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<document id="7E3CCC019B75986F755EA0A69FF39F50" ID-CLB-Dataset="32767" ID-DOI="10.11646/zootaxa.4286.1.1" ID-GBIF-Dataset="a614f1d9-bda9-4df0-a486-aef5af7813fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="828206" ID-ZooBank="6A22EB50-0146-465A-8E2A-523E2F8FE470" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500108241184" checkinUser="plazi" docAuthor="Galvis, William" docDate="2017" docId="0386A719573B9065FF21F9B5FB3DFC97" docLanguage="en" docName="zootaxa.4286.1.1.pdf" docOrigin="Zootaxa 4286 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amphidraus" docType="treatment" docVersion="7" lastPageNumber="22" masterDocId="FFBFDF6157289070FFB6FFCFFF81FFE8" masterDocTitle="Nineteen new species of Amphidraus Simon, 1900 (Salticidae: Euophryini) from Colombia, with comments about their conservation" masterLastPageNumber="40" masterPageNumber="1" pageNumber="20" updateTime="1698465056562" updateUser="ExternalLinkService">
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<mods:title id="55FDA4CB53CD0E4FC7DF0343F254E934">Nineteen new species of Amphidraus Simon, 1900 (Salticidae: Euophryini) from Colombia, with comments about their conservation</mods:title>
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<mods:namePart id="83A8D02F3C2376C47D057DDF2F05D066">Galvis, William</mods:namePart>
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<mods:date id="5CB09AB5DE0471954C742D2CB13DB668">2017</mods:date>
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<mods:identifier id="E4A4DD567FD11FE9729A6471113249C0" type="DOI">10.11646/zootaxa.4286.1.1</mods:identifier>
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<treatment id="0386A719573B9065FF21F9B5FB3DFC97" ID-DOI="http://doi.org/10.5281/zenodo.6045664" ID-GBIF-Taxon="132548496" ID-Zenodo-Dep="6045664" LSID="urn:lsid:plazi:treatment:0386A719573B9065FF21F9B5FB3DFC97" httpUri="http://treatment.plazi.org/id/0386A719573B9065FF21F9B5FB3DFC97" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20">
<subSubSection id="C3354584573B9063FF21F9B5FDC4F95E" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="8B90160F573B9063FF21F9B5FE42F97C" blockId="19.[151,581,1658,1718]" box="[151,451,1658,1684]" pageId="19" pageNumber="20">
<heading id="D0D8A163573B9063FF21F9B5FE42F97C" bold="true" box="[151,451,1658,1684]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
<emphasis id="B95BCA1D573B9063FF21F9B5FE42F97C" bold="true" box="[151,451,1658,1684]" pageId="19" pageNumber="20">
<emphasis id="B95BCA1D573B9063FF21F9B5FEDEF97C" bold="true" box="[151,351,1658,1684]" italics="true" pageId="19" pageNumber="20">
<taxonomicName id="4C2F6D8C573B9063FF21F9B5FEACF97C" ID-CoL="62BBT" ID-ENA="1317342" box="[151,301,1658,1684]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">Amphidraus</taxonomicName>
sua
</emphasis>
sp. nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="8B90160F573B9063FF21F952FDC4F95E" blockId="19.[151,581,1658,1718]" box="[151,581,1693,1718]" pageId="19" pageNumber="20">
<figureCitation id="13140A8A573B9063FF21F952FE94F95E" box="[151,277,1693,1718]" captionStart="FIGURES 12" captionStartId="20.[151,264,1898,1920]" captionTargetBox="[162,1417,458,1867]" captionTargetId="figure@20.[162,1426,445,1883]" captionTargetPageId="20" captionText="FIGURES 12 a – d. A. sua sp. nov., a) left male palp, ventral view; b) same, retrolateral view; c) epigyne, cleared, ventral view; d) same, dorsal view. Abbreviations: cd = copulatory duct, co = copulatory opening, e = embolus, ED = embolic disc, EF = embolic filament, fd = fertilization duct, PED = process on embolic disc, po = pocket, sp = spermathecal, RTA = retrolateral tibial apophysis of the male palp, SD = sperm duct, TL = tegular lobe. Scale bars: 0.20 mm." httpUri="https://zenodo.org/record/828230/files/figure.png" pageId="19" pageNumber="20">Figs 12a–d</figureCitation>
,
<figureCitation id="13140A8A573B9063FE96F952FEE6F95E" box="[288,359,1693,1718]" captionStart="FIGURES 18" captionStartId="29.[151,264,1705,1727]" captionTargetBox="[340,1240,556,1672]" captionTargetId="figure@29.[340,1247,553,1684]" captionTargetPageId="29" captionText="FIGURES 18 a – i. A. somondoco sp. nov., a) female holotype (ANDES – IN 2899). A. sotairensis sp. nov., b) male holotype (ICN – Ar 7981). A. sua sp. nov., c) male holotype (ICN – Ar 8271); d) female paratype (ICN – Ar 8272). A. tanimuca sp. nov., e) male holotype (ICN – Ar 6775). A. tisquesusa sp. nov., f) male holotype (ICN – Ar 8256). A. tundama sp. nov., g) male holotype (ICN – Ar 7839); h) female paratype (ICN – Ar 7845); i) female paratype (ICN – Ar 7846). Scale bars: 0.50 mm." httpUri="https://zenodo.org/record/828242/files/figure.png" pageId="19" pageNumber="20">18c–d</figureCitation>
,
<figureCitation id="13140A8A573B9063FEC5F952FE37F95E" box="[371,438,1693,1718]" captionStart="FIGURES 22" captionStartId="33.[151,264,1900,1922]" captionTargetBox="[195,1386,193,1874]" captionTargetId="figure@33.[162,1426,193,1883]" captionTargetPageId="33" captionText="FIGURES 22 a – k. Amphidraus spp, left male palp. A. sikuani sp. nov., a) ventral view; b) same, retrolateral view. A. sotairensis sp. nov., c) ventral view; d) same, retrolateral view. A. sua sp. nov., e) ventral view; f) same, retrolateral view. A. tisquesusa sp. nov., g) ventral view; h) same, retrolateral view. A. tanimuca sp. nov., i) ventral view of the bulb and cymbium; j) palp without cymbium to show the RTA, retrolateral view; k) same, prolateral view; l) same, ventral view." httpUri="https://zenodo.org/record/828250/files/figure.png" pageId="19" pageNumber="20">22e–f</figureCitation>
,
<figureCitation id="13140A8A573B9063FE77F952FD9CF95E" box="[449,541,1693,1718]" captionStart="FIGURES 25" captionStartId="36.[151,264,1889,1911]" captionTargetBox="[185,1412,194,1864]" captionTargetId="figure@36.[151,1436,193,1867]" captionTargetPageId="36" captionText="FIGURES 25 a – l. Amphidraus spp, epigyne. A. sikuani sp. nov., a) ventral view; g) disposition of the internal structures. A. somondoco sp. nov., b) ventral view; h) disposition of the internal structures. A. sua sp. nov., c) ventral view; i) disposition of the internal structures. A. tundama sp. nov., d) ventral view; j) disposition of the internal structures. A. zaque sp. nov., e) ventral view; k) disposition of the internal structures. A. zipa sp. nov., f) ventral view; l) disposition of the internal structures." httpUri="https://zenodo.org/record/828256/files/figure.png" pageId="19" pageNumber="20">
25c,
<date id="FF9130CF573B9063FE41F952FD9CF95E" box="[503,541,1693,1718]" pageId="19" pageNumber="20" value="1925-01">25i</date>
</figureCitation>
,
<figureCitation id="13140A8A573B9063FD9FF951FDC4F95E" box="[553,581,1694,1718]" captionStart="FIGURE 27" captionStartId="38.[151,250,1317,1339]" captionTargetBox="[138,1418,182,1273]" captionTargetId="figure@38.[137,1422,179,1282]" captionTargetPageId="38" captionText="FIGURE 27. Distribution records of the species of the genus Amphidraus Simon from Colombia treated here. Symbols with a dot inside represent species that have distributions in the Cundiboyacense high-Andean plateau, in the Boyacá and Cundinamarca departments." httpUri="https://zenodo.org/record/828260/files/figure.png" pageId="19" pageNumber="20">27</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3354584573B9063FF21F92AFE41F881" pageId="19" pageNumber="20" type="description">
<paragraph id="8B90160F573B9063FF21F92AFE41F881" blockId="19.[151,1437,1765,2006]" pageId="19" pageNumber="20">
<materialsCitation id="3B471C52573B9063FF21F92AFC81F8C9" ID-GBIF-Occurrence="1572324842" collectingDate="2016-12-07" collectionCode="ICN" collectorName="Reserva Natural La Almenara &amp; R. Anderson" country="Colombia" elevation="1152" latitude="4.88799" location="Reserva Natural La Almenara" longLatPrecision="1" longitude="-73.24989" municipality="Santa Maria" pageId="19" pageNumber="20" specimenCount="2" stateProvince="Boyaca" typeStatus="holotype">
<emphasis id="B95BCA1D573B9063FF21F92AFEE0F916" bold="true" box="[151,353,1765,1790]" pageId="19" pageNumber="20">
<typeStatus id="5494A8AD573B9063FF21F92AFF5EF916" box="[151,223,1765,1790]" pageId="19" pageNumber="20">Types</typeStatus>
.
<typeStatus id="5494A8AD573B9063FF5DF92AFEDDF916" box="[235,348,1765,1790]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
:
</emphasis>
male from
<collectorName id="26DA73D9573B9063FE52F929FCB7F916" box="[484,822,1765,1790]" pageId="19" pageNumber="20">
<location id="8EF040D4573B9063FE52F929FCB7F916" LSID="urn:lsid:plazi:treatment:0386A719573B9065FF21F9B5FB3DFC97:8EF040D4573B9063FE52F929FCB7F916" box="[484,822,1765,1790]" country="Colombia" latitude="4.88799" longLatPrecision="1" longitude="-73.24989" municipality="Santa Maria" name="Reserva Natural La Almenara" pageId="19" pageNumber="20" stateProvince="Boyaca">Reserva Natural La Almenara</location>
</collectorName>
,
<collectingMunicipality id="6BF48C75573B9063FCF4F929FC4FF916" box="[834,974,1766,1790]" pageId="19" pageNumber="20">Santa María</collectingMunicipality>
,
<collectingRegion id="49EBD8ED573B9063FC6FF929FBB1F916" box="[985,1072,1766,1790]" country="Colombia" name="Boyaca" pageId="19" pageNumber="20">Boyacá</collectingRegion>
,
<collectingCountry id="F338569F573B9063FB8DF92AFB2DF916" box="[1083,1196,1765,1790]" name="Colombia" pageId="19" pageNumber="20">Colombia</collectingCountry>
,
<elevation id="0002F13C573B9063FB0EF92AFA8FF915" box="[1208,1294,1765,1790]" metricMagnitude="3" metricUnit="m" metricValue="1.152" pageId="19" pageNumber="20" unit="m" value="1152.0">
<quantity id="4CD7BBEA573B9063FB0EF92AFA8FF915" box="[1208,1294,1765,1790]" metricMagnitude="3" metricUnit="m" metricValue="1.152" pageId="19" pageNumber="20" unit="m" value="1152.0">1152 m</quantity>
</elevation>
,
<geoCoordinate id="EE1B70C8573B9063FAACF929FA19F916" box="[1306,1432,1766,1790]" direction="north" orientation="latitude" pageId="19" pageNumber="20" precision="1" value="4.88799">4.88799°N</geoCoordinate>
,
<geoCoordinate id="EE1B70C8573B9063FF21F8C6FEA7F8C9" box="[151,294,1801,1825]" direction="west" orientation="longitude" pageId="19" pageNumber="20" precision="1" value="-73.24989">73.24989°W</geoCoordinate>
,
<collectingDate id="EFD5C927573B9063FE87F8C6FE2EF8CA" box="[305,431,1801,1826]" pageId="19" pageNumber="20" value="2016-12-07">
<date id="FF9130CF573B9063FE87F8C6FE2EF8CA" box="[305,431,1801,1826]" pageId="19" pageNumber="20" value="2016-12-07">7.XII.2016</date>
</collectingDate>
,
<collectorName id="26DA73D9573B9063FE0CF8C5FDC8F8CA" box="[442,585,1801,1826]" pageId="19" pageNumber="20">R. Anderson</collectorName>
(
<collectionCode id="ED3E8ECA573B9063FDE1F8C6FD09F8C9" box="[599,648,1801,1825]" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="19" pageNumber="20">ICN</collectionCode>
–Ar 8271)
</materialsCitation>
.
<materialsCitation id="3B471C52573B9063FCBAF8C6FE41F881" ID-GBIF-Occurrence="1572324844" collectingDate="2016-12-07" collectionCode="ICN" collectorName="Bosque de Palmas &amp; R. Anderson" country="Colombia" county="Santa Maria" elevation="1163" latitude="4.85251" location="Bosque de Palmas" longLatPrecision="1" longitude="-73.25006" municipality="Vereda Calichana" pageId="19" pageNumber="20" specimenCount="1" stateProvince="Boyaca" typeStatus="paratype">
<emphasis id="B95BCA1D573B9063FCBAF8C6FC03F8CA" bold="true" box="[780,898,1801,1826]" pageId="19" pageNumber="20">
<typeStatus id="5494A8AD573B9063FCBAF8C6FCFCF8CA" box="[780,893,1801,1826]" pageId="19" pageNumber="20" type="paratype">Paratype</typeStatus>
:
</emphasis>
one female from
<collectorName id="26DA73D9573B9063FBFCF8C5FA9AF8CA" box="[1098,1307,1801,1826]" pageId="19" pageNumber="20">
<location id="8EF040D4573B9063FBFCF8C5FA9AF8CA" LSID="urn:lsid:plazi:treatment:0386A719573B9065FF21F9B5FB3DFC97:8EF040D4573B9063FBFCF8C5FA9AF8CA" box="[1098,1307,1801,1826]" country="Colombia" county="Santa Maria" latitude="4.85251" longLatPrecision="1" longitude="-73.25006" municipality="Vereda Calichana" name="Bosque de Palmas" pageId="19" pageNumber="20" stateProvince="Boyaca">Bosque de Palmas</location>
</collectorName>
, way to the antennae,
<collectingMunicipality id="6BF48C75573B9063FEBAF8E2FE57F8AE" box="[268,470,1837,1862]" pageId="19" pageNumber="20">Vereda Calichana</collectingMunicipality>
,
<collectingCounty id="62F16E83573B9063FE53F8E1FDF2F8AE" box="[485,627,1838,1862]" pageId="19" pageNumber="20">Santa María</collectingCounty>
,
<collectingRegion id="49EBD8ED573B9063FD37F8E1FD59F8AE" box="[641,728,1838,1862]" country="Colombia" name="Boyaca" pageId="19" pageNumber="20">Boyacá</collectingRegion>
,
<collectingCountry id="F338569F573B9063FD50F8E2FCD6F8AE" box="[742,855,1837,1862]" name="Colombia" pageId="19" pageNumber="20">Colombia</collectingCountry>
,
<elevation id="0002F13C573B9063FCD0F8E2FC3EF8AD" box="[870,959,1837,1862]" metricMagnitude="3" metricUnit="m" metricValue="1.1629999999999998" pageId="19" pageNumber="20" unit="m" value="1163.0">
<quantity id="4CD7BBEA573B9063FCD0F8E2FC3EF8AD" box="[870,959,1837,1862]" metricMagnitude="3" metricUnit="m" metricValue="1.1629999999999998" pageId="19" pageNumber="20" unit="m" value="1163.0">1163 m</quantity>
</elevation>
,
<geoCoordinate id="EE1B70C8573B9063FC78F8E2FBCAF8AE" box="[974,1099,1837,1862]" direction="north" orientation="latitude" pageId="19" pageNumber="20" precision="1" value="4.85251">4.85251°N</geoCoordinate>
,
<geoCoordinate id="EE1B70C8573B9063FBEFF8E2FB69F8AE" box="[1113,1256,1837,1862]" direction="west" orientation="longitude" pageId="19" pageNumber="20" precision="1" value="-73.25006">73.25006°W</geoCoordinate>
,
<collectingDate id="EFD5C927573B9063FB41F8E2FAF5F8AE" box="[1271,1396,1837,1862]" pageId="19" pageNumber="20" value="2016-12-07">
<date id="FF9130CF573B9063FB41F8E2FAF5F8AE" box="[1271,1396,1837,1862]" pageId="19" pageNumber="20" value="2016-12-07">7.XII.2016</date>
</collectingDate>
,
<collectorName id="26DA73D9573B9063FA35F8E1FE85F882" pageId="19" pageNumber="20">R. Anderson</collectorName>
(
<collectionCode id="ED3E8ECA573B9063FEA4F89EFEC2F881" box="[274,323,1873,1897]" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="19" pageNumber="20">ICN</collectionCode>
–Ar 8272).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3354584573B9063FF71F8BAFCE3F83E" pageId="19" pageNumber="20" type="etymology">
<paragraph id="8B90160F573B9063FF71F8BAFCE3F83E" blockId="19.[151,1437,1765,2006]" pageId="19" pageNumber="20">
<emphasis id="B95BCA1D573B9063FF71F8BAFECFF866" bold="true" box="[199,334,1909,1934]" pageId="19" pageNumber="20">Etymology.</emphasis>
The epithet is a noun in apposition that honors Sua, the goddess of sun (also known as Sue or Xué) in the Muisca mythology, indigenous people that inhabited the region where the species was found, in the central Andean highlands of the present-day Colombia's Eastern Range
</paragraph>
</subSubSection>
<subSubSection id="C3354584573C9064FF71FF58FE75FE60" pageId="20" pageNumber="21" type="diagnosis">
<paragraph id="8B90160F573C9064FF71FF58FE75FE60" blockId="20.[151,1436,151,392]" pageId="20" pageNumber="21">
<emphasis id="B95BCA1D573C9064FF71FF58FEC0FF58" bold="true" box="[199,321,151,176]" pageId="20" pageNumber="21">Diagnosis.</emphasis>
Males of
<emphasis id="B95BCA1D573C9064FE02FF57FE7AFF58" box="[436,507,152,176]" italics="true" pageId="20" pageNumber="21">A. sua</emphasis>
<emphasis id="B95BCA1D573C9064FDB5FF58FDDCFF58" bold="true" box="[515,605,151,176]" pageId="20" pageNumber="21">sp. nov.</emphasis>
are similar to those of
<taxonomicName id="4C2F6D8C573C9064FCD2FF57FC32FF58" box="[868,947,152,176]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="mae" status="sp. nov.">
<emphasis id="B95BCA1D573C9064FCD2FF57FC32FF58" box="[868,947,152,176]" italics="true" pageId="20" pageNumber="21">A. mae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687766573C9064FC0CFF58FB94FF58" box="[954,1045,151,176]" pageId="20" pageNumber="21" rank="species">
<emphasis id="B95BCA1D573C9064FC0CFF58FB94FF58" bold="true" box="[954,1045,151,176]" pageId="20" pageNumber="21">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C2F6D8C573C9064FBFAFF57FB5BFF58" box="[1100,1242,151,176]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="quimbaya" status="sp. nov.">
<emphasis id="B95BCA1D573C9064FBFAFF57FB5BFF58" box="[1100,1242,151,176]" italics="true" pageId="20" pageNumber="21">A. quimbaya</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687766573C9064FB55FF58FABCFF58" box="[1251,1341,151,176]" pageId="20" pageNumber="21" rank="species">
<emphasis id="B95BCA1D573C9064FB55FF58FABCFF58" bold="true" box="[1251,1341,151,176]" pageId="20" pageNumber="21">sp. nov.</emphasis>
</taxonomicNameLabel>
by their RTA shapes, but can be distinguished from those by their combined presence of a RTA with the ventral lobe longer and pointed, and the presence of an embolar disc spur (EDS) (
<figureCitation id="13140A8A573C9064FCEAFF10FC5CFF10" box="[860,989,223,248]" captionStart="FIGURES 12" captionStartId="20.[151,264,1898,1920]" captionTargetBox="[162,1417,458,1867]" captionTargetId="figure@20.[162,1426,445,1883]" captionTargetPageId="20" captionText="FIGURES 12 a – d. A. sua sp. nov., a) left male palp, ventral view; b) same, retrolateral view; c) epigyne, cleared, ventral view; d) same, dorsal view. Abbreviations: cd = copulatory duct, co = copulatory opening, e = embolus, ED = embolic disc, EF = embolic filament, fd = fertilization duct, PED = process on embolic disc, po = pocket, sp = spermathecal, RTA = retrolateral tibial apophysis of the male palp, SD = sperm duct, TL = tegular lobe. Scale bars: 0.20 mm." httpUri="https://zenodo.org/record/828230/files/figure.png" pageId="20" pageNumber="21">Figs 12a–b</figureCitation>
,
<figureCitation id="13140A8A573C9064FC5CFF10FBAFFF10" box="[1002,1070,223,248]" captionStart="FIGURES 22" captionStartId="33.[151,264,1900,1922]" captionTargetBox="[195,1386,193,1874]" captionTargetId="figure@33.[162,1426,193,1883]" captionTargetPageId="33" captionText="FIGURES 22 a – k. Amphidraus spp, left male palp. A. sikuani sp. nov., a) ventral view; b) same, retrolateral view. A. sotairensis sp. nov., c) ventral view; d) same, retrolateral view. A. sua sp. nov., e) ventral view; f) same, retrolateral view. A. tisquesusa sp. nov., g) ventral view; h) same, retrolateral view. A. tanimuca sp. nov., i) ventral view of the bulb and cymbium; j) palp without cymbium to show the RTA, retrolateral view; k) same, prolateral view; l) same, ventral view." httpUri="https://zenodo.org/record/828250/files/figure.png" pageId="20" pageNumber="21">22e–f</figureCitation>
). Females of
<emphasis id="B95BCA1D573C9064FB71FF2FFA8FFF10" box="[1223,1294,224,248]" italics="true" pageId="20" pageNumber="21">A. sua</emphasis>
<emphasis id="B95BCA1D573C9064FAA1FF10FAF3FF10" bold="true" box="[1303,1394,223,248]" pageId="20" pageNumber="21">sp. nov.</emphasis>
are similar to those of
<taxonomicName id="4C2F6D8C573C9064FEC4FECBFE42FEF4" box="[370,451,260,284]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="mae" status="sp. nov.">
<emphasis id="B95BCA1D573C9064FEC4FECBFE42FEF4" box="[370,451,260,284]" italics="true" pageId="20" pageNumber="21">A. mae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687766573C9064FE7AFECBFDA6FEF5" box="[460,551,260,285]" pageId="20" pageNumber="21" rank="species">
<emphasis id="B95BCA1D573C9064FE7AFECBFDA6FEF5" bold="true" box="[460,551,260,285]" pageId="20" pageNumber="21">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<emphasis id="B95BCA1D573C9064FD8EFECBFDF9FEF4" box="[568,632,260,284]" italics="true" pageId="20" pageNumber="21">A. sie</emphasis>
<emphasis id="B95BCA1D573C9064FD34FECBFD5FFEF5" bold="true" box="[642,734,260,285]" pageId="20" pageNumber="21">sp. nov.</emphasis>
and
<taxonomicName id="4C2F6D8C573C9064FCAFFECBFC3BFEF5" box="[793,954,260,285]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="somondoco" status="sp. nov.">
<emphasis id="B95BCA1D573C9064FCAFFECBFC3BFEF5" box="[793,954,260,285]" italics="true" pageId="20" pageNumber="21">A. somondoco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687766573C9064FC75FECBFB9EFEF5" box="[963,1055,260,285]" pageId="20" pageNumber="21" rank="species">
<emphasis id="B95BCA1D573C9064FC75FECBFB9EFEF5" bold="true" box="[963,1055,260,285]" pageId="20" pageNumber="21">sp. nov.</emphasis>
</taxonomicNameLabel>
by the general disposition of the internal structures of the epigyne, but can be distinguished from those by their more anteriorly placed copulatory openings, longer copulatory ducts, anterior rounded spermathecae, and single rounded, medial pocket in the epigyne (
<figureCitation id="13140A8A573C9064FF48FEA0FEFFFE60" box="[254,382,367,392]" captionStart="FIGURES 12" captionStartId="20.[151,264,1898,1920]" captionTargetBox="[162,1417,458,1867]" captionTargetId="figure@20.[162,1426,445,1883]" captionTargetPageId="20" captionText="FIGURES 12 a – d. A. sua sp. nov., a) left male palp, ventral view; b) same, retrolateral view; c) epigyne, cleared, ventral view; d) same, dorsal view. Abbreviations: cd = copulatory duct, co = copulatory opening, e = embolus, ED = embolic disc, EF = embolic filament, fd = fertilization duct, PED = process on embolic disc, po = pocket, sp = spermathecal, RTA = retrolateral tibial apophysis of the male palp, SD = sperm duct, TL = tegular lobe. Scale bars: 0.20 mm." httpUri="https://zenodo.org/record/828230/files/figure.png" pageId="20" pageNumber="21">Figs 12c–d</figureCitation>
,
<figureCitation id="13140A8A573C9064FE3FFEA0FE66FE60" box="[393,487,367,392]" captionStart="FIGURES 25" captionStartId="36.[151,264,1889,1911]" captionTargetBox="[185,1412,194,1864]" captionTargetId="figure@36.[151,1436,193,1867]" captionTargetPageId="36" captionText="FIGURES 25 a – l. Amphidraus spp, epigyne. A. sikuani sp. nov., a) ventral view; g) disposition of the internal structures. A. somondoco sp. nov., b) ventral view; h) disposition of the internal structures. A. sua sp. nov., c) ventral view; i) disposition of the internal structures. A. tundama sp. nov., d) ventral view; j) disposition of the internal structures. A. zaque sp. nov., e) ventral view; k) disposition of the internal structures. A. zipa sp. nov., f) ventral view; l) disposition of the internal structures." httpUri="https://zenodo.org/record/828256/files/figure.png" pageId="20" pageNumber="21">
25c,
<date id="FF9130CF573C9064FE76FEA0FE66FE60" box="[448,487,367,392]" pageId="20" pageNumber="21" value="1925-01">25i</date>
</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF504687573C9064FF21F8A5FCFBF834" httpUri="https://zenodo.org/record/828230/files/figure.png" pageId="20" pageNumber="21" targetBox="[162,1417,458,1867]" targetPageId="20">
<paragraph id="8B90160F573C9064FF21F8A5FCFBF834" blockId="20.[151,1436,1898,2012]" pageId="20" pageNumber="21">
<emphasis id="B95BCA1D573C9064FF21F8A5FED4F868" bold="true" box="[151,341,1898,1920]" pageId="20" pageNumber="21">FIGURES 12a–d.</emphasis>
<emphasis id="B95BCA1D573C9064FEEDF8A4FE1BF868" box="[347,410,1899,1920]" italics="true" pageId="20" pageNumber="21">A. sua</emphasis>
<emphasis id="B95BCA1D573C9064FE16F8A5FE71F868" bold="true" box="[416,496,1898,1920]" pageId="20" pageNumber="21">sp. nov.</emphasis>
,
<emphasis id="B95BCA1D573C9064FE4BF8A5FD90F868" bold="true" box="[509,529,1898,1920]" pageId="20" pageNumber="21">a)</emphasis>
left male palp, ventral view;
<emphasis id="B95BCA1D573C9064FC8DF8A5FCD0F868" bold="true" box="[827,849,1898,1920]" pageId="20" pageNumber="21">b)</emphasis>
same, retrolateral view;
<emphasis id="B95BCA1D573C9064FBFDF8A5FBDEF868" bold="true" box="[1099,1119,1898,1920]" pageId="20" pageNumber="21">c)</emphasis>
epigyne, cleared, ventral view;
<emphasis id="B95BCA1D573C9064FF21F846FF2FF877" bold="true" box="[151,174,1929,1951]" pageId="20" pageNumber="21">d)</emphasis>
same, dorsal view. Abbreviations: cd=copulatory duct, co=copulatory opening, e=embolus, ED=embolic disc, EF=embolic filament, fd=fertilization duct, PED=process on embolic disc, po=pocket, sp=spermathecal, RTA=retrolateral tibial apophysis of the male palp, SD=sperm duct, TL=tegular lobe. Scale bars: 0.20 mm.
</paragraph>
</caption>
<subSubSection id="C3354584573D9065FF71FF58FA15FD18" pageId="21" pageNumber="22" type="description">
<paragraph id="8B90160F573D9065FF71FF58FC2FFE38" blockId="21.[151,1437,151,896]" pageId="21" pageNumber="22">
<emphasis id="B95BCA1D573D9065FF71FF58FDA5FF58" bold="true" box="[199,548,151,176]" pageId="21" pageNumber="22">
Description. Male (
<typeStatus id="5494A8AD573D9065FE1AFF58FD99FF58" box="[428,536,151,176]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
).
</emphasis>
Total length: 2.77. Carapace dark brown with a dorsal lighter mark, 1.46 long, 1.02 wide, 0.77 high (
<figureCitation id="13140A8A573D9065FE3DFF73FE6AFF3C" box="[395,491,188,213]" captionStart="FIGURES 18" captionStartId="29.[151,264,1705,1727]" captionTargetBox="[340,1240,556,1672]" captionTargetId="figure@29.[340,1247,553,1684]" captionTargetPageId="29" captionText="FIGURES 18 a – i. A. somondoco sp. nov., a) female holotype (ANDES – IN 2899). A. sotairensis sp. nov., b) male holotype (ICN – Ar 7981). A. sua sp. nov., c) male holotype (ICN – Ar 8271); d) female paratype (ICN – Ar 8272). A. tanimuca sp. nov., e) male holotype (ICN – Ar 6775). A. tisquesusa sp. nov., f) male holotype (ICN – Ar 8256). A. tundama sp. nov., g) male holotype (ICN – Ar 7839); h) female paratype (ICN – Ar 7845); i) female paratype (ICN – Ar 7846). Scale bars: 0.50 mm." httpUri="https://zenodo.org/record/828242/files/figure.png" pageId="21" pageNumber="22">Fig. 18c</figureCitation>
). OC black, 0.75 long. Anterior eye row 1.15 wide and posterior 1.00 wide. Sternum brown, 0.60 long, 0.48 wide. Labium brown, 0.16 long, 0.19 wide. Chelicerae brown with four or five retromarginal and two promarginal teeth. Palp brown with rounded tegular lobe (TL), long embolic filament (EF), embolar disc spur present (EDS), bilobed RTA, with the ventral lobe long and finger-like (
<figureCitation id="13140A8A573D9065FB32FEE8FA85FEA8" box="[1156,1284,295,320]" captionStart="FIGURES 12" captionStartId="20.[151,264,1898,1920]" captionTargetBox="[162,1417,458,1867]" captionTargetId="figure@20.[162,1426,445,1883]" captionTargetPageId="20" captionText="FIGURES 12 a – d. A. sua sp. nov., a) left male palp, ventral view; b) same, retrolateral view; c) epigyne, cleared, ventral view; d) same, dorsal view. Abbreviations: cd = copulatory duct, co = copulatory opening, e = embolus, ED = embolic disc, EF = embolic filament, fd = fertilization duct, PED = process on embolic disc, po = pocket, sp = spermathecal, RTA = retrolateral tibial apophysis of the male palp, SD = sperm duct, TL = tegular lobe. Scale bars: 0.20 mm." httpUri="https://zenodo.org/record/828230/files/figure.png" pageId="21" pageNumber="22">Figs 12a–b</figureCitation>
,
<figureCitation id="13140A8A573D9065FAB9FEE8FAD2FEA8" box="[1295,1363,295,320]" captionStart="FIGURES 22" captionStartId="33.[151,264,1900,1922]" captionTargetBox="[195,1386,193,1874]" captionTargetId="figure@33.[162,1426,193,1883]" captionTargetPageId="33" captionText="FIGURES 22 a – k. Amphidraus spp, left male palp. A. sikuani sp. nov., a) ventral view; b) same, retrolateral view. A. sotairensis sp. nov., c) ventral view; d) same, retrolateral view. A. sua sp. nov., e) ventral view; f) same, retrolateral view. A. tisquesusa sp. nov., g) ventral view; h) same, retrolateral view. A. tanimuca sp. nov., i) ventral view of the bulb and cymbium; j) palp without cymbium to show the RTA, retrolateral view; k) same, prolateral view; l) same, ventral view." httpUri="https://zenodo.org/record/828250/files/figure.png" pageId="21" pageNumber="22">22e–f</figureCitation>
). Legs 4312, all yellow, I darker in coloration, with femur I dark brown. Leg macrosetae: femur, I d 1 di; II–IV d 2 di, p 1 di; patella, III–IV r 1 me, tibia, I
<date id="FF9130CF573D9065FDBCFEBFFDC6FE60" box="[522,583,368,392]" pageId="21" pageNumber="22">v 2-2</date>
-2; II
<date id="FF9130CF573D9065FD33FEBFFD40FE60" box="[645,705,368,392]" pageId="21" pageNumber="22">v 2-2</date>
-2, p 1 di; III
<date id="FF9130CF573D9065FCE1FEBFFCF9FE60" box="[855,888,368,392]" pageId="21" pageNumber="22">v 1</date>
-0-1, p 1 pr, r 0-1-1; IV
<date id="FF9130CF573D9065FB31FEBFFB29FE60" box="[1159,1192,368,392]" pageId="21" pageNumber="22">v 1</date>
-0-1, p 0-1-1, r 1-1-1; metatarsus, I
<date id="FF9130CF573D9065FE87FE5AFEF0FE44" box="[305,369,404,428]" pageId="21" pageNumber="22">v 2-2</date>
; II
<date id="FF9130CF573D9065FE2CFE5AFE58FE44" box="[410,473,404,428]" pageId="21" pageNumber="22">v 2-2</date>
, p 1 di; III
<date id="FF9130CF573D9065FDD7FE5AFD08FE44" box="[609,649,404,428]" pageId="21" pageNumber="22">v 2</date>
-0-1, p 1-0-2, r 1-0-2; IV
<date id="FF9130CF573D9065FC06FE5AFC52FE44" box="[944,979,404,428]" pageId="21" pageNumber="22">v 1</date>
-0-1, p 1-0-2, r 1-1-2. Abdomen creamcolored with a reticulated pattern of brown marks and spots (
<figureCitation id="13140A8A573D9065FCF6FE78FC20FE38" box="[832,929,439,464]" captionStart="FIGURES 18" captionStartId="29.[151,264,1705,1727]" captionTargetBox="[340,1240,556,1672]" captionTargetId="figure@29.[340,1247,553,1684]" captionTargetPageId="29" captionText="FIGURES 18 a – i. A. somondoco sp. nov., a) female holotype (ANDES – IN 2899). A. sotairensis sp. nov., b) male holotype (ICN – Ar 7981). A. sua sp. nov., c) male holotype (ICN – Ar 8271); d) female paratype (ICN – Ar 8272). A. tanimuca sp. nov., e) male holotype (ICN – Ar 6775). A. tisquesusa sp. nov., f) male holotype (ICN – Ar 8256). A. tundama sp. nov., g) male holotype (ICN – Ar 7839); h) female paratype (ICN – Ar 7845); i) female paratype (ICN – Ar 7846). Scale bars: 0.50 mm." httpUri="https://zenodo.org/record/828242/files/figure.png" pageId="21" pageNumber="22">Fig. 18c</figureCitation>
).
</paragraph>
<paragraph id="8B90160F573D9065FF71FE13FA15FD18" blockId="21.[151,1437,151,896]" pageId="21" pageNumber="22">
<emphasis id="B95BCA1D573D9065FF71FE13FE2DFE1D" bold="true" box="[199,428,476,501]" pageId="21" pageNumber="22">
Female (
<typeStatus id="5494A8AD573D9065FE99FE13FE1EFE1D" box="[303,415,476,501]" pageId="21" pageNumber="22" type="paratype">paratype</typeStatus>
).
</emphasis>
Total length 2.89. Carapace brown with a posterior lighter stripe, 1.50 long, 1.12 wide, 0.72 high (
<figureCitation id="13140A8A573D9065FEA2FE30FEF6FDF0" box="[276,375,511,536]" captionStart="FIGURES 18" captionStartId="29.[151,264,1705,1727]" captionTargetBox="[340,1240,556,1672]" captionTargetId="figure@29.[340,1247,553,1684]" captionTargetPageId="29" captionText="FIGURES 18 a – i. A. somondoco sp. nov., a) female holotype (ANDES – IN 2899). A. sotairensis sp. nov., b) male holotype (ICN – Ar 7981). A. sua sp. nov., c) male holotype (ICN – Ar 8271); d) female paratype (ICN – Ar 8272). A. tanimuca sp. nov., e) male holotype (ICN – Ar 6775). A. tisquesusa sp. nov., f) male holotype (ICN – Ar 8256). A. tundama sp. nov., g) male holotype (ICN – Ar 7839); h) female paratype (ICN – Ar 7845); i) female paratype (ICN – Ar 7846). Scale bars: 0.50 mm." httpUri="https://zenodo.org/record/828242/files/figure.png" pageId="21" pageNumber="22">Fig. 18d</figureCitation>
). OC black, 0.82 long. Anterior eye row 1.14 wide and posterior 1.01 wide. Sternum brown, 0.60 long, 0.48 wide. Labium brown, 0.17 long, 0.22 wide. Chelicerae brown, with four retromarginal and two promarginal teeth. Legs 4312, all yellowish with brown marks. Leg macrosetae: femur, I–IV d 1 di; patellae, III–IV r 1 me; tibiae, I
<date id="FF9130CF573D9065FEFEFDA2FE05FD6C" box="[328,388,620,644]" pageId="21" pageNumber="22">v 2-2</date>
-2; II
<date id="FF9130CF573D9065FE74FDA2FE7CFD6C" box="[450,509,620,644]" pageId="21" pageNumber="22">v 2-2</date>
-2, p 1 di; III
<date id="FF9130CF573D9065FD27FDA2FD35FD6C" box="[657,692,620,644]" pageId="21" pageNumber="22">v 1</date>
pr, p 1-0-1, r 1-0-1; IV
<date id="FF9130CF573D9065FC0BFDA2FC61FD6C" box="[957,992,620,644]" pageId="21" pageNumber="22">v 1</date>
pr, p 1-0-1, r 1-0-1; metatarsus, I
<date id="FF9130CF573D9065FAECFDA2FA19FD6C" box="[1370,1432,620,644]" pageId="21" pageNumber="22">v 2-2</date>
; II
<date id="FF9130CF573D9065FF07FD5FFF6FFD40" box="[177,238,656,680]" pageId="21" pageNumber="22">v 2-2</date>
, p 1 di; III
<date id="FF9130CF573D9065FEDBFD5FFE12FD40" box="[365,403,656,680]" pageId="21" pageNumber="22">v 2</date>
-0-1, p 1-0-2, r 1-0-2; IV
<date id="FF9130CF573D9065FD19FD5FFD50FD40" box="[687,721,656,680]" pageId="21" pageNumber="22">v 1</date>
-0-1, p 1-0-2, r 1-1-2. Abdomen dark brown with dorsal creamcolored marks (
<figureCitation id="13140A8A573D9065FEF1FD7BFE28FD25" box="[327,425,692,717]" captionStart="FIGURES 18" captionStartId="29.[151,264,1705,1727]" captionTargetBox="[340,1240,556,1672]" captionTargetId="figure@29.[340,1247,553,1684]" captionTargetPageId="29" captionText="FIGURES 18 a – i. A. somondoco sp. nov., a) female holotype (ANDES – IN 2899). A. sotairensis sp. nov., b) male holotype (ICN – Ar 7981). A. sua sp. nov., c) male holotype (ICN – Ar 8271); d) female paratype (ICN – Ar 8272). A. tanimuca sp. nov., e) male holotype (ICN – Ar 6775). A. tisquesusa sp. nov., f) male holotype (ICN – Ar 8256). A. tundama sp. nov., g) male holotype (ICN – Ar 7839); h) female paratype (ICN – Ar 7845); i) female paratype (ICN – Ar 7846). Scale bars: 0.50 mm." httpUri="https://zenodo.org/record/828242/files/figure.png" pageId="21" pageNumber="22">Fig. 18d</figureCitation>
). Epigyne (
<figureCitation id="13140A8A573D9065FD9CFD7BFD2AFD25" box="[554,683,692,717]" captionStart="FIGURES 12" captionStartId="20.[151,264,1898,1920]" captionTargetBox="[162,1417,458,1867]" captionTargetId="figure@20.[162,1426,445,1883]" captionTargetPageId="20" captionText="FIGURES 12 a – d. A. sua sp. nov., a) left male palp, ventral view; b) same, retrolateral view; c) epigyne, cleared, ventral view; d) same, dorsal view. Abbreviations: cd = copulatory duct, co = copulatory opening, e = embolus, ED = embolic disc, EF = embolic filament, fd = fertilization duct, PED = process on embolic disc, po = pocket, sp = spermathecal, RTA = retrolateral tibial apophysis of the male palp, SD = sperm duct, TL = tegular lobe. Scale bars: 0.20 mm." httpUri="https://zenodo.org/record/828230/files/figure.png" pageId="21" pageNumber="22">Figs 12c–d</figureCitation>
,
<figureCitation id="13140A8A573D9065FD01FD7BFC92FD25" box="[695,787,692,717]" captionStart="FIGURES 25" captionStartId="36.[151,264,1889,1911]" captionTargetBox="[185,1412,194,1864]" captionTargetId="figure@36.[151,1436,193,1867]" captionTargetPageId="36" captionText="FIGURES 25 a – l. Amphidraus spp, epigyne. A. sikuani sp. nov., a) ventral view; g) disposition of the internal structures. A. somondoco sp. nov., b) ventral view; h) disposition of the internal structures. A. sua sp. nov., c) ventral view; i) disposition of the internal structures. A. tundama sp. nov., d) ventral view; j) disposition of the internal structures. A. zaque sp. nov., e) ventral view; k) disposition of the internal structures. A. zipa sp. nov., f) ventral view; l) disposition of the internal structures." httpUri="https://zenodo.org/record/828256/files/figure.png" pageId="21" pageNumber="22">
25c,
<date id="FF9130CF573D9065FD58FD7BFC92FD25" box="[750,787,692,717]" pageId="21" pageNumber="22" value="1925-01">25i</date>
</figureCitation>
) with anterior and medial copulatory openings, very long copulatory ducts, a posterior single medial pocket, anterior rounded spermathecae, and medial oval spermathecae.
</paragraph>
</subSubSection>
<subSubSection id="C3354584573D9065FF71FD33FE8CFCB5" pageId="21" pageNumber="22" type="discussion">
<paragraph id="8B90160F573D9065FF71FD33FE8CFCB5" blockId="21.[151,1437,151,896]" pageId="21" pageNumber="22">
<emphasis id="B95BCA1D573D9065FF71FD33FECEFCFD" bold="true" box="[199,335,764,789]" pageId="21" pageNumber="22">Comments.</emphasis>
The
<typeStatus id="5494A8AD573D9065FE38FD32FE3FFCFD" box="[398,446,765,789]" pageId="21" pageNumber="22">type</typeStatus>
material was collected in beating low vegetation and in Winkler traps in a secondary Andean forest, in the same locality with the
<typeStatus id="5494A8AD573D9065FD26FCEFFD41FCD0" box="[656,704,800,824]" pageId="21" pageNumber="22">type</typeStatus>
specimens of
<taxonomicName id="4C2F6D8C573D9065FCD2FCEFFC35FCD0" box="[868,948,799,824]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="chie" status="sp. nov.">
<emphasis id="B95BCA1D573D9065FCD2FCEFFC35FCD0" box="[868,948,799,824]" italics="true" pageId="21" pageNumber="22">A. chie</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2687766573D9065FC0AFCD0FB97FCD0" box="[956,1046,799,824]" pageId="21" pageNumber="22" rank="species">
<emphasis id="B95BCA1D573D9065FC0AFCD0FB97FCD0" bold="true" box="[956,1046,799,824]" pageId="21" pageNumber="22">sp. nov.</emphasis>
</taxonomicNameLabel>
, so they are here considered to be sympatric.
</paragraph>
</subSubSection>
<subSubSection id="C3354584573D9065FF71FCA8FB3DFC97" box="[199,1212,871,896]" pageId="21" pageNumber="22" type="distribution">
<paragraph id="8B90160F573D9065FF71FCA8FB3DFC97" blockId="21.[151,1437,151,896]" box="[199,1212,871,896]" pageId="21" pageNumber="22">
<emphasis id="B95BCA1D573D9065FF71FCA8FEE1FC68" bold="true" box="[199,352,871,896]" pageId="21" pageNumber="22">Distribution.</emphasis>
<collectingCountry id="F338569F573D9065FED1FCA8FE54FC68" box="[359,469,871,896]" name="Colombia" pageId="21" pageNumber="22">Colombia</collectingCountry>
(
<collectingRegion id="49EBD8ED573D9065FE52FCA7FDBAFC68" box="[484,571,872,896]" country="Colombia" name="Boyaca" pageId="21" pageNumber="22">Boyacá</collectingRegion>
) (
<figureCitation id="13140A8A573D9065FDE4FCA8FD26FC68" box="[594,679,871,896]" captionStart="FIGURE 27" captionStartId="38.[151,250,1317,1339]" captionTargetBox="[138,1418,182,1273]" captionTargetId="figure@38.[137,1422,179,1282]" captionTargetPageId="38" captionText="FIGURE 27. Distribution records of the species of the genus Amphidraus Simon from Colombia treated here. Symbols with a dot inside represent species that have distributions in the Cundiboyacense high-Andean plateau, in the Boyacá and Cundinamarca departments." httpUri="https://zenodo.org/record/828260/files/figure.png" pageId="21" pageNumber="22">Fig. 27</figureCitation>
). Known altitudinal distribution:
<quantity id="4CD7BBEA573D9065FBAAFCA8FB39FC97" box="[1052,1208,871,896]" metricMagnitude="3" metricUnit="m" metricValue="1.1575" metricValueMax="1.1629999999999998" metricValueMin="1.152" pageId="21" pageNumber="22" unit="m" value="1157.5" valueMax="1163.0" valueMin="1152.0">1152–1163 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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