<documentid="2123370B77FBB5B33D95765E98247185"ID-DOI="10.5852/ejt.2021.737.1259"ID-GBIF-Dataset="4b7a838b-1449-4ae4-90ff-5a7f113868e0"ID-ISSN="2118-9773"ID-Zenodo-Dep="4594357"ID-ZooBank="FE4ED74A-3FE6-4CA6-A116-CB3AF46826F7"IM.materialsCitations_approvedBy="valdenar"IM.metadata_approvedBy="felipe"IM.tables_approvedBy="felipe"IM.taxonomicNames_approvedBy="felipe"IM.treatmentCitations_approvedBy="valdenar"IM.treatments_approvedBy="valdenar"checkinTime="1615409071345"checkinUser="plazi"docAuthor="Goulding, Tricia C., Bourke, Adam J., Comendador, Joseph, Khalil, Munawar, Quang, Ngo Xuan, Tan, Shau Hwai, Tan, Siong Kiat & Dayrat, Benoît"docDate="2021"docId="03A6D248FFD18B05DEDFFD70FB78D369"docLanguage="en"docName="ejt-737_1-133.pdf"docOrigin="European Journal of Taxonomy 737 (1)"docSource="http://dx.doi.org/10.5852/ejt.2021.737.1259"docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1"docStyleId="EF2B578F1D15862ADE45B0C07C620911"docStyleName="EJT.2018-.journal_article.type1"docStyleVersion="14"docTitle="Platevindex applanatus Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat 2021, comb. nov."docType="treatment"docVersion="15"lastPageNumber="88"masterDocId="FF9FAA30FF9C8B52DF38FFAFFFC3D05C"masterDocTitle="Systematic revision of Platevindex Baker, 1938 (Gastropoda: Euthyneura: Onchidiidae)"masterLastPageNumber="133"masterPageNumber="1"pageNumber="78"updateTime="1733327641084"updateUser="valdenar"zenodo-license-document="CC-BY-4.0"zenodo-license-figures="CC-BY-4.0"zenodo-license-treatments="UNSPECIFIED">
<mods:affiliationid="4676F82AEE0D13FA205A6A009A26123B">Department of Biology, Pennsylvania State University, University Park, PA 16802, USA. & Smithsonian Institution, National Museum of Natural History, PO Box 37012, MRC 163, Washington, DC, 20013, USA.</mods:affiliation>
<mods:affiliationid="7436533139A47C4149EE670152F94963">College of Engineering, Information Technology and the Environment, Charles Darwin University, Ellengowan Dr., Casuarina, NT 0810, Australia.</mods:affiliation>
<mods:affiliationid="EE497BF1DFDD311B093D23107185E0BF">National Museum of the Philippines, Taft Ave., Ermita, Manila, 1000 Metro Manila, Philippines.</mods:affiliation>
<mods:affiliationid="65E106F5D98DD4FC71A86A56B5677781">Department of Marine Science, Faculty of Agriculture, Universitas Malikussaleh, Reuleut Main Campus, Kecamatan Muara Batu, North Aceh, Aceh, 24355, Indonesia.</mods:affiliation>
<mods:affiliationid="786A422DF957E2D8BA4A3F5791DED157">Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. & Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam.</mods:affiliation>
<mods:affiliationid="29D7D7FBF4655066EB2763DF83EEDB9A">Centre for Marine and Coastal Studies, Universiti Sains Malaysia, 11800 Minden Penang, Malaysia. & Marine Science Laboratory, School of Biological Sciences, Universiti Sains Malaysia, 11800 Minden Penang, Malaysia.</mods:affiliation>
<mods:affiliationid="CB10C8A1975EE4D847C7ABAE56597CED">Lee Kong Chian Natural History Museum, 2 Conservatory Dr, National University of Singapore, 117377, Singapore.</mods:affiliation>
<mods:affiliationid="FFE3DAFE01491D68C192A25EA8EF47D8">Department of Biology, Pennsylvania State University, University Park, PA 16802, USA.</mods:affiliation>
<taxonomicNameid="4C0F18DDFFD18B1FDEDFFD70FC07D2A5"ID-CoL="872JJ"authority="(Simroth, 1920)"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Simroth"baseAuthorityYear="1920"box="[487,964,735,762]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="77"pageNumber="78"phylum="Mollusca"rank="species"species="applanatus"status="comb. nov.">
<bibRefCitationid="EF9E1EAFFFD18B1FDC29FD70FC7FD2A5"author="Simroth H."box="[785,956,735,761]"pageId="77"pageNumber="78"pagination="259 - 306"refId="ref74467"refString="Simroth H. 1920. Uber einige Nacktschnecken vom Malayischen Archipel von Lombok an ostwarts bis zu den Gesellschafts-Inseln. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 35: 259 - 306. Available from https: // biodiversitylibrary. org / page / 48256899 [accessed 22 Dec. 2018]."type="journal article"year="1920">Simroth, 1920</bibRefCitation>
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<bibRefCitationid="EF9E1EAFFFD18B1FDEE2FCEBFD02D302"author="Simroth H."box="[474,705,836,862]"pageId="77"pageNumber="78"pagination="259 - 306"refId="ref74467"refString="Simroth H. 1920. Uber einige Nacktschnecken vom Malayischen Archipel von Lombok an ostwarts bis zu den Gesellschafts-Inseln. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 35: 259 - 306. Available from https: // biodiversitylibrary. org / page / 48256899 [accessed 22 Dec. 2018]."type="journal article"year="1920">Simroth, 1920: 294</bibRefCitation>
<locationid="8ED03585FFD18B1FDCD3FC78FAC2D3AD"LSID="urn:lsid:plazi:treatment:03A6D248FFD18B05DEDFFD70FB78D369:8ED03585FFD18B1FDCD3FC78FAC2D3AD"box="[1003,1281,983,1010]"country="Indonesia"municipality="Kobroor"name="Strand von Manumbai"pageId="77"pageNumber="78">Strand von Manumbai</location>
<collectingDateid="EFF5BC76FFD18B1FDB28FC55FB71D448"box="[1040,1202,1018,1045]"pageId="77"pageNumber="78"value="1908-03-13">13 Mar. 1908</collectingDate>
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<collectingDateid="EFF5BC76FFD18B1FDDFFF91EFCA8D697"box="[711,875,1713,1739]"pageId="77"pageNumber="78"value="2013-03-10">10 Mar. 2013</collectingDate>
</date>
; station 85;
<collectedFromid="381320A3FFD18B1FDCC7F91FFE65D6B2"pageId="77"pageNumber="78">sand, rocks and pieces of wood outside a mangrove</collectedFrom>
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<collectedFromid="381320A3FFD18B1FDA39F8F3FC64D7C5"pageId="77"pageNumber="78">mangrove with dark brown mud, slugs on rotting petiole of palm frond</collectedFrom>
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<bibRefCitationid="EF9E1EAFFFD28B1CDD7BF978FD2DD6AD"author="Simroth H."box="[579,750,1751,1777]"pageId="78"pageNumber="79"pagination="259 - 306"refId="ref74467"refString="Simroth H. 1920. Uber einige Nacktschnecken vom Malayischen Archipel von Lombok an ostwarts bis zu den Gesellschafts-Inseln. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 35: 259 - 306. Available from https: // biodiversitylibrary. org / page / 48256899 [accessed 22 Dec. 2018]."type="journal article"year="1920">Simroth, 1920</bibRefCitation>
<materialsCitationid="3B676903FFD38B1DDF85FEA6FE18D135"ID-GBIF-Occurrence="3051754359"collectedFrom="limestone rocky intertidal"collectingDate="2012-11-09"collectionCode="MNHN"collectorName="Papua Niugini"country="Papua New Guinea"latitude="-5.0033336"location="South Dumduman Island"longLatPrecision="130"longitude="145.79333"pageId="79"pageNumber="80"specimenCode="MNHN-IM-2013-12493"specimenCount="1"stateProvince="Madang">
<collectingCountryid="F31823CEFFD38B1DDF85FEA6FE1AD17F"box="[189,473,265,291]"name="Papua New Guinea"pageId="79"pageNumber="80">PAPUA NEW GUINEA</collectingCountry>
<collectingRegionid="49CBADBCFFD38B1DDECEFEA6FD9DD17F"box="[502,606,265,291]"country="Papua New Guinea"name="Madang"pageId="79"pageNumber="80">Madang</collectingRegion>
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<collectingRegionid="49CBADBCFFD38B1DDBF0FDE9FAA7D23C"box="[1224,1380,582,608]"country="Papua New Guinea"name="New Ireland"pageId="79"pageNumber="80">New Ireland</collectingRegion>
; MNHN expedition Kavieng 2014 leg.; station KM05;
<collectedFromid="381320A3FFD38B1DDC7EFD23FEEDD295"pageId="79"pageNumber="80">mixed hard platform and seagrass bed at outlet of rivulet</collectedFrom>
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<locationid="8ED03585FFD38B1DDD85FCD1FC52D3C4"LSID="urn:lsid:plazi:treatment:03A6D248FFD18B05DEDFFD70FB78D369:8ED03585FFD38B1DDD85FCD1FC52D3C4"box="[701,913,894,920]"country="Vanuatu"latitude="-15.581667"longLatPrecision="128"longitude="167.04"name="Santo Rose Point"pageId="79"pageNumber="80">Santo Rose Point</location>
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<materialsCitationid="3B676903FFD38B1DDF85FC6BFB0CD382"ID-GBIF-Occurrence="3051754423"box="[189,1231,963,991]"collectedFrom="intertidal, coral sand"collectingDate="2006-09-10"collectionCode="MNHN"country="Vanuatu"latitude="-15.581667"location="Santo Rose Point"longLatPrecision="128"longitude="167.04"pageId="79"pageNumber="80"specimenCode="MNHN-IM-2013-62404"specimenCount="1">
<taxonomicNameid="4C0F18DDFFD38B1DDE1BF8F6FD3ED72F"authority="(Simroth, 1920)"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Simroth"baseAuthorityYear="1920"box="[291,765,1881,1907]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="79"pageNumber="80"phylum="Mollusca"rank="species"species="applanatus"status="comb. nov.">
<bibRefCitationid="EF9E1EAFFFD38B1DDD71F8F6FD36D72F"author="Simroth H."box="[585,757,1881,1907]"pageId="79"pageNumber="80"pagination="259 - 306"refId="ref74467"refString="Simroth H. 1920. Uber einige Nacktschnecken vom Malayischen Archipel von Lombok an ostwarts bis zu den Gesellschafts-Inseln. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 35: 259 - 306. Available from https: // biodiversitylibrary. org / page / 48256899 [accessed 22 Dec. 2018]."type="journal article"year="1920">Simroth, 1920</bibRefCitation>
<emphasisid="B97BBF4CFFCC8B02DFD2FE98FD00D10D"bold="true"box="[234,707,311,337]"pageId="80"pageNumber="81">Color and morphology of live animals</emphasis>
(
<figureCitationid="13347FDBFFCC8B02DDEBFE98FCEAD10E"box="[723,809,311,338]"captionStart="Fig"captionStartId="78.[189,232,1751,1777]"captionTargetBox="[229,1358,266,1727]"captionTargetId="figure-143@78.[229,1358,266,1727]"captionTargetPageId="78"captionText="Fig. 36. Platevindex applanatus (Simroth, 1920) comb. nov., live animals. A. Dorsal view, 16 mm long [2893], Indonesia, Kei Islands (UMIZ 00102). B. Dorsal view, 16 mm long [2895], Indonesia, Kei Islands (UMIZ 00102). C. Dorsal view, 15 mm long [2904], Indonesia, Kei Islands (UMIZ 00103). D. Dorsal view, 16 mm long [3563], Indonesia, Kei Islands (UMIZ 00102). E. Dorsal view, 18 mm long [2757], Indonesia, Ambon (UMIZ 00101). F. Dorsal view, 5 mm long [2925], Indonesia, Kei Islands (UMIZ 00104).G. Ventral view, same as E. H. Ventral view, 12 mm long [2900], Indonesia, Kei Islands (UMIZ 00102). I. Dorsal view, 9 mm long [3152], Mauritius (MNHN-IM-2019-1393). J. Same as I. K. Ventral view, same as I."figureDoi="http://doi.org/10.5281/zenodo.4594426"httpUri="https://zenodo.org/record/4594426/files/figure.png"pageId="80"pageNumber="81">Fig. 36</figureCitation>
)
</paragraph>
<paragraphid="8BB0635EFFCC8B02DF85FEF5FE67D21B"blockId="80.[188,1399,311,583]"pageId="80"pageNumber="81">Live animals are not covered with mud and their natural color can be seen without washing. The notum is bumpy with prominent, distinctly-raised papillae (with or without dorsal eyes). Live animals may become nearly hemispherical when disturbed. The dorsal color is variable although it is typically brown with yellow longitudinal (irregular) lines. Slugs with an entirely dark brown notum or a light beige notum also occur. The hyponotum is light grey and almost translucent in small specimens. The margin of the hyponotum is frequently translucent with the brown color of the notum showing through. The foot is light yellow.</paragraph>
Papillae with dorsal eyes are present. Their exact number is difficult to determine, as they are often retracted, but ranges approximately from 4 to 26, except in the case of very small specimens, in which as few as 2 dorsal eyes have been observed. Each papilla bears one dorsal eye. Dorsal eyes may be distributed across the notum or only present in the middle but are always absent on the margin (i.e., eyes are never <
<figureCitationid="13347FDBFFCC8B02DEF0FC92FDE4D30B"box="[456,551,829,855]"captionStart="Fig"captionStartId="14.[189,232,1714,1740]"captionTargetBox="[193,1395,1067,1681]"captionTargetId="figure-507@14.[189,1398,1056,1711]"captionTargetPageId="14"captionText="Fig. 1. Intestinal loops of type I. Small black arrows show the direction of the intestinal transport. The clockwise loop is in blue. The counterclockwise loop is in yellow. The transitional loop (between clockwise and counterclockwise loops) is in green. Red arrows indicate the orientation of the transitional loop. A. Type I (as defined by Plate 1893), with a transitional loop oriented at 3 o’clock, redrawn from Plate (1893: pl. 8, fig. 29). B. Type I, with a transitional loop oriented at 3 o’clock, Platevindex luteus (Semper, 1880) [5613] (ITBZC IM 00016), Vietnam. C. Type I, with a transitional loop oriented at 1 o’clock,P. applanatus (Simroth, 1920) [5420] (MNHN IM-2013-13765), Papua New Guinea.D. Type I, with a transitional loop oriented at 6 o’clock, P. luteus [2837] (UMIZ 00089), Ambon, Indonesia. Scale bars: B–C = 2 mm; D = 3 mm."figureDoi="http://doi.org/10.5281/zenodo.4594359"httpUri="https://zenodo.org/record/4594359/files/figure.png"pageId="80"pageNumber="81">Figs 1C</figureCitation>
,
<figureCitationid="13347FDBFFCC8B02DD0CFC92FD43D30B"box="[564,640,829,855]"captionStart-0="Fig"captionStart-1="Fig"captionStart-2="Fig"captionStart-3="Fig"captionStartId-0="79.[189,232,1880,1906]"captionStartId-1="80.[188,231,1885,1911]"captionStartId-2="81.[189,232,1822,1848]"captionStartId-3="82.[188,231,1716,1742]"captionTargetBox-0="[189,1398,1039,1855]"captionTargetBox-1="[189,1398,1044,1859]"captionTargetBox-2="[189,1398,978,1796]"captionTargetBox-3="[214,1363,634,1696]"captionTargetId-0="figure-73@79.[189,1398,1039,1863]"captionTargetId-1="figure-329@80.[189,1398,1044,1875]"captionTargetId-2="figure-417@81.[189,1398,978,1804]"captionTargetId-3="figure-299@82.[189,1398,626,1707]"captionTargetPageId-0="79"captionTargetPageId-1="80"captionTargetPageId-2="81"captionTargetPageId-3="82"captionText-0="Fig. 37. Platevindex applanatus (Simroth, 1920) comb. nov., radula, Indonesia, Ambon. A. Rachidian and innermost lateral teeth [2754] (UMIZ 00101). B. lateral teeth with rachidian teeth, same as A. C. Lateral teeth [2755] (UMIZ 00101). D. Outermost lateral teeth, same as A. Scale bars: A, D = 30 μm; B = 50 μm; C = 100 μm."captionText-1="Fig. 38. Platevindex applanatus (Simroth, 1920) comb. nov., radula, Mauritius. A. Left, outermost, lateral teeth [3604] (MNHN-IM-2019-1392). B. Rachidian tooth and innermost lateral teeth, same as A. C. Rachidian tooth and left, innermost lateral teeth [3152] (MNHN-IM-2019-1393). D. Right, lateral teeth [3442] (MNHN-IM-2019-1394). Scale bars: A = 10 μm; B–D = 20 μm."captionText-2="Fig. 39. Platevindex applanatus (Simroth, 1920) comb. nov., radula, Papua New Guinea. A. Rachidian and innermost lateral teeth [5450] (MNHN-IM-2013-14051). B. Outer lateral teeth, same as A. C. Inner lateral teeth, same as A. D. Rachidian and innermost lateral teeth [5424] (MNHN-IM-2013-14043). Scale bars: A = 20 μm; B, D = 30 μm; C = 50 μm."captionText-3="Fig. 40. Platevindex applanatus (Simroth, 1920) comb. nov., digestive system, intestinal loops of type I. A. Dorsal view, type I with a transitional loop oriented at 2 o’clock, Kei, Indonesia [2892] (UMIZ 00102). B. Dorsal view, type I with a transitional loop oriented at 2 o’clock, Mauritius [3152] (MNHN- IM-2019-1393). C. Dorsal view, type I with a transitional loop oriented at 1 o’clock, Papua New Guinea [5420] (MNHN-IM-2013-13765). D. Stomach, ventral view, same as C. Abbreviations: ddg = dorsal digestive gland; e = esophagus; i = intestine; oldg = opening of the lateral lobe of the digestive gland; pdg = posterior digestive gland; rg = rectal gland; st = stomach; st1 = stomach chamber 1; st2 = stomach chamber 2; st3 = stomach chamber 3; st4 = stomach chamber 4. Scale bars: A, D = 3 mm; B = 2 mm; C = 4 mm."figureDoi-0="http://doi.org/10.5281/zenodo.4594428"figureDoi-1="http://doi.org/10.5281/zenodo.4594430"figureDoi-2="http://doi.org/10.5281/zenodo.4594432"figureDoi-3="http://doi.org/10.5281/zenodo.4594434"httpUri-0="https://zenodo.org/record/4594428/files/figure.png"httpUri-1="https://zenodo.org/record/4594430/files/figure.png"httpUri-2="https://zenodo.org/record/4594432/files/figure.png"httpUri-3="https://zenodo.org/record/4594434/files/figure.png"pageId="80"pageNumber="81">37–40</figureCitation>
in length. Examples of radular formulae are presented in
<tableCitationid="C68D56E5FFCC8B02DBE6FCCFFAFBD327"box="[1246,1336,864,891]"captionStart="Table 5"captionStartId="42.[189,258,265,291]"captionTargetBox="[223,1387,323,1985]"captionTargetPageId="42"captionText="Table 5. Radular formulae in species of Platevindex Baker, 1938."httpUri="http://table.plazi.org/id/DF7033D6FFB68B78DF85FEA6FC7DD17F"pageId="80"pageNumber="81"tableUuid="DF7033D6FFB68B78DF85FEA6FC7DD17F">Table 5</tableCitation>
. Due to the high genetic divergence (in COI sequences) between populations of
<taxonomicNameid="4C0F18DDFFCC8B02DB7BFC2BFB21D3C1"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Simroth"baseAuthorityYear="1920"box="[1091,1250,899,925]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="80"pageNumber="81"phylum="Mollusca"rank="species"species="applanatus">
<collectingCountryid="F31823CEFFCC8B02DDB4FC08FCB2D39D"box="[652,881,935,961]"name="Papua New Guinea"pageId="80"pageNumber="81">Papua New Guinea</collectingCountry>
(
<figureCitationid="13347FDBFFCC8B02DCBAFC08FBCED39D"box="[898,1037,935,961]"captionStart-0="Fig"captionStart-1="Fig"captionStart-2="Fig"captionStartId-0="79.[189,232,1880,1906]"captionStartId-1="80.[188,231,1885,1911]"captionStartId-2="81.[189,232,1822,1848]"captionTargetBox-0="[189,1398,1039,1855]"captionTargetBox-1="[189,1398,1044,1859]"captionTargetBox-2="[189,1398,978,1796]"captionTargetId-0="figure-73@79.[189,1398,1039,1863]"captionTargetId-1="figure-329@80.[189,1398,1044,1875]"captionTargetId-2="figure-417@81.[189,1398,978,1804]"captionTargetPageId-0="79"captionTargetPageId-1="80"captionTargetPageId-2="81"captionText-0="Fig. 37. Platevindex applanatus (Simroth, 1920) comb. nov., radula, Indonesia, Ambon. A. Rachidian and innermost lateral teeth [2754] (UMIZ 00101). B. lateral teeth with rachidian teeth, same as A. C. Lateral teeth [2755] (UMIZ 00101). D. Outermost lateral teeth, same as A. Scale bars: A, D = 30 μm; B = 50 μm; C = 100 μm."captionText-1="Fig. 38. Platevindex applanatus (Simroth, 1920) comb. nov., radula, Mauritius. A. Left, outermost, lateral teeth [3604] (MNHN-IM-2019-1392). B. Rachidian tooth and innermost lateral teeth, same as A. C. Rachidian tooth and left, innermost lateral teeth [3152] (MNHN-IM-2019-1393). D. Right, lateral teeth [3442] (MNHN-IM-2019-1394). Scale bars: A = 10 μm; B–D = 20 μm."captionText-2="Fig. 39. Platevindex applanatus (Simroth, 1920) comb. nov., radula, Papua New Guinea. A. Rachidian and innermost lateral teeth [5450] (MNHN-IM-2013-14051). B. Outer lateral teeth, same as A. C. Inner lateral teeth, same as A. D. Rachidian and innermost lateral teeth [5424] (MNHN-IM-2013-14043). Scale bars: A = 20 μm; B, D = 30 μm; C = 50 μm."figureDoi-0="http://doi.org/10.5281/zenodo.4594428"figureDoi-1="http://doi.org/10.5281/zenodo.4594430"figureDoi-2="http://doi.org/10.5281/zenodo.4594432"httpUri-0="https://zenodo.org/record/4594428/files/figure.png"httpUri-1="https://zenodo.org/record/4594430/files/figure.png"httpUri-2="https://zenodo.org/record/4594432/files/figure.png"pageId="80"pageNumber="81">Figs 37–39</figureCitation>
I, with a transitional loop oriented between 1 and 2 o’clock (
<figureCitationid="13347FDBFFCC8B02DC51FC65FC0BD3B8"box="[873,968,970,996]"captionStart="Fig"captionStartId="14.[189,232,1714,1740]"captionTargetBox="[193,1395,1067,1681]"captionTargetId="figure-507@14.[189,1398,1056,1711]"captionTargetPageId="14"captionText="Fig. 1. Intestinal loops of type I. Small black arrows show the direction of the intestinal transport. The clockwise loop is in blue. The counterclockwise loop is in yellow. The transitional loop (between clockwise and counterclockwise loops) is in green. Red arrows indicate the orientation of the transitional loop. A. Type I (as defined by Plate 1893), with a transitional loop oriented at 3 o’clock, redrawn from Plate (1893: pl. 8, fig. 29). B. Type I, with a transitional loop oriented at 3 o’clock, Platevindex luteus (Semper, 1880) [5613] (ITBZC IM 00016), Vietnam. C. Type I, with a transitional loop oriented at 1 o’clock,P. applanatus (Simroth, 1920) [5420] (MNHN IM-2013-13765), Papua New Guinea.D. Type I, with a transitional loop oriented at 6 o’clock, P. luteus [2837] (UMIZ 00089), Ambon, Indonesia. Scale bars: B–C = 2 mm; D = 3 mm."figureDoi="http://doi.org/10.5281/zenodo.4594359"httpUri="https://zenodo.org/record/4594359/files/figure.png"pageId="80"pageNumber="81">Figs 1C</figureCitation>
,
<figureCitationid="13347FDBFFCC8B02DCEEFC65FC36D3B8"box="[982,1013,970,996]"captionStart="Fig"captionStartId="82.[188,231,1716,1742]"captionTargetBox="[214,1363,634,1696]"captionTargetId="figure-299@82.[189,1398,626,1707]"captionTargetPageId="82"captionText="Fig. 40. Platevindex applanatus (Simroth, 1920) comb. nov., digestive system, intestinal loops of type I. A. Dorsal view, type I with a transitional loop oriented at 2 o’clock, Kei, Indonesia [2892] (UMIZ 00102). B. Dorsal view, type I with a transitional loop oriented at 2 o’clock, Mauritius [3152] (MNHN- IM-2019-1393). C. Dorsal view, type I with a transitional loop oriented at 1 o’clock, Papua New Guinea [5420] (MNHN-IM-2013-13765). D. Stomach, ventral view, same as C. Abbreviations: ddg = dorsal digestive gland; e = esophagus; i = intestine; oldg = opening of the lateral lobe of the digestive gland; pdg = posterior digestive gland; rg = rectal gland; st = stomach; st1 = stomach chamber 1; st2 = stomach chamber 2; st3 = stomach chamber 3; st4 = stomach chamber 4. Scale bars: A, D = 3 mm; B = 2 mm; C = 4 mm."figureDoi="http://doi.org/10.5281/zenodo.4594434"httpUri="https://zenodo.org/record/4594434/files/figure.png"pageId="80"pageNumber="81">40</figureCitation>
<taxonomicNameid="4C0F18DDFFCC8B02DE10F8F2FCC9D72B"authority="(Simroth, 1920)"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Simroth"baseAuthorityYear="1920"box="[296,778,1885,1911]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="80"pageNumber="81"phylum="Mollusca"rank="species"species="applanatus"status="comb. nov.">
<bibRefCitationid="EF9E1EAFFFCC8B02DD6BF8F2FCC1D72B"author="Simroth H."box="[595,770,1885,1911]"pageId="80"pageNumber="81"pagination="259 - 306"refId="ref74467"refString="Simroth H. 1920. Uber einige Nacktschnecken vom Malayischen Archipel von Lombok an ostwarts bis zu den Gesellschafts-Inseln. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 35: 259 - 306. Available from https: // biodiversitylibrary. org / page / 48256899 [accessed 22 Dec. 2018]."type="journal article"year="1920">Simroth, 1920</bibRefCitation>
In the posterior part of the reproductive system, the oviduct is wider than the deferent duct (up to twice as wide). Its distal section (distal to the spermatheca) is slightly longer than its proximal section (up to two or three times as wide) or equal to it in length. The deferent duct is longer than the oviduct, loosely attached to it, and varies from slightly convoluted with a few loose loops to tightly coiled with U-shaped loops (
<figureCitationid="13347FDBFFCD8B03DE35FE16FEA5D18F"box="[269,358,441,467]"captionStart="Fig"captionStartId="83.[190,233,1855,1881]"captionTargetBox="[224,1373,611,1829]"captionTargetId="figure-206@83.[189,1398,601,1853]"captionTargetPageId="83"captionText="Fig. 41. Platevindex applanatus (Simroth, 1920) comb. nov., posterior reproductive system. A. Kei, Indonesia [2892] (UMIZ 00102). B. Mauritius [3152] (MNHN-IM-2019-1393). C. Papua New Guinea [5420] (MNHN-IM-2013-13765). Abbreviations: dd = deferent duct; fgm = female gland mass; hd = hermaphroditic duct; hg = hermaphroditic gland; ov = oviduct; rs = receptaculum seminis; sp = spermatheca. Scale bars: A, C = 2 mm; B = 1 mm."figureDoi="http://doi.org/10.5281/zenodo.4594437"httpUri="https://zenodo.org/record/4594437/files/figure.png"pageId="81"pageNumber="82">Fig. 41</figureCitation>
). The position of the female pore varies from being adjacent to the anus to
away from it in large specimens. The flexible, distal region of the penis with hooks measures approximately
<quantityid="4CF7CEBBFFCD8B03DF85FE50FE8FD246"box="[189,332,511,538]"metricMagnitude="-3"metricUnit="m"metricValue="2.25"metricValueMax="3.0"metricValueMin="1.5"pageId="81"pageNumber="82"unit="mm"value="2.25"valueMax="3.0"valueMin="1.5">1.5 to 3 mm</quantity>
<quantityid="4CF7CEBBFFCD8B03DC9FFE50FBE3D246"box="[935,1056,511,538]"metricMagnitude="-3"metricUnit="m"metricValue="2.5"metricValueMax="3.0"metricValueMin="2.0"pageId="81"pageNumber="82"unit="mm"value="2.5"valueMax="3.0"valueMin="2.0">2 to 3 mm</quantity>
<collectingCountryid="F31823CEFFCD8B03DF85FD8CFE60D260"box="[189,419,546,573]"name="Papua New Guinea"pageId="81"pageNumber="82">Papua New Guinea</collectingCountry>
it is approximately
<quantityid="4CF7CEBBFFCD8B03DDACFD8DFCE4D261"box="[660,807,546,573]"metricMagnitude="-3"metricUnit="m"metricValue="1.75"metricValueMax="2.0"metricValueMin="1.5"pageId="81"pageNumber="82"unit="mm"value="1.75"valueMax="2.0"valueMin="1.5">1.5 to 2 mm</quantity>
long. The penial hooks are large and can be seen inside the semi-transparent penis. They are usually
<quantityid="4CF7CEBBFFCD8B03DC21FDE9FC6AD23C"box="[793,937,582,609]"metricMagnitude="-2"metricUnit="m"metricValue="6.75"metricValueMax="9.0"metricValueMin="4.5"pageId="81"pageNumber="82"unit="mm"value="67.5"valueMax="90.0"valueMin="45.0">45 to 90 µm</quantity>
long (
<figureCitationid="13347FDBFFCD8B03DCCCFDE9FB8CD23C"box="[1012,1103,582,608]"captionStart="Fig"captionStartId="85.[191,234,698,724]"captionTargetBox="[189,1398,271,671]"captionTargetId="figure-192@85.[189,1398,265,681]"captionTargetPageId="85"captionText="Fig. 43. Platevindex applanatus (Simroth, 1920) comb. nov., penial hooks. Indonesia, Ambon [2757] (UMIZ 00101). Scale bars: A = 50 μm; B = 60 μm."figureDoi="http://doi.org/10.5281/zenodo.4594441"httpUri="https://zenodo.org/record/4594441/files/figure.png"pageId="81"pageNumber="82">Figs 43</figureCitation>
,
<figureCitationid="13347FDBFFCD8B03DB63FDE9FB51D23C"box="[1115,1170,582,608]"captionStart="Fig"captionStartId="85.[189,232,1926,1952]"captionTargetBox="[189,1398,1192,1901]"captionTargetId="figure-194@85.[189,1398,1192,1918]"captionTargetPageId="85"captionText="Fig. 44. Platevindex applanatus (Simroth, 1920) comb. nov., penial hooks. A. Papua New Guinea [5424] (MNHN-IM-2013-14043). B–C. Mauritius [3152] (MNHN-IM-2019-1393). Scale bars: A = 30 μm; B = 10 μm; C = 60 μm."figureDoi="http://doi.org/10.5281/zenodo.4594443"httpUri="https://zenodo.org/record/4594443/files/figure.png"pageId="81"pageNumber="82">44A</figureCitation>
), but between
<quantityid="4CF7CEBBFFCD8B03DA04FDE9FEC9D2DF"metricMagnitude="-2"metricUnit="m"metricValue="4.0"metricValueMax="6.0"metricValueMin="2.0"pageId="81"pageNumber="82"unit="mm"value="40.0"valueMax="60.0"valueMin="20.0">20 to 60 µm</quantity>
, except in a few specimens in which it inserts approximately ¾ down the length of the visceral cavity. It inserts approximately halfway to ¾ down the length of the visceral cavity in specimens from
<collectingCountryid="F31823CEFFCD8B03DBD4FD7CFED1D34C"name="Papua New Guinea"pageId="81"pageNumber="82">Papua New Guinea</collectingCountry>
(and near the posterior end of the visceral cavity in one specimen). In
, it inserts at the posterior end of the visceral cavity in the largest specimen, and by the heart halfway down the visceral cavity in a smaller specimen. The length of the retractor muscle varies from much shorter than the penial sheath (
<taxonomicNameid="4C0F18DDFFCD8B03DE19F8B1FD3AD764"authority="(Simroth, 1920)"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Simroth"baseAuthorityYear="1920"box="[289,761,1822,1848]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="81"pageNumber="82"phylum="Mollusca"rank="species"species="applanatus"status="comb. nov.">
<bibRefCitationid="EF9E1EAFFFCD8B03DD7DF8B1FD33D764"author="Simroth H."box="[581,752,1822,1848]"pageId="81"pageNumber="82"pagination="259 - 306"refId="ref74467"refString="Simroth H. 1920. Uber einige Nacktschnecken vom Malayischen Archipel von Lombok an ostwarts bis zu den Gesellschafts-Inseln. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 35: 259 - 306. Available from https: // biodiversitylibrary. org / page / 48256899 [accessed 22 Dec. 2018]."type="journal article"year="1920">Simroth, 1920</bibRefCitation>
<tableCitationid="C68D56E5FFCE8B00DD7CFEA6FD5DD17F"box="[580,670,265,291]"captionStart="Table 4"captionStartId="22.[184,253,658,686]"captionTargetBox="[198,1916,827,1626]"captionTargetPageId="22"captionText="Table 4 (continued on next page).Summary of internaland external traits in speciesofPlatevindex Baker, 1938.Traits are described in the corresponding species descriptions and vary among individuals. External traits include the length of the animal, number of dorsal eyes, color of the foot and other distinguishing characters such as the color of the hyponotum. Internal traits include the insertion of the retractor muscle (RM), the length of the retractor muscle compared to that of the penial sheath (PS), the length of the region with penial hooks and the type of the intestinal loops."httpUri="http://table.plazi.org/id/DF7033D6FF8A8B44DF80FD3DF8C6D344"pageId="82"pageNumber="83"tableUuid="DF7033D6FF8A8B44DF80FD3DF8C6D344">Table 4</tableCitation>
<taxonomicNameid="4C0F18DDFFCE8B00DE7EFE98FD9FD10D"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Simroth"baseAuthorityYear="1920"box="[326,604,311,337]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="82"pageNumber="83"phylum="Mollusca"rank="species"species="applanatus">
<taxonomicNameid="4C0F18DDFFCE8B00DCCEFE97FBBFD10D"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[1014,1148,312,337]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="82"pageNumber="83"phylum="Mollusca"rank="species"species="martensi">
<taxonomicNameid="4C0F18DDFFCE8B00DB81FE97FAD0D10D"authority="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat, 2021"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"box="[1209,1299,312,337]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="82"pageNumber="83"phylum="Mollusca"rank="species"species="aptei"status="sp. nov.">
by the light yellow color of its foot. The presence of prominent dorsal papillae helps to distinguish
<taxonomicNameid="4C0F18DDFFCE8B00DF85FED0FE9CD1C4"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Simroth"baseAuthorityYear="1920"box="[189,351,382,408]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="82"pageNumber="83"phylum="Mollusca"rank="species"species="applanatus">
, which is characterized by similar prominent dorsal papillae. The dorsal coloration of
<taxonomicNameid="4C0F18DDFFCE8B00DC23FE0EFBECD1E7"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Simroth"baseAuthorityYear="1920"box="[795,1071,417,443]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="82"pageNumber="83"phylum="Mollusca"rank="species"species="applanatus">
<taxonomicNameid="4C0F18DDFFCE8B00DE89FE47FD91D25D"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Simroth"baseAuthorityYear="1920"box="[433,594,487,513]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="82"pageNumber="83"phylum="Mollusca"rank="species"species="applanatus">
<taxonomicNameid="4C0F18DDFFCE8B00DC48FDA4FBCDD279"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Simroth"baseAuthorityYear="1920"box="[880,1038,523,549]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="82"pageNumber="83"phylum="Mollusca"rank="species"species="applanatus">
also reach a smaller maximum size than individuals of
<taxonomicNameid="4C0F18DDFFCE8B00DEF7FD80FD92D214"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[463,593,559,584]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="82"pageNumber="83"phylum="Mollusca"rank="species"species="martensi">
<taxonomicNameid="4C0F18DDFFCE8B00DDBFFD80FCD4D214"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[647,791,558,584]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="82"pageNumber="83"phylum="Mollusca"rank="species"species="amboinae">
<taxonomicNameid="4C0F18DDFFCE8B00DE1AF91BFD38D692"authority="(Simroth, 1920)"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Simroth"baseAuthorityYear="1920"box="[290,763,1716,1742]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="82"pageNumber="83"phylum="Mollusca"rank="species"species="applanatus"status="comb. nov.">
<bibRefCitationid="EF9E1EAFFFCE8B00DD7FF91BFD30D692"author="Simroth H."box="[583,755,1716,1742]"pageId="82"pageNumber="83"pagination="259 - 306"refId="ref74467"refString="Simroth H. 1920. Uber einige Nacktschnecken vom Malayischen Archipel von Lombok an ostwarts bis zu den Gesellschafts-Inseln. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 35: 259 - 306. Available from https: // biodiversitylibrary. org / page / 48256899 [accessed 22 Dec. 2018]."type="journal article"year="1920">Simroth, 1920</bibRefCitation>
<taxonomicNameid="4C0F18DDFFCF8B01DD5FFEC1FCCDD1D4"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Simroth"baseAuthorityYear="1920"box="[615,782,366,392]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="83"pageNumber="84"phylum="Mollusca"rank="species"species="applanatus">
. However, the orientation of the transitional loop can be used to reliably distinguish
<taxonomicNameid="4C0F18DDFFCF8B01DCB1FE1AFBE8D192"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Simroth"baseAuthorityYear="1920"box="[905,1067,436,462]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="83"pageNumber="84"phylum="Mollusca"rank="species"species="applanatus">
<taxonomicNameid="4C0F18DDFFCF8B01DDECFE78FC2ED1AD"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Simroth"baseAuthorityYear="1920"box="[724,1005,471,497]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="83"pageNumber="84"phylum="Mollusca"rank="species"species="applanatus">
based on reproductive anatomy. The deferent duct in the posterior reproductive system is more closely attached to the oviduct, longer and more highly coiled in
<taxonomicNameid="4C0F18DDFFCF8B01DCF7FDB1FBB3D264"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Simroth"baseAuthorityYear="1920"box="[975,1136,542,568]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="83"pageNumber="84"phylum="Mollusca"rank="species"species="applanatus">
. The distal region of the oviduct (distal to the spermatheca) is also longer in
<taxonomicNameid="4C0F18DDFFC88B06DB33FEA5FB69D17F"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Simroth"baseAuthorityYear="1920"box="[1035,1194,265,291]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="84"pageNumber="85"phylum="Mollusca"rank="species"species="applanatus">
. Finally, the distal region of the penis with hooks is longer in
<taxonomicNameid="4C0F18DDFFC88B06DC43FE82FBDBD11A"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Simroth"baseAuthorityYear="1920"box="[891,1048,300,326]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="84"pageNumber="85"phylum="Mollusca"rank="species"species="applanatus">
). The insertion of the retractor muscle varies greatly within
<taxonomicNameid="4C0F18DDFFC88B06DB15FEFFFB0DD135"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Simroth"baseAuthorityYear="1920"box="[1069,1230,335,361]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="84"pageNumber="85"phylum="Mollusca"rank="species"species="applanatus">
<taxonomicNameid="4C0F18DDFFCF8B01DE1FF890FCC4D706"authority="(Simroth, 1920)"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Simroth"baseAuthorityYear="1920"box="[295,775,1855,1882]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="83"pageNumber="84"phylum="Mollusca"rank="species"species="applanatus"status="comb. nov.">
<bibRefCitationid="EF9E1EAFFFCF8B01DD69F890FD3CD706"author="Simroth H."box="[593,767,1855,1882]"pageId="83"pageNumber="84"pagination="259 - 306"refId="ref74467"refString="Simroth H. 1920. Uber einige Nacktschnecken vom Malayischen Archipel von Lombok an ostwarts bis zu den Gesellschafts-Inseln. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 35: 259 - 306. Available from https: // biodiversitylibrary. org / page / 48256899 [accessed 22 Dec. 2018]."type="journal article"year="1920">Simroth, 1920</bibRefCitation>
<figureCitationid="13347FDBFFC88B06DE5EFE78FE12D1AD"box="[358,465,471,497]"captionStart="Fig"captionStartId="36.[189,232,1716,1742]"captionTargetBox="[189,1398,741,1688]"captionTargetId="figure-232@36.[189,1398,741,1688]"captionTargetPageId="36"captionText="Fig. 10. Geographic distributions of species of Platevindex. Colored areas correspond to hypothetical ranges proposed based on all known records, and the colors are the same as those used for each species in the phylogenetic trees (Figs 1–5). Distinct colors are used for each subspecies and mitochondrial unit in P. coriaceus (Semper, 1880) as well as for each subspecies of P. burnupi (Collinge, 1902). Black dots show sites sampled for this study and red dots (China and Japan) show localities of previously published genetic sequences included in the phylogenetic analyses. A. Platevindex luteus (Semper, 1880), P. applanatus (Simroth, 1920), P. burnupi. B. Platevindex tigrinus (Stoliczka, 1869), P. coriaceus coriaceus (units #1, #2, and #3) and P. coriaceus darwinensis subsp. nov. C. Platevindex martensi (Plate, 1893), P. aptei Goulding & Dayrat sp. nov., P. amboinae (Plate, 1893) and P. latus (Plate, 1893)."figureDoi="http://doi.org/10.5281/zenodo.4594373"httpUri="https://zenodo.org/record/4594373/files/figure.png"pageId="84"pageNumber="85">Fig. 10A</figureCitation>
<collectingCountryid="F31823CEFFC88B06DCCDFDA9FB19D27C"box="[1013,1242,518,544]"name="Papua New Guinea"pageId="84"pageNumber="85">Papua New Guinea</collectingCountry>
:
<collectingRegionid="49CBADBCFFC88B06DBDDFDA9FA86D27C"box="[1253,1349,518,544]"country="Papua New Guinea"name="Madang"pageId="84"pageNumber="85">Madang</collectingRegion>
and
<collectingRegionid="49CBADBCFFC88B06DF84FD86FE91D21F"box="[188,338,553,579]"country="Papua New Guinea"name="New Ireland"pageId="84"pageNumber="85">New Ireland</collectingRegion>
<figureCitationid="13347FDBFFC88B06DE15FDC5FE40D2D9"box="[301,387,618,645]"captionStart="Fig"captionStartId="86.[189,232,1663,1689]"captionTargetBox="[189,1398,264,1630]"captionTargetId="figure-140@86.[189,1398,264,1630]"captionTargetPageId="86"captionText="Fig. 45. Habitats of Platevindex applanatus (Simroth, 1920) comb. nov., Indonesia (A–C), Papua New Guinea (D–E) and Mauritius (F). A. Ambon, log behind a rocky mangrove in the very high intertidal, shaded by trees (station 127). B. Kei Islands, rocks with algae at the edge of the mangrove (station 137). C. Kei Islands, Un, rocks in the high intertidal behind a mangrove, surrounded by dry vegetation, algae growing on the rocks (station 137). D. Madang, Hargun Island, night tide (station PM24). E. New Ireland, Povalval, east coast of New Ireland, mixed hard platform and seagrass bed at outlet of rivulet (station KM05). F. Mahebourg, close up of rotting petiole of palm frond at margin of mangrove where juvenile onchidiids were found."figureDoi="http://doi.org/10.5281/zenodo.4594445"httpUri="https://zenodo.org/record/4594445/files/figure.png"pageId="84"pageNumber="85">Fig. 45</figureCitation>
<taxonomicNameid="4C0F18DDFFC88B06DF85FD36FE0CD2EF"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Simroth"baseAuthorityYear="1920"box="[189,463,665,691]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="84"pageNumber="85"phylum="Mollusca"rank="species"species="applanatus">
is primarily found in the high intertidal, including on limestone, on rocks covered with a thin layer of algae. In the Kei Islands, individuals were found on rocks at the upper margin of the intertidal near a mangrove. In
, individuals were found on logs very high in the intertidal, including at one locality that was not near any mangrove. In mangroves, individuals were typically
<taxonomicNameid="4C0F18DDFFC88B06DE20F8EFFD22D706"authority="(Simroth, 1920)"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Simroth"baseAuthorityYear="1920"box="[280,737,1856,1882]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="84"pageNumber="85"phylum="Mollusca"rank="species"species="applanatus"status="comb. nov.">
<bibRefCitationid="EF9E1EAFFFC88B06DD0BF8EFFD1AD706"author="Simroth H."box="[563,729,1856,1882]"pageId="84"pageNumber="85"pagination="259 - 306"refId="ref74467"refString="Simroth H. 1920. Uber einige Nacktschnecken vom Malayischen Archipel von Lombok an ostwarts bis zu den Gesellschafts-Inseln. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 35: 259 - 306. Available from https: // biodiversitylibrary. org / page / 48256899 [accessed 22 Dec. 2018]."type="journal article"year="1920">Simroth,1920</bibRefCitation>
<taxonomicNameid="4C0F18DDFFC98B07DE1CFD15FCC2D288"authority="(Simroth, 1920)"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Simroth"baseAuthorityYear="1920"box="[292,769,698,724]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="85"pageNumber="86"phylum="Mollusca"rank="species"species="applanatus"status="comb. nov.">
<bibRefCitationid="EF9E1EAFFFC98B07DD74FD15FD3AD288"author="Simroth H."box="[588,761,698,724]"pageId="85"pageNumber="86"pagination="259 - 306"refId="ref74467"refString="Simroth H. 1920. Uber einige Nacktschnecken vom Malayischen Archipel von Lombok an ostwarts bis zu den Gesellschafts-Inseln. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 35: 259 - 306. Available from https: // biodiversitylibrary. org / page / 48256899 [accessed 22 Dec. 2018]."type="journal article"year="1920">Simroth, 1920</bibRefCitation>
, it was found in mangroves, on pieces of wood and wet petioles of palm leaves.
<taxonomicNameid="4C0F18DDFFC98B07DD33FCF9FCDCD32C"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Simroth"baseAuthorityYear="1920"box="[523,799,854,880]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="85"pageNumber="86"phylum="Mollusca"rank="species"species="applanatus">
, the retractor muscle inserts near the heart). Therefore,
<taxonomicNameid="4C0F18DDFFCB8B05DEABFEE0FD6BD135"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Simroth"baseAuthorityYear="1920"box="[403,680,335,361]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="87"pageNumber="88"phylum="Mollusca"rank="species"species="applanatus">
<taxonomicNameid="4C0F18DDFFC98B07DE25F828FD2CD7FD"authority="(Simroth, 1920)"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Simroth"baseAuthorityYear="1920"box="[285,751,1927,1953]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="85"pageNumber="86"phylum="Mollusca"rank="species"species="applanatus"status="comb. nov.">
<bibRefCitationid="EF9E1EAFFFC98B07DD05F828FD25D7FD"author="Simroth H."box="[573,742,1927,1953]"pageId="85"pageNumber="86"pagination="259 - 306"refId="ref74467"refString="Simroth H. 1920. Uber einige Nacktschnecken vom Malayischen Archipel von Lombok an ostwarts bis zu den Gesellschafts-Inseln. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 35: 259 - 306. Available from https: // biodiversitylibrary. org / page / 48256899 [accessed 22 Dec. 2018]."type="journal article"year="1920">Simroth, 1920</bibRefCitation>
<taxonomicNameid="4C0F18DDFFCA8B04DE93F9D0FC40D6C6"authority="(Simroth, 1920)"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Simroth"baseAuthorityYear="1920"box="[427,899,1663,1690]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="86"pageNumber="87"phylum="Mollusca"rank="species"species="applanatus"status="comb. nov.">
<bibRefCitationid="EF9E1EAFFFCA8B04DDF7F9D0FCB8D6C6"author="Simroth H."box="[719,891,1663,1690]"pageId="86"pageNumber="87"pagination="259 - 306"refId="ref74467"refString="Simroth H. 1920. Uber einige Nacktschnecken vom Malayischen Archipel von Lombok an ostwarts bis zu den Gesellschafts-Inseln. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 35: 259 - 306. Available from https: // biodiversitylibrary. org / page / 48256899 [accessed 22 Dec. 2018]."type="journal article"year="1920">Simroth, 1920</bibRefCitation>
<bibRefCitationid="EF9E1EAFFFCB8B05DF85FE14FE2DD18A"author="Hoffmann H."box="[189,494,443,470]"pageId="87"pageNumber="88"pagination="29 - 118"refId="ref72283"refString="Hoffmann H. 1928. Zur Kenntnis der Oncidiiden. Zoologische Jahrbucher (Jena) 55: 29 - 118."type="journal article"year="1928">Hoffmann (1928: 84–85)</bibRefCitation>
and
<bibRefCitationid="EF9E1EAFFFCB8B05DD0AFE14FC91D18A"author="Labbe A."box="[562,850,443,470]"pageId="87"pageNumber="88"pagination="173 - 246"refId="ref72690"refString="Labbe A. 1934. Les Silicodermes (Labbe) du Museum d'Histoire naturelle de Paris. Premiere partie: Classification, formes nouvelles ou peu connues. Annales de l'Institut oceanographique 14: 173 - 246."type="journal article"year="1934">Labbé (1934: 225–226)</bibRefCitation>
<bibRefCitationid="EF9E1EAFFFCB8B05DEF9FE70FD2ED1A5"author="Tapparone-Canefri C."box="[449,749,479,505]"pageId="87"pageNumber="88"pagination="1 - 313"refId="ref75045"refString="Tapparone-Canefri C. 1883. Fauna malacologica delle Nuova Guinea e delle Isole adiacenti. Annali del Museo Civico di Storia Naturale, Genova 19: 1 - 313. Available from https: // biodiversitylibrary. org / page / 10812600 [accessed 22 Dec. 2018]."type="journal article"year="1883">Tapparone-Canefri, 1883</bibRefCitation>
</taxonomicName>
were both junior synonyms of
<taxonomicNameid="4C0F18DDFFCB8B05DB41FE70FE23D241"authority="Quoy & Gaimard, 1825"authorityName="Quoy & Gaimard"authorityYear="1825"class="Gastropoda"family="Onchidiidae"genus="Onchidium"kingdom="Animalia"order="Systellommatophora"pageId="87"pageNumber="88"phylum="Mollusca"rank="species"species="planatum">
<taxonomicNameid="4C0F18DDFFCB8B05DED7FDADFD28D240"authorityName="Quoy & Gaimard"authorityYear="1825"box="[495,747,514,540]"class="Gastropoda"family="Onchidiidae"genus="Onchidium"kingdom="Animalia"order="Systellommatophora"pageId="87"pageNumber="88"phylum="Mollusca"rank="species"species="planatum">
, may not even refer to an onchidiid based on the brief written description. Two pieces of a notum were located recently at the MNHN which are part of the
<taxonomicNameid="4C0F18DDFFCB8B05DDDEFDE6FCBAD23E"authorityName="Quoy & Gaimard"authorityYear="1825"box="[742,889,584,611]"class="Gastropoda"family="Onchidiidae"genus="Onchidium"kingdom="Animalia"order="Systellommatophora"pageId="87"pageNumber="88"phylum="Mollusca"rank="species"species="planatum">
(MNHN-IM-2000-33706). However, these are so poorly preserved that it is not possible to determine whether they were part of an onchidiid slug.
<bibRefCitationid="EF9E1EAFFFCB8B05DF85FD20FDC0D2F5"author="Tapparone-Canefri C."box="[189,515,655,681]"pageId="87"pageNumber="88"pagination="1 - 313"refId="ref75045"refString="Tapparone-Canefri C. 1883. Fauna malacologica delle Nuova Guinea e delle Isole adiacenti. Annali del Museo Civico di Storia Naturale, Genova 19: 1 - 313. Available from https: // biodiversitylibrary. org / page / 10812600 [accessed 22 Dec. 2018]."type="journal article"year="1883">Tapparone-Canefri’s (1883)</bibRefCitation>
<taxonomicNameid="4C0F18DDFFCB8B05DB46FD1DFABBD290"authorityName="Quoy & Gaimard"authorityYear="1825"box="[1150,1400,690,716]"class="Gastropoda"family="Onchidiidae"genus="Onchidium"kingdom="Animalia"order="Systellommatophora"pageId="87"pageNumber="88"phylum="Mollusca"rank="species"species="planatum">
<bibRefCitationid="EF9E1EAFFFCB8B05DC81FD7AFB2DD2AC"author="Labbe A."box="[953,1262,725,752]"pageId="87"pageNumber="88"pagination="173 - 246"refId="ref72690"refString="Labbe A. 1934. Les Silicodermes (Labbe) du Museum d'Histoire naturelle de Paris. Premiere partie: Classification, formes nouvelles ou peu connues. Annales de l'Institut oceanographique 14: 173 - 246."type="journal article"year="1934">Labbé’s (1934: 225–226)</bibRefCitation>
specimens identified as
<taxonomicNameid="4C0F18DDFFCB8B05DE6AFD56FE27D34E"authorityName="Quoy & Gaimard"authorityYear="1825"box="[338,484,760,787]"class="Gastropoda"family="Onchidiidae"genus="Onchidium"kingdom="Animalia"order="Systellommatophora"pageId="87"pageNumber="88"phylum="Mollusca"rank="species"species="planatum">
<taxonomicNameid="4C0F18DDFFCB8B05DF84FCB3FE98D369"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Simroth"baseAuthorityYear="1920"box="[188,347,795,821]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="87"pageNumber="88"phylum="Mollusca"rank="species"species="applanatus">
