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<document id="9CBA10BD1CC3B097CD082EF2C2CCF262" ID="10.11646/zootaxa.4263.1.7SLASH10972" ID-CLB-Dataset="33095" ID-DOI="10.11646/zootaxa.4263.1.7" ID-GBIF-Dataset="707b0d75-86cc-4ddf-a776-608137fc4745" ID-ISSN="1175-5326" ID-Zenodo-Dep="572597" ID-ZooBank="7CE2F6FE-792B-4997-9714-BD02BF230406" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1494240554651" checkinUser="plazi" docAuthor="Mendis Wickramasinghe, L. J., Vidanapathirana, Dulan Ranga, Gehan Rajeev, M. D. &amp; Gower, David J." docDate="2017" docId="03D7345DEA782A45FF50C0A4C0AFF874" docLanguage="en" docName="zootaxa.4263.1.7SLASH10972.pdf" docOrigin="Zootaxa 4263 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhinophis roshanpererai Wickramasinghe, Vidanapathirana, Rajeev &amp; Gower, 2017, sp. nov." docType="treatment" docVersion="8" lastPageNumber="161" masterDocId="FFEE4C25EA7A2A4DFFC7C16AC046FFE1" masterDocTitle="A new species of Rhinophis Hemprich, 1820 (Serpentes: Uropeltidae) from the central hills of Sri Lanka" masterLastPageNumber="164" masterPageNumber="153" pageNumber="155" updateTime="1698439333568" updateUser="ExternalLinkService">
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<mods:title id="6CEB116CE21D39DD3241811674D93692">A new species of Rhinophis Hemprich, 1820 (Serpentes: Uropeltidae) from the central hills of Sri Lanka</mods:title>
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<mods:namePart id="F185F26DC8DAB2695D5ADBBCBEBB4793">Mendis Wickramasinghe, L. J.</mods:namePart>
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<mods:namePart id="4BEA78CAF43909C9CFA6CCD83F5A012B">Vidanapathirana, Dulan Ranga</mods:namePart>
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<treatment id="03D7345DEA782A45FF50C0A4C0AFF874" ID-DOI="http://doi.org/10.5281/zenodo.6028835" ID-GBIF-Taxon="130497598" ID-Zenodo-Dep="6028835" LSID="urn:lsid:plazi:treatment:03D7345DEA782A45FF50C0A4C0AFF874" httpUri="http://treatment.plazi.org/id/03D7345DEA782A45FF50C0A4C0AFF874" lastPageId="8" lastPageNumber="161" pageId="2" pageNumber="155">
<subSubSection id="C364D6C0EA782A4FFF50C0A4C127FDE8" pageId="2" pageNumber="155" type="nomenclature">
<paragraph id="8BC1854BEA782A4FFF50C0A4C26AFE09" blockId="2.[151,556,462,522]" box="[151,556,462,488]" pageId="2" pageNumber="155">
<heading id="D0893227EA782A4FFF50C0A4C26AFE09" bold="true" box="[151,556,462,488]" fontSize="11" level="1" pageId="2" pageNumber="155" reason="1">
<emphasis id="B90A5959EA782A4FFF50C0A4C26AFE09" bold="true" box="[151,556,462,488]" pageId="2" pageNumber="155">
<taxonomicName id="4C7EFEC8EA782A4FFF50C0A4C181FE09" box="[151,455,462,488]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="155" phylum="Chordata" rank="species" species="roshanpererai" status="sp. nov.">
<emphasis id="B90A5959EA782A4FFF50C0A4C181FE09" bold="true" box="[151,455,462,488]" italics="true" pageId="2" pageNumber="155">Rhinophis roshanpererai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A239E422EA782A4FFE09C0A4C26AFE09" box="[462,556,462,488]" pageId="2" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="8BC1854BEA782A4FFF50C09BC127FDE8" blockId="2.[151,556,462,522]" box="[151,353,497,522]" pageId="2" pageNumber="155">
<figureCitation id="134599CEEA782A4FFF50C09BC147FDEB" box="[151,257,497,522]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="1.[151,250,1675,1697]" captionStartId-1="2.[151,248,1319,1341]" captionStartId-2="4.[151,250,1955,1977]" captionStartId-3="5.[151,250,1983,2005]" captionStartId-4="6.[151,250,1953,1975]" captionTargetBox-0="[164,1425,318,1643]" captionTargetBox-1="[151,1436,703,1307]" captionTargetBox-2="[281,1307,193,1932]" captionTargetBox-3="[276,1302,193,1932]" captionTargetBox-4="[214,1373,193,1932]" captionTargetId-0="figure@1.[151,1435,303,1651]" captionTargetId-1="figure@2.[151,1436,703,1307]" captionTargetId-2="figure@4.[281,1307,193,1932]" captionTargetId-3="figure@5.[276,1302,193,1932]" captionTargetId-4="figure@6.[214,1373,193,1932]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="FIGURE 1. Map of Sri Lanka showing the type locality (Galkanda) of Rhinophis roshanpererai sp. nov." captionText-1="FIGURE 2. Dorsolateral view in life of the holotype of Rhinophis roshanpererai sp. nov., NMSL 2016.08. 0 1 NH, 214 mm (total length)." captionText-2="FIGURE 3. Holotype of Rhinophis roshanpererai sp. nov., NMSL 2016.08. 0 1 NH, scale bars in centimeters." captionText-3="FIGURE 4. Head and tail ends of the two paratypes of Rhinophis roshanpererai sp. nov. (A) DWC 2016.05. 0 3, (B) DWC 2016.05. 0 4. Scale bar 2 cm." captionText-4="FIGURE 5. Rhinophis roshanpererai sp. nov., paratype DWC 2016.05. 0 4, posterodorsal view of tail in preserved specimen showing three (two lower, one upper) rather than four major spines on the tail shield (bases of these major spines are surrounded by pale areas on shield)." httpUri-0="https://zenodo.org/record/572598/files/figure.png" httpUri-1="https://zenodo.org/record/572599/files/figure.png" httpUri-2="https://zenodo.org/record/572600/files/figure.png" httpUri-3="https://zenodo.org/record/572601/files/figure.png" httpUri-4="https://zenodo.org/record/572602/files/figure.png" pageId="2" pageNumber="155">Figs. 15</figureCitation>
;
<tableCitation id="C6FCB0F0EA782A4FFEC9C09BC127FDE8" box="[270,353,497,522]" captionStart="TABLE 1" captionStartId="3.[151,239,152,175]" captionText="TABLE 1. Meristic and morphometric data for 52 characters of holotype (*) and paratypes (**) of Rhinophis roshanpererai sp. nov. Specimen numbers have NMSL prefix for holotype and DWC prefix for paratypes. Bilateral scale counts given in left, right order. Character numbers in parentheses, as follows: (1) supralabials, (2) infralabials, (3) diameter of eye, (4) maximum length of ocular, (5) length ocular / eye, (6) distance between eyes, (7) distance between eye-naris, (8) length of frontal, (9) width of frontal, (10) head length = distance between snout tip-posterior edge 4 th supralabial, (11) total length / HL, (12) head width level with corner of mouth, (13) distance between nares, (14) maximum longitudinal length of prefrontal scales, (15) length of parietal scale = distance between posterior tip of parietal and posterior end of suture between frontal-ocular, (16) width of parietal scale = distance between posterior tip of ocular and posterior tip of frontal, (17) length of midline suture between parietals, (18) distance rostral-frontal, (19) distance between tip of snout and posterior edge of rostral, (20) distance between tip of snout and posterior limit of midline suture between parietals, (21) maximum width of rostral, viewed ventrally, (22) distance between snout tip-naris, (23) distance between snout tip-eye, (24) midbody width, (25) midbody depth, (26) midbody circumference, (27) exposed width of first scale row at midbody, (28) exposed width of ventral scales at midbody, (29) number of ventral scale level with which dorsal scale row is lost (i. e. location of change between 19 and 17 dorsal scale rows), (30) dorsal scale rows behind head, at midbody and one head length anterior to vent (excluding ventrals), (31) exposed length of first scale row at midbody, (32) width of ventral / first dorsal scale rows, (33) ventral scales, (34) vertebral scales, (35) snout vent length, (36) width at vent, (37) disposition of two anal scales (l / r) or (r / l), (38) number of small scales overlapped by anals, excluding first subcaudals, (39) subcaudal scales, (40) tail length, (41) total length, (42) tail length as % of total length, (43) total length / midbody width, (44) maximum width of tail shield, (45) maximum diameter of tail shield, (46) number of scales (including subcaudals) surrounding base of tail shield, (47) dorsal base to tip of tail shield (upper spines), (48) dorsal base to tip of tail shield (lower spines), (49) lateral base to tip of tail shield (upper), (50) lateral base to tip of tail shield (lower), (51) ventral base to tip of tail shield (upper), (52) ventral base to tip of tail shield (lower)." pageId="2" pageNumber="155">Table 1</tableCitation>
</paragraph>
</subSubSection>
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<materialsCitation id="3B168F16EA782A4FFF50C358C4C6FD72" ID-GBIF-Occurrence="1500371340" collectingDate="2010-05-10" collectionCode="NMSL" collectorName="L. J. Mendis Wickramasinghe &amp; Dulan Ranga Vidanapathirana &amp; M. D. Gehan Rajeev" country="Sri Lanka" county="Badulla District" elevation="940" latitude="6.752217" location="Galkanda" longLatPrecision="1" longitude="80.95562" municipality="Beragala" pageId="2" pageNumber="155" specimenCode="NMSL 2016.08" specimenCount="1" stateProvince="Uva Province" typeStatus="holotype">
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<typeStatus id="54C53BE9EA782A4FFF50C358C140FDAA" box="[151,262,562,587]" pageId="2" pageNumber="155" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DBD82D30EA782A4FFED4C359C18AFDAA" box="[275,460,562,587]" collectionCode="NMSL" country="Sri Lanka" httpUri="http://grbio.org/cool/9670-f832" name="National Museum of Sri Lanka" pageId="2" pageNumber="155">
<collectionCode id="ED6F1D8EEA782A4FFED4C359C127FDAA" box="[275,353,563,587]" country="Sri Lanka" httpUri="http://grbio.org/cool/9670-f832" name="National Museum of Sri Lanka" pageId="2" pageNumber="155">NMSL</collectionCode>
2016.08
</specimenCode>
.0 1 NH (
<figureCitation id="134599CEEA782A4FFDEEC358C2D1FDAA" box="[553,663,562,587]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="2.[151,248,1319,1341]" captionStartId-1="4.[151,250,1955,1977]" captionTargetBox-0="[151,1436,703,1307]" captionTargetBox-1="[281,1307,193,1932]" captionTargetId-0="figure@2.[151,1436,703,1307]" captionTargetId-1="figure@4.[281,1307,193,1932]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURE 2. Dorsolateral view in life of the holotype of Rhinophis roshanpererai sp. nov., NMSL 2016.08. 0 1 NH, 214 mm (total length)." captionText-1="FIGURE 3. Holotype of Rhinophis roshanpererai sp. nov., NMSL 2016.08. 0 1 NH, scale bars in centimeters." httpUri-0="https://zenodo.org/record/572599/files/figure.png" httpUri-1="https://zenodo.org/record/572600/files/figure.png" pageId="2" pageNumber="155">Figs. 23</figureCitation>
;
<tableCitation id="C6FCB0F0EA782A4FFD63C358C2B1FDAA" box="[676,759,562,587]" captionStart="TABLE 1" captionStartId="3.[151,239,152,175]" captionText="TABLE 1. Meristic and morphometric data for 52 characters of holotype (*) and paratypes (**) of Rhinophis roshanpererai sp. nov. Specimen numbers have NMSL prefix for holotype and DWC prefix for paratypes. Bilateral scale counts given in left, right order. Character numbers in parentheses, as follows: (1) supralabials, (2) infralabials, (3) diameter of eye, (4) maximum length of ocular, (5) length ocular / eye, (6) distance between eyes, (7) distance between eye-naris, (8) length of frontal, (9) width of frontal, (10) head length = distance between snout tip-posterior edge 4 th supralabial, (11) total length / HL, (12) head width level with corner of mouth, (13) distance between nares, (14) maximum longitudinal length of prefrontal scales, (15) length of parietal scale = distance between posterior tip of parietal and posterior end of suture between frontal-ocular, (16) width of parietal scale = distance between posterior tip of ocular and posterior tip of frontal, (17) length of midline suture between parietals, (18) distance rostral-frontal, (19) distance between tip of snout and posterior edge of rostral, (20) distance between tip of snout and posterior limit of midline suture between parietals, (21) maximum width of rostral, viewed ventrally, (22) distance between snout tip-naris, (23) distance between snout tip-eye, (24) midbody width, (25) midbody depth, (26) midbody circumference, (27) exposed width of first scale row at midbody, (28) exposed width of ventral scales at midbody, (29) number of ventral scale level with which dorsal scale row is lost (i. e. location of change between 19 and 17 dorsal scale rows), (30) dorsal scale rows behind head, at midbody and one head length anterior to vent (excluding ventrals), (31) exposed length of first scale row at midbody, (32) width of ventral / first dorsal scale rows, (33) ventral scales, (34) vertebral scales, (35) snout vent length, (36) width at vent, (37) disposition of two anal scales (l / r) or (r / l), (38) number of small scales overlapped by anals, excluding first subcaudals, (39) subcaudal scales, (40) tail length, (41) total length, (42) tail length as % of total length, (43) total length / midbody width, (44) maximum width of tail shield, (45) maximum diameter of tail shield, (46) number of scales (including subcaudals) surrounding base of tail shield, (47) dorsal base to tip of tail shield (upper spines), (48) dorsal base to tip of tail shield (lower spines), (49) lateral base to tip of tail shield (upper), (50) lateral base to tip of tail shield (lower), (51) ventral base to tip of tail shield (upper), (52) ventral base to tip of tail shield (lower)." pageId="2" pageNumber="155">Table 1</tableCitation>
), adult male,
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SVL,
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,
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,
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,
<collectingRegion id="49BA4BA9EA782A4FFF3DC33DC1D0FD8E" box="[250,406,598,623]" country="Sri Lanka" name="Uva" pageId="2" pageNumber="155">Uva Province</collectingRegion>
,
<collectingCountry id="F369C5DBEA782A4FFE64C33CC257FD8E" box="[419,529,598,623]" name="Sri Lanka" pageId="2" pageNumber="155">Sri Lanka</collectingCountry>
(
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,
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,
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elevation
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</elevation>
). Collected by
<collectorName id="268BE09DEA782A4FFAD1C33DC111FD72" pageId="2" pageNumber="155">L.J. Mendis Wickramasinghe</collectorName>
,
<collectorName id="268BE09DEA782A4FFEA6C310C2F3FD72" box="[353,693,634,659]" pageId="2" pageNumber="155">Dulan Ranga Vidanapathirana</collectorName>
, and
<collectorName id="268BE09DEA782A4FFD37C311C39FFD72" box="[752,985,634,659]" pageId="2" pageNumber="155">M. D. Gehan Rajeev</collectorName>
,
<date id="FFC0A38BEA782A4FFC23C311C43AFD72" box="[996,1148,635,659]" pageId="2" pageNumber="155" value="2010-05-10">
<collectingDate id="EF845A63EA782A4FFC23C311C43AFD72" box="[996,1148,635,659]" pageId="2" pageNumber="155" value="2010-05-10">10 May 2010</collectingDate>
</date>
.
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</paragraph>
<caption id="DF01D5C3EA782A4FFF50C44DC0A2FABA" httpUri="https://zenodo.org/record/572599/files/figure.png" pageId="2" pageNumber="155" targetBox="[151,1436,703,1307]" targetPageId="2">
<paragraph id="8BC1854BEA782A4FFF50C44DC0A2FABA" blockId="2.[151,1435,1319,1371]" pageId="2" pageNumber="155">
<emphasis id="B90A5959EA782A4FFF50C44DC157FADD" bold="true" box="[151,273,1319,1341]" pageId="2" pageNumber="155">FIGURE 2.</emphasis>
Dorsolateral view in life of the holotype of
<taxonomicName id="4C7EFEC8EA782A4FFD02C44DC3FAFADC" box="[709,956,1319,1341]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="155" phylum="Chordata" rank="species" species="roshanpererai" status="sp. nov.">
<emphasis id="B90A5959EA782A4FFD02C44DC3FAFADC" box="[709,956,1319,1341]" italics="true" pageId="2" pageNumber="155">Rhinophis roshanpererai</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="A239E422EA782A4FFC05C44DC457FADD" box="[962,1041,1319,1340]" pageId="2" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, NMSL 2016.08.0 1 NH, 214 mm (total length).
</paragraph>
</caption>
<paragraph id="8BC1854BEA782A4FFF00C4E2C2B1FA25" blockId="2.[151,1436,1416,2004]" pageId="2" pageNumber="155">
<emphasis id="B90A5959EA782A4FFF00C4E2C100FA40" bold="true" box="[199,326,1416,1441]" pageId="2" pageNumber="155">Paratypes.</emphasis>
DWC 2016.05.0 3, adult female,
<quantity id="4C8628AEEA782A4FFD79C4E2C376FA41" box="[702,816,1416,1441]" metricMagnitude="-1" metricUnit="m" metricValue="2.052" pageId="2" pageNumber="155" unit="mm" value="205.2">205.2 mm</quantity>
SVL (
<figureCitation id="134599CEEA782A4FFCBAC4E2C387FA40" box="[893,961,1416,1441]" captionStart="FIGURE 4" captionStartId="5.[151,250,1983,2005]" captionTargetBox="[276,1302,193,1932]" captionTargetId="figure@5.[276,1302,193,1932]" captionTargetPageId="5" captionText="FIGURE 4. Head and tail ends of the two paratypes of Rhinophis roshanpererai sp. nov. (A) DWC 2016.05. 0 3, (B) DWC 2016.05. 0 4. Scale bar 2 cm." httpUri="https://zenodo.org/record/572601/files/figure.png" pageId="2" pageNumber="155">Fig. 4</figureCitation>
A);
<materialsCitation id="3B168F16EA782A4FFC2AC4E3C2B1FA25" ID-GBIF-Occurrence="1500371341" collectingDate="2010-05-10" collectionCode="DWC" collectorName="L. J. Mendis Wickramasinghe &amp; Dulan Ranga Vidanapathirana &amp; M. D. Gehan Rajeev" country="Sri Lanka" county="Badulla District" elevation="940" latitude="6.752217" location="Galkanda" longLatPrecision="1" longitude="80.95562" municipality="Beragala" pageId="2" pageNumber="155" specimenCode="DWC 2016.05" specimenCount="1" stateProvince="Uva Province">
<specimenCode id="DBD82D30EA782A4FFC2AC4E3C4D1FA40" box="[1005,1175,1416,1441]" collectionCode="DWC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14734" name="West Chester University" pageId="2" pageNumber="155">
<collectionCode id="ED6F1D8EEA782A4FFC2AC4E3C468FA40" box="[1005,1070,1417,1441]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14734" name="West Chester University" pageId="2" pageNumber="155">DWC</collectionCode>
2016.05
</specimenCode>
.0 4, adult female,
<quantity id="4C8628AEEA782A4FFA9AC4E3C085FA22" metricMagnitude="-1" metricUnit="m" metricValue="2.182" pageId="2" pageNumber="155" unit="mm" value="218.2">218.2 mm</quantity>
SVL (
<figureCitation id="134599CEEA782A4FFEC9C4C1C117FA25" box="[270,337,1451,1476]" captionStart="FIGURE 4" captionStartId="5.[151,250,1983,2005]" captionTargetBox="[276,1302,193,1932]" captionTargetId="figure@5.[276,1302,193,1932]" captionTargetPageId="5" captionText="FIGURE 4. Head and tail ends of the two paratypes of Rhinophis roshanpererai sp. nov. (A) DWC 2016.05. 0 3, (B) DWC 2016.05. 0 4. Scale bar 2 cm." httpUri="https://zenodo.org/record/572601/files/figure.png" pageId="2" pageNumber="155">Fig. 4</figureCitation>
B5). Collection data as for holotype.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C364D6C0EA782A4FFF00C4A4C57BF8C5" pageId="2" pageNumber="155" type="diagnosis">
<paragraph id="8BC1854BEA782A4FFF00C4A4C2A2F9CF" blockId="2.[151,1436,1416,2004]" pageId="2" pageNumber="155">
<materialsCitation id="3B168F16EA782A4FFF00C4A4C2A2F9CF" ID-GBIF-Occurrence="1500371342" collectorName="Diagnosis. A" country="Sri Lanka" location="Central" pageId="2" pageNumber="155" specimenCount="1" stateProvince="Central">
<collectorName id="268BE09DEA782A4FFF00C4A4C11AFA06" box="[199,348,1486,1511]" pageId="2" pageNumber="155">
<emphasis id="B90A5959EA782A4FFF00C4A4C107FA06" bold="true" box="[199,321,1486,1511]" pageId="2" pageNumber="155">Diagnosis.</emphasis>
A
</collectorName>
<taxonomicName id="4C7EFEC8EA782A4FFEA4C4A4C193FA06" box="[355,469,1486,1511]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="155" phylum="Chordata" rank="genus">
<emphasis id="B90A5959EA782A4FFEA4C4A4C193FA06" box="[355,469,1486,1511]" italics="true" pageId="2" pageNumber="155">Rhinophis</emphasis>
</taxonomicName>
restricted to the
<collectingRegion id="49BA4BA9EA782A4FFD57C4A4C2A5FA06" box="[656,739,1486,1511]" country="Sri Lanka" name="Central" pageId="2" pageNumber="155">Central</collectingRegion>
Highlands of
<collectingCountry id="F369C5DBEA782A4FFC46C4A4C3A9FA06" box="[897,1007,1486,1511]" name="Sri Lanka" pageId="2" pageNumber="155">Sri Lanka</collectingCountry>
with 17 dorsal scale rows at midbody, more than 160 and fewer than 175 ventral scales, a small tail shield with spines, three or four of which prominent, and lacking yellowish markings laterally or dorsally.
</materialsCitation>
</paragraph>
<paragraph id="8BC1854BEA782A4FFF00C752C57BF8C5" blockId="2.[151,1436,1416,2004]" pageId="2" pageNumber="155">
<emphasis id="B90A5959EA782A4FFF00C752C136F9B0" bold="true" box="[199,368,1592,1617]" pageId="2" pageNumber="155">Identification.</emphasis>
The new uropeltid species is referred to
<taxonomicName id="4C7EFEC8EA782A4FFC97C752C384F9B0" box="[848,962,1592,1617]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="155" phylum="Chordata" rank="genus">
<emphasis id="B90A5959EA782A4FFC97C752C384F9B0" box="[848,962,1592,1617]" italics="true" pageId="2" pageNumber="155">Rhinophis</emphasis>
</taxonomicName>
because it has an eye that lies within an ocular scale (not so in
<taxonomicName id="4C7EFEC8EA782A4FFE63C731C342F995" authority="Gunther, 1868" authorityName="Gunther" authorityYear="1868" box="[420,772,1627,1652]" class="Reptilia" family="Uropeltidae" genus="Platyplectrurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="155" phylum="Chordata" rank="genus">
<emphasis id="B90A5959EA782A4FFE63C731C214F995" box="[420,594,1627,1652]" italics="true" pageId="2" pageNumber="155">Platyplectrurus</emphasis>
<bibRefCitation id="EFEFF8BAEA782A4FFD9AC731C342F995" author="Gunther" box="[605,772,1627,1652]" pageId="2" pageNumber="155" refString="Gunther, A. C. L. G. (1868) Sixth account of new species of snakes in the collection of the British Museum. Annals and Magazine of Natural History, 1, 413 - 429." type="journal article" year="1868">Günther, 1868</bibRefCitation>
</taxonomicName>
), has a clearly discrete tail shield, lacks a mental groove (present in
<taxonomicName id="4C7EFEC8EA782A4FFEDAC714C2C5F976" authority="Gunther, 1864" authorityName="Gunther" authorityYear="1864" box="[285,643,1662,1687]" class="Reptilia" family="Uropeltidae" genus="Melanophidium" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="155" phylum="Chordata" rank="genus">
<emphasis id="B90A5959EA782A4FFEDAC714C188F976" box="[285,462,1662,1687]" italics="true" pageId="2" pageNumber="155">Melanophidium</emphasis>
Günther, 1864
</taxonomicName>
), lacks supra- or postoculars or temporals (at least one of which is present in
<taxonomicName id="4C7EFEC8EA782A4FFED0C7C8C218F95B" authority="Wall, 1921" authorityName="Wall" authorityYear="1921" box="[279,606,1698,1723]" class="Reptilia" family="Uropeltidae" genus="Brachyophidium" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="155" phylum="Chordata" rank="genus">
<emphasis id="B90A5959EA782A4FFED0C7C8C194F95A" box="[279,466,1698,1723]" italics="true" pageId="2" pageNumber="155">Brachyophidium</emphasis>
<bibRefCitation id="EFEFF8BAEA782A4FFE19C7C8C218F95B" author="Wall" box="[478,606,1698,1723]" pageId="2" pageNumber="155" refString="Wall, F. (1921) Ophidia Taprobanica or the snakes of Ceylon. Governmental Press, Colombo, xxii + 581 pp." type="book" year="1921">Wall, 1921</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C7EFEC8EA782A4FFDB5C7C8C426F95A" authority="Plectrurus Dumeril, 1851" authorityName="Plectrurus Dumeril" authorityYear="1851" box="[626,1120,1698,1723]" class="Reptilia" family="Uropeltidae" genus="Platyplectrurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="155" phylum="Chordata" rank="genus">
<emphasis id="B90A5959EA782A4FFDB5C7C8C3ECF95A" box="[626,938,1698,1723]" italics="true" pageId="2" pageNumber="155">Platyplectrurus, Plectrurus</emphasis>
<bibRefCitation id="EFEFF8BAEA782A4FFC70C7C8C426F95A" author="Dumeril" box="[951,1120,1698,1723]" pageId="2" pageNumber="155" refString="Dumeril, A. M. C. &amp; Dumeril, A. H. A. (1851) Catalogue methodique de la collection des reptiles du Museum d'Histoire naturelle de Paris. Gide et Baudry, Paris, 224 pp." type="book" year="1851">Duméril, 1851</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C7EFEC8EA782A4FFB6EC7C9C095F93F" authority="Beddome, 1886" authorityName="Beddome" authorityYear="1886" class="Reptilia" family="Uropeltidae" genus="Teretrurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="155" phylum="Chordata" rank="genus">
<emphasis id="B90A5959EA782A4FFB6EC7C9C55BF95B" box="[1193,1309,1699,1722]" italics="true" pageId="2" pageNumber="155">Teretrurus</emphasis>
<bibRefCitation id="EFEFF8BAEA782A4FFAEDC7C8C095F93F" author="Beddome" pageId="2" pageNumber="155" refString="Beddome, R. H. (1886) An account of the earth-snakes of the peninsula of India and Ceylon. Annals and Magazine of Natural History, 17, 3 - 33." type="journal article" year="1886">Beddome, 1886</bibRefCitation>
</taxonomicName>
), lacks midline contact between the nasals (present in
<taxonomicName id="4C7EFEC8EA782A4FFC91C7AFC2EEF8E0" authority="Melanophidium, Platyplectrurus, Plectrurus, Pseudoplectrurus Boulenger, 1890" authorityName="Melanophidium, Platyplectrurus, Plectrurus, Pseudoplectrurus Boulenger" authorityYear="1890" class="Reptilia" family="Uropeltidae" genus="Brachyophidium" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="155" phylum="Chordata" rank="genus">
<emphasis id="B90A5959EA782A4FFC91C7AFC1A7F8E0" italics="true" pageId="2" pageNumber="155">Brachyophidium, Melanophidium, Platyplectrurus, Plectrurus, Pseudoplectrurus</emphasis>
<bibRefCitation id="EFEFF8BAEA782A4FFE2DC782C2EEF8E0" author="Boulenger" box="[490,680,1768,1793]" pageId="2" pageNumber="155" refString="Boulenger, G. A. (1890) The Fauna of British India including Ceylon and Burma. Reptilia and Batrachia. Taylor and Francis, London, xviii + 541 pp." type="book" year="1890">Boulenger, 1890</bibRefCitation>
</taxonomicName>
,
<emphasis id="B90A5959EA782A4FFD73C783C369F8E1" box="[692,815,1769,1792]" italics="true" pageId="2" pageNumber="155">Teretrurus,</emphasis>
and
<taxonomicName id="4C7EFEC8EA782A4FFCAFC782C397F8E0" box="[872,977,1768,1793]" class="Reptilia" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="155" phylum="Chordata" rank="genus">
<emphasis id="B90A5959EA782A4FFCAFC782C397F8E0" box="[872,977,1768,1793]" italics="true" pageId="2" pageNumber="155">Uropeltis</emphasis>
</taxonomicName>
), and it has midbody dorsal scales in 17 rows (
<quantity id="4C8628AEEA782A4FFF1AC661C151F8C5" box="[221,279,1803,1828]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="2" pageNumber="155" unit="in" value="15.0">15 in</quantity>
<taxonomicName id="4C7EFEC8EA782A4FFEDAC661C56BF8C5" authority="Melanophidium, Platyplectrurus, Plectrurus, Pseudoplectrurus, Teretrurus" authorityName="Melanophidium, Platyplectrurus, Plectrurus, Pseudoplectrurus, Teretrurus" box="[285,1325,1803,1828]" class="Reptilia" family="Uropeltidae" genus="Brachyophidium" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="155" phylum="Chordata" rank="genus">
<emphasis id="B90A5959EA782A4FFEDAC661C19EF8C5" box="[285,472,1803,1828]" italics="true" pageId="2" pageNumber="155">Brachyophidium</emphasis>
,
<emphasis id="B90A5959EA782A4FFE21C661C2D1F8C5" box="[486,663,1803,1828]" italics="true" pageId="2" pageNumber="155">Melanophidium</emphasis>
,
<emphasis id="B90A5959EA782A4FFD62C661C315F8C5" box="[677,851,1803,1828]" italics="true" pageId="2" pageNumber="155">Platyplectrurus</emphasis>
,
<emphasis id="B90A5959EA782A4FFCA6C661C56BF8C5" box="[865,1325,1803,1828]" italics="true" pageId="2" pageNumber="155">Plectrurus, Pseudoplectrurus, Teretrurus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C364D6C0EA782A45FF00C644C409FBEE" lastPageId="8" lastPageNumber="161" pageId="2" pageNumber="155" type="description">
<paragraph id="8BC1854BEA782A4FFF00C644C381F835" blockId="2.[151,1436,1416,2004]" pageId="2" pageNumber="155">
<taxonomicName id="4C7EFEC8EA782A4FFF00C644C198F8A6" box="[199,478,1838,1863]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="155" phylum="Chordata" rank="species" species="roshanpererai" status="sp. nov.">
<emphasis id="B90A5959EA782A4FFF00C644C198F8A6" box="[199,478,1838,1863]" italics="true" pageId="2" pageNumber="155">Rhinophis roshanpererai</emphasis>
</taxonomicName>
<emphasis id="B90A5959EA782A4FFE21C644C206F8A6" bold="true" box="[486,576,1838,1863]" pageId="2" pageNumber="155">
<taxonomicNameLabel id="A239E422EA782A4FFE21C644C206F8A6" box="[486,576,1838,1863]" pageId="2" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from all four Indian species of
<taxonomicName id="4C7EFEC8EA782A4FFC31C644C42EF8A6" box="[1014,1128,1838,1863]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="155" phylum="Chordata" rank="genus">
<emphasis id="B90A5959EA782A4FFC31C644C42EF8A6" box="[1014,1128,1838,1863]" italics="true" pageId="2" pageNumber="155">Rhinophis</emphasis>
</taxonomicName>
by having a very small tail shield with spines (versus relatively much larger tail shield without spines). It differs further in having a ventral count of 168 or 169 (versus more than
<quantity id="4C8628AEEA782A4FFDBFC61CC282F86F" box="[632,708,1909,1934]" metricMagnitude="0" metricUnit="m" metricValue="5.08" pageId="2" pageNumber="155" unit="in" value="200.0">200 in</quantity>
<emphasis id="B90A5959EA782A4FFD16C61CC302F86C" box="[721,836,1910,1933]" italics="true" pageId="2" pageNumber="155">R. goweri</emphasis>
<bibRefCitation id="EFEFF8BAEA782A4FFC97C61FC4D7F86F" author="Aengals" box="[848,1169,1909,1934]" pageId="2" pageNumber="155" refString="Aengals, R. &amp; Ganesh, S. R. (2013) Rhinophis goweri - a new species of Shieldtail snake from the southern Eastern Ghats, India. Russian Journal of Herpetology, 20 (1), 61 - 65." type="journal article" year="2013">Aengals and Ganesh, 2013</bibRefCitation>
, fewer than
<quantity id="4C8628AEEA782A4FFAECC61FC531F86F" box="[1323,1399,1909,1934]" metricMagnitude="0" metricUnit="m" metricValue="3.81" pageId="2" pageNumber="155" unit="in" value="150.0">150 in</quantity>
<taxonomicName id="4C7EFEC8EA782A4FFA43C61CC247F850" authority="Boulenger, 1893" authorityName="Boulenger" authorityYear="1893" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="155" phylum="Chordata" rank="species" species="travancoricus">
<emphasis id="B90A5959EA782A4FFA43C61CC175F851" italics="true" pageId="2" pageNumber="155">R. travancoricus</emphasis>
Boulenger, 1893
</taxonomicName>
, and more than
<quantity id="4C8628AEEA782A4FFD0DC6F3C351F850" box="[714,791,1944,1969]" metricMagnitude="0" metricUnit="m" metricValue="4.318" pageId="2" pageNumber="155" unit="in" value="170.0">170 in</quantity>
<taxonomicName id="4C7EFEC8EA782A4FFCE4C6F3C4F0F850" authority="Boulenger, 1896" authorityName="Boulenger" authorityYear="1896" box="[803,1206,1944,1969]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="155" phylum="Chordata" rank="species" species="fergusonianus">
<emphasis id="B90A5959EA782A4FFCE4C6F3C3A0F850" box="[803,998,1945,1969]" italics="true" pageId="2" pageNumber="155">R. fergusonianus</emphasis>
Boulenger, 1896
</taxonomicName>
); and in having 17 midbody dorsal scale rows (versus
<quantity id="4C8628AEEA782A4FFDE6C6D1C21CF835" box="[545,602,1979,2004]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="2" pageNumber="155" unit="in" value="15.0">15 in</quantity>
<taxonomicName id="4C7EFEC8EA782A4FFDA6C6D7C3FCF835" authority="Beddome, 1863" authorityName="Beddome" authorityYear="1863" box="[609,954,1979,2004]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="155" phylum="Chordata" rank="species" species="sanguineus">
<emphasis id="B90A5959EA782A4FFDA6C6D7C2B8F835" box="[609,766,1981,2004]" italics="true" pageId="2" pageNumber="155">R. sanguineus</emphasis>
Beddome, 1863
</taxonomicName>
).
</paragraph>
<caption id="DF01D5C3EA792A4EFF50C1F2C530FCAE" pageId="3" pageNumber="156">
<paragraph id="8BC1854BEA792A4EFF50C1F2C530FCAE" blockId="3.[151,1436,152,847]" pageId="3" pageNumber="156">
<emphasis id="B90A5959EA792A4EFF50C1F2C157FF4E" bold="true" box="[151,273,152,175]" pageId="3" pageNumber="156">TABLE 1.</emphasis>
Meristic and morphometric data for 52 characters of holotype (*) and paratypes (**) of
<taxonomicName id="4C7EFEC8EA792A4EFAF7C1F2C16BFF2E" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="156" phylum="Chordata" rank="species" species="roshanpererai" status="sp. nov.">
<emphasis id="B90A5959EA792A4EFAF7C1F2C16BFF2E" italics="true" pageId="3" pageNumber="156">Rhinophis roshanpererai</emphasis>
</taxonomicName>
<emphasis id="B90A5959EA792A4EFEF3C1D2C1CEFF2E" bold="true" box="[308,392,184,207]" pageId="3" pageNumber="156">
<taxonomicNameLabel id="A239E422EA792A4EFEF3C1D2C1CEFF2E" box="[308,392,184,207]" pageId="3" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Specimen numbers have NMSL prefix for holotype and DWC prefix for paratypes. Bilateral scale counts given in left, right order. Character numbers in parentheses, as follows: (1) supralabials, (2) infralabials, (3) diameter of eye, (4) maximum length of ocular, (5) length ocular/eye, (6) distance between eyes, (7) distance between eye-naris, (8) length of frontal, (9) width of frontal, (10) head length = distance between snout tip-posterior edge 4th supralabial, (11) total length/HL, (12) head width level with corner of mouth, (13) distance between nares, (14) maximum longitudinal length of prefrontal scales, (15) length of parietal scale = distance between posterior tip of parietal and posterior end of suture between frontal-ocular, (16) width of parietal scale = distance between posterior tip of ocular and posterior tip of frontal, (17) length of midline suture between parietals, (18) distance rostral-frontal, (19) distance between tip of snout and posterior edge of rostral, (20) distance between tip of snout and posterior limit of midline suture between parietals, (21) maximum width of rostral, viewed ventrally, (22) distance between snout tip-naris, (23) distance between snout tip-eye, (24) midbody width, (25) midbody depth, (26) midbody circumference, (27) exposed width of first scale row at midbody, (28) exposed width of ventral scales at midbody, (29) number of ventral scale level with which dorsal scale row is lost (i.e. location of change between 19 and 17 dorsal scale rows), (30) dorsal scale rows behind head, at midbody and one head length anterior to vent (excluding ventrals), (31) exposed length of first scale row at midbody, (32) width of ventral/first dorsal scale rows, (33) ventral scales, (34) vertebral scales, (35) snout vent length, (36) width at vent, (37) disposition of two anal scales (l/r) or (r/l), (38) number of small scales overlapped by anals, excluding first subcaudals, (39) subcaudal scales, (40) tail length, (41) total length, (42) tail length as % of total length, (43) total length/midbody width, (44) maximum width of tail shield, (45) maximum diameter of tail shield, (46) number of scales (including subcaudals) surrounding base of tail shield, (47) dorsal base to tip of tail shield (upper spines), (48) dorsal base to tip of tail shield (lower spines), (49) lateral base to tip of tail shield (upper), (50) lateral base to tip of tail shield (lower), (51) ventral base to tip of tail shield (upper), (52) ventral base to tip of tail shield (lower).
</paragraph>
</caption>
<paragraph id="8BC1854BEA792A4EFF32C200C540F852" pageId="3" pageNumber="156">
<table id="F97E77EBEA79D5B2FF58C200C534F852" box="[159,1394,874,1971]" gridcols="8" gridrows="27" pageId="3" pageNumber="156">
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<th id="769FEE75EA79D5B2FF32C200C1D9FC61" box="[245,415,874,896]" gridcol="1" gridrow="0" pageId="3" pageNumber="156">
2016.08.0 1
<emphasis id="B90A5959EA792A4EFEAAC200C1D9FC9E" bold="true" box="[365,415,874,895]" pageId="3" pageNumber="156">NH*</emphasis>
</th>
<th id="769FEE75EA79D5B2FE03C200C216FC61" box="[452,592,874,896]" gridcol="2" gridrow="0" pageId="3" pageNumber="156">
2016.05.0 3
<emphasis id="B90A5959EA792A4EFDF1C200C216FC9E" bold="true" box="[566,592,874,895]" pageId="3" pageNumber="156">**</emphasis>
</th>
<th id="769FEE75EA79D5B2FDA2C200C2B7FC61" box="[613,753,874,896]" gridcol="3" gridrow="0" pageId="3" pageNumber="156">
2016.05.0 4
<emphasis id="B90A5959EA792A4EFD10C200C2B7FC9E" bold="true" box="[727,753,874,895]" pageId="3" pageNumber="156">**</emphasis>
</th>
<th id="769FEE75EA79D5B2FC9DC200C443FC61" box="[858,1029,874,896]" gridcol="5" gridrow="0" pageId="3" pageNumber="156">
2016.08.0 1
<emphasis id="B90A5959EA792A4EFC15C200C443FC9E" bold="true" box="[978,1029,874,895]" pageId="3" pageNumber="156">NH*</emphasis>
</th>
<th id="769FEE75EA79D5B2FBE1C200C4F4FC61" box="[1062,1202,874,896]" gridcol="6" gridrow="0" pageId="3" pageNumber="156">
2016.05.0 3
<emphasis id="B90A5959EA792A4EFB5FC200C4F4FC9E" bold="true" box="[1176,1202,874,895]" pageId="3" pageNumber="156">**</emphasis>
</th>
<th id="769FEE75EA79D5B2FB21C200C534FC61" box="[1254,1394,874,896]" gridcol="7" gridrow="0" pageId="3" pageNumber="156">
2016.05.0 4
<emphasis id="B90A5959EA792A4EFA9FC200C534FC9E" bold="true" box="[1368,1394,874,895]" pageId="3" pageNumber="156">**</emphasis>
</th>
</tr>
<tr id="354E8709EA79D5B2FF58C2FEC534FC4B" box="[159,1394,916,938]" gridrow="1" pageId="3" pageNumber="156">
<th id="769FEE75EA79D5B2FF58C2FEC08FFC4B" box="[159,201,916,938]" gridcol="0" gridrow="1" pageId="3" pageNumber="156">(1)</th>
<td id="769FEE75EA79D5B2FF32C2FEC1D9FC4B" box="[245,415,916,938]" gridcol="1" gridrow="1" pageId="3" pageNumber="156">4</td>
<td id="769FEE75EA79D5B2FE03C2FEC216FC4B" box="[452,592,916,938]" gridcol="2" gridrow="1" pageId="3" pageNumber="156">4</td>
<td id="769FEE75EA79D5B2FDA2C2FEC2B7FC4B" box="[613,753,916,938]" gridcol="3" gridrow="1" pageId="3" pageNumber="156">4</td>
<td id="769FEE75EA79D5B2FCC1C2FEC377FC4B" box="[774,817,916,938]" gridcol="4" gridrow="1" pageId="3" pageNumber="156">(27)</td>
<td id="769FEE75EA79D5B2FC9DC2FEC443FC4B" box="[858,1029,916,938]" gridcol="5" gridrow="1" pageId="3" pageNumber="156">1.4</td>
<td id="769FEE75EA79D5B2FBE1C2FEC4F4FC4B" box="[1062,1202,916,938]" gridcol="6" gridrow="1" pageId="3" pageNumber="156">1.5</td>
<td id="769FEE75EA79D5B2FB21C2FEC534FC4B" box="[1254,1394,916,938]" gridcol="7" gridrow="1" pageId="3" pageNumber="156">1.6</td>
</tr>
<tr id="354E8709EA79D5B2FF58C2D7C534FC32" box="[159,1394,957,979]" gridrow="2" pageId="3" pageNumber="156">
<th id="769FEE75EA79D5B2FF58C2D7C08FFC32" box="[159,201,957,979]" gridcol="0" gridrow="2" pageId="3" pageNumber="156">(2)</th>
<td id="769FEE75EA79D5B2FF32C2D7C1D9FC32" box="[245,415,957,979]" gridcol="1" gridrow="2" pageId="3" pageNumber="156">3</td>
<td id="769FEE75EA79D5B2FE03C2D7C216FC32" box="[452,592,957,979]" gridcol="2" gridrow="2" pageId="3" pageNumber="156">3</td>
<td id="769FEE75EA79D5B2FDA2C2D7C2B7FC32" box="[613,753,957,979]" gridcol="3" gridrow="2" pageId="3" pageNumber="156">3</td>
<td id="769FEE75EA79D5B2FCC1C2D7C377FC32" box="[774,817,957,979]" gridcol="4" gridrow="2" pageId="3" pageNumber="156">(28)</td>
<td id="769FEE75EA79D5B2FC9DC2D7C443FC32" box="[858,1029,957,979]" gridcol="5" gridrow="2" pageId="3" pageNumber="156">2.2</td>
<td id="769FEE75EA79D5B2FBE1C2D7C4F4FC32" box="[1062,1202,957,979]" gridcol="6" gridrow="2" pageId="3" pageNumber="156">1.9</td>
<td id="769FEE75EA79D5B2FB21C2D7C534FC32" box="[1254,1394,957,979]" gridcol="7" gridrow="2" pageId="3" pageNumber="156">2.0</td>
</tr>
<tr id="354E8709EA79D5B2FF58C28CC534FC1D" box="[159,1394,998,1020]" gridrow="3" pageId="3" pageNumber="156">
<th id="769FEE75EA79D5B2FF58C28CC08FFC1D" box="[159,201,998,1020]" gridcol="0" gridrow="3" pageId="3" pageNumber="156">(3)</th>
<td id="769FEE75EA79D5B2FF32C28CC1D9FC1D" box="[245,415,998,1020]" gridcol="1" gridrow="3" pageId="3" pageNumber="156">0.5</td>
<td id="769FEE75EA79D5B2FE03C28CC216FC1D" box="[452,592,998,1020]" gridcol="2" gridrow="3" pageId="3" pageNumber="156">0.5</td>
<td id="769FEE75EA79D5B2FDA2C28CC2B7FC1D" box="[613,753,998,1020]" gridcol="3" gridrow="3" pageId="3" pageNumber="156">0.6</td>
<td id="769FEE75EA79D5B2FCC1C28CC377FC1D" box="[774,817,998,1020]" gridcol="4" gridrow="3" pageId="3" pageNumber="156">(29)</td>
<td id="769FEE75EA79D5B2FC9DC28CC443FC1D" box="[858,1029,998,1020]" gridcol="5" gridrow="3" pageId="3" pageNumber="156">30, 30</td>
<td id="769FEE75EA79D5B2FBE1C28CC4F4FC1D" box="[1062,1202,998,1020]" gridcol="6" gridrow="3" pageId="3" pageNumber="156">30, 30</td>
<td id="769FEE75EA79D5B2FB21C28CC534FC1D" box="[1254,1394,998,1020]" gridcol="7" gridrow="3" pageId="3" pageNumber="156">30, 30</td>
</tr>
<tr id="354E8709EA79D5B2FF58C57AC534FBC7" box="[159,1394,1040,1062]" gridrow="4" pageId="3" pageNumber="156">
<th id="769FEE75EA79D5B2FF58C57AC08FFBC7" box="[159,201,1040,1062]" gridcol="0" gridrow="4" pageId="3" pageNumber="156">(4)</th>
<td id="769FEE75EA79D5B2FF32C57AC1D9FBC7" box="[245,415,1040,1062]" gridcol="1" gridrow="4" pageId="3" pageNumber="156">1.7</td>
<td id="769FEE75EA79D5B2FE03C57AC216FBC7" box="[452,592,1040,1062]" gridcol="2" gridrow="4" pageId="3" pageNumber="156">1.7</td>
<td id="769FEE75EA79D5B2FDA2C57AC2B7FBC7" box="[613,753,1040,1062]" gridcol="3" gridrow="4" pageId="3" pageNumber="156">1.8</td>
<td id="769FEE75EA79D5B2FCC1C57AC377FBC7" box="[774,817,1040,1062]" gridcol="4" gridrow="4" pageId="3" pageNumber="156">(30)</td>
<td id="769FEE75EA79D5B2FC9DC57AC443FBC7" box="[858,1029,1040,1062]" gridcol="5" gridrow="4" pageId="3" pageNumber="156">19, 17, 17</td>
<td id="769FEE75EA79D5B2FBE1C57AC4F4FBC7" box="[1062,1202,1040,1062]" gridcol="6" gridrow="4" pageId="3" pageNumber="156">19, 17, 17</td>
<td id="769FEE75EA79D5B2FB21C57AC534FBC7" box="[1254,1394,1040,1062]" gridcol="7" gridrow="4" pageId="3" pageNumber="156">19, 17, 17</td>
</tr>
<tr id="354E8709EA79D5B2FF58C553C534FBAE" box="[159,1394,1081,1103]" gridrow="5" pageId="3" pageNumber="156">
<th id="769FEE75EA79D5B2FF58C553C08FFBAE" box="[159,201,1081,1103]" gridcol="0" gridrow="5" pageId="3" pageNumber="156">(5)</th>
<td id="769FEE75EA79D5B2FF32C553C1D9FBAE" box="[245,415,1081,1103]" gridcol="1" gridrow="5" pageId="3" pageNumber="156">3.2</td>
<td id="769FEE75EA79D5B2FE03C553C216FBAE" box="[452,592,1081,1103]" gridcol="2" gridrow="5" pageId="3" pageNumber="156">3.2</td>
<td id="769FEE75EA79D5B2FDA2C553C2B7FBAE" box="[613,753,1081,1103]" gridcol="3" gridrow="5" pageId="3" pageNumber="156">3.3</td>
<td id="769FEE75EA79D5B2FCC1C553C377FBAE" box="[774,817,1081,1103]" gridcol="4" gridrow="5" pageId="3" pageNumber="156">(31)</td>
<td id="769FEE75EA79D5B2FC9DC553C443FBAE" box="[858,1029,1081,1103]" gridcol="5" gridrow="5" pageId="3" pageNumber="156">1.3</td>
<td id="769FEE75EA79D5B2FBE1C553C4F4FBAE" box="[1062,1202,1081,1103]" gridcol="6" gridrow="5" pageId="3" pageNumber="156">1.1</td>
<td id="769FEE75EA79D5B2FB21C553C534FBAE" box="[1254,1394,1081,1103]" gridcol="7" gridrow="5" pageId="3" pageNumber="156">1.4</td>
</tr>
<tr id="354E8709EA79D5B2FF58C508C534FB99" box="[159,1394,1122,1144]" gridrow="6" pageId="3" pageNumber="156">
<th id="769FEE75EA79D5B2FF58C508C08FFB99" box="[159,201,1122,1144]" gridcol="0" gridrow="6" pageId="3" pageNumber="156">(6)</th>
<td id="769FEE75EA79D5B2FF32C508C1D9FB99" box="[245,415,1122,1144]" gridcol="1" gridrow="6" pageId="3" pageNumber="156">2.9</td>
<td id="769FEE75EA79D5B2FE03C508C216FB99" box="[452,592,1122,1144]" gridcol="2" gridrow="6" pageId="3" pageNumber="156">3.0</td>
<td id="769FEE75EA79D5B2FDA2C508C2B7FB99" box="[613,753,1122,1144]" gridcol="3" gridrow="6" pageId="3" pageNumber="156">2.8</td>
<td id="769FEE75EA79D5B2FCC1C508C377FB99" box="[774,817,1122,1144]" gridcol="4" gridrow="6" pageId="3" pageNumber="156">(32)</td>
<td id="769FEE75EA79D5B2FC9DC508C443FB99" box="[858,1029,1122,1144]" gridcol="5" gridrow="6" pageId="3" pageNumber="156">1.5</td>
<td id="769FEE75EA79D5B2FBE1C508C4F4FB99" box="[1062,1202,1122,1144]" gridcol="6" gridrow="6" pageId="3" pageNumber="156">1.3</td>
<td id="769FEE75EA79D5B2FB21C508C534FB99" box="[1254,1394,1122,1144]" gridcol="7" gridrow="6" pageId="3" pageNumber="156">1.3</td>
</tr>
<tr id="354E8709EA79D5B2FF58C5E6C534FB43" box="[159,1394,1164,1186]" gridrow="7" pageId="3" pageNumber="156">
<th id="769FEE75EA79D5B2FF58C5E6C08FFB43" box="[159,201,1164,1186]" gridcol="0" gridrow="7" pageId="3" pageNumber="156">(7)</th>
<td id="769FEE75EA79D5B2FF32C5E6C1D9FB43" box="[245,415,1164,1186]" gridcol="1" gridrow="7" pageId="3" pageNumber="156">2.1</td>
<td id="769FEE75EA79D5B2FE03C5E6C216FB43" box="[452,592,1164,1186]" gridcol="2" gridrow="7" pageId="3" pageNumber="156">2.1</td>
<td id="769FEE75EA79D5B2FDA2C5E6C2B7FB43" box="[613,753,1164,1186]" gridcol="3" gridrow="7" pageId="3" pageNumber="156">2.2</td>
<td id="769FEE75EA79D5B2FCC1C5E6C377FB43" box="[774,817,1164,1186]" gridcol="4" gridrow="7" pageId="3" pageNumber="156">(33)</td>
<td id="769FEE75EA79D5B2FC9DC5E6C443FB43" box="[858,1029,1164,1186]" gridcol="5" gridrow="7" pageId="3" pageNumber="156">169</td>
<td id="769FEE75EA79D5B2FBE1C5E6C4F4FB43" box="[1062,1202,1164,1186]" gridcol="6" gridrow="7" pageId="3" pageNumber="156">168</td>
<td id="769FEE75EA79D5B2FB21C5E6C534FB43" box="[1254,1394,1164,1186]" gridcol="7" gridrow="7" pageId="3" pageNumber="156">169</td>
</tr>
<tr id="354E8709EA79D5B2FF58C5DFC534FB2A" box="[159,1394,1205,1227]" gridrow="8" pageId="3" pageNumber="156">
<th id="769FEE75EA79D5B2FF58C5DFC08FFB2A" box="[159,201,1205,1227]" gridcol="0" gridrow="8" pageId="3" pageNumber="156">(8)</th>
<td id="769FEE75EA79D5B2FF32C5DFC1D9FB2A" box="[245,415,1205,1227]" gridcol="1" gridrow="8" pageId="3" pageNumber="156">2.1</td>
<td id="769FEE75EA79D5B2FE03C5DFC216FB2A" box="[452,592,1205,1227]" gridcol="2" gridrow="8" pageId="3" pageNumber="156">2.1</td>
<td id="769FEE75EA79D5B2FDA2C5DFC2B7FB2A" box="[613,753,1205,1227]" gridcol="3" gridrow="8" pageId="3" pageNumber="156">2.3</td>
<td id="769FEE75EA79D5B2FCC1C5DFC377FB2A" box="[774,817,1205,1227]" gridcol="4" gridrow="8" pageId="3" pageNumber="156">(34)</td>
<td id="769FEE75EA79D5B2FC9DC5DFC443FB2A" box="[858,1029,1205,1227]" gridcol="5" gridrow="8" pageId="3" pageNumber="156">176</td>
<td id="769FEE75EA79D5B2FBE1C5DFC4F4FB2A" box="[1062,1202,1205,1227]" gridcol="6" gridrow="8" pageId="3" pageNumber="156">175</td>
<td id="769FEE75EA79D5B2FB21C5DFC534FB2A" box="[1254,1394,1205,1227]" gridcol="7" gridrow="8" pageId="3" pageNumber="156">174</td>
</tr>
<tr id="354E8709EA79D5B2FF58C5B4C534FB15" box="[159,1394,1246,1268]" gridrow="9" pageId="3" pageNumber="156">
<th id="769FEE75EA79D5B2FF58C5B4C08FFB15" box="[159,201,1246,1268]" gridcol="0" gridrow="9" pageId="3" pageNumber="156">(9)</th>
<td id="769FEE75EA79D5B2FF32C5B4C1D9FB15" box="[245,415,1246,1268]" gridcol="1" gridrow="9" pageId="3" pageNumber="156">1.6</td>
<td id="769FEE75EA79D5B2FE03C5B4C216FB15" box="[452,592,1246,1268]" gridcol="2" gridrow="9" pageId="3" pageNumber="156">1.6</td>
<td id="769FEE75EA79D5B2FDA2C5B4C2B7FB15" box="[613,753,1246,1268]" gridcol="3" gridrow="9" pageId="3" pageNumber="156">1.8</td>
<td id="769FEE75EA79D5B2FCC1C5B4C377FB15" box="[774,817,1246,1268]" gridcol="4" gridrow="9" pageId="3" pageNumber="156">(35)</td>
<td id="769FEE75EA79D5B2FC9DC5B4C443FB15" box="[858,1029,1246,1268]" gridcol="5" gridrow="9" pageId="3" pageNumber="156">207.9</td>
<td id="769FEE75EA79D5B2FBE1C5B4C4F4FB15" box="[1062,1202,1246,1268]" gridcol="6" gridrow="9" pageId="3" pageNumber="156">205.2</td>
<td id="769FEE75EA79D5B2FB21C5B4C534FB15" box="[1254,1394,1246,1268]" gridcol="7" gridrow="9" pageId="3" pageNumber="156">218.2</td>
</tr>
<tr id="354E8709EA79D5B2FF58C462C534FAFF" box="[159,1394,1288,1310]" gridrow="10" pageId="3" pageNumber="156">
<th id="769FEE75EA79D5B2FF58C462C08FFAFF" box="[159,201,1288,1310]" gridcol="0" gridrow="10" pageId="3" pageNumber="156">(10)</th>
<td id="769FEE75EA79D5B2FF32C462C1D9FAFF" box="[245,415,1288,1310]" gridcol="1" gridrow="10" pageId="3" pageNumber="156">6.2</td>
<td id="769FEE75EA79D5B2FE03C462C216FAFF" box="[452,592,1288,1310]" gridcol="2" gridrow="10" pageId="3" pageNumber="156">6.0</td>
<td id="769FEE75EA79D5B2FDA2C462C2B7FAFF" box="[613,753,1288,1310]" gridcol="3" gridrow="10" pageId="3" pageNumber="156">6.3</td>
<td id="769FEE75EA79D5B2FCC1C462C377FAFF" box="[774,817,1288,1310]" gridcol="4" gridrow="10" pageId="3" pageNumber="156">(36)</td>
<td id="769FEE75EA79D5B2FC9DC462C443FAFF" box="[858,1029,1288,1310]" gridcol="5" gridrow="10" pageId="3" pageNumber="156">4.0</td>
<td id="769FEE75EA79D5B2FBE1C462C4F4FAFF" box="[1062,1202,1288,1310]" gridcol="6" gridrow="10" pageId="3" pageNumber="156">4.1</td>
<td id="769FEE75EA79D5B2FB21C462C534FAFF" box="[1254,1394,1288,1310]" gridcol="7" gridrow="10" pageId="3" pageNumber="156">4.2</td>
</tr>
<tr id="354E8709EA79D5B2FF58C45BC534FAA6" box="[159,1394,1329,1351]" gridrow="11" pageId="3" pageNumber="156">
<th id="769FEE75EA79D5B2FF58C45BC08FFAA6" box="[159,201,1329,1351]" gridcol="0" gridrow="11" pageId="3" pageNumber="156">(11)</th>
<td id="769FEE75EA79D5B2FF32C45BC1D9FAA6" box="[245,415,1329,1351]" gridcol="1" gridrow="11" pageId="3" pageNumber="156">34.8</td>
<td id="769FEE75EA79D5B2FE03C45BC216FAA6" box="[452,592,1329,1351]" gridcol="2" gridrow="11" pageId="3" pageNumber="156">35.3</td>
<td id="769FEE75EA79D5B2FDA2C45BC2B7FAA6" box="[613,753,1329,1351]" gridcol="3" gridrow="11" pageId="3" pageNumber="156">35.6</td>
<td id="769FEE75EA79D5B2FCC1C45BC377FAA6" box="[774,817,1329,1351]" gridcol="4" gridrow="11" pageId="3" pageNumber="156">(37)</td>
<td id="769FEE75EA79D5B2FC9DC45BC443FAA6" box="[858,1029,1329,1351]" gridcol="5" gridrow="11" pageId="3" pageNumber="156">r/l</td>
<td id="769FEE75EA79D5B2FBE1C45BC4F4FAA6" box="[1062,1202,1329,1351]" gridcol="6" gridrow="11" pageId="3" pageNumber="156">r/l</td>
<td id="769FEE75EA79D5B2FB21C45BC534FAA6" box="[1254,1394,1329,1351]" gridcol="7" gridrow="11" pageId="3" pageNumber="156">r/l</td>
</tr>
<tr id="354E8709EA79D5B2FF58C430C534FA91" box="[159,1394,1370,1392]" gridrow="12" pageId="3" pageNumber="156">
<th id="769FEE75EA79D5B2FF58C430C08FFA91" box="[159,201,1370,1392]" gridcol="0" gridrow="12" pageId="3" pageNumber="156">(12)</th>
<td id="769FEE75EA79D5B2FF32C430C1D9FA91" box="[245,415,1370,1392]" gridcol="1" gridrow="12" pageId="3" pageNumber="156">4.4</td>
<td id="769FEE75EA79D5B2FE03C430C216FA91" box="[452,592,1370,1392]" gridcol="2" gridrow="12" pageId="3" pageNumber="156">4.4</td>
<td id="769FEE75EA79D5B2FDA2C430C2B7FA91" box="[613,753,1370,1392]" gridcol="3" gridrow="12" pageId="3" pageNumber="156">4.6</td>
<td id="769FEE75EA79D5B2FCC1C430C377FA91" box="[774,817,1370,1392]" gridcol="4" gridrow="12" pageId="3" pageNumber="156">(38)</td>
<td id="769FEE75EA79D5B2FC9DC430C443FA91" box="[858,1029,1370,1392]" gridcol="5" gridrow="12" pageId="3" pageNumber="156">3, 3</td>
<td id="769FEE75EA79D5B2FBE1C430C4F4FA91" box="[1062,1202,1370,1392]" gridcol="6" gridrow="12" pageId="3" pageNumber="156">3, 3</td>
<td id="769FEE75EA79D5B2FB21C430C534FA91" box="[1254,1394,1370,1392]" gridcol="7" gridrow="12" pageId="3" pageNumber="156">3, 3</td>
</tr>
<tr id="354E8709EA79D5B2FF58C4EEC534FA7B" box="[159,1394,1412,1434]" gridrow="13" pageId="3" pageNumber="156">
<th id="769FEE75EA79D5B2FF58C4EEC08FFA7B" box="[159,201,1412,1434]" gridcol="0" gridrow="13" pageId="3" pageNumber="156">(13)</th>
<td id="769FEE75EA79D5B2FF32C4EEC1D9FA7B" box="[245,415,1412,1434]" gridcol="1" gridrow="13" pageId="3" pageNumber="156">1.4</td>
<td id="769FEE75EA79D5B2FE03C4EEC216FA7B" box="[452,592,1412,1434]" gridcol="2" gridrow="13" pageId="3" pageNumber="156">1.4</td>
<td id="769FEE75EA79D5B2FDA2C4EEC2B7FA7B" box="[613,753,1412,1434]" gridcol="3" gridrow="13" pageId="3" pageNumber="156">1.5</td>
<td id="769FEE75EA79D5B2FCC1C4EEC377FA7B" box="[774,817,1412,1434]" gridcol="4" gridrow="13" pageId="3" pageNumber="156">(39)</td>
<td id="769FEE75EA79D5B2FC9DC4EEC443FA7B" box="[858,1029,1412,1434]" gridcol="5" gridrow="13" pageId="3" pageNumber="156">7, 7</td>
<td id="769FEE75EA79D5B2FBE1C4EEC4F4FA7B" box="[1062,1202,1412,1434]" gridcol="6" gridrow="13" pageId="3" pageNumber="156">7, 7</td>
<td id="769FEE75EA79D5B2FB21C4EEC534FA7B" box="[1254,1394,1412,1434]" gridcol="7" gridrow="13" pageId="3" pageNumber="156">7, 6</td>
</tr>
<tr id="354E8709EA79D5B2FF58C4C7C534FA22" box="[159,1394,1453,1475]" gridrow="14" pageId="3" pageNumber="156">
<th id="769FEE75EA79D5B2FF58C4C7C08FFA22" box="[159,201,1453,1475]" gridcol="0" gridrow="14" pageId="3" pageNumber="156">(14)</th>
<td id="769FEE75EA79D5B2FF32C4C7C1D9FA22" box="[245,415,1453,1475]" gridcol="1" gridrow="14" pageId="3" pageNumber="156">1.6</td>
<td id="769FEE75EA79D5B2FE03C4C7C216FA22" box="[452,592,1453,1475]" gridcol="2" gridrow="14" pageId="3" pageNumber="156">1.5</td>
<td id="769FEE75EA79D5B2FDA2C4C7C2B7FA22" box="[613,753,1453,1475]" gridcol="3" gridrow="14" pageId="3" pageNumber="156">1.6</td>
<td id="769FEE75EA79D5B2FCC1C4C7C377FA22" box="[774,817,1453,1475]" gridcol="4" gridrow="14" pageId="3" pageNumber="156">(40)</td>
<td id="769FEE75EA79D5B2FC9DC4C7C443FA22" box="[858,1029,1453,1475]" gridcol="5" gridrow="14" pageId="3" pageNumber="156">6.1</td>
<td id="769FEE75EA79D5B2FBE1C4C7C4F4FA22" box="[1062,1202,1453,1475]" gridcol="6" gridrow="14" pageId="3" pageNumber="156">5.8</td>
<td id="769FEE75EA79D5B2FB21C4C7C534FA22" box="[1254,1394,1453,1475]" gridcol="7" gridrow="14" pageId="3" pageNumber="156">5.8</td>
</tr>
<tr id="354E8709EA79D5B2FF58C4BCC534FA0D" box="[159,1394,1494,1516]" gridrow="15" pageId="3" pageNumber="156">
<th id="769FEE75EA79D5B2FF58C4BCC08FFA0D" box="[159,201,1494,1516]" gridcol="0" gridrow="15" pageId="3" pageNumber="156">(15)</th>
<td id="769FEE75EA79D5B2FF32C4BCC1D9FA0D" box="[245,415,1494,1516]" gridcol="1" gridrow="15" pageId="3" pageNumber="156">2.2</td>
<td id="769FEE75EA79D5B2FE03C4BCC216FA0D" box="[452,592,1494,1516]" gridcol="2" gridrow="15" pageId="3" pageNumber="156">2.1</td>
<td id="769FEE75EA79D5B2FDA2C4BCC2B7FA0D" box="[613,753,1494,1516]" gridcol="3" gridrow="15" pageId="3" pageNumber="156">2.3</td>
<td id="769FEE75EA79D5B2FCC1C4BCC377FA0D" box="[774,817,1494,1516]" gridcol="4" gridrow="15" pageId="3" pageNumber="156">(41)</td>
<td id="769FEE75EA79D5B2FC9DC4BCC443FA0D" box="[858,1029,1494,1516]" gridcol="5" gridrow="15" pageId="3" pageNumber="156">214.0</td>
<td id="769FEE75EA79D5B2FBE1C4BCC4F4FA0D" box="[1062,1202,1494,1516]" gridcol="6" gridrow="15" pageId="3" pageNumber="156">211.0</td>
<td id="769FEE75EA79D5B2FB21C4BCC534FA0D" box="[1254,1394,1494,1516]" gridcol="7" gridrow="15" pageId="3" pageNumber="156">224.0</td>
</tr>
<tr id="354E8709EA79D5B2FF58C76AC534F9F7" box="[159,1394,1536,1558]" gridrow="16" pageId="3" pageNumber="156">
<th id="769FEE75EA79D5B2FF58C76AC08FF9F7" box="[159,201,1536,1558]" gridcol="0" gridrow="16" pageId="3" pageNumber="156">(16)</th>
<td id="769FEE75EA79D5B2FF32C76AC1D9F9F7" box="[245,415,1536,1558]" gridcol="1" gridrow="16" pageId="3" pageNumber="156">1.8</td>
<td id="769FEE75EA79D5B2FE03C76AC216F9F7" box="[452,592,1536,1558]" gridcol="2" gridrow="16" pageId="3" pageNumber="156">1.8</td>
<td id="769FEE75EA79D5B2FDA2C76AC2B7F9F7" box="[613,753,1536,1558]" gridcol="3" gridrow="16" pageId="3" pageNumber="156">2.1</td>
<td id="769FEE75EA79D5B2FCC1C76AC377F9F7" box="[774,817,1536,1558]" gridcol="4" gridrow="16" pageId="3" pageNumber="156">(42)</td>
<td id="769FEE75EA79D5B2FC9DC76AC443F9F7" box="[858,1029,1536,1558]" gridcol="5" gridrow="16" pageId="3" pageNumber="156">2.8</td>
<td id="769FEE75EA79D5B2FBE1C76AC4F4F9F7" box="[1062,1202,1536,1558]" gridcol="6" gridrow="16" pageId="3" pageNumber="156">2.8</td>
<td id="769FEE75EA79D5B2FB21C76AC534F9F7" box="[1254,1394,1536,1558]" gridcol="7" gridrow="16" pageId="3" pageNumber="156">2.6</td>
</tr>
<tr id="354E8709EA79D5B2FF58C743C534F9DE" box="[159,1394,1577,1599]" gridrow="17" pageId="3" pageNumber="156">
<th id="769FEE75EA79D5B2FF58C743C08FF9DE" box="[159,201,1577,1599]" gridcol="0" gridrow="17" pageId="3" pageNumber="156">(17)</th>
<td id="769FEE75EA79D5B2FF32C743C1D9F9DE" box="[245,415,1577,1599]" gridcol="1" gridrow="17" pageId="3" pageNumber="156">0.5</td>
<td id="769FEE75EA79D5B2FE03C743C216F9DE" box="[452,592,1577,1599]" gridcol="2" gridrow="17" pageId="3" pageNumber="156">0.4</td>
<td id="769FEE75EA79D5B2FDA2C743C2B7F9DE" box="[613,753,1577,1599]" gridcol="3" gridrow="17" pageId="3" pageNumber="156">0.4</td>
<td id="769FEE75EA79D5B2FCC1C743C377F9DE" box="[774,817,1577,1599]" gridcol="4" gridrow="17" pageId="3" pageNumber="156">(43)</td>
<td id="769FEE75EA79D5B2FC9DC743C443F9DE" box="[858,1029,1577,1599]" gridcol="5" gridrow="17" pageId="3" pageNumber="156">30.0</td>
<td id="769FEE75EA79D5B2FBE1C743C4F4F9DE" box="[1062,1202,1577,1599]" gridcol="6" gridrow="17" pageId="3" pageNumber="156">29.8</td>
<td id="769FEE75EA79D5B2FB21C743C534F9DE" box="[1254,1394,1577,1599]" gridcol="7" gridrow="17" pageId="3" pageNumber="156">30.9</td>
</tr>
<tr id="354E8709EA79D5B2FF58C738C534F989" box="[159,1394,1618,1640]" gridrow="18" pageId="3" pageNumber="156">
<th id="769FEE75EA79D5B2FF58C738C08FF989" box="[159,201,1618,1640]" gridcol="0" gridrow="18" pageId="3" pageNumber="156">(18)</th>
<td id="769FEE75EA79D5B2FF32C738C1D9F989" box="[245,415,1618,1640]" gridcol="1" gridrow="18" pageId="3" pageNumber="156">0.6</td>
<td id="769FEE75EA79D5B2FE03C738C216F989" box="[452,592,1618,1640]" gridcol="2" gridrow="18" pageId="3" pageNumber="156">0.4</td>
<td id="769FEE75EA79D5B2FDA2C738C2B7F989" box="[613,753,1618,1640]" gridcol="3" gridrow="18" pageId="3" pageNumber="156">0.6</td>
<td id="769FEE75EA79D5B2FCC1C738C377F989" box="[774,817,1618,1640]" gridcol="4" gridrow="18" pageId="3" pageNumber="156">(44)</td>
<td id="769FEE75EA79D5B2FC9DC738C443F989" box="[858,1029,1618,1640]" gridcol="5" gridrow="18" pageId="3" pageNumber="156">1.6</td>
<td id="769FEE75EA79D5B2FBE1C738C4F4F989" box="[1062,1202,1618,1640]" gridcol="6" gridrow="18" pageId="3" pageNumber="156">1.4</td>
<td id="769FEE75EA79D5B2FB21C738C534F989" box="[1254,1394,1618,1640]" gridcol="7" gridrow="18" pageId="3" pageNumber="156">1.6</td>
</tr>
<tr id="354E8709EA79D5B2FF58C716C534F973" box="[159,1394,1660,1682]" gridrow="19" pageId="3" pageNumber="156">
<th id="769FEE75EA79D5B2FF58C716C08FF973" box="[159,201,1660,1682]" gridcol="0" gridrow="19" pageId="3" pageNumber="156">(19)</th>
<td id="769FEE75EA79D5B2FF32C716C1D9F973" box="[245,415,1660,1682]" gridcol="1" gridrow="19" pageId="3" pageNumber="156">2.0</td>
<td id="769FEE75EA79D5B2FE03C716C216F973" box="[452,592,1660,1682]" gridcol="2" gridrow="19" pageId="3" pageNumber="156">2.1</td>
<td id="769FEE75EA79D5B2FDA2C716C2B7F973" box="[613,753,1660,1682]" gridcol="3" gridrow="19" pageId="3" pageNumber="156">2.3</td>
<td id="769FEE75EA79D5B2FCC1C716C377F973" box="[774,817,1660,1682]" gridcol="4" gridrow="19" pageId="3" pageNumber="156">(45)</td>
<td id="769FEE75EA79D5B2FC9DC716C443F973" box="[858,1029,1660,1682]" gridcol="5" gridrow="19" pageId="3" pageNumber="156">2.3</td>
<td id="769FEE75EA79D5B2FBE1C716C4F4F973" box="[1062,1202,1660,1682]" gridcol="6" gridrow="19" pageId="3" pageNumber="156">2.1</td>
<td id="769FEE75EA79D5B2FB21C716C534F973" box="[1254,1394,1660,1682]" gridcol="7" gridrow="19" pageId="3" pageNumber="156">2.4</td>
</tr>
<tr id="354E8709EA79D5B2FF58C7CFC534F95A" box="[159,1394,1701,1723]" gridrow="20" pageId="3" pageNumber="156">
<th id="769FEE75EA79D5B2FF58C7CFC08FF95A" box="[159,201,1701,1723]" gridcol="0" gridrow="20" pageId="3" pageNumber="156">(20)</th>
<td id="769FEE75EA79D5B2FF32C7CFC1D9F95A" box="[245,415,1701,1723]" gridcol="1" gridrow="20" pageId="3" pageNumber="156">5.4</td>
<td id="769FEE75EA79D5B2FE03C7CFC216F95A" box="[452,592,1701,1723]" gridcol="2" gridrow="20" pageId="3" pageNumber="156">5.2</td>
<td id="769FEE75EA79D5B2FDA2C7CFC2B7F95A" box="[613,753,1701,1723]" gridcol="3" gridrow="20" pageId="3" pageNumber="156">5.7</td>
<td id="769FEE75EA79D5B2FCC1C7CFC377F95A" box="[774,817,1701,1723]" gridcol="4" gridrow="20" pageId="3" pageNumber="156">(46)</td>
<td id="769FEE75EA79D5B2FC9DC7CFC443F95A" box="[858,1029,1701,1723]" gridcol="5" gridrow="20" pageId="3" pageNumber="156">8</td>
<td id="769FEE75EA79D5B2FBE1C7CFC4F4F95A" box="[1062,1202,1701,1723]" gridcol="6" gridrow="20" pageId="3" pageNumber="156">9</td>
<td id="769FEE75EA79D5B2FB21C7CFC534F95A" box="[1254,1394,1701,1723]" gridcol="7" gridrow="20" pageId="3" pageNumber="156">8</td>
</tr>
<tr id="354E8709EA79D5B2FF58C7A4C534F905" box="[159,1394,1742,1764]" gridrow="21" pageId="3" pageNumber="156">
<th id="769FEE75EA79D5B2FF58C7A4C08FF905" box="[159,201,1742,1764]" gridcol="0" gridrow="21" pageId="3" pageNumber="156">(21)</th>
<td id="769FEE75EA79D5B2FF32C7A4C1D9F905" box="[245,415,1742,1764]" gridcol="1" gridrow="21" pageId="3" pageNumber="156">1.3</td>
<td id="769FEE75EA79D5B2FE03C7A4C216F905" box="[452,592,1742,1764]" gridcol="2" gridrow="21" pageId="3" pageNumber="156">1.1</td>
<td id="769FEE75EA79D5B2FDA2C7A4C2B7F905" box="[613,753,1742,1764]" gridcol="3" gridrow="21" pageId="3" pageNumber="156">1.3</td>
<td id="769FEE75EA79D5B2FCC1C7A4C377F905" box="[774,817,1742,1764]" gridcol="4" gridrow="21" pageId="3" pageNumber="156">(47)</td>
<td id="769FEE75EA79D5B2FC9DC7A4C443F905" box="[858,1029,1742,1764]" gridcol="5" gridrow="21" pageId="3" pageNumber="156">1.4</td>
<td id="769FEE75EA79D5B2FBE1C7A4C4F4F905" box="[1062,1202,1742,1764]" gridcol="6" gridrow="21" pageId="3" pageNumber="156">1.5</td>
<td id="769FEE75EA79D5B2FB21C7A4C534F905" box="[1254,1394,1742,1764]" gridcol="7" gridrow="21" pageId="3" pageNumber="156">1.3</td>
</tr>
<tr id="354E8709EA79D5B2FF58C792C534F8EF" box="[159,1394,1784,1806]" gridrow="22" pageId="3" pageNumber="156">
<th id="769FEE75EA79D5B2FF58C792C08FF8EF" box="[159,201,1784,1806]" gridcol="0" gridrow="22" pageId="3" pageNumber="156">(22)</th>
<td id="769FEE75EA79D5B2FF32C792C1D9F8EF" box="[245,415,1784,1806]" gridcol="1" gridrow="22" pageId="3" pageNumber="156">1.1</td>
<td id="769FEE75EA79D5B2FE03C792C216F8EF" box="[452,592,1784,1806]" gridcol="2" gridrow="22" pageId="3" pageNumber="156">1.1</td>
<td id="769FEE75EA79D5B2FDA2C792C2B7F8EF" box="[613,753,1784,1806]" gridcol="3" gridrow="22" pageId="3" pageNumber="156">1.1</td>
<td id="769FEE75EA79D5B2FCC1C792C377F8EF" box="[774,817,1784,1806]" gridcol="4" gridrow="22" pageId="3" pageNumber="156">(48)</td>
<td id="769FEE75EA79D5B2FC9DC792C443F8EF" box="[858,1029,1784,1806]" gridcol="5" gridrow="22" pageId="3" pageNumber="156">2.0</td>
<td id="769FEE75EA79D5B2FBE1C792C4F4F8EF" box="[1062,1202,1784,1806]" gridcol="6" gridrow="22" pageId="3" pageNumber="156">2.2</td>
<td id="769FEE75EA79D5B2FB21C792C534F8EF" box="[1254,1394,1784,1806]" gridcol="7" gridrow="22" pageId="3" pageNumber="156">2.1</td>
</tr>
<tr id="354E8709EA79D5B2FF58C64BC534F8D6" box="[159,1394,1825,1847]" gridrow="23" pageId="3" pageNumber="156">
<th id="769FEE75EA79D5B2FF58C64BC08FF8D6" box="[159,201,1825,1847]" gridcol="0" gridrow="23" pageId="3" pageNumber="156">(23)</th>
<td id="769FEE75EA79D5B2FF32C64BC1D9F8D6" box="[245,415,1825,1847]" gridcol="1" gridrow="23" pageId="3" pageNumber="156">3.3</td>
<td id="769FEE75EA79D5B2FE03C64BC216F8D6" box="[452,592,1825,1847]" gridcol="2" gridrow="23" pageId="3" pageNumber="156">3.1</td>
<td id="769FEE75EA79D5B2FDA2C64BC2B7F8D6" box="[613,753,1825,1847]" gridcol="3" gridrow="23" pageId="3" pageNumber="156">3.3</td>
<td id="769FEE75EA79D5B2FCC1C64BC377F8D6" box="[774,817,1825,1847]" gridcol="4" gridrow="23" pageId="3" pageNumber="156">(49)</td>
<td id="769FEE75EA79D5B2FC9DC64BC443F8D6" box="[858,1029,1825,1847]" gridcol="5" gridrow="23" pageId="3" pageNumber="156">0.8</td>
<td id="769FEE75EA79D5B2FBE1C64BC4F4F8D6" box="[1062,1202,1825,1847]" gridcol="6" gridrow="23" pageId="3" pageNumber="156">0.8</td>
<td id="769FEE75EA79D5B2FB21C64BC534F8D6" box="[1254,1394,1825,1847]" gridcol="7" gridrow="23" pageId="3" pageNumber="156">0.9</td>
</tr>
<tr id="354E8709EA79D5B2FF58C620C534F881" box="[159,1394,1866,1888]" gridrow="24" pageId="3" pageNumber="156">
<th id="769FEE75EA79D5B2FF58C620C08FF881" box="[159,201,1866,1888]" gridcol="0" gridrow="24" pageId="3" pageNumber="156">(24)</th>
<td id="769FEE75EA79D5B2FF32C620C1D9F881" box="[245,415,1866,1888]" gridcol="1" gridrow="24" pageId="3" pageNumber="156">7.1</td>
<td id="769FEE75EA79D5B2FE03C620C216F881" box="[452,592,1866,1888]" gridcol="2" gridrow="24" pageId="3" pageNumber="156">7.1</td>
<td id="769FEE75EA79D5B2FDA2C620C2B7F881" box="[613,753,1866,1888]" gridcol="3" gridrow="24" pageId="3" pageNumber="156">7.2</td>
<td id="769FEE75EA79D5B2FCC1C620C377F881" box="[774,817,1866,1888]" gridcol="4" gridrow="24" pageId="3" pageNumber="156">(50)</td>
<td id="769FEE75EA79D5B2FC9DC620C443F881" box="[858,1029,1866,1888]" gridcol="5" gridrow="24" pageId="3" pageNumber="156">0.9</td>
<td id="769FEE75EA79D5B2FBE1C620C4F4F881" box="[1062,1202,1866,1888]" gridcol="6" gridrow="24" pageId="3" pageNumber="156">1.0</td>
<td id="769FEE75EA79D5B2FB21C620C534F881" box="[1254,1394,1866,1888]" gridcol="7" gridrow="24" pageId="3" pageNumber="156">1.0</td>
</tr>
<tr id="354E8709EA79D5B2FF58C61EC534F86B" box="[159,1394,1908,1930]" gridrow="25" pageId="3" pageNumber="156">
<th id="769FEE75EA79D5B2FF58C61EC08FF86B" box="[159,201,1908,1930]" gridcol="0" gridrow="25" pageId="3" pageNumber="156">(25)</th>
<td id="769FEE75EA79D5B2FF32C61EC1D9F86B" box="[245,415,1908,1930]" gridcol="1" gridrow="25" pageId="3" pageNumber="156">6.6</td>
<td id="769FEE75EA79D5B2FE03C61EC216F86B" box="[452,592,1908,1930]" gridcol="2" gridrow="25" pageId="3" pageNumber="156">6.4</td>
<td id="769FEE75EA79D5B2FDA2C61EC2B7F86B" box="[613,753,1908,1930]" gridcol="3" gridrow="25" pageId="3" pageNumber="156">6.2</td>
<td id="769FEE75EA79D5B2FCC1C61EC377F86B" box="[774,817,1908,1930]" gridcol="4" gridrow="25" pageId="3" pageNumber="156">(51)</td>
<td id="769FEE75EA79D5B2FC9DC61EC443F86B" box="[858,1029,1908,1930]" gridcol="5" gridrow="25" pageId="3" pageNumber="156">1.0</td>
<td id="769FEE75EA79D5B2FBE1C61EC4F4F86B" box="[1062,1202,1908,1930]" gridcol="6" gridrow="25" pageId="3" pageNumber="156">1.2</td>
<td id="769FEE75EA79D5B2FB21C61EC534F86B" box="[1254,1394,1908,1930]" gridcol="7" gridrow="25" pageId="3" pageNumber="156">1.3</td>
</tr>
<tr id="354E8709EA79D5B2FF58C6F7C534F852" box="[159,1394,1949,1971]" gridrow="26" pageId="3" pageNumber="156">
<th id="769FEE75EA79D5B2FF58C6F7C08FF852" box="[159,201,1949,1971]" gridcol="0" gridrow="26" pageId="3" pageNumber="156">(26)</th>
<td id="769FEE75EA79D5B2FF32C6F7C1D9F852" box="[245,415,1949,1971]" gridcol="1" gridrow="26" pageId="3" pageNumber="156">22.2</td>
<td id="769FEE75EA79D5B2FE03C6F7C216F852" box="[452,592,1949,1971]" gridcol="2" gridrow="26" pageId="3" pageNumber="156">24.0</td>
<td id="769FEE75EA79D5B2FDA2C6F7C2B7F852" box="[613,753,1949,1971]" gridcol="3" gridrow="26" pageId="3" pageNumber="156">25.4</td>
<td id="769FEE75EA79D5B2FCC1C6F7C377F852" box="[774,817,1949,1971]" gridcol="4" gridrow="26" pageId="3" pageNumber="156">(52)</td>
<td id="769FEE75EA79D5B2FC9DC6F7C443F852" box="[858,1029,1949,1971]" gridcol="5" gridrow="26" pageId="3" pageNumber="156">0.5</td>
<td id="769FEE75EA79D5B2FBE1C6F7C4F4F852" box="[1062,1202,1949,1971]" gridcol="6" gridrow="26" pageId="3" pageNumber="156">0.4</td>
<td id="769FEE75EA79D5B2FB21C6F7C534F852" box="[1254,1394,1949,1971]" gridcol="7" gridrow="26" pageId="3" pageNumber="156">0.5</td>
</tr>
</table>
</paragraph>
<caption id="DF01D5C3EA7E2A49FF50C6C9C4A8F858" box="[151,1262,1955,1977]" httpUri="https://zenodo.org/record/572600/files/figure.png" pageId="4" pageNumber="157" targetBox="[281,1307,193,1932]" targetPageId="4">
<paragraph id="8BC1854BEA7E2A49FF50C6C9C4A8F858" blockId="4.[151,1262,1955,1977]" box="[151,1262,1955,1977]" pageId="4" pageNumber="157">
<emphasis id="B90A5959EA7E2A49FF50C6C9C154F859" bold="true" box="[151,274,1955,1977]" pageId="4" pageNumber="157">FIGURE 3.</emphasis>
Holotype of
<taxonomicName id="4C7EFEC8EA7E2A49FE5EC6C9C2D2F858" box="[409,660,1955,1977]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="157" phylum="Chordata" rank="species" species="roshanpererai" status="sp. nov.">
<emphasis id="B90A5959EA7E2A49FE5EC6C9C2D2F858" box="[409,660,1955,1977]" italics="true" pageId="4" pageNumber="157">Rhinophis roshanpererai</emphasis>
</taxonomicName>
<emphasis id="B90A5959EA7E2A49FD5CC6CEC2ADF858" bold="true" box="[667,747,1956,1977]" pageId="4" pageNumber="157">
<taxonomicNameLabel id="A239E422EA7E2A49FD5CC6CEC2ADF858" box="[667,747,1956,1977]" pageId="4" pageNumber="157" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, NMSL 2016.08.0 1 NH, scale bars in centimeters.
</paragraph>
</caption>
<caption id="DF01D5C3EA7F2A48FF50C6D5C1F4F815" httpUri="https://zenodo.org/record/572601/files/figure.png" pageId="5" pageNumber="158" targetBox="[276,1302,193,1932]" targetPageId="5">
<paragraph id="8BC1854BEA7F2A48FF50C6D5C1F4F815" blockId="5.[151,1436,1983,2036]" pageId="5" pageNumber="158">
<emphasis id="B90A5959EA7F2A48FF50C6D5C153F835" bold="true" box="[151,277,1983,2005]" pageId="5" pageNumber="158">FIGURE 4.</emphasis>
Head and tail ends of the two paratypes of
<taxonomicName id="4C7EFEC8EA7F2A48FD23C6D5C3A4F834" box="[740,994,1983,2005]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="158" phylum="Chordata" rank="species" species="roshanpererai" status="sp. nov.">
<emphasis id="B90A5959EA7F2A48FD23C6D5C3A4F834" box="[740,994,1983,2005]" italics="true" pageId="5" pageNumber="158">Rhinophis roshanpererai</emphasis>
</taxonomicName>
<emphasis id="B90A5959EA7F2A48FC2CC6AAC47BF834" bold="true" box="[1003,1085,1984,2005]" pageId="5" pageNumber="158">
<taxonomicNameLabel id="A239E422EA7F2A48FC2CC6AAC47BF834" box="[1003,1085,1984,2005]" pageId="5" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) DWC 2016.05.0 3, (B) DWC 2016.05.0 4. Scale bar 2 cm.
</paragraph>
</caption>
<caption id="DF01D5C3EA7C2A4BFF50C6CBC243F815" httpUri="https://zenodo.org/record/572602/files/figure.png" pageId="6" pageNumber="159" targetBox="[214,1373,193,1932]" targetPageId="6">
<paragraph id="8BC1854BEA7C2A4BFF50C6CBC243F815" blockId="6.[151,1436,1953,2036]" pageId="6" pageNumber="159">
<emphasis id="B90A5959EA7C2A4BFF50C6CBC152F856" bold="true" box="[151,276,1953,1975]" pageId="6" pageNumber="159">FIGURE 5.</emphasis>
<taxonomicName id="4C7EFEC8EA7C2A4BFEDBC6CBC25CF856" box="[284,538,1953,1975]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="159" phylum="Chordata" rank="species" species="roshanpererai" status="sp. nov.">
<emphasis id="B90A5959EA7C2A4BFEDBC6CBC25CF856" box="[284,538,1953,1975]" italics="true" pageId="6" pageNumber="159">Rhinophis roshanpererai</emphasis>
</taxonomicName>
<emphasis id="B90A5959EA7C2A4BFDE5C6C8C233F856" bold="true" box="[546,629,1954,1975]" pageId="6" pageNumber="159">
<taxonomicNameLabel id="A239E422EA7C2A4BFDE5C6C8C233F856" box="[546,629,1954,1975]" pageId="6" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype DWC 2016.05.0 4, posterodorsal view of tail in preserved specimen showing three (two lower, one upper) rather than four major spines on the tail shield (bases of these major spines are surrounded by pale areas on shield).
</paragraph>
</caption>
<paragraph id="8BC1854BEA7D2A4AFF00C1F2C33AFDDC" blockId="7.[151,1437,151,1725]" pageId="7" pageNumber="160">
Among
<collectingRegion id="49BA4BA9EA7D2A4AFEE4C1FDC102FF51" box="[291,324,151,176]" country="United Kingdom" name="Surrey" pageId="7" pageNumber="160">Sri</collectingRegion>
Lankan congeners,
<taxonomicName id="4C7EFEC8EA7D2A4AFDEEC1FDC307FF51" box="[553,833,151,176]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="160" phylum="Chordata" rank="species" species="roshanpererai" status="sp. nov.">
<emphasis id="B90A5959EA7D2A4AFDEEC1FDC307FF51" box="[553,833,151,176]" italics="true" pageId="7" pageNumber="160">Rhinophis roshanpererai</emphasis>
</taxonomicName>
<emphasis id="B90A5959EA7D2A4AFC8DC1FDC3E3FF51" bold="true" box="[842,933,151,176]" pageId="7" pageNumber="160">
<taxonomicNameLabel id="A239E422EA7D2A4AFC8DC1FDC3E3FF51" box="[842,933,151,176]" pageId="7" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C7EFEC8EA7D2A4AFBFBC1F3C5DAFF51" authority="Kelaart, 1853" authorityName="Kelaart" authorityYear="1853" box="[1084,1436,151,176]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="160" phylum="Chordata" rank="species" species="saffragamus">
<emphasis id="B90A5959EA7D2A4AFBFBC1F3C4A0FF51" box="[1084,1254,152,176]" italics="true" pageId="7" pageNumber="160">R. saffragamus</emphasis>
(
<bibRefCitation id="EFEFF8BAEA7D2A4AFB30C1FDC5D2FF51" author="Kelaart" box="[1271,1428,151,176]" pageId="7" pageNumber="160" refString="Kelaart, E. F. (1853) Description of new or little-known species of reptiles collected in Ceylon. Annals and Magazine of Natural History, Series 2, 13, pp. 25 - 30." type="journal article" year="1853">Kelaart, 1853</bibRefCitation>
)
</taxonomicName>
in not having a large and flat tail shield or midline contact between the opposite nasal shields, and by having dorsal scales in 17 rather than 19 rows at midbody. The new species differs from
<taxonomicName id="4C7EFEC8EA7D2A4AFC23C18BC5D3FF19" authority="Deraniyagala, 1941" authorityName="Deraniyagala" authorityYear="1941" box="[996,1429,223,248]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="160" phylum="Chordata" rank="species" species="dorsimaculatus">
<emphasis id="B90A5959EA7D2A4AFC23C18BC4F7FF19" box="[996,1201,223,248]" italics="true" pageId="7" pageNumber="160">R. dorsimaculatus</emphasis>
Deraniyagala, 1941
</taxonomicName>
,
<taxonomicName id="4C7EFEC8EA7D2A4AFF50C06FC248FEFD" authority="Hemprich, 1820" authorityName="Hemprich" authorityYear="1820" box="[151,526,260,285]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="160" phylum="Chordata" rank="species" species="homolepis">
<emphasis id="B90A5959EA7D2A4AFF50C06FC176FEFC" box="[151,304,260,285]" italics="true" pageId="7" pageNumber="160">R. homolepis</emphasis>
(Hemprich, 1820)
</taxonomicName>
,
<emphasis id="B90A5959EA7D2A4AFDE4C06FC31AFEFC" box="[547,860,260,285]" italics="true" pageId="7" pageNumber="160">
<taxonomicName id="4C7EFEC8EA7D2A4AFDE4C06FC2E2FEFC" box="[547,676,260,285]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="160" phylum="Chordata" rank="species" species="lineatus">R. lineatus</taxonomicName>
, R. oxyrynchus
</emphasis>
(Schneider, 1801),
<taxonomicName id="4C7EFEC8EA7D2A4AFB88C06FC529FEFD" authority="Wall, 1921" authorityName="Wall" authorityYear="1921" box="[1103,1391,260,285]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="160" phylum="Chordata" rank="species" species="porrectus">
<emphasis id="B90A5959EA7D2A4AFB88C06FC4A6FEFD" box="[1103,1248,261,284]" italics="true" pageId="7" pageNumber="160">R. porrectus</emphasis>
<bibRefCitation id="EFEFF8BAEA7D2A4AFB2AC06EC529FEFD" author="Wall" box="[1261,1391,260,285]" pageId="7" pageNumber="160" refString="Wall, F. (1921) Ophidia Taprobanica or the snakes of Ceylon. Governmental Press, Colombo, xxii + 581 pp." type="book" year="1921">Wall, 1921</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C7EFEC8EA7D2A4AFA43C06FC1E5FEA1" authority="Muller, 1832" authorityName="Muller" authorityYear="1832" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="160" phylum="Chordata" rank="species" species="punctatus">
<emphasis id="B90A5959EA7D2A4AFA43C06FC142FEA1" italics="true" pageId="7" pageNumber="160">R. punctatus</emphasis>
<bibRefCitation id="EFEFF8BAEA7D2A4AFEC9C04DC1E5FEA1" author="Muller" box="[270,419,295,320]" pageId="7" pageNumber="160" refString="Muller, J. (1832) Beitrage zur Anatomie und Naturgeschichte der Amphibia. Zeitschrift fur Physiologie, 4, 190 - 275." type="journal article" year="1832">Müller, 1832</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C7EFEC8EA7D2A4AFE18C043C20CFEA1" box="[479,586,297,320]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="160" phylum="Chordata" rank="species" species="zigzag">
<emphasis id="B90A5959EA7D2A4AFE18C043C20CFEA1" box="[479,586,297,320]" italics="true" pageId="7" pageNumber="160">R. zigzag</emphasis>
</taxonomicName>
by having fewer than 175 ventral scales (versus more than 180), and by having a very small tail shield with spines (versus relatively much larger tail shield without spines).
<taxonomicName id="4C7EFEC8EA7D2A4AFAEDC026C173FE69" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="160" phylum="Chordata" rank="species" species="roshanpererai" status="sp. nov.">
<emphasis id="B90A5959EA7D2A4AFAEDC026C173FE69" italics="true" pageId="7" pageNumber="160">Rhinophis roshanpererai</emphasis>
</taxonomicName>
<emphasis id="B90A5959EA7D2A4AFEFAC005C1DEFE69" bold="true" box="[317,408,367,392]" pageId="7" pageNumber="160">
<taxonomicNameLabel id="A239E422EA7D2A4AFEFAC005C1DEFE69" box="[317,408,367,392]" pageId="7" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C7EFEC8EA7D2A4AFDE8C01BC32BFE69" authority="Nicholls, 1929" authorityName="Nicholls" authorityYear="1929" box="[559,877,367,392]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="160" phylum="Chordata" rank="species" species="phillipsi">
<emphasis id="B90A5959EA7D2A4AFDE8C01BC2EAFE69" box="[559,684,367,392]" italics="true" pageId="7" pageNumber="160">R. phillipsi</emphasis>
(
<bibRefCitation id="EFEFF8BAEA7D2A4AFD7BC005C322FE69" author="Nicholls" box="[700,868,367,392]" pageId="7" pageNumber="160" refString="Nicholls, L. (1929) A new species of earth snake of the genus Silybura (Silybura phillipsi). Ceylon Journal of Science, 15, 153 - 155." type="journal article" year="1929">Nicholls, 1929</bibRefCitation>
)
</taxonomicName>
in having fewer than 190 ventrals and in lacking yellow lines on the dorsum.
<taxonomicName id="4C7EFEC8EA7D2A4AFE26C0FEC2BCFE4C" box="[481,762,404,429]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="160" phylum="Chordata" rank="species" species="roshanpererai" status="sp. nov.">
<emphasis id="B90A5959EA7D2A4AFE26C0FEC2BCFE4C" box="[481,762,404,429]" italics="true" pageId="7" pageNumber="160">Rhinophis roshanpererai</emphasis>
</taxonomicName>
<emphasis id="B90A5959EA7D2A4AFCC3C0FEC326FE4C" bold="true" box="[772,864,404,429]" pageId="7" pageNumber="160">
<taxonomicNameLabel id="A239E422EA7D2A4AFCC3C0FEC326FE4C" box="[772,864,404,429]" pageId="7" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles
<taxonomicName id="4C7EFEC8EA7D2A4AFC24C0FFC4DBFE4C" box="[995,1181,404,429]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="160" phylum="Chordata" rank="species" species="melanogaster">
<emphasis id="B90A5959EA7D2A4AFC24C0FFC4DBFE4C" box="[995,1181,404,429]" italics="true" pageId="7" pageNumber="160">R. melanogaster</emphasis>
</taxonomicName>
in having a small tail shield, but the new species has a shield surface with four (or three) notably prominent spines, one pair above the other (versus two slightly larger spines ventrally); absence of yellowish lines laterally (versus present); perhaps more ventral scales (168169 versus 152166); and a distinct geographical distribution (central highlands of Badulla District vs Knuckles Range, Matale and Kandy Districts).
</paragraph>
<paragraph id="8BC1854BEA7D2A4AFF00C32DC33AFCF4" blockId="7.[151,1437,151,1725]" pageId="7" pageNumber="160">
The ventral scale count in
<taxonomicName id="4C7EFEC8EA7D2A4AFDC4C323C282FD81" box="[515,708,583,608]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="160" phylum="Chordata" rank="species" species="roshanpererai" status="sp. nov.">
<emphasis id="B90A5959EA7D2A4AFDC4C323C252FD81" box="[515,532,585,608]" italics="true" pageId="7" pageNumber="160">R</emphasis>
.
<emphasis id="B90A5959EA7D2A4AFDE1C32DC282FD81" box="[550,708,583,608]" italics="true" pageId="7" pageNumber="160">roshanpererai</emphasis>
</taxonomicName>
<emphasis id="B90A5959EA7D2A4AFD08C32DC36BFD81" bold="true" box="[719,813,583,608]" pageId="7" pageNumber="160">
<taxonomicNameLabel id="A239E422EA7D2A4AFD08C32DC36BFD81" box="[719,813,583,608]" pageId="7" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to or overlapping with those for
<taxonomicName id="4C7EFEC8EA7D2A4AFAF6C323C16AFD64" authority="Kelaart 1853" authorityName="Kelaart" authorityYear="1853" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="160" phylum="Chordata" rank="species" species="blythii">
<emphasis id="B90A5959EA7D2A4AFAF6C323C5DAFD81" box="[1329,1436,583,608]" italics="true" pageId="7" pageNumber="160">R. blythii</emphasis>
<bibRefCitation id="EFEFF8BAEA7D2A4AFF50C306C16AFD64" author="Kelaart" box="[151,300,620,645]" pageId="7" pageNumber="160" refString="Kelaart, E. F. (1853) Description of new or little-known species of reptiles collected in Ceylon. Annals and Magazine of Natural History, Series 2, 13, pp. 25 - 30." type="journal article" year="1853">Kelaart 1853</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C7EFEC8EA7D2A4AFEF0C307C2C3FD65" authority="Wall, 1921" authorityName="Wall" authorityYear="1921" box="[311,645,620,645]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="160" phylum="Chordata" rank="species" species="drummondhayi">
<emphasis id="B90A5959EA7D2A4AFEF0C307C247FD64" box="[311,513,620,645]" italics="true" pageId="7" pageNumber="160">R. drummondhayi</emphasis>
<bibRefCitation id="EFEFF8BAEA7D2A4AFDCFC306C2C3FD65" author="Wall" box="[520,645,620,645]" pageId="7" pageNumber="160" refString="Wall, F. (1921) Ophidia Taprobanica or the snakes of Ceylon. Governmental Press, Colombo, xxii + 581 pp." type="book" year="1921">Wall, 1921</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C7EFEC8EA7D2A4AFD56C307C39BFD65" authority="Cuvier, 1829" authorityName="Cuvier" authorityYear="1829" box="[657,989,620,645]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="160" phylum="Chordata" rank="species" species="philippinus">
<emphasis id="B90A5959EA7D2A4AFD56C307C369FD64" box="[657,815,620,645]" italics="true" pageId="7" pageNumber="160">R. philippinus</emphasis>
(Cuvier, 1829)
</taxonomicName>
, and
<taxonomicName id="4C7EFEC8EA7D2A4AFBDEC307C5DEFD64" authority="Deraniyagala, 1975" authorityName="Deraniyagala" authorityYear="1975" box="[1049,1432,620,645]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="160" phylum="Chordata" rank="species" species="tricolorata">
<emphasis id="B90A5959EA7D2A4AFBDEC307C4F7FD64" box="[1049,1201,620,645]" italics="true" pageId="7" pageNumber="160">R. tricolorata</emphasis>
<bibRefCitation id="EFEFF8BAEA7D2A4AFB7EC306C5DEFD64" author="Deraniyagala" box="[1209,1432,620,645]" pageId="7" pageNumber="160" refString="Deraniyagala, P. E P. (1975) A new fossorial snake of the genus Rhinophis. Spolia Zeylanica, 33, 535 - 536." type="journal article" year="1975">Deraniyagala, 1975</bibRefCitation>
</taxonomicName>
, but the new species differs from these four
<collectingRegion id="49BA4BA9EA7D2A4AFDB8C3E5C2E6FD49" box="[639,672,655,680]" country="United Kingdom" name="Surrey" pageId="7" pageNumber="160">Sri</collectingRegion>
Lankan congeners by having a smaller tail shield with spines, three or four of which are prominent (versus large tail shield without notable spines). The new species differs from
<taxonomicName id="4C7EFEC8EA7D2A4AFA43C3DFC165FD11" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="160" phylum="Chordata" rank="species" species="erangaviraji">
<emphasis id="B90A5959EA7D2A4AFA43C3DFC165FD11" italics="true" pageId="7" pageNumber="160">R. erangaviraji</emphasis>
</taxonomicName>
by having more than 165 ventrals (versus fewer than 155), a smaller tail shield, and by lacking substantial yellow areas on the lateral surface of the body and tail.
</paragraph>
<paragraph id="8BC1854BEA7D2A4AFF00C275C48BF9E9" blockId="7.[151,1437,151,1725]" pageId="7" pageNumber="160">
<emphasis id="B90A5959EA7D2A4AFF00C275C1AAFCD9" bold="true" box="[199,492,799,824]" pageId="7" pageNumber="160">
Description of
<typeStatus id="54C53BE9EA7D2A4AFEB9C275C1A1FCD9" box="[382,487,799,824]" pageId="7" pageNumber="160" type="holotype">holotype</typeStatus>
.
</emphasis>
See
<tableCitation id="C6FCB0F0EA7D2A4AFDEDC275C2C6FCD9" box="[554,640,799,824]" captionStart="TABLE 1" captionStartId="3.[151,239,152,175]" captionText="TABLE 1. Meristic and morphometric data for 52 characters of holotype (*) and paratypes (**) of Rhinophis roshanpererai sp. nov. Specimen numbers have NMSL prefix for holotype and DWC prefix for paratypes. Bilateral scale counts given in left, right order. Character numbers in parentheses, as follows: (1) supralabials, (2) infralabials, (3) diameter of eye, (4) maximum length of ocular, (5) length ocular / eye, (6) distance between eyes, (7) distance between eye-naris, (8) length of frontal, (9) width of frontal, (10) head length = distance between snout tip-posterior edge 4 th supralabial, (11) total length / HL, (12) head width level with corner of mouth, (13) distance between nares, (14) maximum longitudinal length of prefrontal scales, (15) length of parietal scale = distance between posterior tip of parietal and posterior end of suture between frontal-ocular, (16) width of parietal scale = distance between posterior tip of ocular and posterior tip of frontal, (17) length of midline suture between parietals, (18) distance rostral-frontal, (19) distance between tip of snout and posterior edge of rostral, (20) distance between tip of snout and posterior limit of midline suture between parietals, (21) maximum width of rostral, viewed ventrally, (22) distance between snout tip-naris, (23) distance between snout tip-eye, (24) midbody width, (25) midbody depth, (26) midbody circumference, (27) exposed width of first scale row at midbody, (28) exposed width of ventral scales at midbody, (29) number of ventral scale level with which dorsal scale row is lost (i. e. location of change between 19 and 17 dorsal scale rows), (30) dorsal scale rows behind head, at midbody and one head length anterior to vent (excluding ventrals), (31) exposed length of first scale row at midbody, (32) width of ventral / first dorsal scale rows, (33) ventral scales, (34) vertebral scales, (35) snout vent length, (36) width at vent, (37) disposition of two anal scales (l / r) or (r / l), (38) number of small scales overlapped by anals, excluding first subcaudals, (39) subcaudal scales, (40) tail length, (41) total length, (42) tail length as % of total length, (43) total length / midbody width, (44) maximum width of tail shield, (45) maximum diameter of tail shield, (46) number of scales (including subcaudals) surrounding base of tail shield, (47) dorsal base to tip of tail shield (upper spines), (48) dorsal base to tip of tail shield (lower spines), (49) lateral base to tip of tail shield (upper), (50) lateral base to tip of tail shield (lower), (51) ventral base to tip of tail shield (upper), (52) ventral base to tip of tail shield (lower)." pageId="7" pageNumber="160">Table 1</tableCitation>
for morphometric and meristic data. A preserved specimen in good condition;
<quantity id="4C8628AEEA7D2A4AFED7C22EC119FCBD" box="[272,351,836,860]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="160" unit="mm" value="20.0">20 mm</quantity>
long left of ventral incision into coelom extending anteriorly from
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anterior to vent; outer layer of scales loose and missing in parts; a few flank scales more profoundly damaged on left at approximately midbody. Head small, snout pointed (
<figureCitation id="134599CEEA7D2A4AFDF8C2E6C2EAFC45" box="[575,684,908,933]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="2.[151,248,1319,1341]" captionStartId-1="4.[151,250,1955,1977]" captionTargetBox-0="[151,1436,703,1307]" captionTargetBox-1="[281,1307,193,1932]" captionTargetId-0="figure@2.[151,1436,703,1307]" captionTargetId-1="figure@4.[281,1307,193,1932]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURE 2. Dorsolateral view in life of the holotype of Rhinophis roshanpererai sp. nov., NMSL 2016.08. 0 1 NH, 214 mm (total length)." captionText-1="FIGURE 3. Holotype of Rhinophis roshanpererai sp. nov., NMSL 2016.08. 0 1 NH, scale bars in centimeters." httpUri-0="https://zenodo.org/record/572599/files/figure.png" httpUri-1="https://zenodo.org/record/572600/files/figure.png" pageId="7" pageNumber="160">Figs. 23</figureCitation>
). Rostral pointed, longer than wide, without dorsal crest; widest at level of anterior superior corner of first supralabials. Rostral several times longer (in dorsal view) than rostralfrontal gap (
<figureCitation id="134599CEEA7D2A4AFEE3C2BEC12DFC0D" box="[292,363,980,1005]" captionStart="FIGURE 3" captionStartId="4.[151,250,1955,1977]" captionTargetBox="[281,1307,193,1932]" captionTargetId="figure@4.[281,1307,193,1932]" captionTargetPageId="4" captionText="FIGURE 3. Holotype of Rhinophis roshanpererai sp. nov., NMSL 2016.08. 0 1 NH, scale bars in centimeters." httpUri="https://zenodo.org/record/572600/files/figure.png" pageId="7" pageNumber="160">Fig. 3</figureCitation>
). Frontal irregularly hexagonal, longer than wide, lateral (ocular) margins slightly converging posteriorly, posterolateral margins straight to very slightly concave; lateral (ocular) margin shortest, posterolateral edges longest. Frontal longer, wider than rostral. A pair of nasals, separated from each other by posterior half of rostral. External naris small, subcircular, slightly countersunk within small depression, located in anteroventral corner of nasal. Nasal in contact with first and second supralabials. Prefrontals (for most of their length) in contact with each other along midline (left overlapping right), separating frontal from rostral. Prefrontals wider than long, shorter than frontal. Supralabials four, first smallest, making the least contribution to margin of mouth; fourth much the largest. Ocular in contact with third and fourth supralabials. Eye distinct, diameter approximately one third length of ocular, located near anteroventral corner of ocular, bulging slightly from ocular surface, pupil circular. Paired parietals longer than wide, shorter, very slightly wider than frontal, posteriorly broadly rounded, angle between postermedial and posterolateral edges approximately 90°. Opposite parietals in brief midline contact, left overlapping right. Each parietal contacts four scales other than head shields. No mental groove; mental wider than long, smaller than infralabials, contacting first infralabials and single postmental (= first ventral); three pairs of infralabials, second largest, first smallest. First and second ventrals longer than wide, third approximately as long as wide, fourth and subsequent ventrals wider than long. Six or seven maxillary and approximately seven mandibular teeth on each side. Teeth simple, pointed, distinctly retrorse, straight, evenly spaced.
</paragraph>
<paragraph id="8BC1854BEA7D2A4AFF00C77EC12FF95C" blockId="7.[151,1437,151,1725]" pageId="7" pageNumber="160">Body cylindrical. Body scales generally evenly sized on dorsum and along body except for those involved in dorsal scale row reductions. Midline ventral scales between mental and anal of even size though anteriormost ones gradually narrow. Ventrals 168, posteriormost ventral notably smaller, penultimate ventral paired. Dorsal scale rows 19 anteriorly, reducing to 17 by level with 30th ventral and maintained along most of body; scale row reduction formula:</paragraph>
<paragraph id="8BC1854BEA7D2A4AFD25C786C317F8E5" blockId="7.[692,902,1772,1868]" box="[738,849,1772,1796]" pageId="7" pageNumber="160">3 + 4 (30)</paragraph>
<paragraph id="8BC1854BEA7D2A4AFD73C67AC3C0F8C9" blockId="7.[692,902,1772,1868]" box="[692,902,1808,1832]" pageId="7" pageNumber="160">19 --------------- 17</paragraph>
<paragraph id="8BC1854BEA7D2A4AFD25C65EC317F8AD" blockId="7.[692,902,1772,1868]" box="[738,849,1844,1868]" pageId="7" pageNumber="160">3 + 4 (30)</paragraph>
<paragraph id="8BC1854BEA7D2A45FF00C616C162FE4D" blockId="7.[151,1436,1916,2013]" lastBlockId="8.[151,1437,151,1941]" lastPageId="8" lastPageNumber="161" pageId="7" pageNumber="160">
Dorsal scale rows approximately 14 at base of tail. Head and body scales macroscopically smooth, lacking keels. Inconspicuous keels on scales on posteriormost portion of body and on tail, increasingly prominent posteriorly, more obvious ventrally (including on anals) and ventrolaterally. Paired anal scales (right overlying left) considerably larger than posteriormost ventrals and subcaudals. Distal margin of each anal overlaps three other scales in addition to anteriormost subcaudals. Seven right and seven left subcaudals. Tail 'shield' mildly conical, forming tip of tail, small, longer than wide in dorsal view, shorter than the frontal in dorsal view, visible from below and especially above, base (much narrower than base of tail) surrounded by last pair of subcaudals and 6 other scales. In posterior view shield oval to slightly egg-shaped, wider ventrally than dorsally (
<figureCitation id="134599CEEA722A45FB33C04DC57AFEA1" box="[1268,1340,295,320]" captionStart="FIGURE 3" captionStartId="4.[151,250,1955,1977]" captionTargetBox="[281,1307,193,1932]" captionTargetId="figure@4.[281,1307,193,1932]" captionTargetPageId="4" captionText="FIGURE 3. Holotype of Rhinophis roshanpererai sp. nov., NMSL 2016.08. 0 1 NH, scale bars in centimeters." httpUri="https://zenodo.org/record/572600/files/figure.png" pageId="8" pageNumber="161">Fig. 3</figureCitation>
). Shield surface sparsely spinose, most spines small, inconspicuous but four (arranged in two pairs, one above the other) much longer and substantial, pointing straight backwards; ventral pair of larger spines notably longer than dorsal pair (
<figureCitation id="134599CEEA722A45FF15C0FEC150FE4D" box="[210,278,404,429]" captionStart="FIGURE 3" captionStartId="4.[151,250,1955,1977]" captionTargetBox="[281,1307,193,1932]" captionTargetId="figure@4.[281,1307,193,1932]" captionTargetPageId="4" captionText="FIGURE 3. Holotype of Rhinophis roshanpererai sp. nov., NMSL 2016.08. 0 1 NH, scale bars in centimeters." httpUri="https://zenodo.org/record/572600/files/figure.png" pageId="8" pageNumber="161">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8BC1854BEA722A45FF00C0DDC285FD81" blockId="8.[151,1437,151,1941]" pageId="8" pageNumber="161">
<emphasis id="B90A5959EA722A45FF00C0DDC129FE31" bold="true" box="[199,367,439,464]" pageId="8" pageNumber="161">Colour in life.</emphasis>
Dorsum and lateral background uniform black, with sparse, very small yellow flecks (
<figureCitation id="134599CEEA722A45FA8FC0DDC5C9FE31" box="[1352,1423,439,464]" captionStart="FIGURE 2" captionStartId="2.[151,248,1319,1341]" captionTargetBox="[151,1436,703,1307]" captionTargetId="figure@2.[151,1436,703,1307]" captionTargetPageId="2" captionText="FIGURE 2. Dorsolateral view in life of the holotype of Rhinophis roshanpererai sp. nov., NMSL 2016.08. 0 1 NH, 214 mm (total length)." httpUri="https://zenodo.org/record/572599/files/figure.png" pageId="8" pageNumber="161">Fig. 2</figureCitation>
). Ventral background dark brown for most of length, gradually paler anteriorly, darker posteriorly (similar to dark colour of dorsum). Anterior and underside of snout paler than rest of head. Venter with conspicuous yellow blotching, blotches notably larger than on dorsum and lateral surfaces of body; ventral blotching absent on tail, head and anteriormost and posteriormost of body.
</paragraph>
<paragraph id="8BC1854BEA722A45FF00C306C1B8FD49" blockId="8.[151,1437,151,1941]" pageId="8" pageNumber="161">
<emphasis id="B90A5959EA722A45FF00C306C1E6FD64" bold="true" box="[199,416,620,645]" pageId="8" pageNumber="161">Colour in alcohol.</emphasis>
Colour pattern remains with a little fading, black to dark brown, yellow to off white and brown to a paler brown (
<figureCitation id="134599CEEA722A45FE6CC3E5C1B6FD49" box="[427,496,655,680]" captionStart="FIGURE 3" captionStartId="4.[151,250,1955,1977]" captionTargetBox="[281,1307,193,1932]" captionTargetId="figure@4.[281,1307,193,1932]" captionTargetPageId="4" captionText="FIGURE 3. Holotype of Rhinophis roshanpererai sp. nov., NMSL 2016.08. 0 1 NH, scale bars in centimeters." httpUri="https://zenodo.org/record/572600/files/figure.png" pageId="8" pageNumber="161">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8BC1854BEA722A45FF00C3DEC409FBEE" blockId="8.[151,1437,151,1941]" pageId="8" pageNumber="161">
<emphasis id="B90A5959EA722A45FF00C3DEC1B5FD2C" bold="true" box="[199,499,692,717]" pageId="8" pageNumber="161">Paratypes and variation.</emphasis>
Paratype DWC 2016.05.0 3 is slightly longer (
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SVL) than the holotype and the other paratype (DWC 2016.05.04) slightly shorter (
<quantity id="4C8628AEEA722A45FCFBC3BDC3F4FD0E" box="[828,946,727,752]" metricMagnitude="-1" metricUnit="m" metricValue="2.052" pageId="8" pageNumber="161" unit="mm" value="205.2">205.2 mm</quantity>
SVL), both are female. The two paratypes are very similar to the holotype with respect to the description presented above, including identical scale row reductions (19 to 17 rows by level of 30th ventral). DWC 2016.05.0 3 differs from the holotype in having: parietals more notably wider than frontal (
<figureCitation id="134599CEEA722A45FDCEC22EC20AFCBD" box="[521,588,836,861]" captionStart="FIGURE 4" captionStartId="5.[151,250,1983,2005]" captionTargetBox="[276,1302,193,1932]" captionTargetId="figure@5.[276,1302,193,1932]" captionTargetPageId="5" captionText="FIGURE 4. Head and tail ends of the two paratypes of Rhinophis roshanpererai sp. nov. (A) DWC 2016.05. 0 3, (B) DWC 2016.05. 0 4. Scale bar 2 cm." httpUri="https://zenodo.org/record/572601/files/figure.png" pageId="8" pageNumber="161">Fig. 4</figureCitation>
A); seven rather than six scales plus last pair of subcaudals surround base of tail shield; posteriormost ventral paired; supernumerary scale between second pair of subcaudals (
<figureCitation id="134599CEEA722A45FB2CC20DC569FC61" box="[1259,1327,871,896]" captionStart="FIGURE 4" captionStartId="5.[151,250,1983,2005]" captionTargetBox="[276,1302,193,1932]" captionTargetId="figure@5.[276,1302,193,1932]" captionTargetPageId="5" captionText="FIGURE 4. Head and tail ends of the two paratypes of Rhinophis roshanpererai sp. nov. (A) DWC 2016.05. 0 3, (B) DWC 2016.05. 0 4. Scale bar 2 cm." httpUri="https://zenodo.org/record/572601/files/figure.png" pageId="8" pageNumber="161">Fig. 4</figureCitation>
A). DWC 2016.05.0 4 differs from the holotype in having six rather than seven subcaudals on right side, and in having three rather than four major spines on the tail shield, two posteroventrally and one posterodorsally (
<figureCitation id="134599CEEA722A45FAC6C2C5C50DFC29" box="[1281,1355,943,968]" captionStart="FIGURE 5" captionStartId="6.[151,250,1953,1975]" captionTargetBox="[214,1373,193,1932]" captionTargetId="figure@6.[214,1373,193,1932]" captionTargetPageId="6" captionText="FIGURE 5. Rhinophis roshanpererai sp. nov., paratype DWC 2016.05. 0 4, posterodorsal view of tail in preserved specimen showing three (two lower, one upper) rather than four major spines on the tail shield (bases of these major spines are surrounded by pale areas on shield)." httpUri="https://zenodo.org/record/572602/files/figure.png" pageId="8" pageNumber="161">Fig. 5</figureCitation>
). Both paratypes appear to have seven maxillary teeth on each side; mandibular counts are more difficult but are estimated at six or seven on each ramus. Paratypes closely resemble holotype in colour pattern.
</paragraph>
</subSubSection>
<subSubSection id="C364D6C0EA722A45FF00C576C41CFB09" pageId="8" pageNumber="161" type="etymology">
<paragraph id="8BC1854BEA722A45FF00C576C1F3FB41" blockId="8.[151,1437,151,1941]" pageId="8" pageNumber="161">
<emphasis id="B90A5959EA722A45FF00C576C108FBD4" bold="true" box="[199,334,1052,1077]" pageId="8" pageNumber="161">Etymology.</emphasis>
The species epithet
<taxonomicName id="4C7EFEC8EA722A45FDF1C576C295FBD4" box="[566,723,1052,1077]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="161" phylum="Chordata" rank="species" species="roshanpererai">
<emphasis id="B90A5959EA722A45FDF1C576C295FBD4" box="[566,723,1052,1077]" italics="true" pageId="8" pageNumber="161">roshanpererai</emphasis>
</taxonomicName>
is named for the late Roshan Perera, who was an Instructor of the Reptiles group of the Young Zoologists Association of
<collectingCountry id="F369C5DBEA722A45FCEAC555C3D8FBB9" box="[813,926,1087,1112]" name="Sri Lanka" pageId="8" pageNumber="161">
<collectingRegion id="49BA4BA9EA722A45FCEAC555C308FBB9" box="[813,846,1087,1112]" country="United Kingdom" name="Surrey" pageId="8" pageNumber="161">Sri</collectingRegion>
Lanka
</collectingCountry>
, Department of National Zoological Gardens, in recognition of his dedicated services to wildlife conservation in
<collectingCountry id="F369C5DBEA722A45FC42C50EC3B1FB9C" box="[901,1015,1124,1149]" name="Sri Lanka" pageId="8" pageNumber="161">
<collectingRegion id="49BA4BA9EA722A45FC42C50EC3E0FB9C" box="[901,934,1124,1149]" country="United Kingdom" name="Surrey" pageId="8" pageNumber="161">Sri</collectingRegion>
Lanka
</collectingCountry>
. The species name
<taxonomicName id="4C7EFEC8EA722A45FB17C50EC528FB9C" box="[1232,1390,1124,1149]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="161" phylum="Chordata" rank="species" species="roshanpererai">
<emphasis id="B90A5959EA722A45FB17C50EC528FB9C" box="[1232,1390,1124,1149]" italics="true" pageId="8" pageNumber="161">roshanpererai</emphasis>
</taxonomicName>
is a noun in the genitive case.
</paragraph>
<paragraph id="8BC1854BEA722A45FF00C5C6C41CFB09" blockId="8.[151,1437,151,1941]" pageId="8" pageNumber="161">
<emphasis id="B90A5959EA722A45FF00C5C6C262FB24" bold="true" box="[199,548,1196,1221]" pageId="8" pageNumber="161">Suggested vernacular names.</emphasis>
Roshan Pererage thudulla, Roshan Pereravin nilakael pambu, Roshan Pereras sheildtail (or Roshan Pereras
<taxonomicName id="4C7EFEC8EA722A45FE2FC5A5C21BFB09" box="[488,605,1231,1256]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="161" phylum="Chordata" rank="genus">Rhinophis</taxonomicName>
) in Sinhala, Tamil, and English, respectively.
</paragraph>
</subSubSection>
<subSubSection id="C364D6C0EA722A45FF00C59EC0AFF874" pageId="8" pageNumber="161" type="distribution">
<paragraph id="8BC1854BEA722A45FF00C59EC53CF8C9" blockId="8.[151,1437,151,1941]" pageId="8" pageNumber="161">
<emphasis id="B90A5959EA722A45FF00C59EC219FAEC" bold="true" box="[199,607,1268,1293]" pageId="8" pageNumber="161">Distribution, habitat and threats.</emphasis>
The first author first encountered the new species as a single roadkill specimen at the
<typeStatus id="54C53BE9EA722A45FE9BC473C1CAFAD0" box="[348,396,1305,1329]" pageId="8" pageNumber="161">type</typeStatus>
locality in 1999. In five or six subsequent visits to the
<typeStatus id="54C53BE9EA722A45FBF2C473C423FAD0" box="[1077,1125,1305,1329]" pageId="8" pageNumber="161">type</typeStatus>
locality approximately 30 individuals of
<taxonomicName id="4C7EFEC8EA722A45FEFAC456C210FAB4" box="[317,598,1340,1365]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="161" phylum="Chordata" rank="species" species="roshanpererai" status="sp. nov.">
<emphasis id="B90A5959EA722A45FEFAC456C210FAB4" box="[317,598,1340,1365]" italics="true" pageId="8" pageNumber="161">Rhinophis roshanpererai</emphasis>
</taxonomicName>
<emphasis id="B90A5959EA722A45FD98C456C2FDFAB4" bold="true" box="[607,699,1340,1365]" pageId="8" pageNumber="161">
<taxonomicNameLabel id="A239E422EA722A45FD98C456C2FDFAB4" box="[607,699,1340,1365]" pageId="8" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
have been observed, including a second roadkill specimen. The
<typeStatus id="54C53BE9EA722A45FF50C40BC081FA98" box="[151,199,1377,1401]" pageId="8" pageNumber="161">type</typeStatus>
series of
<taxonomicName id="4C7EFEC8EA722A45FEF1C40BC1B3FA99" box="[310,501,1375,1400]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="161" phylum="Chordata" rank="species" species="roshanpererai" status="sp. nov.">
<emphasis id="B90A5959EA722A45FEF1C40BC1B3FA99" box="[310,501,1375,1400]" italics="true" pageId="8" pageNumber="161">R. roshanpererai</emphasis>
</taxonomicName>
<emphasis id="B90A5959EA722A45FE3AC435C211FA99" bold="true" box="[509,599,1375,1400]" pageId="8" pageNumber="161">
<taxonomicNameLabel id="A239E422EA722A45FE3AC435C211FA99" box="[509,599,1375,1400]" pageId="8" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was found within a
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radius, dug during the day from soil ca.
<quantity id="4C8628AEEA722A45FAF9C435C5DBFA99" box="[1342,1437,1375,1400]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="161" unit="mm" value="150.0">150 mm</quantity>
deep among banana plants in a home garden. Other specimens have been seen at or close to (within a couple of kilometers) of this site in a wide range of habitats, including shaded patches of grassland, tea plantations, and disturbed riverine forest, always dug from soil or leaf litter during the day. A few specimens have been seen moving on the surface, only at night. Several other individuals of the new species were dug from soil in disturbed riverine forest in 1999 from Uda Diyaluma, approximately
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away (
<geoCoordinate id="EE4AE38CEA722A45FC3DC77EC4FAF9CD" box="[1018,1212,1556,1581]" direction="north" orientation="latitude" pageId="8" pageNumber="161" precision="1" value="6.7357087">6° 44 08.55” N</geoCoordinate>
,
<geoCoordinate id="EE4AE38CEA722A45FB0BC77EC5D1F9CD" box="[1228,1431,1556,1581]" direction="east" orientation="longitude" pageId="8" pageNumber="161" precision="1" value="81.03253">81° 01 57.10” E</geoCoordinate>
, elevation
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), and from Haldummulla, approximately
<quantity id="4C8628AEEA722A45FCF6C75DC32AF9B1" box="[817,876,1591,1616]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="161" unit="km" value="6.0">6 km</quantity>
from the
<typeStatus id="54C53BE9EA722A45FC26C753C457F9B0" box="[993,1041,1593,1617]" pageId="8" pageNumber="161">type</typeStatus>
locality (
<geoCoordinate id="EE4AE38CEA722A45FB47C75DC57DF9B1" box="[1152,1339,1591,1616]" direction="north" orientation="latitude" pageId="8" pageNumber="161" precision="1" value="6.7610974">6° 45 39.95” N</geoCoordinate>
,
<geoCoordinate id="EE4AE38CEA722A45FA8EC752C0B9F995" direction="east" orientation="longitude" pageId="8" pageNumber="161" precision="1" value="80.901596">80° 54 05.73” E</geoCoordinate>
, elevation
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). Despite a substantial amount of fieldwork (including digging through soil and leaf litter) at similar altitudes, we have not observed this species outside this region, including at, for example, Haputale, less than
<quantity id="4C8628AEEA722A45FE46C7CEC1F9F95C" box="[385,447,1700,1725]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="161" unit="km" value="2.0">2 km</quantity>
North of the
<typeStatus id="54C53BE9EA722A45FDADC7CFC2DCF95C" box="[618,666,1701,1725]" pageId="8" pageNumber="161">type</typeStatus>
locality but approximately
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higher in elevation.
<taxonomicName id="4C7EFEC8EA722A45FAEDC7CEC173F901" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="161" phylum="Chordata" rank="species" species="roshanpererai" status="sp. nov.">
<emphasis id="B90A5959EA722A45FAEDC7CEC173F901" italics="true" pageId="8" pageNumber="161">Rhinophis roshanpererai</emphasis>
</taxonomicName>
<emphasis id="B90A5959EA722A45FEF9C7ADC1DDF901" bold="true" box="[318,411,1735,1760]" pageId="8" pageNumber="161">
<taxonomicNameLabel id="A239E422EA722A45FEF9C7ADC1DDF901" box="[318,411,1735,1760]" pageId="8" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has not been found in sympatry with other uropeltid species. The nearest observation of other species that we know of (L.J.M.W., pers. obs.) is for
<taxonomicName id="4C7EFEC8EA722A45FC81C787C454F8E4" box="[838,1042,1772,1797]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="161" phylum="Chordata" rank="species" species="drummondhayi">
<emphasis id="B90A5959EA722A45FC81C787C454F8E4" box="[838,1042,1772,1797]" italics="true" pageId="8" pageNumber="161">R. drummondhayi</emphasis>
</taxonomicName>
at
<quantity id="4C8628AEEA722A45FBFEC786C4C5F8E5" box="[1081,1155,1772,1797]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="8" pageNumber="161" unit="m" value="960.0">960 m</quantity>
elevation at Kubalwela, approximately
<quantity id="4C8628AEEA722A45FEF8C665C1C1F8C9" box="[319,391,1807,1832]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="161" unit="km" value="16.0">16 km</quantity>
to the northeast by north (bearing of 30°) of the
<typeStatus id="54C53BE9EA722A45FC62C67BC393F8C8" box="[933,981,1809,1833]" pageId="8" pageNumber="161">type</typeStatus>
locality of
<taxonomicName id="4C7EFEC8EA722A45FB92C67BC554F8C9" box="[1109,1298,1807,1832]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="161" phylum="Chordata" rank="species" species="roshanpererai" status="sp. nov.">
<emphasis id="B90A5959EA722A45FB92C67BC554F8C9" box="[1109,1298,1807,1832]" italics="true" pageId="8" pageNumber="161">R. roshanpererai</emphasis>
</taxonomicName>
<emphasis id="B90A5959EA722A45FADDC665C535F8C9" bold="true" box="[1306,1395,1807,1832]" pageId="8" pageNumber="161">
<taxonomicNameLabel id="A239E422EA722A45FADDC665C535F8C9" box="[1306,1395,1807,1832]" pageId="8" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
.
</paragraph>
<paragraph id="8BC1854BEA722A45FF00C65FC0AFF874" blockId="8.[151,1437,151,1941]" pageId="8" pageNumber="161">We suspect that the vertical and horizontal distributional range of the new species is small, and that substantial human disturbance in the form of intensive agriculture and urbanization represent the likely greatest conservation threats.</paragraph>
</subSubSection>
</treatment>
</document>