471 lines
42 KiB
XML
471 lines
42 KiB
XML
<document ID-DOI="10.25221/fee.440.2" ID-ISSN="2713-2196" ID-Zenodo-Dep="7166541" approvalRequired="37" approvalRequired_for_textStreams="37" checkinTime="1665255068009" checkinUser="felipe" docAuthor="Makarov, K. V. & Sundukov, Yu. N." docDate="2021" docId="03920E453456333486A4FC68496CF9B1" docLanguage="en" docName="FarEastEntomol.440.13-24.pdf" docOrigin="Far Eastern Entomologist 440" docSource="http://dx.doi.org/10.25221/fee.440.2" docStyle="DocumentStyle:68641B8F4A21450A5F9A74164B730F50.1:FarEastEntomol.2017-.journal_article" docStyleId="68641B8F4A21450A5F9A74164B730F50" docStyleName="FarEastEntomol.2017-.journal_article" docStyleVersion="1" docTitle="Bembidion (Peryphanes) sanatum subsp. iwanai Makarov et Sundukov 2021, ssp. n." docType="treatment" docVersion="2" lastPageNumber="19" masterDocId="FFAB763D345433328647FD574D63FFE4" masterDocTitle="A new subspecies of Bembidion sanatum (Coleoptera: Carabidae) endemic to the Mendeleev Volcano (Kunashir Island, Russia)" masterLastPageNumber="24" masterPageNumber="13" pageNumber="15" updateTime="1665270205595" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new subspecies of Bembidion sanatum (Coleoptera: Carabidae) endemic to the Mendeleev Volcano (Kunashir Island, Russia)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Makarov, K. V.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sundukov, Yu. N.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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||
<mods:titleInfo>
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<mods:title>Far Eastern Entomologist</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-11-02</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>440</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>13</mods:start>
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<mods:end>24</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.25221/fee.440.2</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.25221/fee.440.2</mods:identifier>
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<mods:identifier type="ISSN">2713-2196</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7166541</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:03920E453456333486A4FC68496CF9B1" httpUri="http://treatment.plazi.org/id/03920E453456333486A4FC68496CF9B1" lastPageId="6" lastPageNumber="19" pageId="2" pageNumber="15">
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||
<subSubSection pageId="2" pageNumber="15" type="nomenclature">
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||
<paragraph blockId="2.[227,948,319,341]" box="[227,948,319,341]" pageId="2" pageNumber="15">
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||
<heading box="[227,948,319,341]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="15" reason="2">
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<emphasis bold="true" box="[227,948,319,341]" pageId="2" pageNumber="15">
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<taxonomicName authority="Makarov et Sundukov" authorityName="Makarov et Sundukov" authorityYear="2021" box="[227,874,319,341]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="15" phylum="Arthropoda" rank="subSpecies" species="sanatum" status="ssp. n." subGenus="Peryphanes" subSpecies="iwanai">
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<emphasis bold="true" box="[227,335,319,341]" italics="true" pageId="2" pageNumber="15">Bembidion</emphasis>
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(
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<emphasis bold="true" box="[349,465,319,341]" italics="true" pageId="2" pageNumber="15">Peryphanes</emphasis>
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)
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<emphasis bold="true" box="[479,637,319,341]" italics="true" pageId="2" pageNumber="15">sanatum iwanai</emphasis>
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Makarov et Sundukov
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</taxonomicName>
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,
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<taxonomicNameLabel box="[884,948,320,341]" pageId="2" pageNumber="15" rank="subSpecies">ssp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="2.[227,1077,352,374]" box="[227,1077,352,374]" pageId="2" pageNumber="15">
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http://zoobank.org/NomenclaturalActs/
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<uuid box="[610,1077,352,374]" pageId="2" pageNumber="15">AB33AD02-E50F-4FBC-BC41-A0F9533AC812</uuid>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection box="[227,706,385,407]" pageId="2" pageNumber="15" type="description">
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<paragraph blockId="2.[227,706,385,407]" box="[227,706,385,407]" pageId="2" pageNumber="15">
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<figureCitation box="[227,312,385,407]" captionStart="Figs 1–4" captionStartId="3.[265,306,1540,1562]" captionTargetBox="[228,1105,288,1476]" captionTargetId="figure-2@3.[227,1122,285,1480]" captionTargetPageId="3" captionText="Figs 1–4. Bembidion sanatum, dorsal habitus: 1, 2 – B. sanatum iwanai, ssp. n.: 1 – male," figureDoi="http://doi.org/10.5281/zenodo.7166543" httpUri="https://zenodo.org/record/7166543/files/figure.png" pageId="2" pageNumber="15">Figs 1–2</figureCitation>
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,
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<figureCitation box="[322,492,385,407]" captionStart="Figs 5–16" captionStartId="4.[265,306,1525,1547]" captionTargetBox="[275,1078,455,1485]" captionTargetId="figure-75@4.[275,1078,448,1520]" captionTargetPageId="4" captionText="Figs 5–16. Bembidion sanatum, detals: 5, 6, 9, 10, 13,14 – B. sanatum iwanai, ssp. n.; 7," figureDoi="http://doi.org/10.5281/zenodo.7166547" httpUri="https://zenodo.org/record/7166547/files/figure.png" pageId="2" pageNumber="15">5–6, 9–10, 13–14</figureCitation>
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,
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<figureCitation box="[502,706,385,407]" captionStart="Figs 17–31" captionStartId="6.[265,306,1543,1565]" captionTargetBox="[233,1122,290,1465]" captionTargetId="figure-0@6.[227,1122,288,1483]" captionTargetPageId="6" captionText="Figs 17–31. Male and female genitals of Bembidion sanatum: 17–19, 23–25, 28–29 – B." figureDoi="http://doi.org/10.5281/zenodo.7166549" httpUri="https://zenodo.org/record/7166549/files/figure.png" pageId="2" pageNumber="15">17–19, 23–25, 28–29</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="15" type="materials_examined">
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<paragraph blockId="2.[227,1127,440,1593]" box="[265,1127,440,462]" pageId="2" pageNumber="15">
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<materialsCitation box="[265,1127,440,462]" collectionCode="TYPE, MATERIAL" country="Russia" location="Kunashir Island" municipality="Kislaya River" pageId="2" pageNumber="15" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<collectionCode box="[265,462,440,461]" pageId="2" pageNumber="15">TYPE MATERIAL</collectionCode>
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.
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<typeStatus box="[475,564,440,462]" pageId="2" pageNumber="15">Holotype</typeStatus>
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–
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<specimenCount box="[595,615,440,461]" pageId="2" pageNumber="15" type="male">♂</specimenCount>
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,
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<emphasis bold="true" box="[629,697,440,462]" pageId="2" pageNumber="15">
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<collectingCountry box="[629,697,440,462]" name="Russia" pageId="2" pageNumber="15">Russia</collectingCountry>
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</emphasis>
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:
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<location LSID="urn:lsid:plazi:treatment:03920E453456333486A4FC68496CF9B1:8EE4E98834563330848EFCEF4E0BFE2A" box="[713,872,440,462]" country="Russia" municipality="Kislaya River" name="Kunashir Island" pageId="2" pageNumber="15">Kunashir Island</location>
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, valley of
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<collectingMunicipality box="[984,1123,440,462]" pageId="2" pageNumber="15">Kislaya River</collectingMunicipality>
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,
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</materialsCitation>
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</paragraph>
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<paragraph blockId="2.[227,1127,440,1593]" pageId="2" pageNumber="15">
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<materialsCitation box="[227,786,468,490]" collectingDate="2011-06-18" collectionCode="VI, ZISP" collectorName="A. Matalin" pageId="2" pageNumber="15" specimenCount="1">
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below hot springs,
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<date box="[423,539,468,489]" pageId="2" pageNumber="15" value="2011-06-18">
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<collectingDate box="[423,539,468,489]" pageId="2" pageNumber="15" value="2011-06-18">
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18.
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<collectionCode box="[450,479,468,489]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="2" pageNumber="15">VI</collectionCode>
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2011
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</collectingDate>
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</date>
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, leg.
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<collectorName box="[599,708,468,490]" pageId="2" pageNumber="15">A. Matalin</collectorName>
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(
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<collectionCode box="[725,779,468,489]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="2" pageNumber="15">ZISP</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation collectionCode="MPU" pageId="2" pageNumber="15" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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<typeStatus box="[800,897,468,489]" pageId="2" pageNumber="15">Paratypes</typeStatus>
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: the same label as the
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<typeStatus box="[227,314,495,517]" pageId="2" pageNumber="15">holotype</typeStatus>
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,
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<specimenCount box="[324,361,495,517]" pageId="2" pageNumber="15" type="female">1 ♀</specimenCount>
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(
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<collectionCode box="[375,432,495,516]" country="France" lsid="urn:lsid:biocol.org:col:15519" name="Université Montpellier 2" pageId="2" pageNumber="15" type="Herbarium">MPU</collectionCode>
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)
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</materialsCitation>
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;
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<materialsCitation collectingDate="2016-08-03" collectionCode="FEB" collectorName="Yu. & L. Sundukov" county="Kislaya River" location="Kunashir Island" municipality="Mendeleev Volcano" pageId="2" pageNumber="15" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
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<location LSID="urn:lsid:plazi:treatment:03920E453456333486A4FC68496CF9B1:8EE4E988345633308785FCB84F3DFDE1" box="[450,606,495,517]" county="Kislaya River" municipality="Mendeleev Volcano" name="Kunashir Island" pageId="2" pageNumber="15">Kunashir Island</location>
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,
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<collectingMunicipality box="[617,814,495,517]" pageId="2" pageNumber="15">Mendeleev Volcano</collectingMunicipality>
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, source of
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<collectingCounty box="[921,1058,495,517]" pageId="2" pageNumber="15">Kislaya River</collectingCounty>
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, below the solfatara field,
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<date box="[412,529,523,545]" pageId="2" pageNumber="15" value="2016-08-03">
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<collectingDate box="[412,529,523,545]" pageId="2" pageNumber="15" value="2016-08-03">3.VIII 2016</collectingDate>
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</date>
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,
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<specimenCount box="[540,580,523,544]" pageId="2" pageNumber="15" type="male">1 ♂</specimenCount>
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,
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<specimenCount box="[591,628,523,544]" pageId="2" pageNumber="15" type="female">1 ♀</specimenCount>
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, leg.
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<collectorName box="[685,879,523,545]" pageId="2" pageNumber="15">Yu. & L. Sundukov</collectorName>
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(
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<collectionCode box="[894,943,523,544]" pageId="2" pageNumber="15">FEB</collectionCode>
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||
)
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</materialsCitation>
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;
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<materialsCitation collectingDate="2016-09-22" collectionCode="ZISP, ZMMU, MPU, FEB" collectorName="Yu. Sundukov" county="Kislaya River" location="Kunashir Island" municipality="Mendeleev Volcano" pageId="2" pageNumber="15" specimenCount="32" specimenCount-female="17" specimenCount-male="15" typeStatus="paratype">
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the same locality but
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<date box="[268,384,550,572]" pageId="2" pageNumber="15" value="2016-09-22">
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<collectingDate box="[268,384,550,572]" pageId="2" pageNumber="15" value="2016-09-22">22.IX 2016</collectingDate>
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</date>
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,
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<specimenCount box="[398,453,550,572]" pageId="2" pageNumber="15" type="male">15 ♂</specimenCount>
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,
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<specimenCount box="[467,519,551,572]" pageId="2" pageNumber="15" type="female">17 ♀</specimenCount>
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, leg.
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<collectorName box="[581,724,550,572]" pageId="2" pageNumber="15">Yu. Sundukov</collectorName>
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(
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<collectionCode box="[742,795,551,572]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="2" pageNumber="15">ZISP</collectionCode>
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,
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<collectionCode box="[809,886,551,572]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="2" pageNumber="15" type="Museum">ZMMU</collectionCode>
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||
,
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<collectionCode box="[900,954,551,572]" country="France" lsid="urn:lsid:biocol.org:col:15519" name="Université Montpellier 2" pageId="2" pageNumber="15" type="Herbarium">MPU</collectionCode>
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,
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<collectionCode box="[968,1012,551,572]" pageId="2" pageNumber="15">FEB</collectionCode>
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)
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</materialsCitation>
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||
;
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<materialsCitation collectingDate="2017-07-15" collectionCode="FEB" collectorName="Yu. & L. Sundukov" county="Kislaya River" location="Kunashir Island" municipality="Mendeleev Volcano" pageId="2" pageNumber="15" specimenCount="15" specimenCount-female="9" specimenCount-male="6" typeStatus="paratype">
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the same locality but
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<date box="[342,461,578,600]" pageId="2" pageNumber="15" value="2017-07-15">
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<collectingDate box="[342,461,578,600]" pageId="2" pageNumber="15" value="2017-07-15">15.VII 2017</collectingDate>
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</date>
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,
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<specimenCount box="[471,509,578,600]" pageId="2" pageNumber="15" type="male">6 ♂</specimenCount>
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,
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<specimenCount box="[519,555,578,600]" pageId="2" pageNumber="15" type="female">9 ♀</specimenCount>
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, leg.
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<collectorName box="[608,798,578,600]" pageId="2" pageNumber="15">Yu. & L. Sundukov</collectorName>
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(
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<collectionCode box="[811,856,578,599]" pageId="2" pageNumber="15">FEB</collectionCode>
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||
)
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</materialsCitation>
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.
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</paragraph>
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<paragraph blockId="2.[227,1127,440,1593]" box="[265,1118,605,627]" pageId="2" pageNumber="15">
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COMPARATIVE
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||
<collectionCode box="[452,579,606,627]" pageId="2" pageNumber="15">MATERIAL</collectionCode>
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||
.
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<emphasis bold="true" box="[597,1118,605,627]" pageId="2" pageNumber="15">
|
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<taxonomicName authority="Bates" box="[597,1114,605,627]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="15" phylum="Arthropoda" rank="subSpecies" species="sanatum" subGenus="Peryphanes" subSpecies="sanatum">
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||
<emphasis bold="true" box="[597,705,605,627]" italics="true" pageId="2" pageNumber="15">Bembidion</emphasis>
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||
(
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||
<emphasis bold="true" box="[727,843,605,627]" italics="true" pageId="2" pageNumber="15">Peryphanes</emphasis>
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||
)
|
||
<emphasis bold="true" box="[865,1050,606,627]" italics="true" pageId="2" pageNumber="15">sanatum sanatum</emphasis>
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Bates
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</taxonomicName>
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,
|
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</emphasis>
|
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="5" lastPageNumber="18" pageId="2" pageNumber="15" type="description">
|
||
<paragraph blockId="2.[227,1127,440,1593]" box="[227,1127,633,655]" pageId="2" pageNumber="15">
|
||
<emphasis bold="true" box="[227,275,633,654]" pageId="2" pageNumber="15">1883</emphasis>
|
||
.
|
||
<materialsCitation box="[288,1104,633,655]" collectingDate="1993-09-18" country="Japan" location="Mount Hikageyama Jyoushu" municipality="Honshu Island" pageId="2" pageNumber="15" specimenCount="2" specimenCount-male="2" stateProvince="Gunma">
|
||
<emphasis bold="true" box="[288,351,633,654]" pageId="2" pageNumber="15">
|
||
<collectingCountry box="[288,351,633,654]" name="Japan" pageId="2" pageNumber="15">Japan</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<collectingMunicipality box="[364,506,633,655]" pageId="2" pageNumber="15">Honshu Island</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:03920E453456333486A4FC68496CF9B1:8EE4E988345633308442FF2D4E7FFD6B" box="[517,796,633,655]" country="Japan" municipality="Honshu Island" name="Mount Hikageyama Jyoushu" pageId="2" pageNumber="15" stateProvince="Gunma">Mount Hikageyama Jyoushu</location>
|
||
,
|
||
<collectingRegion box="[807,933,633,655]" country="Japan" name="Gunma" pageId="2" pageNumber="15">Gunma Pref.</collectingRegion>
|
||
,
|
||
<date box="[943,1054,633,654]" pageId="2" pageNumber="15" value="1993-09-18">
|
||
<collectingDate box="[943,1054,633,654]" pageId="2" pageNumber="15" value="1993-09-18">18.IX 1993</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount box="[1065,1104,633,655]" pageId="2" pageNumber="15" type="male">2 ♂</specimenCount>
|
||
</materialsCitation>
|
||
, 2
|
||
</paragraph>
|
||
<paragraph blockId="2.[227,1127,440,1593]" box="[227,531,661,683]" pageId="2" pageNumber="15">
|
||
<materialsCitation box="[227,528,661,683]" collectionCode="ZISP, MPU" collectorName="S. Morita" pageId="2" pageNumber="15" specimenCount="1" specimenCount-female="1">
|
||
<specimenCount box="[227,245,661,682]" pageId="2" pageNumber="15" type="female">♀</specimenCount>
|
||
, leg.
|
||
<collectorName box="[298,389,661,683]" pageId="2" pageNumber="15">S. Morita</collectorName>
|
||
(
|
||
<collectionCode box="[402,455,661,682]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="2" pageNumber="15">ZISP</collectionCode>
|
||
,
|
||
<collectionCode box="[464,520,661,682]" country="France" lsid="urn:lsid:biocol.org:col:15519" name="Université Montpellier 2" pageId="2" pageNumber="15" type="Herbarium">MPU</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="2.[227,1127,440,1593]" box="[265,733,688,710]" pageId="2" pageNumber="15">
|
||
Measured:
|
||
<specimenCount box="[372,410,689,710]" pageId="2" pageNumber="15" type="male">8 ♂</specimenCount>
|
||
,
|
||
<specimenCount box="[420,457,689,710]" pageId="2" pageNumber="15" type="female">7 ♀</specimenCount>
|
||
; genital preparate:
|
||
<specimenCount box="[642,680,688,710]" pageId="2" pageNumber="15" type="male">5 ♂</specimenCount>
|
||
;
|
||
<materialsCitation box="[691,728,688,710]" isTailEnumeration="true" pageId="2" pageNumber="15" specimenCount="5" specimenCount-female="5">
|
||
<specimenCount box="[691,728,688,710]" pageId="2" pageNumber="15" type="female">5 ♀</specimenCount>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="2.[227,1127,440,1593]" box="[265,1054,716,738]" pageId="2" pageNumber="15">
|
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DESCRIPTION. Body moderately convex; TL =
|
||
<quantity box="[742,860,716,738]" metricMagnitude="-3" metricUnit="m" metricValue="5.699999999999999" metricValueMax="6.3" metricValueMin="5.1" pageId="2" pageNumber="15" unit="mm" value="5.699999999999999" valueMax="6.3" valueMin="5.1">5.1–6.3 mm</quantity>
|
||
, EW =
|
||
<quantity box="[932,1050,716,737]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.7" metricValueMin="2.2" pageId="2" pageNumber="15" unit="mm" value="2.45" valueMax="2.7" valueMin="2.2">2.2–2.7 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="2.[227,1127,440,1593]" pageId="2" pageNumber="15">Head and pronotum dark brown, with intense blue shine; elytra resin-brown or brownish with slightly darkened apices; mandibles brown, with blackened apices and middle part;</paragraph>
|
||
<paragraph blockId="2.[227,1127,440,1593]" box="[227,1024,799,821]" pageId="2" pageNumber="15">
|
||
palps, antennae and legs pale brown; ventral surface black or tar-brown (
|
||
<figureCitation box="[925,1012,799,821]" captionStart="Figs 1–4" captionStartId="3.[265,306,1540,1562]" captionTargetBox="[228,1105,288,1476]" captionTargetId="figure-2@3.[227,1122,285,1480]" captionTargetPageId="3" captionText="Figs 1–4. Bembidion sanatum, dorsal habitus: 1, 2 – B. sanatum iwanai, ssp. n.: 1 – male," figureDoi="http://doi.org/10.5281/zenodo.7166543" httpUri="https://zenodo.org/record/7166543/files/figure.png" pageId="2" pageNumber="15">Figs 1, 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="2.[227,1127,440,1593]" pageId="2" pageNumber="15">
|
||
Dorsal surface shiny, elytral microsculpture (better visible in females) formed by thin transverse lines (
|
||
<figureCitation box="[394,498,854,876]" captionStart="Figs 5–16" captionStartId="4.[265,306,1525,1547]" captionTargetBox="[275,1078,455,1485]" captionTargetId="figure-75@4.[275,1078,448,1520]" captionTargetPageId="4" captionText="Figs 5–16. Bembidion sanatum, detals: 5, 6, 9, 10, 13,14 – B. sanatum iwanai, ssp. n.; 7," figureDoi="http://doi.org/10.5281/zenodo.7166547" httpUri="https://zenodo.org/record/7166547/files/figure.png" pageId="2" pageNumber="15">Figs 9, 10</figureCitation>
|
||
), barely visible at x56. Head and pronotum with scattered fine punctures; on the elytra, only striae are punctured.
|
||
</paragraph>
|
||
<paragraph blockId="2.[227,1127,440,1593]" box="[265,1127,909,931]" pageId="2" pageNumber="15">
|
||
Basic measurements (
|
||
<specimenCount box="[475,612,909,931]" pageId="2" pageNumber="15" type="generic">30 specimens</specimenCount>
|
||
): HW 1.08–1.22, M 1.13; HL 0.83–0.97, M 0.91; PA
|
||
</paragraph>
|
||
<paragraph blockId="2.[227,1127,440,1593]" box="[227,1127,937,959]" pageId="2" pageNumber="15">0.98–1.12, M 1.05; PW 1.36–1.62, M 1.46; PB 1.03–1.22, M 1.13; PLt 1.13–1.35, M 1.23;</paragraph>
|
||
<paragraph blockId="2.[227,1127,440,1593]" box="[227,1126,964,986]" pageId="2" pageNumber="15">PLm 1.09–1.21, M 1.16; EW 2.28–2.60, M 2.42; EL 3.43–3.98, M 3.66; TL 5.43–5.94, M</paragraph>
|
||
<paragraph blockId="2.[227,1127,440,1593]" box="[227,276,992,1014]" pageId="2" pageNumber="15">5.65.</paragraph>
|
||
<paragraph blockId="2.[227,1127,440,1593]" box="[265,1127,1019,1041]" pageId="2" pageNumber="15">Head normal, moderately convex, slightly transverse (HW/HL = 1.17–1.33, M 1.26).</paragraph>
|
||
<paragraph blockId="2.[227,1127,440,1593]" pageId="2" pageNumber="15">Eyes small, moderately convex. Antennae slender, long, 0.85–0.93 times as long as elytra in both sexes; 2nd antennomere shortest; 3rd to 10th 1.2–1.6 times as long as 2nd one; scape</paragraph>
|
||
<paragraph blockId="2.[227,1127,440,1593]" box="[227,1127,1102,1124]" pageId="2" pageNumber="15">
|
||
1.2–1.3 times thicker than other antennomeres; ratio of antennomeres in
|
||
<specimenCount box="[940,1029,1102,1124]" pageId="2" pageNumber="15" type="male">one male</specimenCount>
|
||
= 19: 14:
|
||
</paragraph>
|
||
<paragraph blockId="2.[227,1127,440,1593]" box="[227,1127,1130,1152]" pageId="2" pageNumber="15">
|
||
21.5: 21.5: 21: 19: 18: 18: 17.5: 17: 22.5, in
|
||
<specimenCount box="[653,759,1130,1152]" pageId="2" pageNumber="15" type="female">one female</specimenCount>
|
||
= 17: 14: 22: 21: 20: 19: 18: 17: 17.5:
|
||
</paragraph>
|
||
<paragraph blockId="2.[227,1127,440,1593]" box="[227,1127,1157,1180]" pageId="2" pageNumber="15">17: 20.5. Mandibles relatively short, slender, with pointed apices. Labrum weakly transverse,</paragraph>
|
||
<paragraph blockId="2.[227,1127,440,1593]" pageId="2" pageNumber="15">with six setae along a straight anterior margin. Clypeus slightly transverse, hexagonal, with two lateral setae in anterior third. Tooth of mentum rather large, wide, narrowly rounded at apex, with two basal setae. Submentum on each side with one long and one short seta at posterior margin, and with one long seta at anterior angle. Gula smooth. Temples short, about</paragraph>
|
||
<paragraph blockId="2.[227,1127,440,1593]" pageId="2" pageNumber="15">1/3 diameter of eye, smoothly merging into a wide neck. Two supraorbital setae in the middle and at posterior margin of eye. Frontal grooves deep and wide, slightly projecting beyond anterior supraorbital setae; weakly converging anteriorly from setae to clypeus,</paragraph>
|
||
<paragraph blockId="2.[227,1127,440,1593]" pageId="2" pageNumber="15">sharply diverging on clypeus to its lateral margins; grooves and area behind apex of frontal grooves with scattered, but rather coarse punctures.</paragraph>
|
||
<paragraph blockId="2.[227,1127,440,1593]" box="[265,1126,1433,1456]" pageId="2" pageNumber="15">
|
||
Pronotum (
|
||
<figureCitation box="[379,479,1433,1456]" captionStart="Figs 5–16" captionStartId="4.[265,306,1525,1547]" captionTargetBox="[275,1078,455,1485]" captionTargetId="figure-75@4.[275,1078,448,1520]" captionTargetPageId="4" captionText="Figs 5–16. Bembidion sanatum, detals: 5, 6, 9, 10, 13,14 – B. sanatum iwanai, ssp. n.; 7," figureDoi="http://doi.org/10.5281/zenodo.7166547" httpUri="https://zenodo.org/record/7166547/files/figure.png" pageId="2" pageNumber="15">Figs 5, 6</figureCitation>
|
||
) cordate, regularly and moderately convex, slightly transverse
|
||
</paragraph>
|
||
<paragraph blockId="2.[227,1127,440,1593]" pageId="2" pageNumber="15">(PW/PLm = 1.21–1.35, M 1.26), wider than head (PW/HW = 1.23–1.34, M 1.30), with maximum width at 3/5 off base. Anterior margin straight, distinctly bordered from anterior angles towards 2/3 to midline. Anterior angles not projecting forward, with shortly pointed apices. Lateral sides rather strongly and uniformly convex, long and rectilinear before posterior angles. Lateral margins narrow throughout; with two lateral setae on each side:</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7166543" ID-Zenodo-Dep="7166543" box="[265,1127,1540,1562]" httpUri="https://zenodo.org/record/7166543/files/figure.png" pageId="3" pageNumber="16" startId="3.[265,306,1540,1562]" targetBox="[228,1105,288,1476]" targetPageId="3">
|
||
<paragraph blockId="3.[227,1127,1540,1590]" box="[265,1127,1540,1562]" pageId="3" pageNumber="16">
|
||
Figs 1–4.
|
||
<taxonomicName authorityName="Bates" authorityYear="1883" box="[361,552,1540,1562]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="16" phylum="Arthropoda" rank="species" species="sanatum">
|
||
<emphasis box="[361,552,1540,1562]" italics="true" pageId="3" pageNumber="16">Bembidion sanatum</emphasis>
|
||
</taxonomicName>
|
||
, dorsal habitus: 1, 2 –
|
||
<taxonomicName authorityName="Makarov et Sundukov" authorityYear="2021" box="[772,951,1541,1562]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="16" phylum="Arthropoda" rank="subSpecies" species="sanatum" subSpecies="iwanai">
|
||
<emphasis box="[772,787,1541,1562]" italics="true" pageId="3" pageNumber="16">B</emphasis>
|
||
.
|
||
<emphasis box="[799,951,1541,1562]" italics="true" pageId="3" pageNumber="16">sanatum iwanai</emphasis>
|
||
</taxonomicName>
|
||
, ssp. n.: 1 – male,
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="3.[227,1127,1540,1590]" box="[227,818,1568,1590]" pageId="3" pageNumber="16">
|
||
2 – female; 3, 4 –
|
||
<taxonomicName box="[407,602,1568,1589]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="16" phylum="Arthropoda" rank="subSpecies" species="sanatum" subSpecies="sanatum">
|
||
<emphasis box="[407,422,1568,1589]" italics="true" pageId="3" pageNumber="16">B</emphasis>
|
||
.
|
||
<emphasis box="[434,602,1568,1589]" italics="true" pageId="3" pageNumber="16">sanatum sanatum</emphasis>
|
||
</taxonomicName>
|
||
: 3 – male, 4 – female.
|
||
</paragraph>
|
||
<paragraph blockId="4.[227,1127,260,420]" pageId="4" pageNumber="17">before maximum width and at posterior angles. Base slightly and uniformly convex throughout, slightly wider than anterior margin (PB/PA = 1.04–1.11, M 1.08). Posterior angles distinct, rectangular, pointed at apices. Midline thin, distinct from anterior margin to a basal transverse bulge. Anterior transverse impression very weak or absent, often distinctly punctate in middle part. Basal fovea large, rounded, deep; their basal half and area between them rather coarsely punctured.</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7166547" ID-Zenodo-Dep="7166547" box="[265,1126,1525,1547]" httpUri="https://zenodo.org/record/7166547/files/figure.png" pageId="4" pageNumber="17" startId="4.[265,306,1525,1547]" targetBox="[275,1078,455,1485]" targetPageId="4">
|
||
<paragraph blockId="4.[227,1127,1525,1630]" box="[265,1126,1525,1547]" pageId="4" pageNumber="17">
|
||
Figs 5–16.
|
||
<taxonomicName authorityName="Bates" authorityYear="1883" box="[373,564,1525,1547]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="species" species="sanatum">
|
||
<emphasis box="[373,564,1525,1547]" italics="true" pageId="4" pageNumber="17">Bembidion sanatum</emphasis>
|
||
</taxonomicName>
|
||
, detals: 5, 6, 9, 10, 13,14 –
|
||
<taxonomicName authorityName="Makarov et Sundukov" authorityYear="2021" box="[835,1015,1526,1547]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="sanatum" subSpecies="iwanai">
|
||
<emphasis box="[835,850,1526,1547]" italics="true" pageId="4" pageNumber="17">B</emphasis>
|
||
.
|
||
<emphasis box="[862,1015,1526,1547]" italics="true" pageId="4" pageNumber="17">sanatum iwanai</emphasis>
|
||
</taxonomicName>
|
||
, ssp. n.; 7,
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="4.[227,1127,1525,1630]" box="[227,1127,1553,1575]" pageId="4" pageNumber="17">
|
||
8, 11, 12, 15, 16 –
|
||
<taxonomicName box="[420,619,1553,1575]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="sanatum" subSpecies="sanatum">
|
||
<emphasis box="[420,435,1554,1575]" italics="true" pageId="4" pageNumber="17">B</emphasis>
|
||
.
|
||
<emphasis box="[449,619,1554,1575]" italics="true" pageId="4" pageNumber="17">sanatum sanatum</emphasis>
|
||
</taxonomicName>
|
||
; 5–8 – pronotum; 9–12 – elytral striation; 13–16 –
|
||
</paragraph>
|
||
<paragraph blockId="4.[227,1127,1525,1630]" box="[227,1127,1580,1602]" pageId="4" pageNumber="17">microsculpture of elytra near series umbilicata; 5, 7, 9, 11, 13, 15 – males; 6, 8, 10, 12, 14, 16 –</paragraph>
|
||
<paragraph blockId="4.[227,1127,1525,1630]" box="[227,309,1608,1630]" pageId="4" pageNumber="17">females.</paragraph>
|
||
<paragraph blockId="5.[227,1128,263,1609]" box="[265,1127,263,285]" pageId="5" pageNumber="18">
|
||
Elytra (
|
||
<figureCitation box="[338,427,263,285]" captionStart="Figs 1–4" captionStartId="3.[265,306,1540,1562]" captionTargetBox="[228,1105,288,1476]" captionTargetId="figure-2@3.[227,1122,285,1480]" captionTargetPageId="3" captionText="Figs 1–4. Bembidion sanatum, dorsal habitus: 1, 2 – B. sanatum iwanai, ssp. n.: 1 – male," figureDoi="http://doi.org/10.5281/zenodo.7166543" httpUri="https://zenodo.org/record/7166543/files/figure.png" pageId="5" pageNumber="18">Figs 1, 2</figureCitation>
|
||
,
|
||
<figureCitation box="[439,489,263,284]" captionStart="Figs 5–16" captionStartId="4.[265,306,1525,1547]" captionTargetBox="[275,1078,455,1485]" captionTargetId="figure-75@4.[275,1078,448,1520]" captionTargetPageId="4" captionText="Figs 5–16. Bembidion sanatum, detals: 5, 6, 9, 10, 13,14 – B. sanatum iwanai, ssp. n.; 7," figureDoi="http://doi.org/10.5281/zenodo.7166547" httpUri="https://zenodo.org/record/7166547/files/figure.png" pageId="5" pageNumber="18">9, 10</figureCitation>
|
||
) oval, wide (EL/EW = 1.45–1.62, M 1.53; EW/PW = 1.56–1.73,
|
||
</paragraph>
|
||
<paragraph blockId="5.[227,1128,263,1609]" pageId="5" pageNumber="18">M 1.66), long (EL/PLt = 2.87–3.18, M 2.98), rather strongly convex, with maximum width near 2/3 basal part. Shoulders prominent regularly rounded. Basal border reaching the apex of 5th groove, smoothly passing into lateral margin. Lateral sides in anterior half slightly convex, in posterior half regularly and arcuately narrowed towards apices; preapical sinuation absent. Apical angle rounded, forming a short gap between elytra. 1st–6th grooves small, superficial at apex, finely punctured throughout; 7th groove smoothed, outlined only by punctures in basal half, at apex with one apical and one preapical pore. Parascutellar stria rather long finely punctured, parallel to suture. Elytral intervals flat throughout or 1st–5th</paragraph>
|
||
<paragraph blockId="5.[227,1128,263,1609]" pageId="5" pageNumber="18">barely convex in basal half. Scutellar pore rather large, joining with 1st and 2nd striae at base; 3rd interval with two small discal pores placed near 3rd stria at 1/3 or 2/3 off base,</paragraph>
|
||
<paragraph blockId="5.[227,1128,263,1609]" pageId="5" pageNumber="18">respectively. Series umbilicata consisting of eight setae: four in humeral group, two at 2/3 off base, and two near apex. Hind wings well-developed.</paragraph>
|
||
<paragraph blockId="5.[227,1128,263,1609]" pageId="5" pageNumber="18">
|
||
Ventral surface smooth, impunctate. Metasternal process pointed at apex, sharply bordered. Metepisterna long, along outer margin approximately twice as long as wide along anterior margin (measured in
|
||
<specimenCount box="[520,620,677,699]" pageId="5" pageNumber="18" type="male">two males</specimenCount>
|
||
and
|
||
<specimenCount box="[671,783,677,699]" pageId="5" pageNumber="18" type="female">one female</specimenCount>
|
||
). Abdominal sternites simple, each with one pair of setae, with neither additional hairs nor setae. Anal sternite in males with microsculpture of fine transverse lines, with two apical setae; in females with microsculpture of barely noticeable, strongly transverse cells, with four apical setae.
|
||
</paragraph>
|
||
<paragraph blockId="5.[227,1128,263,1609]" pageId="5" pageNumber="18">Legs long, slender; hind tarsus 0.79–0.89 times as long as hind tibia; ratio of metatarsomeres 1–5 as 14: 6: 6: 4: 13; in males, the 1st protarsomere strongly enlarged, and the 2nd moderately enlarged.</paragraph>
|
||
<paragraph blockId="5.[227,1128,263,1609]" box="[265,1126,870,892]" pageId="5" pageNumber="18">
|
||
Male genitalia. Aedeagus (
|
||
<figureCitation box="[520,630,870,892]" captionStart="Figs 17–31" captionStartId="6.[265,306,1543,1565]" captionTargetBox="[233,1122,290,1465]" captionTargetId="figure-0@6.[227,1122,288,1483]" captionTargetPageId="6" captionText="Figs 17–31. Male and female genitals of Bembidion sanatum: 17–19, 23–25, 28–29 – B." figureDoi="http://doi.org/10.5281/zenodo.7166549" httpUri="https://zenodo.org/record/7166549/files/figure.png" pageId="5" pageNumber="18">Figs 17, 18</figureCitation>
|
||
) large, about half as long as elytra (AL/EL = 0.53–
|
||
</paragraph>
|
||
<paragraph blockId="5.[227,1128,263,1609]" pageId="5" pageNumber="18">
|
||
0.55), with prominent basal sclerites. Lamella short, with a rounded apex; ventral surface of aedeagus without projection. Ribbon brush large, entirely protruding from basal part of aedeagus. Basal chitin platelet well-developed, extended anteriorly; ventral chitin platelet barely visible. Main sclerite long, its distal part a long flagellum running side by side with spur of lamina 1 (lama paracopulatrice); base of lamina 1 rounded (
|
||
<figureCitation box="[809,921,1008,1030]" captionStart="Figs 17–31" captionStartId="6.[265,306,1543,1565]" captionTargetBox="[233,1122,290,1465]" captionTargetId="figure-0@6.[227,1122,288,1483]" captionTargetPageId="6" captionText="Figs 17–31. Male and female genitals of Bembidion sanatum: 17–19, 23–25, 28–29 – B." figureDoi="http://doi.org/10.5281/zenodo.7166549" httpUri="https://zenodo.org/record/7166549/files/figure.png" pageId="5" pageNumber="18">Figs 23–25</figureCitation>
|
||
). Left paramere (
|
||
<figureCitation box="[1089,1128,1008,1030]" captionStart="Figs 17–31" captionStartId="6.[265,306,1543,1565]" captionTargetBox="[233,1122,290,1465]" captionTargetId="figure-0@6.[227,1122,288,1483]" captionTargetPageId="6" captionText="Figs 17–31. Male and female genitals of Bembidion sanatum: 17–19, 23–25, 28–29 – B." figureDoi="http://doi.org/10.5281/zenodo.7166549" httpUri="https://zenodo.org/record/7166549/files/figure.png" pageId="5" pageNumber="18">Fig.</figureCitation>
|
||
</paragraph>
|
||
<paragraph blockId="5.[227,1128,263,1609]" pageId="5" pageNumber="18">19) wide, triangular, with a small, trapezoidal, outer lobe; right paramere narrow, with a large outer blade; their chaetotaxy variable: left paramere with two (rarely a single) apical and two subapical setae, right paramere with three (rarely two) apical and one or two subapical setae.</paragraph>
|
||
<paragraph blockId="5.[227,1128,263,1609]" pageId="5" pageNumber="18">
|
||
Female genitalia (
|
||
<figureCitation box="[440,555,1118,1140]" captionStart="Figs 17–31" captionStartId="6.[265,306,1543,1565]" captionTargetBox="[233,1122,290,1465]" captionTargetId="figure-0@6.[227,1122,288,1483]" captionTargetPageId="6" captionText="Figs 17–31. Male and female genitals of Bembidion sanatum: 17–19, 23–25, 28–29 – B." figureDoi="http://doi.org/10.5281/zenodo.7166549" httpUri="https://zenodo.org/record/7166549/files/figure.png" pageId="5" pageNumber="18">Figs 28, 29</figureCitation>
|
||
). Spermatheca duct long, well-sclerotized, with five or six revolutions. Spermatheca with two chambers subequal in length; basal chamber noticeably asymmetric, convex on one side with gland opening at apex, apical chamber slightly curved.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="5" pageNumber="18" type="diagnosis">
|
||
<paragraph blockId="5.[227,1128,263,1609]" pageId="5" pageNumber="18">
|
||
DIAGNOSIS. By the size, colouration, and proportions,
|
||
<taxonomicName authorityName="Makarov et Sundukov" authorityYear="2021" box="[838,1025,1202,1223]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="subSpecies" species="sanatum" subSpecies="iwanai">
|
||
<emphasis box="[838,1025,1202,1223]" italics="true" pageId="5" pageNumber="18">B. sanatum iwanai</emphasis>
|
||
</taxonomicName>
|
||
ssp. n. is similar to the nominative subspecies, but the pronotum is slightly larger (PLm/EL 0.32,
|
||
<emphasis box="[1100,1127,1229,1250]" italics="true" pageId="5" pageNumber="18">vs.</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="5.[227,1128,263,1609]" pageId="5" pageNumber="18">
|
||
0.31; PW/EW 0.61,
|
||
<emphasis box="[422,449,1257,1278]" italics="true" pageId="5" pageNumber="18">vs.</emphasis>
|
||
0.69). The microsculpture of the outer elytral intervals is less strongly developed than in the nominative subspecies, while the 7
|
||
<superScript attach="left" box="[778,790,1279,1293]" fontSize="6" pageId="5" pageNumber="18">th</superScript>
|
||
and 8
|
||
<superScript attach="left" box="[850,862,1279,1293]" fontSize="6" pageId="5" pageNumber="18">th</superScript>
|
||
elytral intervals in the new subspecies are usually narrower than 6
|
||
<superScript attach="left" box="[606,618,1306,1320]" fontSize="6" pageId="5" pageNumber="18">th</superScript>
|
||
(0.9–1.1,
|
||
<emphasis box="[719,745,1312,1333]" italics="true" pageId="5" pageNumber="18">vs.</emphasis>
|
||
1.0–
|
||
<quantity box="[795,851,1312,1334]" metricMagnitude="-2" metricUnit="m" metricValue="3.048" pageId="5" pageNumber="18" unit="in" value="1.2">1.2 in</quantity>
|
||
the nominative subspecies).
|
||
</paragraph>
|
||
<paragraph blockId="5.[227,1128,263,1609]" box="[227,1127,1339,1361]" pageId="5" pageNumber="18">
|
||
The lamella of the aedeagus is somewhat narrower; the basal part of lamina 1 is rounded (
|
||
<emphasis box="[1100,1127,1340,1361]" italics="true" pageId="5" pageNumber="18">vs.</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="5.[227,1128,263,1609]" pageId="5" pageNumber="18">
|
||
angular in the nominative subspecies). The spermatheca duct shows a lower number of convolutions (5–6,
|
||
<emphasis box="[413,440,1395,1416]" italics="true" pageId="5" pageNumber="18">vs.</emphasis>
|
||
<quantity box="[445,506,1394,1416]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.032" metricValueMin="1.778" pageId="5" pageNumber="18" unit="in" value="7.5" valueMax="8.0" valueMin="7.0">7–8 in</quantity>
|
||
the nominative subspecies).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="5" pageNumber="18" type="distribution">
|
||
<paragraph blockId="5.[227,1128,263,1609]" box="[265,1124,1422,1444]" pageId="5" pageNumber="18">
|
||
DISTRIBUTION AND HABITAT. Known only from the
|
||
<typeStatus box="[835,876,1423,1444]" pageId="5" pageNumber="18">type</typeStatus>
|
||
locality, i.e. the valley of
|
||
</paragraph>
|
||
<paragraph blockId="5.[227,1128,263,1609]" pageId="5" pageNumber="18">
|
||
Kislaya that stems from the northeastern solfatara field of the Mendeleyev Volcano. Beetles are encountered under riverside stones in the middle flow part of the river where water temperatures are already lower, but both mineralization and acidity remain considerable (
|
||
<figureCitation box="[934,1042,1505,1527]" captionStart="Figs 32–33" captionStartId="7.[265,306,659,681]" captionTargetBox="[227,1122,257,626]" captionTargetId="figure-0@7.[227,1122,257,626]" captionTargetPageId="7" captionText="Figs 32–33. Habitat of Bembidion sanatum iwanai, ssp. n.: 32 – upstream of Kislaya River; 33 – the bank of Kislaya River in the middle reaches." figureDoi="http://doi.org/10.5281/zenodo.7166551" httpUri="https://zenodo.org/record/7166551/files/figure.png" pageId="5" pageNumber="18">Figs 32, 33</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="6" lastPageNumber="19" pageId="5" pageNumber="18" type="etymology">
|
||
<paragraph blockId="5.[227,1128,263,1609]" box="[265,1123,1532,1554]" pageId="5" pageNumber="18">
|
||
ETYMOLOGY. The
|
||
<typeStatus box="[480,521,1533,1554]" pageId="5" pageNumber="18">type</typeStatus>
|
||
series of the new subspecies was collected in the valley of
|
||
</paragraph>
|
||
<paragraph blockId="5.[227,1128,263,1609]" box="[227,1123,1560,1582]" pageId="5" pageNumber="18">Kislaya River where sulfur had previously been mined. Thus, the epithet comes from the</paragraph>
|
||
<paragraph blockId="5.[227,1128,263,1609]" box="[227,910,1587,1609]" pageId="5" pageNumber="18">
|
||
Ainu “
|
||
<emphasis box="[292,357,1588,1609]" italics="true" pageId="5" pageNumber="18">iwanai</emphasis>
|
||
”, meaning “a valley with sulphur” (Batchelor, 1905: 42).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7166549" ID-Zenodo-Dep="7166549" box="[265,1126,1543,1565]" httpUri="https://zenodo.org/record/7166549/files/figure.png" pageId="6" pageNumber="19" startId="6.[265,306,1543,1565]" targetBox="[233,1122,290,1465]" targetPageId="6">
|
||
<paragraph blockId="6.[227,1127,1543,1621]" box="[265,1126,1543,1565]" pageId="6" pageNumber="19">
|
||
Figs 17–31. Male and female genitals of
|
||
<taxonomicName authorityName="Bates" authorityYear="1883" box="[667,859,1543,1565]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="19" phylum="Arthropoda" rank="species" species="sanatum">
|
||
<emphasis box="[667,859,1543,1565]" italics="true" pageId="6" pageNumber="19">Bembidion sanatum</emphasis>
|
||
</taxonomicName>
|
||
: 17–19, 23–25, 28–29 –
|
||
<emphasis box="[1106,1121,1544,1565]" italics="true" pageId="6" pageNumber="19">B</emphasis>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="6.[227,1127,1543,1621]" box="[227,1127,1571,1593]" pageId="6" pageNumber="19">
|
||
<emphasis box="[227,384,1571,1593]" italics="true" pageId="6" pageNumber="19">
|
||
<taxonomicName authorityName="Bates" authorityYear="1883" box="[227,308,1572,1593]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="19" phylum="Arthropoda" rank="species" species="sanatum">sanatum</taxonomicName>
|
||
iwanai
|
||
</emphasis>
|
||
, ssp. n.; 20–22, 26–27, 30–31 –
|
||
<taxonomicName box="[733,938,1571,1593]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="19" phylum="Arthropoda" rank="subSpecies" species="sanatum" subSpecies="sanatum">
|
||
<emphasis box="[733,748,1572,1593]" italics="true" pageId="6" pageNumber="19">B</emphasis>
|
||
.
|
||
<emphasis box="[765,938,1572,1593]" italics="true" pageId="6" pageNumber="19">sanatum sanatum</emphasis>
|
||
</taxonomicName>
|
||
; 17, 18, 20, 21 –
|
||
</paragraph>
|
||
<paragraph blockId="6.[227,1127,1543,1621]" box="[227,1039,1599,1621]" pageId="6" pageNumber="19">aedeagus; 19, 22 – parameres; 23–27 – basal part of lamina 1; 28–31 – spermatheca.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |