="29.[142,218,1650,1669]" captionTargetBox="[243,1320,193,1620]" captionTargetId="figure-0@29.[243,1320,193,1620]" captionTargetPageId="29" captionText="Figure 19. Fossil and recent Catherinum. C–K, N–Q, T, V, Catherinum anglicum sp. nov., Upper Campanian, Belemnitella mucronata zone, Keswick, Norwich, UK; (O) is the holotype, the other specimens are paratypes. C, D, F, G, carinolatera in external (C, D, F) and internal (G, H) aspects (C, NHMUK IC 1203; D, 1204; F–G, 1206; H, 1208). E, transverse section of carina (NHMUK IC 1205); I, T, dorsal view of carinae (NHMUK IC 1209, 1216); P, lateral view of carina (NHMUK IC 1214); V, internal view of base of carina (NHMUK IC 1217); J, K, O, internal views of scuta, to show large pit for dwarf males (NHMUK IC 1210, 1211, 1213); N, external view of scutum (NHMUK IC 1212); Q, external view of tergum (NHMUK IC 1215). A, B, L, M, R, S, U, Catherinum hispidum (G.O. Sars, 1890). Recent, Norwegian Sea. A, B, carinolatus in external and internal views. L, M, external and internal views of scutum. R, external view of tergum. S, U, dorsal and internal views of carina. Scale bars: S, U, V, 1 mm; A–D, F, G, K–O, Q, R, T, 0.5 mm; E, H, P, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5357846" httpUri="https://zenodo.org/record/5357846/files/figure.png" pageId="35" pageNumber="301">Fig. 19A–D, F–H</figureCitation>
|
|||
|
|
), but the tergal margin is longer in
|
|||
|
|
<taxonomicName authority="Gale, 2016" authorityName="Gale" authorityYear="2016" box="[1052,1205,923,945]" class="Maxillopoda" family="Scalpellidae" genus="Catherinum" kingdom="Animalia" order="Pedunculata" pageId="35" pageNumber="301" phylum="Arthropoda" rank="species" species="anglicum" status="sp. nov.">
|
|||
|
|
<emphasis box="[1052,1205,923,945]" italics="true" pageId="35" pageNumber="301">Ca. anglicum</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel box="[1211,1296,924,945]" pageId="35" pageNumber="301" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
It can be distinguished from small carinae of
|
|||
|
|
<taxonomicName class="Maxillopoda" family="Scalpellidae" genus="Diotascalpellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="35" pageNumber="301" phylum="Arthropoda" rank="species" species="fossula">
|
|||
|
|
<emphasis italics="true" pageId="35" pageNumber="301">Diotascalpellum fossula</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by the absence of intraparietes. A further undescribed species of
|
|||
|
|
<taxonomicName authorityName="ZEVINA" authorityYear="1978" box="[1098,1242,1015,1036]" class="Maxillopoda" family="Scalpellidae" genus="Catherinum" kingdom="Animalia" order="Pedunculata" pageId="35" pageNumber="301" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis box="[1098,1242,1015,1036]" italics="true" pageId="35" pageNumber="301">Catherinum</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is present in the Norwich chalk assemblage; this will be named subsequently.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph blockId="35.[804,1420,434,1313]" pageId="35" pageNumber="301">
|
|||
|
|
This is a very small species: the scuta have a maximum height of
|
|||
|
|
<quantity box="[1037,1103,1138,1159]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="35" pageNumber="301" unit="mm" value="2.0">2 mm</quantity>
|
|||
|
|
, and the total height of the individuals was probably not more than
|
|||
|
|
<quantity box="[1269,1336,1168,1190]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="35" pageNumber="301" unit="mm" value="5.0">5 mm</quantity>
|
|||
|
|
. It was possibly attached to living crustaceans or pycnogonids, by analogy with the small extant species
|
|||
|
|
<taxonomicName authority="(Hoek, 1883)" baseAuthorityName="Hoek" baseAuthorityYear="1883" class="Maxillopoda" family="Scalpellidae" genus="Weltnerium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="35" pageNumber="301" phylum="Arthropoda" rank="species" species="nymphocola">
|
|||
|
|
<emphasis italics="true" pageId="35" pageNumber="301">Weltnerium nymphocola</emphasis>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation author="Hoek PPC" box="[957,1088,1260,1282]" pageId="35" pageNumber="301" pagination="169" refId="ref23201" refString="Hoek PPC. 1883. Report on the Cirripedia. Report on Scientific Results from the exploratory Voyages of the Challenger. Zoology 8: 169." type="journal article" year="1883">Hoek, 1883</bibRefCitation>
|
|||
|
|
)
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName authority="Weltner" authorityName="Weltner" class="Maxillopoda" family="Scalpellidae" genus="Verum" kingdom="Animalia" order="Pedunculata" pageId="35" pageNumber="301" phylum="Arthropoda" rank="species" species="brachiumcancri">
|
|||
|
|
<emphasis box="[1157,1419,1260,1282]" italics="true" pageId="35" pageNumber="301">Verum brachiumcancri</emphasis>
|
|||
|
|
Weltner
|
|||
|
|
</taxonomicName>
|
|||
|
|
(see
|
|||
|
|
<bibRefCitation author="Buhl-Mortensen L & Hoeg JT" box="[959,1318,1291,1313]" pageId="35" pageNumber="301" pagination="545 - 557" refId="ref22423" refString="Buhl-Mortensen L, Hoeg JT. 2012. Reproductive strategy of two deep-sea scalpellid barnacles (Crustacea: Cirripedia: Thoracica) associated with pycnogonids and decapods and the first description of a penis in dwarf males. Organisms, Diversity and Evolution 13: 545 - 557." type="journal article" year="2012">Buhl-Mortensen & Høeg, 2012</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|