treatments-xml/data/7E/06/FF/7E06FF1D7E3259A8AAF8E92363C22043.xml
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<document ID-DOI="http://dx.doi.org/10.3897/nl.47.114772" ID-Pensoft-Pub="2367-5365-47-57" ID-Pensoft-UUID="4A77F35768A45842A1B27D54D5B50724" ID-ZooBank="18BDAA9F5AE5479ABC3D1998C8193791" ModsDocID="2367-5365-47-57" checkinTime="1710546606512" checkinUser="pensoft" docAuthor="Shovkoon, Dmitry F. &amp; Trofimova, Tatyana A." docDate="2024" docId="7E06FF1D7E3259A8AAF8E92363C22043" docLanguage="en" docName="NotaLepidopt 47: 57-79" docOrigin="Nota Lepidopterologica 47" docPubDate="2024-03-15" docSource="http://dx.doi.org/10.3897/nl.47.114772" docTitle="Carriola saturnioides" docType="treatment" docVersion="1" id="4A77F35768A45842A1B27D54D5B50724" lastPageNumber="57" masterDocId="4A77F35768A45842A1B27D54D5B50724" masterDocTitle="Review of the genus Carriola Swinhoe, 1922 (Lepidoptera, Erebidae, Lymantriinae), with descriptions of four new species" masterLastPageNumber="79" masterPageNumber="57" pageNumber="57" updateTime="1710546606512" updateUser="pensoft">
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<mods:title id="481EED2D92C3947135B0DE8B6FA3898B">Review of the genus Carriola Swinhoe, 1922 (Lepidoptera, Erebidae, Lymantriinae), with descriptions of four new species</mods:title>
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<mods:namePart id="EEB06023757B5A0D4E1CD74B3BC39498">Shovkoon, Dmitry F.</mods:namePart>
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<mods:affiliation id="9E2E83D31BD61FC575C78DD033B48133">Faculty of Science, University of South Bohemia, Branisovska 31, 370 05 Ceske Budejovice, Czech Republic; shovkoon. d @ googlemail. com &amp; Institute of Entomology, Biology Centre CAS, Branisovska 1760, 370 05 Ceske Budejovice, Czech Republic</mods:affiliation>
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<mods:namePart id="F46D4F14FCEBC19FAFC48F1ADFA4F419">Trofimova, Tatyana A.</mods:namePart>
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<mods:affiliation id="F083E2449BBED8A37EE82EB9E876280E">Laboratory of Animal Systematic and Faunistic Samara State University, ul. Ac. Pavlova 1, Samara, 443011, Russia; apamea @ mail. ru</mods:affiliation>
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<mods:title id="8316DE7ACC68A3061C4CB7955E3799F6">Nota Lepidopterologica</mods:title>
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<mods:date id="C27709F4E0549E3DE7E82B18D4EC0A1D">2024</mods:date>
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<mods:number id="7705320E0721745B059CC5E6C272AA14">47</mods:number>
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<treatment id="7E06FF1D7E3259A8AAF8E92363C22043" LSID="urn:lsid:plazi:treatment:7E06FF1D7E3259A8AAF8E92363C22043" httpUri="http://treatment.plazi.org/id/7E06FF1D7E3259A8AAF8E92363C22043" lastPageNumber="57" pageId="0" pageNumber="57">
<subSubSection id="4B906EB6482FC9DA2A047F5204D2A9A4" pageId="0" pageNumber="57" type="nomenclature">
<paragraph id="77CAE44EDE93E46912228F80995D9B95" pageId="0" pageNumber="57">
<taxonomicName id="52711122F5A80459F5835945EAA09D23" LSID="7E06FF1D-7E32-59A8-AAF8-E92363C22043" authority="(Snellen, 1879)" baseAuthorityName="Snellen" baseAuthorityYear="1879" class="Insecta" family="Erebidae" genus="Carriola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carriola saturnioides" order="Lepidoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="saturnioides">Carriola saturnioides (Snellen, 1879)</taxonomicName>
</paragraph>
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<subSubSection id="0839FBEFB67C50B27E539D9FD46F9263" pageId="0" pageNumber="57" type="description">
<paragraph id="13C28E2792A7CDDF361B7A19594B2CF0" pageId="0" pageNumber="57">
<figureCitation id="903A23999DE8414B2942C92DE7054259" captionStart="Figures 1018" captionStartId="F3" captionText="Figures 10 - 18. The wing venation and hyaline window mapping of Carriola species. 10. C. ecnomoda (Swinhoe, 1907); 11. C. thyridophora (Hampson, [1893]); 12. C. witti sp. nov.; 13. C. seminsula (Strand, 1914); 14. C. saturnioides (Snellen, 1879); 15. C. polyakovi sp. nov.; 16. C. shorokhovi sp. nov.; 17. C. fenestrata (Hampson, [1893]); 18. C. zolotuhini sp. nov." figureDoi="10.3897/nl.47.114772.figures10-18" httpUri="https://binary.pensoft.net/fig/1003322" pageId="0" pageNumber="57">Figs 14</figureCitation>
<figureCitation id="979CD64EAD7D8D6D92B532FC36789918" captionStart="Figures 1927" captionStartId="F4" captionText="Figures 19 - 27. Adults of Carriola species. a. Male; b. Underside wings of male; c. Female. 19. C. ecnomoda (Swinhoe, 1907); a, b. East Java, Mt. Baluran (GU 26.698; ZSM); c. East Java, Mt. Baluran, (GU 26.699; ZSM); 20. C. thyridophora (Hampson, [1893]); a, b. NE India. Assam (GU 27.442; ZSM); c. SW Kampuchea, Kirirom Nat. Park (GU 28.073; ZSM); 21 C. witti sp. nov.; a, b. HT (GU 25.852; ZSM); c. PT (GU 25.853; ZSM); 22. C. seminsula (Strand, 1914); a, b. Sud Vietnam, Bao Loc (GU 27.431; ZSM); c. Nord Vietnam, Mai Chau (GU 27.445; ZSM); 23. C. saturnioides (Snellen, 1879); a, b. N. Sulawesi, (GU 21.966; ZSM); c. Sulawesi, Namo (GU 25.850; ZSM); 24. C. polyakovi sp. nov.; a, b. HT (GU 28.779; ZSM); c. PT (GU 28.780; ZSM); 25. C. shorokhovi sp. nov.; a, b. HT (GU 26.961; ZSM); c. PT (GU 28.784; ZSM); 26. C. fenestrata (Hampson, [1893]); a, b. Vietnam, prov. Thai Nguyen (GU 27.435; ZSM); c. Vietnam, prov. Thai Nguyen, (GU 27.436; ZSM); 27. C. zolotuhini sp. nov.; a, b. HT (GU 28.069; ZSM). Scale bar represents 10 mm, all specimens to scale." figureDoi="10.3897/nl.47.114772.figures19-27" httpUri="https://binary.pensoft.net/fig/1003323" pageId="0" pageNumber="57">, 23</figureCitation>
<figureCitation id="8C9A113DD9A43871A89F22E5C5CB6E13" captionStart="Figures 2836" captionStartId="F5" captionText="Figures 28 - 36. Male genitalia of Carriola species. a. Genitalia; b. Aedeagus with everted vesica; c. Superuncus (VIII tergum). 28. C. ecnomoda (Swinhoe, 1907) (GU 26.698; ZSM); 29. C. thyridophora (Hampson, [1893]) (GU 27.442; ZSM); 30. C. witti sp. nov. HT (GU 25.852; ZSM); 31. C. seminsula (Strand, 1914): HT (prep. NO 597; SMF); 32. C. saturnioides (Snellen, 1879) (GU 21.966; ZSM); 33. C. polyakovi sp. nov. HT (GU 28.779; ZSM); 34. C. shorokhovi sp. nov. HT (GU 26.961; ZSM); 35. C. fenestrata (Hampson, [1893]) (GU 27.435; ZSM); 36. C. zolotuhini sp. nov. HT (GU 28.069; ZSM). Scale bar represents 1 mm, all genitalia to scale." figureDoi="10.3897/nl.47.114772.figures28-36" httpUri="https://binary.pensoft.net/fig/1003324" pageId="0" pageNumber="57">, 32</figureCitation>
<figureCitation id="F5744B40A2CE8C8874DF24A7AA64CA7C" captionStart="Figures 3744" captionStartId="F6" captionText="Figures 37 - 44. Female genitalia of Carriola species. 37. C. ecnomoda (Swinhoe, 1907) (GU 26.699; ZSM); 38. C. thyridophora (Hampson, [1893]) (GU 28.073; ZSM); 39. C. witti sp. nov. PT (GU 25.853; ZSM); 40. C. seminsula (Strand, 1914) (GU 27.445; ZSM); 41. C. saturnioides (Snellen, 1879) (GU 25.850; ZSM); 42. C. polyakovi sp. nov. PT (GU 28.780; ZSM); 43. C. shorokhovi sp. nov. PT (GU 28.784; ZSM); 44. C. fenestrata (Hampson, [1893]) (GU 27.436; ZSM). Scale bar represents 1 mm, all genitalia to scale." figureDoi="10.3897/nl.47.114772.figures37-44" httpUri="https://binary.pensoft.net/fig/1003325" pageId="0" pageNumber="57">, 41</figureCitation>
<figureCitation id="383017EE0FBB6244E3DAFAB03543F6FD" captionStart="Figures 4553" captionStartId="F7" captionText="Figures 45 - 53. Distribution maps of Carriola species. 45. C. ecnomoda (Swinhoe, 1907); 46. C. thyridophora (Hampson, [1893]); 47. C. witti sp. nov.; 48. C. seminsula (Strand, 1914); 49. C. saturnioides (Snellen, 1879); 50. C. polyakovi sp. nov.; 51. C. shorokhovi sp. nov.; 52. C. fenestrata (Hampson, [1893]); 53. C. zolotuhini sp. nov." figureDoi="10.3897/nl.47.114772.figures45-53" httpUri="https://binary.pensoft.net/fig/1003326" pageId="0" pageNumber="57">, 49</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="57860B27D2E47F9BE791C7BF548F78AA" pageId="0" pageNumber="57" type="reference_group">
<paragraph id="82D32EDF1D57D17B9890CC472C8F9077" pageId="0" pageNumber="57">
<taxonomicName id="78210554C4BDC55FEB8E00DDFE5536AC" class="Insecta" family="Erebidae" genus="Laelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laelia saturnioides" order="Lepidoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="saturnioides">Laelia saturnioides</taxonomicName>
Snellen, 1879, Tijdschrift voor entomologie 22: 105 pl. 8, figs. 7, 7a-c. Type locality: Celebes, Takalar [Indonesia, Sulawesi].
</paragraph>
</subSubSection>
<subSubSection id="72EC190432EA734CF3B25E9E452F692C" pageId="0" pageNumber="57" type="materials_examined">
<paragraph id="53090A3E8F6978845B6408EDCF744507" pageId="0" pageNumber="57">Material examined.</paragraph>
<paragraph id="D0ACDBED0D511726F95AB3D4ABFE8F2E" pageId="0" pageNumber="57">
<materialsCitation id="3FE3B57A1740A7B3EEB303D69A34F046" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" country="Indonesia" location="Sulawesi" specimenCount="1" stateProvince="Sulawesi">
<collectingCountry id="2B9648C71A09D685555F5D265E958EFC" name="Indonesia">Indonesia</collectingCountry>
:
<collectingRegion id="E1BE189A3D7B55CEF6A3EA47A333E3CA" country="Indonesia" name="Sulawesi Utara">Sulawesi</collectingRegion>
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<materialsCitation id="D975D2CE93892CF43A65D933E48ACABA" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" country="Indonesia" location="Prov." specimenCount="7" specimenCount-male="7" stateProvince="Sulawesi Selatan">
<specimenCount id="3DE1C33DC069550354F84577B0F4484D" count="7" type="male">7 ♂</specimenCount>
,
<location id="21FC9CAEDB71AAD3F67C700312C8CB34" LSID="urn:lsid:plazi:treatment:7E06FF1D7E3259A8AAF8E92363C22043:21FC9CAEDB71AAD3F67C700312C8CB34" country="Indonesia" name="Prov." stateProvince="Sulawesi Selatan">Prov.</location>
<collectingRegion id="DC23E5CEB6627C4F4EA30F8C284431FC" country="Indonesia" name="Sulawesi Selatan">Sulawesi Selatan</collectingRegion>
</materialsCitation>
,
<materialsCitation id="611270A70D8ED590FA45CECBEE111A62" collectingDate="1984-05-02" collectingDateMax="1984-05-09" collectingDateMin="1984-05-02" collectorName="Talkessel, A. Schintlmeister" country="Indonesia" elevation="200" latitude="-4.9333334" location="Prov." longLatPrecision="1306" longitude="119.65" municipality="Bantimurung" specimenCount="10" specimenCount-female="1" specimenCount-male="9" stateProvince="Sulawesi Selatan">
<collectingCountry id="0ABCEF7B429E92D089BC3145605AE00F" name="Indonesia">Indonesia</collectingCountry>
,
<collectingMunicipality id="B204080A58ECF91E6769F6A9A741864C">Bantimurung</collectingMunicipality>
,
<locationDeviation id="609A34AC32EE0FA5878014062E4B85A9" location="Ujung Padang">
<quantity id="2BCE1138778FCA45406460563F19F556" metricMagnitude="4" metricUnit="m" metricValue="4.0" unit="km" value="40.0">40 km</quantity>
NNE Ujung Padang
</locationDeviation>
,
<geoCoordinate id="0063EF32D767CB7A44319480C73DC111" degrees="4" direction="south" minutes="56" orientation="latitude" precision="925" value="-4.9333334">4°56'S</geoCoordinate>
,
<geoCoordinate id="C11580A81EDC71064B1C2AC6E302ED8A" degrees="119" direction="east" minutes="39" orientation="longitude" precision="925" value="119.65">119°39'E</geoCoordinate>
,
<collectorName id="B2DB1BFD6DF48FC20D518C25676E5C6A">Talkessel</collectorName>
,
<quantity id="B55A842D9D8A7C7C381DD5A4E1FF5007" metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">
<elevation id="9EC947A66A4B04C4174E99CF8296E8E7" metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">200 m</elevation>
</quantity>
, leg.
<collectorName id="D062D71539213DABFE943C9A43F87455">A. Schintlmeister</collectorName>
<collectingDate id="35CD72F6286294E3D5FB230EC0DFEA1C" value="1984-05-02" valueMax="1984-05-09" valueMin="1984-05-02">2.-9.v.1984</collectingDate>
; ZSM •
<specimenCount id="A2EDA23512C0F3187A1F0CE3690B9A3A" count="9" type="male">9 ♂</specimenCount>
,
<specimenCount id="9CDEEA88F078089AFF9BBDE612D339B3" count="1" type="female">1 ♀</specimenCount>
,
<location id="601659A7182FE42A70E263070DE5FE12" LSID="urn:lsid:plazi:treatment:7E06FF1D7E3259A8AAF8E92363C22043:601659A7182FE42A70E263070DE5FE12" country="Indonesia" latitude="-4.9333334" longLatPrecision="1306" longitude="119.65" municipality="Bantimurung" name="Prov." stateProvince="Sulawesi Selatan">Prov.</location>
<collectingRegion id="21DA3A2A962401400B2B73F1426ED4DC" country="Indonesia" name="Sulawesi Selatan">Sulawesi Selatan</collectingRegion>
</materialsCitation>
,
<materialsCitation id="5F4467C352508856D30E0E32B3E1ADC0" collectingDate="1984-05-06" collectingDateMax="1984-05-07" collectingDateMin="1984-05-06" collectorName="Bergpass-Suedhang, Urwald, Kalk, A. Schintlmeister" country="Indonesia" elevation="250" location="Sulawesi" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Sulawesi">
<collectingCountry id="3AE73C69015AAD2D82869256122FBCFA" name="Indonesia">Indonesia</collectingCountry>
,
<normalizedToken id="01D95155FD87C4D15057358961AAC09C" originalValue="Bergpass-Südhang">
<collectorName id="0C6DFE9761EEA4F3FF13BCCC069CAA3A">Bergpass-Suedhang</collectorName>
</normalizedToken>
,
<quantity id="872CAFCC7EF01D6A11772086CA475082" metricMagnitude="2" metricUnit="m" metricValue="2.5" unit="m" value="250.0">
<elevation id="8BD813773B4B3ACAF38BBF1A5C2A400D" metricMagnitude="2" metricUnit="m" metricValue="2.5" unit="m" value="250.0">250 m</elevation>
</quantity>
,
<collectorName id="4398C9CEE434B532F7A5F42D34BF799E">Urwald</collectorName>
auf
<collectorName id="06A87AE13B7DB360339DD8F162BA64A4">Kalk</collectorName>
, leg.
<collectorName id="4DCF92D1CECC606CACCA40D9415517DD">A. Schintlmeister</collectorName>
<collectingDate id="805BDB8CF40A77A46E2809E0ED8918D4" value="1984-05-06" valueMax="1984-05-07" valueMin="1984-05-06">6.-7.v.1984</collectingDate>
(GU 25.849); ZSM •
<specimenCount id="C38719D7CFB08CA9A4F4B647C30B74A9" count="3" type="male">3 ♂</specimenCount>
,
<specimenCount id="43E39B3E6A217FEBCAE3D1ECE9F8187B" count="2" type="female">2 ♀</specimenCount>
,
<collectingRegion id="EF1E748DCD67215D05EDA4B742B8EE79" country="Indonesia" name="Sulawesi Utara">Sulawesi</collectingRegion>
</materialsCitation>
,
<materialsCitation id="4C697823B206E83EDEC7E7DCCE9C0291" collectingDate="1997-03" collectorName="Dr. Ronald Brechlin" country="Indonesia" county="Selatan" elevation="1100" latitude="-3.0" location="Palopo" longLatPrecision="1307" longitude="120.166664" municipality="Puncak" specimenCount="1" specimenCount-female="1" stateProvince="Sulawesi Utara">
<collectingCounty id="5DA2E8192E5BD8FEA8558E413F2C8DB9">Selatan</collectingCounty>
,
<collectingMunicipality id="88C474E80C7550CEE805F8BF6EB6ED72">Puncak</collectingMunicipality>
,
<location id="AAEFD5F8B31A9EBDFD9B4B3CB36F452E" LSID="urn:lsid:plazi:treatment:7E06FF1D7E3259A8AAF8E92363C22043:AAEFD5F8B31A9EBDFD9B4B3CB36F452E" country="Indonesia" county="Selatan" latitude="-3.0" longLatPrecision="1307" longitude="120.166664" municipality="Puncak" name="Palopo" stateProvince="Sulawesi Utara">Palopo</location>
,
<quantity id="38930FC3F434AF0336C9D39B6E346234" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.3" metricValueMin="0.9" unit="m" value="1100.0" valueMax="1300.0" valueMin="900.0">
<elevation id="5EC92065670AE2C8A3726BD623877F1F" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.3" metricValueMin="0.9" unit="m" value="1100.0" valueMax="1300.0" valueMin="900.0">900-1300 m</elevation>
</quantity>
,
<geoCoordinate id="97E2551E1F19F64490FCA9BB6E86D6E1" degrees="3" direction="south" minutes="00" orientation="latitude" precision="925" value="-3.0">3°00'S</geoCoordinate>
;
<geoCoordinate id="F5294AA18CACD083D662FA5CB2415EF9" degrees="120" direction="east" minutes="10" orientation="longitude" precision="925" value="120.166664">120°10'E</geoCoordinate>
,
<collectingDate id="3DD1EF0B8801ACB82EB2905160D5FD50" value="1997-03">iii.1997</collectingDate>
, ex coll.
<collectorName id="A518A5EDD9FDD1A7F26C9CF5776B9EC9">Dr. Ronald Brechlin</collectorName>
; ZSM •
<specimenCount id="D969465BBE5B036860CAD9642219B275" count="1" type="female">1 ♀</specimenCount>
, the
</materialsCitation>
<materialsCitation id="5B1953A8090D9FDF0A32CC759070799F" collectingDate="1998-04" collectorName="Dr. Ronald Brechlin" country="Indonesia" county="Selatan" elevation="950" latitude="-3.0" location="Palopo" longLatPrecision="1307" longitude="120.166664" municipality="Puncak" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Sulawesi Utara">
same data, but
<collectingDate id="1AC7D0FAE464F834B3308F1ADA75B0C8" value="1998-04">iv.1998</collectingDate>
,
<quantity id="2B7468DDE595CFF2AAE880A7FE6103B7" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">
<elevation id="9A39A74435F4ED8D05B868ECB2E986FA" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">900-1000 m</elevation>
</quantity>
, ex coll.
<collectorName id="80750177A266E7133826B1A99249EF7C">Dr. Ronald Brechlin</collectorName>
; ZSM •
<specimenCount id="8EF9F1EBAD8D73DA7DB2CA85065949A1" count="2" type="male">2 ♂</specimenCount>
,
<specimenCount id="486CEE1CAD8B38BD5D73BE5617F79562" count="1" type="female">1 ♀</specimenCount>
, the
</materialsCitation>
<materialsCitation id="AD6F59093CA93DA555308EA202E3CAD5" collectingDate="1997-01" collectorName="Dr. Ronald Brechlin" country="Indonesia" county="Selatan" elevation="1100" latitude="-2.9166667" location="Palopo" longLatPrecision="1307" longitude="120.083336" municipality="Puncak" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Sulawesi Utara">
same data,
<geoCoordinate id="21718620DD03389BA4DC1CD699EF330E" degrees="2" direction="south" minutes="55" orientation="latitude" precision="925" value="-2.9166667">2°55'S</geoCoordinate>
,
<geoCoordinate id="D191D694DC8D322D552CBA5E1F9FC73E" degrees="120" direction="east" minutes="05" orientation="longitude" precision="925" value="120.083336">120°05'E</geoCoordinate>
,
<collectingDate id="57DCD65F61ABC017D6284CF05A69C653" value="1997-01">i.1997</collectingDate>
,
<quantity id="AD67F393B060653076ED674B87514226" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.3" metricValueMin="0.9" unit="m" value="1100.0" valueMax="1300.0" valueMin="900.0">
<elevation id="4D442C07E4CF8C9A1D948FD7BD766564" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.3" metricValueMin="0.9" unit="m" value="1100.0" valueMax="1300.0" valueMin="900.0">900-1300 m</elevation>
</quantity>
, leg. local collector; ZSM •
<specimenCount id="5F33EAD191ECC0D11808A3BEBBC9E2B3" count="1" type="male">1 ♂</specimenCount>
,
<specimenCount id="CD95467D2273F7E66E4BC299FD8C8FEB" count="1" type="female">1 ♀</specimenCount>
</materialsCitation>
,
<materialsCitation id="40B0F06B975ECF495D2661B6C4EEF043" collectingDate="1997-09" collectorName="Einh. Sammer" country="Indonesia" county="Selatan" elevation="1100" latitude="-2.9166667" location="Puncak" longLatPrecision="1307" longitude="120.083336" municipality="Puncak" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi Utara">
same data, but
<collectingDate id="CFC08D48C4F0C7C583DF1B55FE3F9144" value="1997-09">ix.1997</collectingDate>
leg.
<collectorName id="069611E8AF91F75A0A44D0725C0212BB">Einh. Sammer</collectorName>
; ZSM •
<specimenCount id="7D65025910335C177496FF38175A4568" count="1" type="male">1 ♂</specimenCount>
,
<location id="AB2B7E5DD6C07D4C8ED2F9B612005C44" LSID="urn:lsid:plazi:treatment:7E06FF1D7E3259A8AAF8E92363C22043:AB2B7E5DD6C07D4C8ED2F9B612005C44" country="Indonesia" county="Selatan" latitude="-2.9166667" longLatPrecision="1307" longitude="120.083336" name="Puncak" stateProvince="Sulawesi Utara">Puncak</location>
</materialsCitation>
,
<materialsCitation id="2BD2932C4423DD8FBD8665EF913E152E" collectingDate="1984-05-10" collectingDateMax="1984-05-19" collectingDateMin="1984-05-10" collectorName="Bergwald, A. Schintlmeister" country="Indonesia" elevation="1100" latitude="-2.9833333" location="27 km W Palopo" longLatPrecision="1307" longitude="120.0" specimenCount="2" specimenCount-male="2" stateProvince="Sulawesi">
<location id="745321C12CFAA9D3200108599105BCDA" LSID="urn:lsid:plazi:treatment:7E06FF1D7E3259A8AAF8E92363C22043:745321C12CFAA9D3200108599105BCDA" country="Indonesia" latitude="-2.9833333" longLatPrecision="1307" longitude="120.0" name="27 km W Palopo" stateProvince="Sulawesi">
<quantity id="41CD8333C45E177C1CAFB13420E43E9B" metricMagnitude="4" metricUnit="m" metricValue="2.7" unit="km" value="27.0">27 km</quantity>
W Palopo
</location>
,
<collectingDate id="6CA8AA375439741CD174A57226A1611A" value="1984-05-10" valueMax="1984-05-19" valueMin="1984-05-10">10-19.v.1984</collectingDate>
,
<geoCoordinate id="6D406D27F75027A779E58881521401C1" degrees="2" direction="south" minutes="59" orientation="latitude" precision="925" value="-2.9833333">2°59'S</geoCoordinate>
,
<geoCoordinate id="85428A6287947674A311CD289ACA0555" degrees="120" direction="east" minutes="00" orientation="longitude" precision="925" value="120.0">120°00'E</geoCoordinate>
,
<collectorName id="A57BE005D9E3C1A7B8B61C55A85918F6">Bergwald</collectorName>
,
<quantity id="A3B1588294B78CCE706BC5CC53BEC615" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" unit="m" value="1100.0" valueMax="1200.0" valueMin="1000.0">
<elevation id="0047B73229AAFF47DD0F2A9285953394" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" unit="m" value="1100.0" valueMax="1200.0" valueMin="1000.0">1000-1200 m</elevation>
</quantity>
, leg.
<collectorName id="FBCC0BD849D7A7B4CDEB7E338B90CD6A">A. Schintlmeister</collectorName>
; ZSM •
<specimenCount id="5056A114DE06BADCB8F42270B288D78A" count="2" type="male">2 ♂</specimenCount>
; Z.
<collectingRegion id="51FB78EE429300DC2C9744911021A275" country="Indonesia" name="Sulawesi Utara">Sulawesi</collectingRegion>
</materialsCitation>
,
<materialsCitation id="06CA68C022CEE34D6005374DA398F459" collectingDate="1995-02" collectorName="Sinjaev, Tarasov" country="Indonesia" elevation="1000" latitude="-1.4333334" location="Kulawi" longLatPrecision="1308" longitude="120.0" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi">
<location id="A4083423D5D243FE1964EA679093E3C6" LSID="urn:lsid:plazi:treatment:7E06FF1D7E3259A8AAF8E92363C22043:A4083423D5D243FE1964EA679093E3C6" country="Indonesia" latitude="-1.4333334" longLatPrecision="1308" longitude="120.0" name="Kulawi" stateProvince="Sulawesi">Kulawi</location>
,
<geoCoordinate id="BF6BF0B9898390F48CE446B2672F38A3" degrees="1" direction="south" minutes="26" orientation="latitude" precision="925" value="-1.4333334">1°26'S</geoCoordinate>
,
<geoCoordinate id="3031E5054602AE2550BBF7E07FC9DE92" degrees="120" direction="east" minutes="00" orientation="longitude" precision="925" value="120.0">120°00'E</geoCoordinate>
, 78.
<collectingDate id="6C89F64BC7F1C4AC23A8AF222B2DC710" value="1995-02">ii.1995</collectingDate>
,
<quantity id="4F8B96DB3B0D6831515ECB8213E80A47" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">
<elevation id="22F537B03E1C0876FA7580EFAF7C27EB" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">1000 m</elevation>
</quantity>
, leg.
<collectorName id="F8112426D7FF3C5B0EE038BFD8309080">Sinjaev</collectorName>
&amp;
<collectorName id="B4B7B2E07BB67FA7FC03349B72E1C170">Tarasov</collectorName>
; ZSM •
<specimenCount id="613405417C548888BFF0FEB076770CD4" count="1" type="male">1 ♂</specimenCount>
, N.
<collectingRegion id="F304632F7DA6DCFF4CE65F30194BD887" country="Indonesia" name="Sulawesi Utara">Sulawesi</collectingRegion>
</materialsCitation>
,
<materialsCitation id="71B40AA25367DD39CB30E0194BB4770E" collectingDate="1988-12" collectorName="Nr. Tondano, Mt. Makaweiben" country="Indonesia" elevation="1000" location="Sulawesi" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Sulawesi">
<collectorName id="A4419CD7DC0C6C6DA60A0ACC75124494">Nr. Tondano</collectorName>
,
<collectorName id="1BE936DB83E6A4D33C38E6FBF208DAB4" hasSafeAffix="true">Mt. Makaweiben</collectorName>
,
<quantity id="8E45632496A64498363B41318F361815" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">
<elevation id="922DA290D33AC9BD0EE4E9CE46D5F51D" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<collectingDate id="FE4D5E59EF0920BED8FE2537DC953BB3" value="1988-12">xii.1988</collectingDate>
(GU 21.966); ZSM •
<specimenCount id="CEADFD1EAB48B722582E26D1C12D5925" count="3" type="male">3 ♂</specimenCount>
,
<specimenCount id="197B898AB1E2EBB3524A0F8B54ABFD4F" count="3" type="female">3 ♀</specimenCount>
, C.
<collectingRegion id="DE935F606670187D1354AC730E950B4C" country="Indonesia" name="Sulawesi Utara">Sulawesi</collectingRegion>
</materialsCitation>
,
<materialsCitation id="BC3C71B6214C1A05DB63C4A0BB1254B5" collectingDate="1995-09-21" collectingDateMax="1995-09-22" collectingDateMin="1995-09-21" collectorName="Namo" country="Indonesia" elevation="650" location="St. Naumann" municipality="Palu-Gimpu" specimenCount="1" stateProvince="Sulawesi Utara">
<collectorName id="AAE64F546DB4CC434EF3400568653A26">Namo</collectorName>
,
<quantity id="0C188C2D44E111A367C6B644953EBEC1" metricMagnitude="2" metricUnit="m" metricValue="6.5" unit="m" value="650.0">
<elevation id="220DE6354915494744DB66A90F0C5114" metricMagnitude="2" metricUnit="m" metricValue="6.5" unit="m" value="650.0">650 m</elevation>
</quantity>
,
<collectingDate id="1E3C49C1FB39178BEEFBCF3ECD52566F" value="1995-09-21" valueMax="1995-09-22" valueMin="1995-09-21">21-22.ix.1995</collectingDate>
,
<normalizedToken id="ECFA8F8C6764BA5ED668AB9724025719" originalValue="Straβe">Straβe</normalizedToken>
<collectingMunicipality id="264776D3A90CE6862E7A3C707B716063">Palu-Gimpu</collectingMunicipality>
, leg
<location id="6B75D4C8B06D7EDCEA3B1EFB6D82E50C" LSID="urn:lsid:plazi:treatment:7E06FF1D7E3259A8AAF8E92363C22043:6B75D4C8B06D7EDCEA3B1EFB6D82E50C" country="Indonesia" municipality="Palu-Gimpu" name="St. Naumann" stateProvince="Sulawesi Utara">St. Naumann</location>
(GU 25.850); ZSM
</materialsCitation>
.
</paragraph>
<paragraph id="DA2BE626CC5A2BACC05E1C0E7F5878F2" pageId="0" pageNumber="57">
Peleng Island •
<specimenCount id="C10B162D9B8EF36F9FC1E0CE5B5EE09D" count="8" type="male">8 ♂</specimenCount>
,
<quantity id="FF1E3C428B51736011000CA48DE33903" metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="km" value="2.0">2 km</quantity>
W Sambuit,
<quantity id="C2AF9C0E2F2C088401B05B81CAC70F9D" metricMagnitude="2" metricUnit="m" metricValue="1.5" unit="m" value="150.0">150 m</quantity>
,
<date id="56AEEA7E54911BDC7D904E8E39FDF8B9" value="1998-07">vii.1998</date>
, leg. local collector (GU 25.851); ZSM.
</paragraph>
</subSubSection>
<subSubSection id="16909690E5A3DD0F227EB219F262E596" pageId="0" pageNumber="57" type="diagnosis">
<paragraph id="162C199B2162DAA6381EBA6628AAA5B8" pageId="0" pageNumber="57">Diagnosis.</paragraph>
<paragraph id="17A64E721EBF81307E35550C47BBD448" pageId="0" pageNumber="57">
Externally, the male of
<taxonomicName id="5EB6F9CBC35E4510969A9EAE5E8BD340" lsidName="C. saturnioides" pageId="0" pageNumber="57" rank="species" species="saturnioides">
<emphasis id="12E434B09A057BC45D03B62E7F51770A" italics="true" pageId="0" pageNumber="57">C. saturnioides</emphasis>
</taxonomicName>
is distinguished by the absence of the hyaline satellite spot Rs-M1 on the hindwing (Figs
<figureCitation id="91D589FB5B805976638908B68D9CA388" captionStart="Figures 1018" captionStartId="F3" captionText="Figures 10 - 18. The wing venation and hyaline window mapping of Carriola species. 10. C. ecnomoda (Swinhoe, 1907); 11. C. thyridophora (Hampson, [1893]); 12. C. witti sp. nov.; 13. C. seminsula (Strand, 1914); 14. C. saturnioides (Snellen, 1879); 15. C. polyakovi sp. nov.; 16. C. shorokhovi sp. nov.; 17. C. fenestrata (Hampson, [1893]); 18. C. zolotuhini sp. nov." figureDoi="10.3897/nl.47.114772.figures10-18" httpUri="https://binary.pensoft.net/fig/1003322" pageId="0" pageNumber="57">14</figureCitation>
,
<figureCitation id="F9D4832373059ADE008387F1FAABC085" captionStart="Figures 1927" captionStartId="F4" captionText="Figures 19 - 27. Adults of Carriola species. a. Male; b. Underside wings of male; c. Female. 19. C. ecnomoda (Swinhoe, 1907); a, b. East Java, Mt. Baluran (GU 26.698; ZSM); c. East Java, Mt. Baluran, (GU 26.699; ZSM); 20. C. thyridophora (Hampson, [1893]); a, b. NE India. Assam (GU 27.442; ZSM); c. SW Kampuchea, Kirirom Nat. Park (GU 28.073; ZSM); 21 C. witti sp. nov.; a, b. HT (GU 25.852; ZSM); c. PT (GU 25.853; ZSM); 22. C. seminsula (Strand, 1914); a, b. Sud Vietnam, Bao Loc (GU 27.431; ZSM); c. Nord Vietnam, Mai Chau (GU 27.445; ZSM); 23. C. saturnioides (Snellen, 1879); a, b. N. Sulawesi, (GU 21.966; ZSM); c. Sulawesi, Namo (GU 25.850; ZSM); 24. C. polyakovi sp. nov.; a, b. HT (GU 28.779; ZSM); c. PT (GU 28.780; ZSM); 25. C. shorokhovi sp. nov.; a, b. HT (GU 26.961; ZSM); c. PT (GU 28.784; ZSM); 26. C. fenestrata (Hampson, [1893]); a, b. Vietnam, prov. Thai Nguyen (GU 27.435; ZSM); c. Vietnam, prov. Thai Nguyen, (GU 27.436; ZSM); 27. C. zolotuhini sp. nov.; a, b. HT (GU 28.069; ZSM). Scale bar represents 10 mm, all specimens to scale." figureDoi="10.3897/nl.47.114772.figures19-27" httpUri="https://binary.pensoft.net/fig/1003323" pageId="0" pageNumber="57">23a</figureCitation>
). The underside of the forewing has a pale, monotonous background without a pronounced banding pattern. The black border around the hyaline windows is also very faint or absent (Fig.
<figureCitation id="DAB1616049B062E0B7A9E1E8A772E1CF" captionStart="Figures 1927" captionStartId="F4" captionText="Figures 19 - 27. Adults of Carriola species. a. Male; b. Underside wings of male; c. Female. 19. C. ecnomoda (Swinhoe, 1907); a, b. East Java, Mt. Baluran (GU 26.698; ZSM); c. East Java, Mt. Baluran, (GU 26.699; ZSM); 20. C. thyridophora (Hampson, [1893]); a, b. NE India. Assam (GU 27.442; ZSM); c. SW Kampuchea, Kirirom Nat. Park (GU 28.073; ZSM); 21 C. witti sp. nov.; a, b. HT (GU 25.852; ZSM); c. PT (GU 25.853; ZSM); 22. C. seminsula (Strand, 1914); a, b. Sud Vietnam, Bao Loc (GU 27.431; ZSM); c. Nord Vietnam, Mai Chau (GU 27.445; ZSM); 23. C. saturnioides (Snellen, 1879); a, b. N. Sulawesi, (GU 21.966; ZSM); c. Sulawesi, Namo (GU 25.850; ZSM); 24. C. polyakovi sp. nov.; a, b. HT (GU 28.779; ZSM); c. PT (GU 28.780; ZSM); 25. C. shorokhovi sp. nov.; a, b. HT (GU 26.961; ZSM); c. PT (GU 28.784; ZSM); 26. C. fenestrata (Hampson, [1893]); a, b. Vietnam, prov. Thai Nguyen (GU 27.435; ZSM); c. Vietnam, prov. Thai Nguyen, (GU 27.436; ZSM); 27. C. zolotuhini sp. nov.; a, b. HT (GU 28.069; ZSM). Scale bar represents 10 mm, all specimens to scale." figureDoi="10.3897/nl.47.114772.figures19-27" httpUri="https://binary.pensoft.net/fig/1003323" pageId="0" pageNumber="57">23b</figureCitation>
). The adults are the largest of the genus
<taxonomicName id="161BC283D627119115F92C21E405918F" authorityName="Swinhoe" authorityYear="1922" class="Insecta" family="Erebidae" genus="Carriola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carriola" order="Lepidoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="AF90FE1C1B446E4CD8BB7EB2B116B0A7" italics="true" pageId="0" pageNumber="57">Carriola</emphasis>
</taxonomicName>
. The length of the male forewing is 15-25.5 mm, that of the female forewing 18-30.2 mm. The male genitalia (Fig.
<figureCitation id="ECF599D43036BC3D3E2F3496AC25D112" captionStart="Figures 2836" captionStartId="F5" captionText="Figures 28 - 36. Male genitalia of Carriola species. a. Genitalia; b. Aedeagus with everted vesica; c. Superuncus (VIII tergum). 28. C. ecnomoda (Swinhoe, 1907) (GU 26.698; ZSM); 29. C. thyridophora (Hampson, [1893]) (GU 27.442; ZSM); 30. C. witti sp. nov. HT (GU 25.852; ZSM); 31. C. seminsula (Strand, 1914): HT (prep. NO 597; SMF); 32. C. saturnioides (Snellen, 1879) (GU 21.966; ZSM); 33. C. polyakovi sp. nov. HT (GU 28.779; ZSM); 34. C. shorokhovi sp. nov. HT (GU 26.961; ZSM); 35. C. fenestrata (Hampson, [1893]) (GU 27.435; ZSM); 36. C. zolotuhini sp. nov. HT (GU 28.069; ZSM). Scale bar represents 1 mm, all genitalia to scale." figureDoi="10.3897/nl.47.114772.figures28-36" httpUri="https://binary.pensoft.net/fig/1003324" pageId="0" pageNumber="57">32</figureCitation>
) are characterised by the conical superuncus (Fig.
<figureCitation id="11B5B56B257CABD34BA727B056C28428" captionStart="Figures 1927" captionStartId="F4" captionText="Figures 19 - 27. Adults of Carriola species. a. Male; b. Underside wings of male; c. Female. 19. C. ecnomoda (Swinhoe, 1907); a, b. East Java, Mt. Baluran (GU 26.698; ZSM); c. East Java, Mt. Baluran, (GU 26.699; ZSM); 20. C. thyridophora (Hampson, [1893]); a, b. NE India. Assam (GU 27.442; ZSM); c. SW Kampuchea, Kirirom Nat. Park (GU 28.073; ZSM); 21 C. witti sp. nov.; a, b. HT (GU 25.852; ZSM); c. PT (GU 25.853; ZSM); 22. C. seminsula (Strand, 1914); a, b. Sud Vietnam, Bao Loc (GU 27.431; ZSM); c. Nord Vietnam, Mai Chau (GU 27.445; ZSM); 23. C. saturnioides (Snellen, 1879); a, b. N. Sulawesi, (GU 21.966; ZSM); c. Sulawesi, Namo (GU 25.850; ZSM); 24. C. polyakovi sp. nov.; a, b. HT (GU 28.779; ZSM); c. PT (GU 28.780; ZSM); 25. C. shorokhovi sp. nov.; a, b. HT (GU 26.961; ZSM); c. PT (GU 28.784; ZSM); 26. C. fenestrata (Hampson, [1893]); a, b. Vietnam, prov. Thai Nguyen (GU 27.435; ZSM); c. Vietnam, prov. Thai Nguyen, (GU 27.436; ZSM); 27. C. zolotuhini sp. nov.; a, b. HT (GU 28.069; ZSM). Scale bar represents 10 mm, all specimens to scale." figureDoi="10.3897/nl.47.114772.figures19-27" httpUri="https://binary.pensoft.net/fig/1003323" pageId="0" pageNumber="57">23c</figureCitation>
) and the rhomboid shape of the valva, which is widened in the medial part and narrows strongly towards the curved crochet tip of the valva. The width of the basal part of the valva is in a ratio of 1:2.5 to the length (Fig.
<figureCitation id="1CB86EF74EB871FF4142BC733419A96B" captionStart="Figures 2836" captionStartId="F5" captionText="Figures 28 - 36. Male genitalia of Carriola species. a. Genitalia; b. Aedeagus with everted vesica; c. Superuncus (VIII tergum). 28. C. ecnomoda (Swinhoe, 1907) (GU 26.698; ZSM); 29. C. thyridophora (Hampson, [1893]) (GU 27.442; ZSM); 30. C. witti sp. nov. HT (GU 25.852; ZSM); 31. C. seminsula (Strand, 1914): HT (prep. NO 597; SMF); 32. C. saturnioides (Snellen, 1879) (GU 21.966; ZSM); 33. C. polyakovi sp. nov. HT (GU 28.779; ZSM); 34. C. shorokhovi sp. nov. HT (GU 26.961; ZSM); 35. C. fenestrata (Hampson, [1893]) (GU 27.435; ZSM); 36. C. zolotuhini sp. nov. HT (GU 28.069; ZSM). Scale bar represents 1 mm, all genitalia to scale." figureDoi="10.3897/nl.47.114772.figures28-36" httpUri="https://binary.pensoft.net/fig/1003324" pageId="0" pageNumber="57">32a</figureCitation>
). The female genitalia are characterised by the narrow ductus bursae and the clearly regular triangular shape of the signum. In the centre, the signum is 2.5 times as thick as the width of the lobes (Fig.
<figureCitation id="8AD72EC44D17F4238257B3DCD43BC10F" captionStart="Figures 3744" captionStartId="F6" captionText="Figures 37 - 44. Female genitalia of Carriola species. 37. C. ecnomoda (Swinhoe, 1907) (GU 26.699; ZSM); 38. C. thyridophora (Hampson, [1893]) (GU 28.073; ZSM); 39. C. witti sp. nov. PT (GU 25.853; ZSM); 40. C. seminsula (Strand, 1914) (GU 27.445; ZSM); 41. C. saturnioides (Snellen, 1879) (GU 25.850; ZSM); 42. C. polyakovi sp. nov. PT (GU 28.780; ZSM); 43. C. shorokhovi sp. nov. PT (GU 28.784; ZSM); 44. C. fenestrata (Hampson, [1893]) (GU 27.436; ZSM). Scale bar represents 1 mm, all genitalia to scale." figureDoi="10.3897/nl.47.114772.figures37-44" httpUri="https://binary.pensoft.net/fig/1003325" pageId="0" pageNumber="57">41</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="1A61134E181E2471B27437C70F97E70E" pageId="0" pageNumber="57" type="biology_ecology">
<paragraph id="22CC38CF83DF79BC15D166ACBD026412" pageId="0" pageNumber="57">Biology.</paragraph>
<paragraph id="8EB45151F5FCCA2872F4D56DF46F8D13" pageId="0" pageNumber="57">Unknown. The flight period is from February to December. The species has been collected in forests at elevations of up to 1300 m.a.s.l.</paragraph>
</subSubSection>
<subSubSection id="1F2E0F5FD6FE96053432585F5C8D3348" pageId="0" pageNumber="57" type="distribution">
<paragraph id="A8D655A199055A96E50F98FCC0EFD411" pageId="0" pageNumber="57">Distribution.</paragraph>
<paragraph id="EC3155F69FF21688ECB679FD95712EEB" pageId="0" pageNumber="57">
(Fig.
<figureCitation id="8434CAE3F273C45193FA0FBD1AF3E5AC" captionStart="Figures 4553" captionStartId="F7" captionText="Figures 45 - 53. Distribution maps of Carriola species. 45. C. ecnomoda (Swinhoe, 1907); 46. C. thyridophora (Hampson, [1893]); 47. C. witti sp. nov.; 48. C. seminsula (Strand, 1914); 49. C. saturnioides (Snellen, 1879); 50. C. polyakovi sp. nov.; 51. C. shorokhovi sp. nov.; 52. C. fenestrata (Hampson, [1893]); 53. C. zolotuhini sp. nov." figureDoi="10.3897/nl.47.114772.figures45-53" httpUri="https://binary.pensoft.net/fig/1003326" pageId="0" pageNumber="57">49</figureCitation>
). Indonesia (Sulawesi, and Peleng Island off the east coast of Sulawesi).
</paragraph>
</subSubSection>
</treatment>
</document>