253 lines
49 KiB
XML
253 lines
49 KiB
XML
<document id="D0C27B99BA14076FBFDAA45EC76F0979" ID-DOI="10.26107/RBZ-2023-0047" ID-ISSN="2345-7600" ID-Zenodo-Dep="10271307" ID-ZooBank="821BC4EC-5AF9-4727-84A3-C44839DFBE28" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1701819361795" checkinUser="felipe" docAuthor="Ng, Peter K. L., Ahyong, Shane T. & Castro, Peter" docDate="2023" docId="038EC92D79627E4CFCED697F26E1FC32" docLanguage="en" docName="RafflesBZool.71.606-631.pdf" docOrigin="Raffles Bulletin of Zoology 71" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Calocarcinidae Stevcic 2005, new status" docType="treatment" docVersion="2" lastPageNumber="624" masterDocId="FFB7B15579727E5EFFBA6E582531FFA7" masterDocTitle="Re-appraisal of the families and subfamilies of Trapezioidea Miers, 1886, with establishment of a new family, Ectaesthesiidae (Crustacea: Decapoda: Brachyura)" masterLastPageNumber="631" masterPageNumber="606" pageNumber="622" updateTime="1701819819889" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="466C5CA7F05437F6F444DD9EADC07EFD">Re-appraisal of the families and subfamilies of Trapezioidea Miers, 1886, with establishment of a new family, Ectaesthesiidae (Crustacea: Decapoda: Brachyura)</mods:title>
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<mods:namePart id="39F2A561EC13D3F7727134173ADB7EDB">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="A4352FD89DFC0EA38E373F609FAE8E2E">Ahyong, Shane T.</mods:namePart>
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<mods:namePart id="6E9E56E78D65F1E59A29CEE162457EC9">Castro, Peter</mods:namePart>
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<mods:title id="6696F9DA5421B532C10E55573A42F091">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="228A68445B2D6CDD0B806D59B57A3DBD">2023</mods:date>
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<mods:number id="EE0B6FC517CA1179FC8477D08D4EBA09">2023-10-30</mods:number>
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<mods:number id="F85A72B62DD4A443549304A3C217DC36">71</mods:number>
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<mods:classification id="CB052A9BD8AD43EE323BAA689B807AB5">journal article</mods:classification>
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<mods:identifier id="52822EE8ED7D6CF40C15204C840CE4E2" type="DOI">10.26107/RBZ-2023-0047</mods:identifier>
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<mods:identifier id="9BC0AEDDB645226AB88E1D61F3120D5C" type="ISSN">2345-7600</mods:identifier>
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<treatment id="038EC92D79627E4CFCED697F26E1FC32" LSID="urn:lsid:plazi:treatment:038EC92D79627E4CFCED697F26E1FC32" httpUri="http://treatment.plazi.org/id/038EC92D79627E4CFCED697F26E1FC32" lastPageId="18" lastPageNumber="624" pageId="16" pageNumber="622">
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<subSubSection id="C33D2BB079627E4EFCED697F2156F8DC" pageId="16" pageNumber="622" type="nomenclature">
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<paragraph id="8B98783B79627E4EFCED697F20B0F898" blockId="16.[855,1409,1831,1857]" box="[855,1409,1831,1857]" pageId="16" pageNumber="622">
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<heading id="D0D0CF5779627E4EFCED697F20B0F898" bold="true" box="[855,1409,1831,1857]" centered="true" fontSize="10" level="2" pageId="16" pageNumber="622" reason="2">
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<emphasis id="B953A42979627E4EFCED697F20B0F898" bold="true" box="[855,1409,1831,1857]" pageId="16" pageNumber="622">
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Family
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<taxonomicName id="4C2703B879627E4EFC15697F21CBF898" ID-CoL="BV5SW" authority="Stevcic, 2005" authorityName="Stevcic" authorityYear="2005" box="[943,1274,1831,1857]" class="Malacostraca" family="Calocarcinidae" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="622" phylum="Arthropoda" rank="family" status="new status">
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Calocarcinidae
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<bibRefCitation id="EFB605CA79627E4EFBDE697F21CBF898" author="Stevcic Z" box="[1124,1274,1831,1857]" pageId="16" pageNumber="622" pagination="1 - 159" refId="ref21857" refString="Stevcic Z (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Stevcic Brachyura). Natura Croatica (Fauna Croatica), 14 (1): 1 - 159." type="journal article" year="2005">Števčić, 2005</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A260195279627E4EFAB2697F20B0F898" box="[1288,1409,1831,1855]" pageId="16" pageNumber="622">new status</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B98783B79627E4EFC97693E2156F8DC" blockId="16.[813,1170,1893,1946]" box="[813,1127,1893,1917]" pageId="16" pageNumber="622">
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Calocarcinini
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<bibRefCitation id="EFB605CA79627E4EFC0C693E2155F8DC" author="Stevcic Z" box="[950,1124,1893,1917]" pageId="16" pageNumber="622" pagination="1 - 159" refId="ref21857" refString="Stevcic Z (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Stevcic Brachyura). Natura Croatica (Fauna Croatica), 14 (1): 1 - 159." type="journal article" year="2005">Števčić, 2005: 40</bibRefCitation>
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.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C33D2BB079627E4EFC9769DA21A3F830" box="[813,1170,1922,1946]" pageId="16" pageNumber="622" type="description">
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<paragraph id="8B98783B79627E4EFC9769DA21A3F830" blockId="16.[813,1170,1893,1946]" box="[813,1170,1922,1946]" pageId="16" pageNumber="622">
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Philippicarcinini
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<bibRefCitation id="EFB605CA79627E4EFC6E69DB21BFF830" author="Stevcic Z" box="[980,1166,1922,1946]" pageId="16" pageNumber="622" pagination="125 - 139" refId="ref21892" refString="Stevcic Z (2011) Addition to the reclassification of the brachyuran crabs (Crustacea: Decapoda: Brachyura) part I. New taxa. Natura Croatica, 20: 125 - 139." type="journal article" year="2011">Števčić, 2011: 131</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33D2BB079627E4CFC976998273BFAB2" lastPageId="18" lastPageNumber="624" pageId="16" pageNumber="622" type="diagnosis">
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<paragraph id="8B98783B79627E4CFC97699824ABFB12" blockId="16.[813,1453,1984,2105]" lastBlockId="18.[112,754,189,1205]" lastPageId="18" lastPageNumber="624" pageId="16" pageNumber="622">
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<emphasis id="B953A42979627E4EFC9769982690F87F" bold="true" box="[813,929,1984,2008]" pageId="16" pageNumber="622">Diagnosis.</emphasis>
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Carapace hexagonal; antero- and posterolateral margins clearly demarcated, anterolateral margin with 1 or 2 teeth; front weakly bilobed, almost smooth; posterolateral margin gently convex to sinuous (
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<figureCitation id="131C64BE79627E4EFB7E6678201EF79E" box="[1220,1327,2080,2105]" captionStart="Fig" captionStartId="19.[136,171,1920,1942]" captionTargetBox="[174,1436,189,1904]" captionTargetId="figure-7@19.[174,1436,189,1904]" captionTargetPageId="19" captionText="Fig. 11. Dorsal habitus (A, B, E, F, I–K); cephalothorax anterior view (C, G, H); D, left P4, ventral view. A, Trapezia cymodoce, male (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; B, Quadrella maculosa, male (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; C, D, Hexagonaloides bathyalis, holotype male (3.8 × 5.5 mm) (CBM-ZC 9935), Japan); E, G, Calocarcinus africanus, male (12.6 × 9.3 mm) (ZRC 2008.1281), New Caledonia; F, H, Philippicarcinus oviformis, male (16.0 × 11.3 mm) (ZRC 2009.0058), Philippines; I, Domecia acanthophora, male (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; J, Cherusius triunguiculatus (6.4 × 4.6 mm) (ZRC 2016.0121), Easter Island; K, Palmyria palmyrensis (4.9 × 3.4 mm) (UF 12897), Réunion. Scale = 0.5 mm. (C, after Komai et al., 2010: fig. 2B; D, courtesy of T. Komai)." figureDoi="http://doi.org/10.5281/zenodo.10271337" httpUri="https://zenodo.org/record/10271337/files/figure.png" pageId="16" pageNumber="622">Fig. 11E</figureCitation>
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); carapace relatively low to high, dorsal surface gently or distinctly convex in frontal view (
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<figureCitation id="131C64BE79607E4CFEC56E852735FF52" box="[383,516,221,245]" captionStart="Fig" captionStartId="19.[136,171,1920,1942]" captionTargetBox="[174,1436,189,1904]" captionTargetId="figure-7@19.[174,1436,189,1904]" captionTargetPageId="19" captionText="Fig. 11. Dorsal habitus (A, B, E, F, I–K); cephalothorax anterior view (C, G, H); D, left P4, ventral view. A, Trapezia cymodoce, male (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; B, Quadrella maculosa, male (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; C, D, Hexagonaloides bathyalis, holotype male (3.8 × 5.5 mm) (CBM-ZC 9935), Japan); E, G, Calocarcinus africanus, male (12.6 × 9.3 mm) (ZRC 2008.1281), New Caledonia; F, H, Philippicarcinus oviformis, male (16.0 × 11.3 mm) (ZRC 2009.0058), Philippines; I, Domecia acanthophora, male (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; J, Cherusius triunguiculatus (6.4 × 4.6 mm) (ZRC 2016.0121), Easter Island; K, Palmyria palmyrensis (4.9 × 3.4 mm) (UF 12897), Réunion. Scale = 0.5 mm. (C, after Komai et al., 2010: fig. 2B; D, courtesy of T. Komai)." figureDoi="http://doi.org/10.5281/zenodo.10271337" httpUri="https://zenodo.org/record/10271337/files/figure.png" pageId="18" pageNumber="624">Fig. 11E–H</figureCitation>
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); posterior margin of epistome with median lobe small, lateral margin relatively short, with 2 triangular lobes, separated by V-shaped cleft from pterygostomial lobe (
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<figureCitation id="131C64BE79607E4CFE2D6F6524DEFEF2" box="[407,495,317,341]" captionStart="Fig" captionStartId="8.[113,148,1170,1192]" captionTargetBox="[113,1451,189,1144]" captionTargetId="figure-13@8.[113,1451,189,1144]" captionTargetPageId="8" captionText="Fig. 5. Frontal view of cephalothorax showing epistome, orbits, antennules, and antenna. A, Tetralia cinctipes, male (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; B, Trapezia cymodoce, male (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; C, Sphenomerides trapezioides, male (7.8 × 5.9 mm) (ZRC 2000.2103), Madagascar; D, Calocarcinus africanus, male (9.3 × 7.2 mm) (ZRC 2008.1281), New Caledonia; E, Domecia acanthophora, male (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; F, Cherusius triunguiculatus, female (6.4 × 4.6 mm) (ZRC 2016.0121), Easter Island." figureDoi="http://doi.org/10.5281/zenodo.10271321" httpUri="https://zenodo.org/record/10271321/files/figure.png" pageId="18" pageNumber="624">Fig. 5D</figureCitation>
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). Basal antennal article quadrate, mobile, peduncle excluded from orbit. Maxilliped 3 merus quadrate, wider than long, anteroexternal part not auriculiform, as wide as rectangular ischium (at midlength); distal edge of exopod reaching to distal edge of merus (
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<figureCitation id="131C64BE79607E4CFD7F6FE525A2FE52" captionStart="Fig" captionStartId="8.[113,148,1170,1192]" captionTargetBox="[113,1451,189,1144]" captionTargetId="figure-13@8.[113,1451,189,1144]" captionTargetPageId="8" captionText="Fig. 5. Frontal view of cephalothorax showing epistome, orbits, antennules, and antenna. A, Tetralia cinctipes, male (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; B, Trapezia cymodoce, male (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; C, Sphenomerides trapezioides, male (7.8 × 5.9 mm) (ZRC 2000.2103), Madagascar; D, Calocarcinus africanus, male (9.3 × 7.2 mm) (ZRC 2008.1281), New Caledonia; E, Domecia acanthophora, male (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; F, Cherusius triunguiculatus, female (6.4 × 4.6 mm) (ZRC 2016.0121), Easter Island." figureDoi="http://doi.org/10.5281/zenodo.10271321" httpUri="https://zenodo.org/record/10271321/files/figure.png" pageId="18" pageNumber="624">Fig. 5E</figureCitation>
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). Chelipeds long, heterochelous, larger cheliped about twice carapace width; merus quadrate in dorsal view, flexor and extensor margins smooth; carpus inner angle with or without short spine, propodus outer surface smooth, without tubercles (
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<figureCitation id="131C64BE79607E4CFF586C052475FDD2" box="[226,324,605,629]" captionStart="Fig" captionStartId="19.[136,171,1920,1942]" captionTargetBox="[174,1436,189,1904]" captionTargetId="figure-7@19.[174,1436,189,1904]" captionTargetPageId="19" captionText="Fig. 11. Dorsal habitus (A, B, E, F, I–K); cephalothorax anterior view (C, G, H); D, left P4, ventral view. A, Trapezia cymodoce, male (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; B, Quadrella maculosa, male (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; C, D, Hexagonaloides bathyalis, holotype male (3.8 × 5.5 mm) (CBM-ZC 9935), Japan); E, G, Calocarcinus africanus, male (12.6 × 9.3 mm) (ZRC 2008.1281), New Caledonia; F, H, Philippicarcinus oviformis, male (16.0 × 11.3 mm) (ZRC 2009.0058), Philippines; I, Domecia acanthophora, male (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; J, Cherusius triunguiculatus (6.4 × 4.6 mm) (ZRC 2016.0121), Easter Island; K, Palmyria palmyrensis (4.9 × 3.4 mm) (UF 12897), Réunion. Scale = 0.5 mm. (C, after Komai et al., 2010: fig. 2B; D, courtesy of T. Komai)." figureDoi="http://doi.org/10.5281/zenodo.10271337" httpUri="https://zenodo.org/record/10271337/files/figure.png" pageId="18" pageNumber="624">Fig. 11E</figureCitation>
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). P2–5 short, longest pereopod slightly longer than carapace width; merus relatively stout; merus, carpus and propodus smooth, unarmed (
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<figureCitation id="131C64BE79607E4CFD986CC527B2FD12" box="[546,643,669,693]" captionStart="Fig" captionStartId="19.[136,171,1920,1942]" captionTargetBox="[174,1436,189,1904]" captionTargetId="figure-7@19.[174,1436,189,1904]" captionTargetPageId="19" captionText="Fig. 11. Dorsal habitus (A, B, E, F, I–K); cephalothorax anterior view (C, G, H); D, left P4, ventral view. A, Trapezia cymodoce, male (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; B, Quadrella maculosa, male (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; C, D, Hexagonaloides bathyalis, holotype male (3.8 × 5.5 mm) (CBM-ZC 9935), Japan); E, G, Calocarcinus africanus, male (12.6 × 9.3 mm) (ZRC 2008.1281), New Caledonia; F, H, Philippicarcinus oviformis, male (16.0 × 11.3 mm) (ZRC 2009.0058), Philippines; I, Domecia acanthophora, male (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; J, Cherusius triunguiculatus (6.4 × 4.6 mm) (ZRC 2016.0121), Easter Island; K, Palmyria palmyrensis (4.9 × 3.4 mm) (UF 12897), Réunion. Scale = 0.5 mm. (C, after Komai et al., 2010: fig. 2B; D, courtesy of T. Komai)." figureDoi="http://doi.org/10.5281/zenodo.10271337" httpUri="https://zenodo.org/record/10271337/files/figure.png" pageId="18" pageNumber="624">Fig. 11E</figureCitation>
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); dactylus flexor margin with robust setae; without transverse, comb-like rows of setae. Sternopleonal cavity reaching anteriorly to level of midlength of P2 coxae (
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<figureCitation id="131C64BE79607E4CFE786CA52703FCB2" box="[450,562,765,789]" captionStart="Fig" captionStartId="11.[136,171,1561,1583]" captionTargetBox="[136,1474,189,1546]" captionTargetId="figure-8@11.[136,1474,189,1546]" captionTargetPageId="11" captionText="Fig.7. Male anterior thoracic sternum and pleon.A,Tetralia cinctipes (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; B, Tetraloides heterodactylus (7.6 × 7.1 mm) (ZRC 2019.0699), Japan; C, Trapezia cymodoce (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; D, Quadrella maculosa (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; E, Sphenomerides trapezioides (7.8 × 5.9 mm) (ZRC 2000.2103), Madagascar; F, Calocarcinus africanus (9.3 × 7.2 mm) (ZRC 2008.1281), New Caledonia; G, Philippicarcinus oviformis (16.0 × 11.3 mm) (ZRC 2009.0058), Philippines; H, Domecia acanthophora (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; I, Cherusius triunguiculatus (4.7 × 3.9 mm) (ZRC 2016.0121), Easter Island." figureDoi="http://doi.org/10.5281/zenodo.10271327" httpUri="https://zenodo.org/record/10271327/files/figure.png" pageId="18" pageNumber="624">Fig. 7F, G</figureCitation>
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); tubercle of male pleonal-locking mechanism distinct, on posterior one-third of thoracic sternite 5, may be near suture with sternite 6 (
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<figureCitation id="131C64BE79607E4CFD796D6525A6FCD2" captionStart="Fig" captionStartId="13.[136,171,1937,1959]" captionTargetBox="[144,1464,189,1923]" captionTargetId="figure-8@13.[144,1464,189,1923]" captionTargetPageId="13" captionText="Fig. 8. Posterior part of male thoracic sternum and pleon (A–F); sternopleonal cavity showing tubercle of pleonal-locking mechanism on sternite 5 (G–L). A, G, Tetralia cinctipes, male (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; B, Trapezia cymodoce (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; C, I, Quadrella maculosa (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; D, J, Sphenomerides trapezioides (7.8 × 5.9 mm) (ZRC 2000.2103), Madagascar; E, Calocarcinus africanus (9.3 × 7.2 mm) (ZRC 2008.1281), New Caledonia; F, L. Domecia acanthophora (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; H, Tetraloides heterodactylus (7.6 × 7.1 mm) (ZRC 2019.0699), Japan; K, Calocarcinus africanus (12.6 × 9.3 mm) (ZRC 2008.1281), New Caledonia." figureDoi="http://doi.org/10.5281/zenodo.10271329" httpUri="https://zenodo.org/record/10271329/files/figure.png" pageId="18" pageNumber="624">Fig. 8K</figureCitation>
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).
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<collectingRegion id="49E3B6D979607E4CFF146D0525D4FCD2" box="[174,229,861,885]" country="Maldives" name="Maale" pageId="18" pageNumber="624">Male</collectingRegion>
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pleon subquadrate; linguiform, lateral margins of somites 3–6 subparallel, somites 3–5 fused with sutures visible medially (
|
||
<figureCitation id="131C64BE79607E4CFE886DC52482FC12" box="[306,435,925,949]" captionStart="Fig" captionStartId="11.[136,171,1561,1583]" captionTargetBox="[136,1474,189,1546]" captionTargetId="figure-8@11.[136,1474,189,1546]" captionTargetPageId="11" captionText="Fig.7. Male anterior thoracic sternum and pleon.A,Tetralia cinctipes (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; B, Tetraloides heterodactylus (7.6 × 7.1 mm) (ZRC 2019.0699), Japan; C, Trapezia cymodoce (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; D, Quadrella maculosa (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; E, Sphenomerides trapezioides (7.8 × 5.9 mm) (ZRC 2000.2103), Madagascar; F, Calocarcinus africanus (9.3 × 7.2 mm) (ZRC 2008.1281), New Caledonia; G, Philippicarcinus oviformis (16.0 × 11.3 mm) (ZRC 2009.0058), Philippines; H, Domecia acanthophora (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; I, Cherusius triunguiculatus (4.7 × 3.9 mm) (ZRC 2016.0121), Easter Island." figureDoi="http://doi.org/10.5281/zenodo.10271327" httpUri="https://zenodo.org/record/10271327/files/figure.png" pageId="18" pageNumber="624">Figs. 7F, G</figureCitation>
|
||
,
|
||
<figureCitation id="131C64BE79607E4CFE7A6DC524D0FC12" box="[448,481,925,949]" captionStart="Fig" captionStartId="13.[136,171,1937,1959]" captionTargetBox="[144,1464,189,1923]" captionTargetId="figure-8@13.[144,1464,189,1923]" captionTargetPageId="13" captionText="Fig. 8. Posterior part of male thoracic sternum and pleon (A–F); sternopleonal cavity showing tubercle of pleonal-locking mechanism on sternite 5 (G–L). A, G, Tetralia cinctipes, male (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; B, Trapezia cymodoce (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; C, I, Quadrella maculosa (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; D, J, Sphenomerides trapezioides (7.8 × 5.9 mm) (ZRC 2000.2103), Madagascar; E, Calocarcinus africanus (9.3 × 7.2 mm) (ZRC 2008.1281), New Caledonia; F, L. Domecia acanthophora (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; H, Tetraloides heterodactylus (7.6 × 7.1 mm) (ZRC 2019.0699), Japan; K, Calocarcinus africanus (12.6 × 9.3 mm) (ZRC 2008.1281), New Caledonia." figureDoi="http://doi.org/10.5281/zenodo.10271329" httpUri="https://zenodo.org/record/10271329/files/figure.png" pageId="18" pageNumber="624">8E</figureCitation>
|
||
). G1 strongly sinuous or curved, relatively slender to stout, distal margins lined with short or long spinules (
|
||
<figureCitation id="131C64BE79607E4CFECB6D8524C1FC52" box="[369,496,989,1013]" captionStart="Fig" captionStartId="15.[136,171,1520,1542]" captionTargetBox="[142,1479,192,1505]" captionTargetId="figure-8@15.[142,1491,186,1505]" captionTargetPageId="15" captionText="Fig. 9. Left G1 and G2, pleonal view. A, B, Tetralia cinctipes (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; C, D, Tetraloides heterodactylus (7.6 × 7.1 mm) (ZRC 2019.0699), Japan; E, F,Trapezia cymodoce (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; G, H, Quadrella maculosa (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; I, J, Sphenomerides trapezioides (7.8 × 5.9 mm) (ZRC 2000.2103), Madagascar; K, L, Calocarcinus africanus (12.6 × 9.3 mm) (ZRC 2008.1281), New Caledonia; M, N, Philippicarcinus oviformis (16.0 × 11.3 mm) (ZRC 2009.0058), Philippines; O, P, Domecia acanthophora (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; Q, R, Cherusius triunguiculatus (4.7 × 3.9 mm) (ZRC 2016.0121), Easter Island. A, C, E, G, I, K, M, O, Q, left G1; B, D, F, H, J, L, N, P, R, left G2." figureDoi="http://doi.org/10.5281/zenodo.10271331" httpUri="https://zenodo.org/record/10271331/files/figure.png" pageId="18" pageNumber="624">Fig. 9K, M</figureCitation>
|
||
). G2 relatively slender, three-quarters to subequal to length of G1, distal portion elongated, filiform (
|
||
<figureCitation id="131C64BE79607E4CFEE26A4524E6FB92" box="[344,471,1053,1077]" captionStart="Fig" captionStartId="15.[136,171,1520,1542]" captionTargetBox="[142,1479,192,1505]" captionTargetId="figure-8@15.[142,1491,186,1505]" captionTargetPageId="15" captionText="Fig. 9. Left G1 and G2, pleonal view. A, B, Tetralia cinctipes (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; C, D, Tetraloides heterodactylus (7.6 × 7.1 mm) (ZRC 2019.0699), Japan; E, F,Trapezia cymodoce (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; G, H, Quadrella maculosa (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; I, J, Sphenomerides trapezioides (7.8 × 5.9 mm) (ZRC 2000.2103), Madagascar; K, L, Calocarcinus africanus (12.6 × 9.3 mm) (ZRC 2008.1281), New Caledonia; M, N, Philippicarcinus oviformis (16.0 × 11.3 mm) (ZRC 2009.0058), Philippines; O, P, Domecia acanthophora (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; Q, R, Cherusius triunguiculatus (4.7 × 3.9 mm) (ZRC 2016.0121), Easter Island. A, C, E, G, I, K, M, O, Q, left G1; B, D, F, H, J, L, N, P, R, left G2." figureDoi="http://doi.org/10.5281/zenodo.10271331" httpUri="https://zenodo.org/record/10271331/files/figure.png" pageId="18" pageNumber="624">Fig. 9L, N</figureCitation>
|
||
). Vulva relatively large, ovate to triangular, positioned on anterior half of sternite 6, adjacent to suture with sternite 5, vulvae positioned close to median line, directed inwards; large lateral, flexible sternal vulvar cover (
|
||
<figureCitation id="131C64BE79607E4CFEB06AC524BFFB12" box="[266,398,1181,1205]" captionStart="Fig" captionStartId="17.[136,171,1911,1933]" captionTargetBox="[136,1474,227,1896]" captionTargetId="figure-115@17.[136,1474,227,1896]" captionTargetPageId="17" captionText="Fig. 10. Vulvae. A, Tetralia nigrolineata (9.8 × 8.6 mm) (ZRC 2019.0701), Japan; B, Tetraloides heterodactylus (8.3 × 6.7 mm) (ZRC 2019.0699), Japan; C, Trapezia cymodoce (12.2 × 10.1 mm) (ZRC 2007.0079), Singapore; D, Quadrella maculosa (5.9 × 5.0 mm) (ZRC 2003.0352), Sulawesi, Indonesia; E, Sphenomerides trapezioides (7.5 × 5.7 mm) (ZRC 2000.2103), Madagascar; F, Calocarcinus africanus (12.0 × 10.1 mm) (ZRC 2008.1281), New Caledonia; G, Philippicarcinus oviformis (14.5 × 10.0 mm) (ZRC 2009.0058), Philippines; H, Domecia acanthophora (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; I, Cherusius triunguiculatus (6.4 × 4.6 mm) (ZRC 2016.0121), Easter Island; J, Palmyria palmyrensis (4.9 × 3.4 mm) (UF 12897), Réunion." figureDoi="http://doi.org/10.5281/zenodo.10271335" httpUri="https://zenodo.org/record/10271335/files/figure.png" pageId="18" pageNumber="624">Fig. 10F, G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF5828B379637E4FFF32692F26CAF7BA" ID-DOI="http://doi.org/10.5281/zenodo.10271335" ID-Zenodo-Dep="10271335" httpUri="https://zenodo.org/record/10271335/files/figure.png" pageId="17" pageNumber="623" startId="17.[136,171,1911,1933]" targetBox="[136,1474,227,1896]" targetPageId="17" targetType="figure">
|
||
<paragraph id="8B98783B79637E4FFF32692F26CAF7BA" blockId="17.[136,1474,1911,2077]" pageId="17" pageNumber="623">
|
||
Fig. 10. Vulvae. A,
|
||
<taxonomicName id="4C2703B879637E4FFEF5692F272AF82B" authorityName="Serene & Pham" authorityYear="1957" box="[335,539,1911,1932]" class="Malacostraca" family="Tetraliidae" genus="Tetralia" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="623" phylum="Arthropoda" rank="species" species="nigrolineata">
|
||
<emphasis id="B953A42979637E4FFEF5692F272AF82B" box="[335,539,1911,1932]" italics="true" pageId="17" pageNumber="623">Tetralia nigrolineata</emphasis>
|
||
</taxonomicName>
|
||
(9.8 × 8.6 mm) (ZRC 2019.0701), Japan; B,
|
||
<taxonomicName id="4C2703B879637E4FFC5B692F21D5F82B" baseAuthorityName="Heller" baseAuthorityYear="1861" box="[993,1252,1911,1932]" class="Malacostraca" family="Tetraliidae" genus="Tetraloides" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="623" phylum="Arthropoda" rank="species" species="heterodactylus">
|
||
<emphasis id="B953A42979637E4FFC5B692F21D5F82B" box="[993,1252,1911,1932]" italics="true" pageId="17" pageNumber="623">Tetraloides heterodactylus</emphasis>
|
||
</taxonomicName>
|
||
(8.3 × 6.7 mm) (ZRC 2019.0699), Japan; C,
|
||
<taxonomicName id="4C2703B879637E4FFEDC69CD2713F80E" baseAuthorityName="Herbst" baseAuthorityYear="1801" box="[358,546,1940,1962]" class="Malacostraca" family="Trapeziidae" genus="Trapezia" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="623" phylum="Arthropoda" rank="species" species="cymodoce">
|
||
<emphasis id="B953A42979637E4FFEDC69CD2713F80E" box="[358,546,1940,1962]" italics="true" pageId="17" pageNumber="623">Trapezia cymodoce</emphasis>
|
||
</taxonomicName>
|
||
(12.2 × 10.1 mm) (ZRC 2007.0079), Singapore; D,
|
||
<taxonomicName id="4C2703B879637E4FFB9E69CC21DBF80E" authorityName="Alcock" authorityYear="1898" box="[1060,1258,1940,1961]" class="Malacostraca" family="Trapeziidae" genus="Quadrella" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="623" phylum="Arthropoda" rank="species" species="maculosa">
|
||
<emphasis id="B953A42979637E4FFB9E69CC21DBF80E" box="[1060,1258,1940,1961]" italics="true" pageId="17" pageNumber="623">Quadrella maculosa</emphasis>
|
||
</taxonomicName>
|
||
(5.9 × 5.0 mm) (ZRC 2003.0352), Sulawesi, Indonesia; E,
|
||
<taxonomicName id="4C2703B879637E4FFE5269E927C4F861" baseAuthorityName="Wood-Mason in Wood-Mason & Alcock" baseAuthorityYear="1891" box="[488,757,1969,1990]" class="Malacostraca" family="Trapeziidae" genus="Sphenomerides" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="623" phylum="Arthropoda" rank="species" species="trapezioides">
|
||
<emphasis id="B953A42979637E4FFE5269E927C4F861" box="[488,757,1969,1990]" italics="true" pageId="17" pageNumber="623">Sphenomerides trapezioides</emphasis>
|
||
</taxonomicName>
|
||
(7.5 × 5.7 mm) (ZRC 2000.2103), Madagascar; F,
|
||
<taxonomicName id="4C2703B879637E4FFB5A69E920F3F861" authorityName=", Calman" authorityYear="1909" box="[1248,1474,1969,1990]" class="Malacostraca" family="Trapeziidae" genus="Calocarcinus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="623" phylum="Arthropoda" rank="species" species="africanus">
|
||
<emphasis id="B953A42979637E4FFB5A69E920F3F861" box="[1248,1474,1969,1990]" italics="true" pageId="17" pageNumber="623">Calocarcinus africanus</emphasis>
|
||
</taxonomicName>
|
||
(12.0 × 10.1 mm) (ZRC 2008.1281), New Caledonia; G,
|
||
<taxonomicName id="4C2703B879637E4FFD006995268CF844" authorityName="Garth & Kim" authorityYear="1983" box="[698,957,1997,2019]" class="Malacostraca" family="Trapeziidae" genus="Philippicarcinus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="623" phylum="Arthropoda" rank="species" species="oviformis">
|
||
<emphasis id="B953A42979637E4FFD006995268CF844" box="[698,957,1997,2019]" italics="true" pageId="17" pageNumber="623">Philippicarcinus oviformis</emphasis>
|
||
</taxonomicName>
|
||
(14.5 × 10.0 mm) (ZRC 2009.0058), Philippines; H,
|
||
<taxonomicName id="4C2703B879637E4FFF3369B32441F858" baseAuthorityName="Desbonne in Desbonne & Schramm" baseAuthorityYear="1867" box="[137,368,2026,2048]" class="Malacostraca" family="Domeciidae" genus="Domecia" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="623" phylum="Arthropoda" rank="species" species="acanthophora">
|
||
<emphasis id="B953A42979637E4FFF3369B32441F858" box="[137,368,2026,2048]" italics="true" pageId="17" pageNumber="623">Domecia acanthophora</emphasis>
|
||
</taxonomicName>
|
||
(7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; I,
|
||
<taxonomicName id="4C2703B879637E4FFC5E69B221D0F858" baseAuthorityName="Borradaile" baseAuthorityYear="1902" box="[996,1249,2026,2047]" class="Malacostraca" family="Domeciidae" genus="Cherusius" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="623" phylum="Arthropoda" rank="species" species="triunguiculatus">
|
||
<emphasis id="B953A42979637E4FFC5E69B221D0F858" box="[996,1249,2026,2047]" italics="true" pageId="17" pageNumber="623">Cherusius triunguiculatus</emphasis>
|
||
</taxonomicName>
|
||
(6.4 × 4.6 mm) (ZRC 2016.0121), Easter Island; J,
|
||
<taxonomicName id="4C2703B879637E4FFE13665F274FF7BB" baseAuthorityName="Rathbun in Edmondson" baseAuthorityYear="1923" box="[425,638,2055,2076]" class="Malacostraca" family="Domeciidae" genus="Palmyria" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="623" phylum="Arthropoda" rank="species" species="palmyrensis">
|
||
<emphasis id="B953A42979637E4FFE13665F274FF7BB" box="[425,638,2055,2076]" italics="true" pageId="17" pageNumber="623">Palmyria palmyrensis</emphasis>
|
||
</taxonomicName>
|
||
(4.9 × 3.4 mm) (UF 12897), Réunion.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B98783B79607E4CFFCB6A85273BFAB2" blockId="18.[113,751,1245,1301]" pageId="18" pageNumber="624">
|
||
<emphasis id="B953A42979607E4CFFCB6A85247FFB52" bold="true" box="[113,334,1245,1269]" pageId="18" pageNumber="624">Genera included.</emphasis>
|
||
<taxonomicName id="4C2703B879607E4CFEE16A852784FB52" authority="Calman, 1909" authorityName="Calman" authorityYear="1909" box="[347,693,1245,1269]" class="Malacostraca" family="Trapeziidae" genus="Calocarcinus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="624" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B953A42979607E4CFEE16A8524CAFB52" box="[347,507,1245,1269]" italics="true" pageId="18" pageNumber="624">Calocarcinus</emphasis>
|
||
<bibRefCitation id="EFB605CA79607E4CFDB26A852784FB52" author="Calman WT" box="[520,693,1245,1269]" pageId="18" pageNumber="624" pagination="30 - 33" refId="ref18287" refString="Calman WT (1909) On a new crab taken from a deep-sea telegraphcable in the Indian Ocean. Annals and Magazine of Natural History, Series 8, 3 (13): 30 - 33, 1 fig." type="journal article" year="1909">Calman, 1909</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C2703B879607E4CFFCB6AA52737FAB2" authority="Garth & Kim, 1983" authorityName="Garth & Kim" authorityYear="1983" box="[113,518,1277,1301]" class="Malacostraca" family="Trapeziidae" genus="Philippicarcinus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="624" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B953A42979607E4CFFCB6AA52412FAB2" box="[113,291,1277,1301]" italics="true" pageId="18" pageNumber="624">Philippicarcinus</emphasis>
|
||
<bibRefCitation id="EFB605CA79607E4CFE966AA52737FAB2" author="Garth JS & Kim HS" box="[300,518,1277,1301]" pageId="18" pageNumber="624" pagination="663 - 729" refId="ref19847" refString="Garth JS & Kim HS (1983) Crabs of the family Xanthidae (Crustacea: Brachyura) from the Philippine Islands and adjacent waters based largely on collections of the U. S. Fish Commission steamer Albatross in 1908 - 1909. Journal of Natural History, 17: 663 - 729." type="journal article" year="1983">Garth & Kim, 1983</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D2BB079607E4CFFCB6B6526E1FC32" pageId="18" pageNumber="624" type="discussion">
|
||
<paragraph id="8B98783B79607E4CFFCB6B6525EFF892" blockId="18.[113,753,1341,1845]" pageId="18" pageNumber="624">
|
||
<emphasis id="B953A42979607E4CFFCB6B6525D3FAF2" bold="true" box="[113,226,1341,1365]" pageId="18" pageNumber="624">Remarks.</emphasis>
|
||
In their discussion of the higher classification of trapezioids,
|
||
<bibRefCitation id="EFB605CA79607E4CFE936B05271EFAD2" author="Lai JCY & Ahyong ST & Jeng M-S & Ng PKL" box="[297,559,1373,1397]" pageId="18" pageNumber="624" pagination="402 - 408" refId="ref20310" refString="Lai JCY, Ahyong ST, Jeng M-S & Ng PKL (2009) Are coraldwelling crabs monophyletic? A phylogeny of the Trapezioidea (Crustacea: Decapoda: Brachyura). Invertebrate Systematics, 23: 402 - 408." type="journal article" year="2009">Lai et al. (2009: 407)</bibRefCitation>
|
||
opined that the “phylogenetic position of Calocarcinines raises additional issues”. They noted that while the Calocarcininae is a stable monophyletic clade, the support for its affinity to
|
||
<taxonomicName id="4C2703B879607E4CFDC96BE527C1FA72" authorityName="Miers" authorityYear="1886" box="[627,752,1469,1493]" pageId="18" pageNumber="620" rank="subFamily" subFamily="Trapeziinae">Trapeziinae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2703B879607E4CFF1B6B85241AFA52" authorityName="Stevcic" authorityYear="2005" box="[161,299,1501,1525]" class="Anthozoa" family="Pocilloporidae" kingdom="Animalia" order="Scleractinia" pageId="18" pageNumber="620" phylum="Cnidaria" rank="subFamily" subFamily="Quadrellinae">Quadrellinae</taxonomicName>
|
||
is not high in MP analyses (though well supported in BI, PP = 1.0). This lends some support to
|
||
<bibRefCitation id="EFB605CA79607E4CFFCB68452478F992" author="Schweitzer C" box="[113,329,1565,1589]" pageId="18" pageNumber="624" pagination="625 - 636" refId="ref21582" refString="Schweitzer C (2005) The Trapeziidae and Domeciidae (Decapoda: Brachyura: Xanthoidea) in the fossil record and a new Eocene genus from Baja California Sur, Mexico. Journal of Crustacean Biology, 25: 625 - 636." type="journal article" year="2005">Schweitzer (2005)</bibRefCitation>
|
||
, who suggested that Calocarcininae might not belong in
|
||
<taxonomicName id="4C2703B879607E4CFEEB686524E3F9F2" authorityName="Miers" authorityYear="1886" box="[337,466,1597,1621]" class="Malacostraca" family="Trapeziidae" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="624" phylum="Arthropoda" rank="family">Trapeziidae</taxonomicName>
|
||
. A similar hypothesis was proposed by
|
||
<bibRefCitation id="EFB605CA79607E4CFEB768052715F9D2" author="Clark PF & Guerao G" box="[269,548,1629,1653]" pageId="18" pageNumber="624" pagination="475 - 500" refId="ref18820" refString="Clark PF & Guerao G (2008) A description of Calocarcinus africanus Calman, 1909 (Brachyura, Xanthoidea) first zoeal stage morphology with implications for Trapeziidae systematics. Proceedings of the Biological Society of Washington, 121: 475 - 500." type="journal article" year="2008">Clark & Guerao (2008)</bibRefCitation>
|
||
, who commented that calocarcinine larvae are very different from those of trapeziines. Although these hypotheses might be correct, an alternative possibility is that the more typically ‘xanthoid’ carapace of the calocarcinines, as well as their xanthoid or eriphioid-like larvae, merely reflects their plesiomorphic condition.
|
||
</paragraph>
|
||
<paragraph id="8B98783B79607E4CFFCB690526C2FDB2" blockId="18.[112,754,1885,2102]" lastBlockId="18.[811,1454,189,533]" pageId="18" pageNumber="624">
|
||
The morphological reappraisal herein supports the suggestions of
|
||
<bibRefCitation id="EFB605CA79607E4CFF346925240EF832" author="Lai JCY & Ahyong ST & Jeng M-S & Ng PKL" box="[142,319,1917,1941]" pageId="18" pageNumber="624" pagination="402 - 408" refId="ref20310" refString="Lai JCY, Ahyong ST, Jeng M-S & Ng PKL (2009) Are coraldwelling crabs monophyletic? A phylogeny of the Trapezioidea (Crustacea: Decapoda: Brachyura). Invertebrate Systematics, 23: 402 - 408." type="journal article" year="2009">Lai et al. (2009)</bibRefCitation>
|
||
that
|
||
<taxonomicName id="4C2703B879607E4CFEC369252738F832" authorityName="Calman" authorityYear="1909" box="[377,521,1917,1941]" class="Malacostraca" family="Trapeziidae" genus="Calocarcinus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="624" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B953A42979607E4CFEC369252738F832" box="[377,521,1917,1941]" italics="true" pageId="18" pageNumber="624">Calocarcinus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2703B879607E4CFDFA692527C3F832" authorityName="Garth & Kim" authorityYear="1983" box="[576,754,1917,1941]" class="Malacostraca" family="Trapeziidae" genus="Philippicarcinus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="624" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B953A42979607E4CFDFA692527C3F832" box="[576,754,1917,1941]" italics="true" pageId="18" pageNumber="624">Philippicarcinus</emphasis>
|
||
</taxonomicName>
|
||
should not be placed in
|
||
<taxonomicName id="4C2703B879607E4CFE3669C5273EF812" authorityName="Miers" authorityYear="1886" box="[396,527,1949,1973]" class="Malacostraca" family="Trapeziidae" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="624" phylum="Arthropoda" rank="family">Trapeziidae</taxonomicName>
|
||
and should instead be treated as a separate family. As discussed earlier for
|
||
<taxonomicName id="4C2703B879607E4CFFCB6985241BF852" authorityName="Stevcic" authorityYear="2005" box="[113,298,2013,2037]" pageId="18" pageNumber="621" rank="subFamily" subFamily="Sphenomeridinae">Sphenomeridinae</taxonomicName>
|
||
, these genera do not have comb-like rows of feeding setae on their P2–5 dactyli. Most significantly, their G2 is very long, sometimes subequal to the length of G1 (
|
||
<figureCitation id="131C64BE79607E4CFD7F66452641FF72" box="[709,880,189,2101]" captionStart="Fig" captionStartId="15.[136,171,1520,1542]" captionTargetBox="[142,1479,192,1505]" captionTargetId="figure-8@15.[142,1491,186,1505]" captionTargetPageId="15" captionText="Fig. 9. Left G1 and G2, pleonal view. A, B, Tetralia cinctipes (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; C, D, Tetraloides heterodactylus (7.6 × 7.1 mm) (ZRC 2019.0699), Japan; E, F,Trapezia cymodoce (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; G, H, Quadrella maculosa (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; I, J, Sphenomerides trapezioides (7.8 × 5.9 mm) (ZRC 2000.2103), Madagascar; K, L, Calocarcinus africanus (12.6 × 9.3 mm) (ZRC 2008.1281), New Caledonia; M, N, Philippicarcinus oviformis (16.0 × 11.3 mm) (ZRC 2009.0058), Philippines; O, P, Domecia acanthophora (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; Q, R, Cherusius triunguiculatus (4.7 × 3.9 mm) (ZRC 2016.0121), Easter Island. A, C, E, G, I, K, M, O, Q, left G1; B, D, F, H, J, L, N, P, R, left G2." figureDoi="http://doi.org/10.5281/zenodo.10271331" httpUri="https://zenodo.org/record/10271331/files/figure.png" pageId="18" pageNumber="624">Fig. 9L, N</figureCitation>
|
||
). In all other trapeziids and tetraliids, G2 is at most half the length of G1 (
|
||
<figureCitation id="131C64BE79607E4CFB996E8521C7FF52" box="[1059,1270,221,245]" captionStart="Fig" captionStartId="15.[136,171,1520,1542]" captionTargetBox="[142,1479,192,1505]" captionTargetId="figure-8@15.[142,1491,186,1505]" captionTargetPageId="15" captionText="Fig. 9. Left G1 and G2, pleonal view. A, B, Tetralia cinctipes (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; C, D, Tetraloides heterodactylus (7.6 × 7.1 mm) (ZRC 2019.0699), Japan; E, F,Trapezia cymodoce (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; G, H, Quadrella maculosa (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; I, J, Sphenomerides trapezioides (7.8 × 5.9 mm) (ZRC 2000.2103), Madagascar; K, L, Calocarcinus africanus (12.6 × 9.3 mm) (ZRC 2008.1281), New Caledonia; M, N, Philippicarcinus oviformis (16.0 × 11.3 mm) (ZRC 2009.0058), Philippines; O, P, Domecia acanthophora (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; Q, R, Cherusius triunguiculatus (4.7 × 3.9 mm) (ZRC 2016.0121), Easter Island. A, C, E, G, I, K, M, O, Q, left G1; B, D, F, H, J, L, N, P, R, left G2." figureDoi="http://doi.org/10.5281/zenodo.10271331" httpUri="https://zenodo.org/record/10271331/files/figure.png" pageId="18" pageNumber="624">Fig. 9B, D, F, H, J</figureCitation>
|
||
). In
|
||
<taxonomicName id="4C2703B879607E4CFA906E85209AFF52" authorityName="Ortmann" authorityYear="1893" box="[1322,1451,221,245]" class="Malacostraca" family="Domeciidae" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="624" phylum="Arthropoda" rank="family">Domeciidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2703B879607E4CFCE56EA52137FEB2" authorityName="Ng & Ahyong & Castro" authorityYear="2023" box="[863,1030,253,277]" class="Malacostraca" family="Ectaesthesiidae" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="624" phylum="Arthropoda" rank="family">Ectaesthesiidae</taxonomicName>
|
||
, G2 is two-thirds to three-quarters the length of G1, but their G1 forms are completely different (
|
||
<figureCitation id="131C64BE79607E4CFC8F6F652688FEF2" box="[821,953,317,341]" captionStart="Fig" captionStartId="15.[136,171,1520,1542]" captionTargetBox="[142,1479,192,1505]" captionTargetId="figure-8@15.[142,1491,186,1505]" captionTargetPageId="15" captionText="Fig. 9. Left G1 and G2, pleonal view. A, B, Tetralia cinctipes (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; C, D, Tetraloides heterodactylus (7.6 × 7.1 mm) (ZRC 2019.0699), Japan; E, F,Trapezia cymodoce (11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; G, H, Quadrella maculosa (7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; I, J, Sphenomerides trapezioides (7.8 × 5.9 mm) (ZRC 2000.2103), Madagascar; K, L, Calocarcinus africanus (12.6 × 9.3 mm) (ZRC 2008.1281), New Caledonia; M, N, Philippicarcinus oviformis (16.0 × 11.3 mm) (ZRC 2009.0058), Philippines; O, P, Domecia acanthophora (7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; Q, R, Cherusius triunguiculatus (4.7 × 3.9 mm) (ZRC 2016.0121), Easter Island. A, C, E, G, I, K, M, O, Q, left G1; B, D, F, H, J, L, N, P, R, left G2." figureDoi="http://doi.org/10.5281/zenodo.10271331" httpUri="https://zenodo.org/record/10271331/files/figure.png" pageId="18" pageNumber="624">Figs. 9O–R</figureCitation>
|
||
). As noted above,
|
||
<bibRefCitation id="EFB605CA79607E4CFB376F66201FFEF2" author="Stevcic Z" box="[1165,1326,317,344]" pageId="18" pageNumber="624" pagination="1 - 159" refId="ref21857" refString="Stevcic Z (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Stevcic Brachyura). Natura Croatica (Fauna Croatica), 14 (1): 1 - 159." type="journal article" year="2005">Števčić (2005)</bibRefCitation>
|
||
recognised a separate trapeziid tribe, Philippicarcinini
|
||
<bibRefCitation id="EFB605CA79607E4CFAB56F062096FED2" author="Stevcic Z" box="[1295,1447,349,376]" pageId="18" pageNumber="624" pagination="1 - 159" refId="ref21857" refString="Stevcic Z (2005) The reclassification of Brachyuran Crabs (Crustacea: Decapoda: Stevcic Brachyura). Natura Croatica (Fauna Croatica), 14 (1): 1 - 159." type="journal article" year="2005">Števčić, 2005</bibRefCitation>
|
||
, for
|
||
<taxonomicName id="4C2703B879607E4CFCED6F25213EFE32" authorityName="Garth & Kim" authorityYear="1983" box="[855,1039,381,405]" class="Malacostraca" family="Trapeziidae" genus="Philippicarcinus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="624" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B953A42979607E4CFCED6F25213EFE32" box="[855,1039,381,405]" italics="true" pageId="18" pageNumber="624">Philippicarcinus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C2703B879607E4CFB986F252187FE32" authorityName="Calman" authorityYear="1909" box="[1058,1206,381,405]" class="Malacostraca" family="Trapeziidae" genus="Calocarcinus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="624" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B953A42979607E4CFB986F252187FE32" box="[1058,1206,381,405]" italics="true" pageId="18" pageNumber="624">Calocarcinus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2703B879607E4CFB496F25209AFE32" authorityName="Garth & Kim" authorityYear="1983" box="[1267,1451,381,405]" class="Malacostraca" family="Trapeziidae" genus="Philippicarcinus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="624" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B953A42979607E4CFB496F25209AFE32" box="[1267,1451,381,405]" italics="true" pageId="18" pageNumber="624">Philippicarcinus</emphasis>
|
||
</taxonomicName>
|
||
are so similar that there is no need to separate them into two subfamilies. As a result, on the morphological evidence available, Philippicarcinini is a junior synonym of
|
||
<taxonomicName id="4C2703B879607E4CFCF16FA526DEFDB2" authorityName="Stevcic" authorityYear="2005" box="[843,1007,509,533]" class="Malacostraca" family="Calocarcinidae" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="624" phylum="Arthropoda" rank="family">Calocarcinidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B98783B79607E4CFC976C652059FD51" blockId="18.[812,1452,573,758]" pageId="18" pageNumber="624">
|
||
<bibRefCitation id="EFB605CA79607E4CFC976C652130FDF2" author="Castro P" box="[813,1025,573,597]" pageId="18" pageNumber="624" pagination="59 - 107" refId="ref18418" refString="Castro P (1997 b) Trapeziid crabs (Brachyura: Xanthoidea: Trapeziidae) of New Caledonia, eastern Australia, and the Coral Sea. In: Richer de Forges B (ed.) Les fonds meubles des lagons de Nouvelle-Caledonie (sedimentologie, benthos). Etudes & Theses, 3: 59 - 107, pls. 1 - 7." type="journal article" year="1997">Castro (1997b: 62)</bibRefCitation>
|
||
indicated that
|
||
<taxonomicName id="4C2703B879607E4CFB166C652139FDD2" authority="Galil & Clark, 1990" authorityName="Galil & Clark" authorityYear="1990" class="Malacostraca" family="Trapeziidae" genus="Calocarcinus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="624" phylum="Arthropoda" rank="species" species="crosnieri">
|
||
<emphasis id="B953A42979607E4CFB166C65209AFDF2" box="[1196,1451,573,597]" italics="true" pageId="18" pageNumber="624">Calocarcinus crosnieri</emphasis>
|
||
<bibRefCitation id="EFB605CA79607E4CFC976C052139FDD2" author="Galil BS & Clark PF" box="[813,1032,605,629]" pageId="18" pageNumber="624" pagination="369 - 388" refId="ref19702" refString="Galil BS & Clark PF (1990) Crustacea Decapoda: Notes on trapeziid crabs from New Caledonia including descriptions of two new species. In: Crosnier A (ed.) Resultats des campagnes MUSORSTOM, Volume 6. Memoires du Museum national d'Histoire naturelle, Serie A, 145: 369 - 388." type="journal article" year="1990">Galil & Clark, 1990</bibRefCitation>
|
||
</taxonomicName>
|
||
was probably a junior synonym of
|
||
<taxonomicName id="4C2703B879607E4CFA296C062105FD32" authority=", Calman, 1909" authorityName=", Calman" authorityYear="1909" class="Malacostraca" family="Trapeziidae" genus="Calocarcinus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="624" phylum="Arthropoda" rank="species" species="africanus">
|
||
<emphasis id="B953A42979607E4CFA296C0626A3FD32" italics="true" pageId="18" pageNumber="624">C. africanus</emphasis>
|
||
,
|
||
<bibRefCitation id="EFB605CA79607E4CFC256C252105FD32" author="Calman WT" box="[927,1076,637,661]" pageId="18" pageNumber="624" pagination="30 - 33" refId="ref18287" refString="Calman WT (1909) On a new crab taken from a deep-sea telegraphcable in the Indian Ocean. Annals and Magazine of Natural History, Series 8, 3 (13): 30 - 33, 1 fig." type="journal article" year="1909">Calman, 1909</bibRefCitation>
|
||
</taxonomicName>
|
||
as there were no major differences observed between the two taxa other than the proportionally longer adult cheliped and proportionately shorter and stouter G1. They are here regarded as subjective synonyms.
|
||
</paragraph>
|
||
<paragraph id="8B98783B79607E4CFC976D4526E1FC32" blockId="18.[813,1450,797,917]" pageId="18" pageNumber="624">
|
||
Calocarcinids seem to utilise a variety of hosts, from azooxanthellate (ahermatypic)scleractinian corals, antipatharians, alcyonaceans, gorgonians, to sponges (
|
||
<bibRefCitation id="EFB605CA79607E4CFC8E6D2526F5FC32" author="Castro P" box="[820,964,893,917]" pageId="18" pageNumber="624" pagination="543 - 581" refId="ref18613" refString="Castro P (2015) Symbiotic Brachyura. In: Castro P, Davie P, Guinot D, Schram FR & von Vaupel Klein JC (eds.) Treatise on zoology - anatomy, taxonomy, biology. The Crustacea. Decapoda: Brachyura (Part 2), 9 C-II. Brill, Leiden, The Netherlands, pp. 543 - 581." type="book chapter" year="2015">Castro, 2015</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |