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dc:creator "Isasmendi, Erik; Cuesta, Elena; Díaz-Martínez, Ignacio; Company, Julio; Sáez-Benito, Patxi; Viera, Luis I.; Torices, Angelica; Pereda-Suberbiola, Xabier" ;
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dc:title "Riojavenatrix lacustris Isasmendi, Cuesta, Díaz-Martínez, Company, Sáez-Benito, Viera, Torices & Pereda-Suberbiola, 2024, sp.nov." ;
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bibo:journal "Zoological Journal of the Linnean Society" ;
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bibo:pubDate "2024-02-19" ;
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bibo:series "Zool. J. Linn. Soc." ;
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bibo:startPage "1" ;
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bibo:volume "202" ;
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dc:creator "Isasmendi, Erik; Cuesta, Elena; Díaz-Martínez, Ignacio; Company, Julio; Sáez-Benito, Patxi; Viera, Luis I.; Torices, Angelica; Pereda-Suberbiola, Xabier" ;
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dc:date "2024" ;
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dc:title "Increasing the theropod record of Europe: a new basal spinosaurid from the Enciso Group of the Cameros Basin (La Rioja, Spain). Evolutionary implications and palaeobiodiversity" ;
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dwc:authorityYear "2024" ;
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dwc:class "Reptilia" ;
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dwc:family "Spinosauridae" ;
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dwc:genus "Riojavenatrix" ;
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dwc:scientificNameAuthorship "Isasmendi, Cuesta, Díaz-Martínez, Company, Sáez-Benito, Viera, Torices & Pereda-Suberbiola, 2024" ;
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dc:description "Figure 3.Riojavenatrix lacustris, mid- to posterior dorsal vertebra (CPI 1677). A, anterior view. B, left lateral view. C, posterior view. Abbreviations:hy, hyposphene; ils, interspinous ligament scar; nc, neural canal; ns, neural spine; pcdl, posterior centrodiapophyseal lamina; pocdp, postzygocentrodiapophyseal fossa; sdf, spinodiapophyseal fossa; spof, spinopostzygapophyseal fossa; spol, spinopostzygapophyseal laminae; sprf; spinoprezygapophyseal fossa; sprl, spinoprezygapophyseal laminae; tp, transverse process.Scale bar: 5 cm." ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.14422925>
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dc:description "Figure 4. Riojavenatrix lacustris, left pubis (CPI 1675A–B) and right ischium (CPI 1641A–B) in anterior (A, F, K, O), medial (B, G, L, P), lateral (C, I, M, R), posterior (D, J, N, S), proximal (E, Q), and distal (H, T) views.A–E, proximal left pubis. F–J, left pubic shaft and distal boot. K–N, proximal right ischium. O–T, right ischial shaft and distal boot. Abbreviations: ap, anterior process of the ischial boot; as, acetabular surface; ilp, iliac peduncle; isb, ischial boot; isp, ischial peduncle; on, obturator notch; pa, pubic apron; pb, pubic boot; pbg, posterior bulge. Scale bars:5 cm." ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.14422927>
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dc:description "Figure 5. Riojavenatrix lacustris, left (CPI 1637) and right (CPI 1643) femora in anterior (A, E), lateral (B, F), posterior (C, G), medial (D, H), distal (I), and proximal (J) views. A–D, J, proximal right femur.E–I, shaft and distal epiphysis of the left femur. Abbreviations: ctf, crista tibiofibularis; exg, extensor groove; fh, femoral head; ft, fourth trochanter; fxg, flexor groove; gt, greater trochanter; lc, lateral condyle; mc, medial condyle; me, medial epicondyle; pag, proximal articular groove.Scale bars: 10 cm." ;
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fabio:hasRepresentation <https://zenodo.org/record/14422927/files/figure.png> ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.14422930>
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dc:description "Figure 6. Riojavenatrix lacustris, right tibia (CPI 1642), left tibia and astragalus (CPI 1638), and left fibula (CPI 1639A–B) in anterior (A, E, K, P), lateral (B, F, L, Q), posterior (C, G, M, R), medial (D, H, N, S), proximal (I, J, O), and distal (T) views.A–D, I, proximal end of the right tibia. E–H, J, left tibia and astragalus.K–O, shaft and proximal end of the left fibula.P–T, distal end of the left fibula.Abbreviations:asf, astragalar facet; ast, astragalus; cn, cnemial crest; fc, fibular crest; icn, intercondylar notch; ift, iliofibularis tubercle; it, incisura tibialis; lc, lateral condyle; lm, lateral malleolus; mf, medial fossa; mm, medial malleolus; sab, supraastragalar buttress. Scale bars: 10 cm in A–O; 5 cm in P–T." ;
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fabio:hasRepresentation <https://zenodo.org/record/14422930/files/figure.png> ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.14422936>
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dc:description "Figure 7. Riojavenatrix lacustris, A–G, left astragalus and distal tibia (CPI 1638), and H–K, left calcaneum (CPI 1676) in anterior (A, H), lateral (B, I), posterior (C, J), medial (D, K), proximal (F), and distal (G) views.Abbreviations:af, astragalar facet; ag, anterior groove; apa, ascending process of the astragalus; ff, fibular facet; fo, foramen; lc, lateral condyle; mc, medial condyle; r, ridge; tf, tibial facet; vg, ventral groove. Scale bars: 5 cm." ;
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fabio:hasRepresentation <https://zenodo.org/record/14422936/files/figure.png> ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.14422938>
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dc:description "Figure 8. Riojavenatrix lacustris, left pedal elements.Metatarsal III (CPI 1640), I-2 phalanx (CPI 1647), phalanx III-1 (CPI 1645), phalanx III-3 (CPI 1646), and phalanx V-2 (or IV-3) (CPI 1648) in distal (A, L, R), dorsal (B, G, M, S, X), lateral (C, H, N, T, Y), ventral (D, I, P, V, AA), medial (E, J, Q, W, AB), and proximal (F, K, O, U, Z) views.A–F, left metatarsal III. G–K, left pedal phalanx I-2. L–Q, left pedal phalanx III-1. R–W, left pedal phalanx III-3. X–AB, left pedal phalanx V-2 (or IV-3). Abbreviations:ef, extensor fossa; ft, flexor tuberosity; k, keel; lc, lateral condyle; lclp, lateral collateral ligament pit; lg, longitudinal groove; lpvf; lateral proximoventral fossa; mc, medial condyle; mclp, medial collateral ligament pit; mpvf; medial proximoventral fossa; pdl, proximodorsal lip.Scale bars:5 cm." ;
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a fabio:Figure .
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dc:creator "Isasmendi, Erik; Cuesta, Elena; Díaz-Martínez, Ignacio; Company, Julio; Sáez-Benito, Patxi; Viera, Luis I.; Torices, Angelica; Pereda-Suberbiola, Xabier" ;
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dc:title "Riojavenatrix lacustris Isasmendi, Cuesta, Díaz-Martínez, Company, Sáez-Benito, Viera, Torices & Pereda-Suberbiola, 2024, sp.nov." ;
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bibo:journal "Zoological Journal of the Linnean Society" ;
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bibo:pubDate "2024-02-19" ;
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bibo:series "Zool. J. Linn. Soc." ;
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bibo:startPage "1" ;
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bibo:volume "202" ;
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dc:creator "Isasmendi, Erik; Cuesta, Elena; Díaz-Martínez, Ignacio; Company, Julio; Sáez-Benito, Patxi; Viera, Luis I.; Torices, Angelica; Pereda-Suberbiola, Xabier" ;
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dc:date "2024" ;
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dc:title "Increasing the theropod record of Europe: a new basal spinosaurid from the Enciso Group of the Cameros Basin (La Rioja, Spain). Evolutionary implications and palaeobiodiversity" ;
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dc:description "Figure 3.Riojavenatrix lacustris, mid- to posterior dorsal vertebra (CPI 1677). A, anterior view. B, left lateral view. C, posterior view. Abbreviations:hy, hyposphene; ils, interspinous ligament scar; nc, neural canal; ns, neural spine; pcdl, posterior centrodiapophyseal lamina; pocdp, postzygocentrodiapophyseal fossa; sdf, spinodiapophyseal fossa; spof, spinopostzygapophyseal fossa; spol, spinopostzygapophyseal laminae; sprf; spinoprezygapophyseal fossa; sprl, spinoprezygapophyseal laminae; tp, transverse process.Scale bar: 5 cm." ;
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dc:description "Figure 4. Riojavenatrix lacustris, left pubis (CPI 1675A–B) and right ischium (CPI 1641A–B) in anterior (A, F, K, O), medial (B, G, L, P), lateral (C, I, M, R), posterior (D, J, N, S), proximal (E, Q), and distal (H, T) views.A–E, proximal left pubis. F–J, left pubic shaft and distal boot. K–N, proximal right ischium. O–T, right ischial shaft and distal boot. Abbreviations: ap, anterior process of the ischial boot; as, acetabular surface; ilp, iliac peduncle; isb, ischial boot; isp, ischial peduncle; on, obturator notch; pa, pubic apron; pb, pubic boot; pbg, posterior bulge. Scale bars:5 cm." ;
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fabio:hasRepresentation <https://zenodo.org/record/14422925/files/figure.png> ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.14422927>
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dc:description "Figure 5. Riojavenatrix lacustris, left (CPI 1637) and right (CPI 1643) femora in anterior (A, E), lateral (B, F), posterior (C, G), medial (D, H), distal (I), and proximal (J) views. A–D, J, proximal right femur.E–I, shaft and distal epiphysis of the left femur. Abbreviations: ctf, crista tibiofibularis; exg, extensor groove; fh, femoral head; ft, fourth trochanter; fxg, flexor groove; gt, greater trochanter; lc, lateral condyle; mc, medial condyle; me, medial epicondyle; pag, proximal articular groove.Scale bars: 10 cm." ;
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fabio:hasRepresentation <https://zenodo.org/record/14422927/files/figure.png> ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.14422930>
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dc:description "Figure 6. Riojavenatrix lacustris, right tibia (CPI 1642), left tibia and astragalus (CPI 1638), and left fibula (CPI 1639A–B) in anterior (A, E, K, P), lateral (B, F, L, Q), posterior (C, G, M, R), medial (D, H, N, S), proximal (I, J, O), and distal (T) views.A–D, I, proximal end of the right tibia. E–H, J, left tibia and astragalus.K–O, shaft and proximal end of the left fibula.P–T, distal end of the left fibula.Abbreviations:asf, astragalar facet; ast, astragalus; cn, cnemial crest; fc, fibular crest; icn, intercondylar notch; ift, iliofibularis tubercle; it, incisura tibialis; lc, lateral condyle; lm, lateral malleolus; mf, medial fossa; mm, medial malleolus; sab, supraastragalar buttress. Scale bars: 10 cm in A–O; 5 cm in P–T." ;
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fabio:hasRepresentation <https://zenodo.org/record/14422930/files/figure.png> ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.14422936>
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dc:description "Figure 7. Riojavenatrix lacustris, A–G, left astragalus and distal tibia (CPI 1638), and H–K, left calcaneum (CPI 1676) in anterior (A, H), lateral (B, I), posterior (C, J), medial (D, K), proximal (F), and distal (G) views.Abbreviations:af, astragalar facet; ag, anterior groove; apa, ascending process of the astragalus; ff, fibular facet; fo, foramen; lc, lateral condyle; mc, medial condyle; r, ridge; tf, tibial facet; vg, ventral groove. Scale bars: 5 cm." ;
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fabio:hasRepresentation <https://zenodo.org/record/14422936/files/figure.png> ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.14422938>
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dc:description "Figure 8. Riojavenatrix lacustris, left pedal elements.Metatarsal III (CPI 1640), I-2 phalanx (CPI 1647), phalanx III-1 (CPI 1645), phalanx III-3 (CPI 1646), and phalanx V-2 (or IV-3) (CPI 1648) in distal (A, L, R), dorsal (B, G, M, S, X), lateral (C, H, N, T, Y), ventral (D, I, P, V, AA), medial (E, J, Q, W, AB), and proximal (F, K, O, U, Z) views.A–F, left metatarsal III. G–K, left pedal phalanx I-2. L–Q, left pedal phalanx III-1. R–W, left pedal phalanx III-3. X–AB, left pedal phalanx V-2 (or IV-3). Abbreviations:ef, extensor fossa; ft, flexor tuberosity; k, keel; lc, lateral condyle; lclp, lateral collateral ligament pit; lg, longitudinal groove; lpvf; lateral proximoventral fossa; mc, medial condyle; mclp, medial collateral ligament pit; mpvf; medial proximoventral fossa; pdl, proximodorsal lip.Scale bars:5 cm." ;
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fabio:hasRepresentation <https://zenodo.org/record/14422938/files/figure.png> ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.14422915>
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dc:description "Figure 1. Geographical and geological location of northeastern Cameros Basin.A, geological map showing the location of Virgen del Villar-1 site in Igea (La Rioja, Spain) (modified from Suarez-Gonzalez et al. 2013). B, geographical map of the Iberian Peninsula." ;
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fabio:hasRepresentation <https://zenodo.org/record/14422915/files/figure.png> ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.14422942>
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dc:description "Figure 9. Phylogenetic results showing the phylogenetic position of Riojavenatrix based on the matrix of Rauhut and Pol (2021). Reduced consensus by iterPCR obtained from the 10 000 most parsimonious trees. Alternative placements of Riojavenatrix and other pruned taxa are indicated by letters and explained in the key (top right). The coloured nodes indicate the main clades (key at bottom left)." ;
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fabio:hasRepresentation <https://zenodo.org/record/14422942/files/figure.png> ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.14422919>
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dc:description "Figure 2. Skeletal reconstruction of Riojavenatrix lacustris holotype recovered at Virgen del Villar-1 site (La Rioja, Spain). The preserved material is coloured in fuchsia. Courtesy of Scott Hartman." ;
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fabio:hasRepresentation <https://zenodo.org/record/14422919/files/figure.png> ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.14422946>
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dc:description "Figure 10.Phylogenetic results showing the phylogenetic position of Riojavenatrix based on the matrix of Mateus and Estraviz-López (2022). A, strict consensus obtained from 307 most parsimonious trees. Bremer support values are above the nodes, and jackknife and bootstrap values are below the nodes. B, reduced consensus by agreement subtree methodology with 16 taxa; jackknife and bootstrap values are below the nodes.The coloured nodes indicate the main clades (key at bottom of figure)." ;
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fabio:hasRepresentation <https://zenodo.org/record/14422946/files/figure.png> ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.14422950>
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dc:description "Figure 11. Comparison of spinosaurid elements with Riojavenatrix lacustris. A–E, distal view of left pubic boot of Riojavenatrix (CPI 1675A–B) (A), right pubic boot of Ichthyovenator (MDS-Savannakhet BK10-11; based on Allain et al. 2012) (B), left pubic boot of Baryonyx (NHMUK VP R9951) (C), left pubic boot of Suchomimus (MNN GDF500) (D), and right pubic boot of FSAC-KK 11888 (E). F–J, lateral view of right ischial boot of Riojavenatrix (CPI 1641A–B) (F), right ischial boot of Vallibonavenatrix (MSMCa-1–3) (G), left ischial boot of Suchomimus (MNN GDF500) (H), left ischial boot of FSAC-KK 11888 (I), and right ischial boot of Ichthyovenator (MDS-Savannakhet BK10- 13; based on Allain et al. 2012) (J). Note that the drawings of the pubic boot of Ichthyovenator (MDS-Savannakhet BK10-11) and FSAC-KK 11888 and the ischial boot of Suchomimus (MNN GDF500) are mirror images of the element. Also note that the arrows point anteriorly.Scale bars:5 cm." ;
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fabio:hasRepresentation <https://zenodo.org/record/14422950/files/figure.png> ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.14422954>
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dc:description "Figure 12. Comparison of spinosaurid elements with Riojavenatrix lacustris. Megalosauroid femora in distal (A–F) and proximal (G–J) views.A, E, I, J, left femur of Riojavenatrix (CPI 1637) (A), Suchomimus (MNN GDF500) (E, J), and Baryonyx (NHMUK VP R9951) (I). B–D, F–H, right femur of CMP-2b/21 (B), Megalosaurus (NHMUK PV 31806) (C), Baryonyx (NHMUK VP R9951) (D), FSAC-KK 11888 (F), Riojavenatrix (CPI 1643) (G), and CMP-MS-0/22 (H). Abbreviations; cft, crita tibiofibularis; exg, extensor groove; fh, femoral head; fxg, flexor groove; gt, greater trochanter; lc, lateral condyle; mc, medial condyle; pag, proximal articular groove. Note that the drawings of Baryonyx (NHMUK VP R9951), Suchomimus (MNN GDF500) in proximal view, and Baryonyx (NHMUK VP R9951), Megalosaurus (NHMUK PV 31806), CMP-2b/211, and FSAC-KK 11888 are mirror images of the elements. Scale bars: 5 cm." ;
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fabio:hasRepresentation <https://zenodo.org/record/14422954/files/figure.png> ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.14422956>
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dc:description "Figure 13. Comparison of spinosaurid elements with Riojavenatrix lacustris. Left tibia of Riojavenatrix (CPI 1638) (A) and Suchomimus (MNN GDF500) (D), and right tibia of Riojavenatrix (CPI 1642) (B), Camarillasaurus (MPG-KPC8) (C), and FSAC-KK 11888 (E) in proximal view. Abbreviations: cn, cnemial crest; icn, intercondylar notch; it, incisura tibialis; lc, lateral condyle; mc, medial condyle. Note that the drawings of Camarillasaurus (MPG-KPC8), Riojavenatrix (CPI 1638), and FSAC-KK 11888 are mirror images of the elements. Scale bars: 10 cm." ;
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fabio:hasRepresentation <https://zenodo.org/record/14422956/files/figure.png> ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.14422958>
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dc:description "Figure 14. Comparison of spinosaurid elements with Riojavenatrix lacustris. Left calcaneum of Riojavenatrix (CPI 1676) (A) and Iberospinus (ML1190-31; based on Mateus and Estraviz-López 2022) (B), and right calcaneum of Baryonyx (NHMUK VP R9951) (C) in lateral view.Abbreviations:ff,fibular facet; fo, foramen. Note that the drawing of Baryonyx (NHMUK VP R9951) is a mirror image of the element. Scale bars: 5 cm." ;
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fabio:hasRepresentation <https://zenodo.org/record/14422958/files/figure.png> ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.14422962>
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dc:description "Figure 15. Comparison of spinosaurid elements with Riojavenatrix lacustris. Left pedal phalanx I-2 of Riojavenatrix (CPI 1647) (A) and Deinonychus (YPM 5205) (F), and right pedal phalanx I-2 of Carcharodontosauria indet.(SHN.019) (B), Tyrannosaurus (FMNH PR2081; based on Brochu 2003) (C), Bambiraptor (AMNH 30556) (D), and Alectrosaurus (AMNH 6554) (E) in medial view.Abbreviation:lg, longitudinal groove. Note that the drawings of Alectrosaurus (AMNH 6554), Bambiraptor (AMNH 30556), Tyrannosaurus (FMNH PR2081), and the indeterminate carcharodontosaurian (SHN.019) are mirror images of the elements.Scale bars: 2 cm, except for D, which is 1 cm." ;
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fabio:hasRepresentation <https://zenodo.org/record/14422962/files/figure.png> ;
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a fabio:Figure .
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<http://dx.doi.org/10.5281/zenodo.14422970>
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dc:description "Figure 16. Histological preparations of Riojavenatrix lacustris material. A, polished cross-section of a possible fragment of the proximal right femur (CPI 1643) in ventral view, showing a thin cortex enclosing a medullary cavity invaded by bony trabeculae of cancellous bone. B, transverse polished section of a fragmentary vertebral centrum showing a pneumatized internal structure.Note the presence of a couple of lines of arrested growth visible even to the naked eye (blue arrowheads). C, transverse thin section of the cortical bone of the same specimen, showing a clear pattern of stratification produced by the cyclical presence of lines of arrested growth (blue arrowheads). D, transverse thin section of the outermost cortex of a fragmentary diaphyseal limb bone fragment, showing two well-preserved lines of arrested grow (blue arrowheads), several isolated secondary osteons, and oval erosion rooms (note that these resorption cavities are surrounded by only a single coat of lamellar bone tissue, remaining as open, incipient secondary osteons).The bone surface is at the top. E, same thin section showing intense remodelling in the innermost cortex.Only two generations of secondary osteons are observed. Although densely packed, secondary osteons overlap each other very little.Abbreviations:er, erosion room; pc, pneumatic cavity; Q, quartz cement; so, secondary osteons. Photographs B–D were taken with plane-polarized light. Scale bars: 1 cm in A; 0.1 mm in B, C; 1 mm in D." ;
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fabio:hasRepresentation <https://zenodo.org/record/14422970/files/figure.png> ;
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a fabio:Figure .
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<http://tb.plazi.org/GgServer/dwcaRecords/03AB87A8FFD17E7A5422C2B2FD345BBF.mc.3B6A3CE3FFD17E6456BAC4CAFA1159A4>
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dwc:locality "La Rioja" ;
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dwc:typeStatus "holotype" ;
|
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trt:httpUri <https://treatment.plazi.org/id/03AB87A8FFD17E7A5422C2B2FD345BBF#3B6A3CE3FFD17E6456BAC4CAFA1159A4> ;
|
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a dwc:MaterialCitation .
|
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