treatments-xml/data/53/3C/87/533C879D2A53FF8ECDBAFA32C529B5BE.xml

<document id="538592D6D1EFDB626D018854831E84E9" ID-DOI="10.5281/zenodo.194925" ID-GBIF-Dataset="d8bca122-7661-4684-bc08-b2075d2f0232" ID-ISSN="1175-5326" ID-Zenodo-Dep="194925" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460219720266" checkinUser="plazi" docAuthor="Nguyen, Truong Quang, Nguyen, Tao Thien, Schmitz, Andreas, Orlov, Nikolai L. &amp; Ziegler, Thomas" docDate="2010" docId="533C879D2A53FF8ECDBAFA32C529B5BE" docLanguage="en" docName="zt02439p068.pdf" docOrigin="Zootaxa 2439" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tropidophorus boehmei Nguyen, Nguyen, Schmitz, Orlov &amp; Ziegler, 2010, sp. nov." docType="treatment" docVersion="7" lastPageNumber="65" masterDocId="AF05FFE52A57FF83CD2DFFB7C144B542" masterDocTitle="A new species of the genus Tropidophorus Duméril &amp; Bibron, 1839 (Squamata: Sauria: Scincidae) from Vietnam" masterLastPageNumber="68" masterPageNumber="53" pageNumber="57" updateTime="1698237492434" updateUser="plazi">
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<mods:title id="0C251A0B481A5E0D512C1D0273070A9D">A new species of the genus Tropidophorus Duméril &amp; Bibron, 1839 (Squamata: Sauria: Scincidae) from Vietnam</mods:title>
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<mods:namePart id="919B26182083EFC27B2899CC8AED7724">Nguyen, Truong Quang</mods:namePart>
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<mods:namePart id="8C5B9BD828519815E2A9F273687BA158">Nguyen, Tao Thien</mods:namePart>
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<mods:namePart id="24FF46B164C25294F7B7F7583F110C9B">Schmitz, Andreas</mods:namePart>
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<mods:namePart id="5EF258597EE1E5D2C38880F84B4D5E9B">Orlov, Nikolai L.</mods:namePart>
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<mods:namePart id="4B54EBB122EDBF8B2EF60DE77F1C9B97">Ziegler, Thomas</mods:namePart>
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<paragraph id="DB2A368B2A53FF87CDBAFA32C359B0DD" blockId="4.[151,541,1413,1474]" box="[151,541,1413,1439]" pageId="4" pageNumber="57">
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<emphasis id="E9E1EA992A53FF87CDBAFA32C359B0DD" bold="true" box="[151,541,1413,1439]" pageId="4" pageNumber="57">
<taxonomicName id="1C954D082A53FF87CDBAFA32C0FDB0DD" box="[151,441,1413,1439]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="57" phylum="Chordata" rank="species" species="boehmei" status="sp. nov.">
<emphasis id="E9E1EA992A53FF87CDBAFA32C0FDB0DD" bold="true" box="[151,441,1413,1439]" italics="true" pageId="4" pageNumber="57">Tropidophorus boehmei</emphasis>
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<taxonomicNameLabel id="F2D257E22A53FF87CCEDFA32C359B0DD" box="[448,541,1413,1439]" pageId="4" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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<paragraph id="DB2A368B2A53FF87CDBAFA1FC05CB080" blockId="4.[151,541,1413,1474]" box="[151,280,1448,1474]" pageId="4" pageNumber="57">
(
<figureCitation id="43AE2A0E2A53FF87CDB2FA1FC1B7B080" box="[159,243,1448,1474]" captionStart="FIGURE 2" captionStartId="5.[151,256,1715,1739]" captionTargetBox="[151,1434,418,1694]" captionTargetId="figure@5.[151,1436,418,1694]" captionTargetPageId="5" captionText="FIGURE 2. Dorsolateral (A) and ventral (B) views of the holotype (VNMN 822) of Tropidophorus boehmei sp. nov. from Sa Pa, Lao Cai Province, northern Vietnam. Photo T. T. Nguyen." httpUri="https://zenodo.org/record/194927/files/figure.png" pageId="4" pageNumber="57">Figs. 2</figureCitation>
, 
<figureCitation id="43AE2A0E2A53FF87CDD2FA1FC04AB080" box="[255,270,1448,1474]" captionStart="FIGURE 3" captionStartId="7.[151,256,1960,1984]" captionTargetBox="[167,1413,199,1911]" captionTargetId="figure@7.[167,1432,170,1938]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal (A), ventral (B), and lateral (c) views of the head of the holotype (VNMN 822) of Tropidophorus boehmei sp. nov. from Sa Pa, Lao Cai Province, northern Vietnam. Photo T. Ziegler." httpUri="https://zenodo.org/record/194928/files/figure.png" pageId="4" pageNumber="57">3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="938F65002A53FF87CDBAFA45C521B27F" pageId="4" pageNumber="57" type="materials_examined">
<paragraph id="DB2A368B2A53FF87CDBAFA45C2CCB318" blockId="4.[151,1437,1522,2002]" pageId="4" pageNumber="57">
<emphasis id="E9E1EA992A53FF87CDBAFA45C054B34E" bold="true" box="[151,272,1522,1548]" pageId="4" pageNumber="57">
<typeStatus id="042E88292A53FF87CDBAFA45C04FB34E" box="[151,267,1522,1548]" pageId="4" pageNumber="57" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="BD84AE4E2A53FF87CC35FA45C036B34E" box="[280,370,1522,1548]" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="4" pageNumber="57">VNMN</collectionCode>
822, adult male, collected on 
<date id="AF2B104B2A53FF87CFF8FA45C236B34E" box="[725,882,1522,1548]" pageId="4" pageNumber="57" value="2009-06-10">10 June 2009</date>
by Nguyen Thien Tao and Nguyen Van Quang in the forest of Hoang Lien Mountain, near Ban Khoang (
<geoCoordinate id="BEA1504C2A53FF87CE7BF9ADC2F8B376" box="[854,956,1560,1588]" direction="north" orientation="latitude" pageId="4" pageNumber="57" precision="925" value="22.4">22o24’N</geoCoordinate>
and 
<geoCoordinate id="BEA1504C2A53FF87CED4F9ADC528B376" box="[1017,1132,1560,1588]" direction="east" orientation="longitude" pageId="4" pageNumber="57" precision="925" value="103.78333">103o47’E</geoCoordinate>
), Sa Pa District, 
<collectingCountry id="A382761B2A53FF87C816F9ADC42EB376" box="[1339,1386,1562,1588]" name="Laos" pageId="4" pageNumber="57">Lao</collectingCountry>
Cai Province, northern 
<collectingCountry id="A382761B2A53FF87CC55F9F7C09AB318" box="[376,478,1600,1626]" name="Vietnam" pageId="4" pageNumber="57">Vietnam</collectingCountry>
, at an altitude of 
<quantity id="1C6D9B6E2A53FF87CF85F9F7C20EB318" box="[680,842,1600,1626]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="4" pageNumber="57" unit="m" value="1250.0" valueMax="1300.0" valueMin="1200.0">1200–1300 m</quantity>
a.s.l.
</paragraph>
<paragraph id="DB2A368B2A53FF87CDE9F9D2C521B27F" blockId="4.[151,1437,1522,2002]" pageId="4" pageNumber="57">
<emphasis id="E9E1EA992A53FF87CDE9F9D2C00CB33D" bold="true" box="[196,328,1637,1663]" pageId="4" pageNumber="57">
<typeStatus id="042E88292A53FF87CDE9F9D2C007B33D" box="[196,323,1637,1663]" pageId="4" pageNumber="57" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="BD84AE4E2A53FF87CC7DF9D2C0D7B33D" box="[336,403,1637,1663]" pageId="4" pageNumber="57">IEBR</collectionCode>
A.0922 (adult male), 
<collectionCode id="BD84AE4E2A53FF87CFB4F9D2C398B33D" box="[665,732,1637,1663]" pageId="4" pageNumber="57">IEBR</collectionCode>
A.0923 (adult female), 
<collectionCode id="BD84AE4E2A53FF87CED5F9D2C516B33D" box="[1016,1106,1637,1663]" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="4" pageNumber="57">VNMN</collectionCode>
820 (adult female), 
<collectionCode id="BD84AE4E2A53FF87C86EF9D2C4D9B33D" box="[1347,1437,1637,1663]" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="4" pageNumber="57">VNMN</collectionCode>
909 (adult male), 
<collectionCode id="BD84AE4E2A53FF87CC44F93DC087B3E6" box="[361,451,1674,1700]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="4" pageNumber="57">MHNG</collectionCode>
2721.49 (adult male), 
<collectionCode id="BD84AE4E2A53FF87CFFDF93DC266B3E6" LSID="urn:lsid:biocol.org:col:34613" box="[720,802,1674,1700]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="4" pageNumber="57">ZFMK</collectionCode>
89136 (adult male), 
<collectionCode id="BD84AE4E2A53FF87C934F93DC52FB3E6" LSID="urn:lsid:biocol.org:col:34613" box="[1049,1131,1674,1700]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="4" pageNumber="57">ZFMK</collectionCode>
89137 (adult female), the same data as the 
<typeStatus id="042E88292A53FF87CC73F907C08CB388" box="[350,456,1712,1738]" pageId="4" pageNumber="57" type="holotype">holotype</typeStatus>
; 
<collectionCode id="BD84AE4E2A53FF87CCF9F907C354B388" box="[468,528,1712,1738]" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="4" pageNumber="57">ZISP</collectionCode>
25745 (adult female) collected on 
<date id="AF2B104B2A53FF87CE87F907C57CB388" box="[938,1080,1712,1738]" pageId="4" pageNumber="57" value="2004-05-06">6 May 2004</date>
by Nikolai Orlov in the forest of Hoang Lien Mountain, near Na Nheo (
<geoCoordinate id="BEA1504C2A53FF87CFA7F96FC3B5B3B0" box="[650,753,1749,1778]" direction="north" orientation="latitude" pageId="4" pageNumber="57" precision="925" value="21.983334">21o59’N</geoCoordinate>
, 
<geoCoordinate id="BEA1504C2A53FF87CFD3F96FC234B3B0" box="[766,880,1749,1778]" direction="east" orientation="longitude" pageId="4" pageNumber="57" precision="925" value="104.333336">104o20’E</geoCoordinate>
), Khanh Yen Ha Commune, Van Ban District, 
<collectingCountry id="A382761B2A53FF87CDBAF949C181B25A" box="[151,197,1790,1816]" name="Laos" pageId="4" pageNumber="57">Lao</collectingCountry>
Cai Province, 
<collectingCountry id="A382761B2A53FF87CC59F949C09EB25A" box="[372,474,1790,1816]" name="Vietnam" pageId="4" pageNumber="57">Vietnam</collectingCountry>
, at an altitude of 
<quantity id="1C6D9B6E2A53FF87CF85F949C245B25A" box="[680,769,1790,1816]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="57" unit="m" value="1000.0">1000 m</quantity>
a.s.l.; and 
<collectionCode id="BD84AE4E2A53FF87CEAFF949C293B25A" box="[898,983,1790,1816]" pageId="4" pageNumber="57">VNUH</collectionCode>
S.162 (Bourret’s collection in Hanoi), collected from “Chapa, 
<collectingCountry id="A382761B2A53FF87CC82F894C341B27F" box="[431,517,1827,1853]" name="Vietnam" pageId="4" pageNumber="57">Tonkin</collectingCountry>
” (now Sa Pa, 
<collectingCountry id="A382761B2A53FF87CF84F894C393B27F" box="[681,727,1827,1853]" name="Laos" pageId="4" pageNumber="57">Lao</collectingCountry>
Cai Province, northern 
<collectingCountry id="A382761B2A53FF87CEDDF894C51DB27F" box="[1008,1113,1827,1853]" name="Vietnam" pageId="4" pageNumber="57">Vietnam</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="938F65002A53FF86CDE9F8FFC197B42E" lastPageId="5" lastPageNumber="58" pageId="4" pageNumber="57" type="diagnosis">
<paragraph id="DB2A368B2A53FF86CDE9F8FFC197B42E" blockId="4.[151,1437,1522,2002]" lastBlockId="5.[151,1436,152,364]" lastPageId="5" lastPageNumber="58" pageId="4" pageNumber="57">
<emphasis id="E9E1EA992A53FF87CDE9F8FFC007B220" bold="true" box="[196,323,1864,1890]" pageId="4" pageNumber="57">Diagnosis.</emphasis>
Size large, maximum SVL up to 
<quantity id="1C6D9B6E2A53FF87CFFEF8FFC209B220" box="[723,845,1864,1890]" metricMagnitude="-1" metricUnit="m" metricValue="1.035" pageId="4" pageNumber="57" unit="mm" value="103.5">103.5 mm</quantity>
; upper head scales smooth; frontonasal undivided; prefrontals small, widely separated from each other; nuchals 1–4; supralabials 6; supraciliaries 7–8, supraciliary row completed along the entire length of the lateral edge of supraoculars; external ear opening present, tympanum diameter smaller than eye length; midbody scale rows 30–32; dorsal and lateral scales smooth; paravertebral scales 60–69, not widened; ventral scales 56–66; scale rows at position of tenth subcaudal 17–18; medial subcaudals divided from first to fifth, remaining ones approximately 1.5 times wider than neighboring scales; limbs well developed, dorsal surface covered by smooth scales; lamellae smooth, 16–19 under fourth toe; dorsum and tail base dark grey with light transverse bands and two dorsolateral rows of white dots from neck to tail base; body sides dark grey with small white dots from behind the eye to tail base.
</paragraph>
</subSubSection>
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<paragraph id="DB2A368B2A52FF86CDBAF904C2C7B3A9" blockId="5.[151,1436,1715,1771]" pageId="5" pageNumber="58">
<emphasis id="E9E1EA992A52FF86CDBAF904C05EB389" bold="true" box="[151,282,1715,1739]" pageId="5" pageNumber="58">FIGURE 2.</emphasis>
Dorsolateral (A) and ventral (B) views of the holotype (VNMN 822) of 
<taxonomicName id="1C954D082A52FF86C911F903C47AB389" box="[1084,1342,1716,1739]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="58" phylum="Chordata" rank="species" species="boehmei" status="sp. nov.">
<emphasis id="E9E1EA992A52FF86C911F903C47AB389" box="[1084,1342,1716,1739]" italics="true" pageId="5" pageNumber="58">Tropidophorus boehmei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2D257E22A52FF86C868F904C4D8B389" box="[1349,1436,1715,1739]" pageId="5" pageNumber="58" rank="species">
<emphasis id="E9E1EA992A52FF86C868F904C4D8B389" bold="true" box="[1349,1436,1715,1739]" pageId="5" pageNumber="58">sp. nov.</emphasis>
</taxonomicNameLabel>
from Sa Pa, Lao Cai Province, northern Vietnam. Photo T. T. Nguyen.
</paragraph>
</caption>
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<paragraph id="DB2A368B2A52FF88CDE9F8A4C59CB7BD" blockId="5.[151,1436,1811,2025]" lastBlockId="11.[151,1437,152,2034]" lastPageId="11" lastPageNumber="64" pageId="5" pageNumber="58">
<emphasis id="E9E1EA992A52FF86CDE9F8A4C0AAB26F" bold="true" box="[196,494,1811,1837]" pageId="5" pageNumber="58">
Description of 
<typeStatus id="042E88292A52FF86CC51F8A4C0AEB26F" box="[380,490,1811,1837]" pageId="5" pageNumber="58" type="holotype">holotype</typeStatus>
.
</emphasis>
Size large (SVL 
<quantity id="1C6D9B6E2A52FF86CFEFF8A3C26AB26C" box="[706,814,1812,1838]" metricMagnitude="-2" metricUnit="m" metricValue="9.91" pageId="5" pageNumber="58" unit="mm" value="99.1">99.1 mm</quantity>
, TaL 
<quantity id="1C6D9B6E2A52FF86CE42F8A3C2AFB26C" box="[879,1003,1812,1838]" metricMagnitude="-1" metricUnit="m" metricValue="1.388" pageId="5" pageNumber="58" unit="mm" value="138.8">138.8 mm</quantity>
), for further measurements see Table 2; head longer than wide; upper head scales smooth; snout obtuse, rounded anteriorly; rostral wider than high, visible from above; supranasals absent; frontonasal as long as wide, in contact with rostral, nasals, anterior loreals, frontal, and prefrontals; prefrontals very small, widely separated from each other; frontal narrowing posteriorly, 1.5 times longer than the distance to the snout, in contact with frontonasal, prefrontals, first to third supraoculars, and frontoparietals; frontoparietals in contact anteriorly, and bordered by frontal, third and fourth supraoculars, parietals and interparietal; interparietal small, without small transparent spot; parietals in contact posteriorly, posterolateral border surrounded by 4 scales on each side; nuchal scales in 2 pairs; nostril in centre of nasal; loreals 2, anterior loreal higher but narrower than posterior one; anterior loreal undivided, posterior loreal in contact with anterior loreal, prefrontal, first supraciliary, preocular, upper presubocular, and second supralabial; preoculars 2; presuboculars 2, anterior large, posterior one in contact with third and fourth supralabials; supraciliaries 8, first largest, supraciliary row completed along the entire length of the lateral edge of the supraoculars; supraoculars 4, second widest, followed by a small postsupraocular; postocular single; pretemporal single; postsuboculars 4, lower one in contact with fourth and fifth supralabials; primary temporals 2, lower one in contact with fifth and sixth supralabials; secondary temporals 3, upper one very large and in contact with parietal; lower eyelid moveable, with four enlarged opaque scales, separated from supralabials by two rows of small scales; supralabials 6, fourth largest and below the eye; a shallow groove on loreal-labial border, from posterior corner of nasal across subocular obliquely downward to the end of fourth supralabial; external ear opening ovoid, without lobules; tympanum superficial, tympanum diameter much smaller than eye length; mental wider than long, rounded anteriorly, in contact with first infralabial on each side and postmental; infralabials 5, first longest; postmental undivided, in contact with mental, first infralabial on both sides, and anterior pair of chinshields; chinshields in 3 pairs, first pair in contact anteriorly, second pair separated by a gular scale, third pair very small and separated by three scales; first and second pairs of chinshields in contact with infralabials, third pair separated from infralabials by small scales.
</paragraph>
<caption id="8FEA66032A50FF84CDBAF81FC55FB2A2" httpUri="https://zenodo.org/record/194928/files/figure.png" pageId="7" pageNumber="60" targetBox="[167,1413,199,1911]" targetPageId="7">
<paragraph id="DB2A368B2A50FF84CDBAF81FC55FB2A2" blockId="7.[151,1436,1960,2016]" pageId="7" pageNumber="60">
<emphasis id="E9E1EA992A50FF84CDBAF81FC05EB282" bold="true" box="[151,282,1960,1984]" pageId="7" pageNumber="60">FIGURE 3.</emphasis>
Dorsal (A), ventral (B), and lateral (c) views of the head of the holotype (VNMN 822) of 
<taxonomicName id="1C954D082A50FF84C9D0F81EC1B5B2A2" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="60" phylum="Chordata" rank="species" species="boehmei" status="sp. nov.">
<emphasis id="E9E1EA992A50FF84C9D0F81EC1B5B2A2" italics="true" pageId="7" pageNumber="60">Tropidophorus boehmei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2D257E22A50FF84CDD5F87FC00AB2A2" box="[248,334,1992,2016]" pageId="7" pageNumber="60" rank="species">
<emphasis id="E9E1EA992A50FF84CDD5F87FC00AB2A2" bold="true" box="[248,334,1992,2016]" pageId="7" pageNumber="60">sp. nov.</emphasis>
</taxonomicNameLabel>
from Sa Pa, Lao Cai Province, northern Vietnam. Photo T. Ziegler.
</paragraph>
</caption>
<caption id="8FEA66032A5FFF8BCDBAF803C1BCB2A9" httpUri="https://zenodo.org/record/194929/files/figure.png" pageId="8" pageNumber="61" targetBox="[204,1380,186,1951]" targetPageId="8">
<paragraph id="DB2A368B2A5FFF8BCDBAF803C1BCB2A9" blockId="8.[151,1436,1972,2027]" pageId="8" pageNumber="61">
<emphasis id="E9E1EA992A5FFF8BCDBAF803C05DB28E" bold="true" box="[151,281,1972,1996]" pageId="8" pageNumber="61">FIGURE 4.</emphasis>
Map showing the distribution (red circle) of 
<taxonomicName id="1C954D082A5FFF8BCFD2F803C2B8B289" box="[767,1020,1972,1995]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="61" phylum="Chordata" rank="species" species="boehmei" status="sp. nov.">
<emphasis id="E9E1EA992A5FFF8BCFD2F803C2B8B289" box="[767,1020,1972,1995]" italics="true" pageId="8" pageNumber="61">Tropidophorus boehmei</emphasis>
</taxonomicName>
<emphasis id="E9E1EA992A5FFF8BC929F803C51DB28E" bold="true" box="[1028,1113,1972,1996]" pageId="8" pageNumber="61">
<taxonomicNameLabel id="F2D257E22A5FFF8BC929F803C51DB28E" box="[1028,1113,1972,1996]" pageId="8" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in Lao Cai Province, northern Vietnam.
</paragraph>
</caption>
<paragraph id="DB2A368B2A5CFF88CDE9FCBFC03DB69E" blockId="11.[151,1437,152,2034]" pageId="11" pageNumber="64">Midbody scales in 30 rows; dorsal scales smooth, subequal to lateral and ventral scales, in 8 rows across the back; paravertebral scales 64, not widened; lateral scales smooth; ventral scales smooth, in 65 rows; precloacals 2+2, inner precloacals overlapping outer ones, central two enlarged, right one overlapped by left one; partially everted hemipenis bifurcated apically; tail thick at base, 17 scale rows at position of tenth subcaudal; medial subcaudals divided from first to third, remaining ones about 1.5 timer wider than neighboring scales.</paragraph>
<paragraph id="DB2A368B2A5CFF88CDE9FC5FC3A6B10E" blockId="11.[151,1437,152,2034]" pageId="11" pageNumber="64">Limbs well developed, pentadactyl; third and fourth fingers equal in length; scales on dorsal surface of basal digit IV in two rows and terminal three scales in single row; subdigital lamellae smooth, numbering 12/ 12 under fourth finger and 16/17 under fourth toe.</paragraph>
<paragraph id="DB2A368B2A5CFF88CDE9FBE0C579B06E" blockId="11.[151,1437,152,2034]" pageId="11" pageNumber="64">
<emphasis id="E9E1EA992A5CFF88CDE9FBE0C091B133" bold="true" box="[196,469,1111,1137]" pageId="11" pageNumber="64">Coloration in alcohol.</emphasis>
Dorsal head, back, and tail base dark grey with indistinct light transverse bands, faded from anterior to posterior: one between anterior corners of eye, two on neck, seven between limb insertions, three on tail base; dorsum with two rows of white dots from neck to the point above vent; body sides dark grey with irregular white dots from behind the eye to groin; supralabials and infralabials with light dots; limbs dorsally dark grey with white dots; chin and throat grey; venter and underside of tail base brownish cream; underside of tail tip dark grey. For coloration in life see 
<figureCitation id="43AE2A0E2A5CFF88CEDFFAA5C57DB06E" box="[1010,1081,1298,1324]" captionStart="FIGURE 2" captionStartId="5.[151,256,1715,1739]" captionTargetBox="[151,1434,418,1694]" captionTargetId="figure@5.[151,1436,418,1694]" captionTargetPageId="5" captionText="FIGURE 2. Dorsolateral (A) and ventral (B) views of the holotype (VNMN 822) of Tropidophorus boehmei sp. nov. from Sa Pa, Lao Cai Province, northern Vietnam. Photo T. T. Nguyen." httpUri="https://zenodo.org/record/194927/files/figure.png" pageId="11" pageNumber="64">Fig. 2</figureCitation>
.
</paragraph>
<paragraph id="DB2A368B2A5CFF88CDE9FA80C37FB080" blockId="11.[151,1437,152,2034]" pageId="11" pageNumber="64">
<emphasis id="E9E1EA992A5CFF88CDE9FA80C0A2B013" bold="true" box="[196,486,1335,1361]" pageId="11" pageNumber="64">
Variation of 
<typeStatus id="042E88292A5CFF88CC4EFA80C0A5B013" box="[355,481,1335,1361]" pageId="11" pageNumber="64" type="paratype">paratypes</typeStatus>
.
</emphasis>
Measurements, pholidosis and color pattern of the 
<typeStatus id="042E88292A5CFF88C965FA8FC5FDB010" box="[1096,1209,1336,1362]" pageId="11" pageNumber="64" type="paratype">paratypes</typeStatus>
are summarized in Tables 2 and 3. The specimen from Van Ban (ZISP 25745) has parietals separated by a small scale posteriorly, and midbody scales in 32 rows. Males are somewhat larger than females (SVL 89.7 ± 
<quantity id="1C6D9B6E2A5CFF88C9BEFA35C5BBB0DE" box="[1171,1279,1410,1436]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="11" pageNumber="64" unit="mm" value="14.3">14.3 mm</quantity>
, n = 6 versus 72.9 ± 
<quantity id="1C6D9B6E2A5CFF88CDC4FA1FC002B080" box="[233,326,1448,1474]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="11" pageNumber="64" unit="mm" value="9.4">9.4 mm</quantity>
, n = 4, respectively).
</paragraph>
</subSubSection>
<subSubSection id="938F65002A5CFF88CDE9FA7AC258B385" pageId="11" pageNumber="64" type="distribution">
<paragraph id="DB2A368B2A5CFF88CDE9FA7AC22DB34E" blockId="11.[151,1437,152,2034]" pageId="11" pageNumber="64">
<emphasis id="E9E1EA992A5CFF88CDE9FA7AC020B0A5" bold="true" box="[196,356,1485,1511]" pageId="11" pageNumber="64">Distribution.</emphasis>
<taxonomicName id="1C954D082A5CFF88CC41FA7AC3C0B0A4" box="[364,644,1485,1510]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="64" phylum="Chordata" rank="species" species="boehmei" status="sp. nov.">
<emphasis id="E9E1EA992A5CFF88CC41FA7AC3C0B0A4" box="[364,644,1485,1510]" italics="true" pageId="11" pageNumber="64">Tropidophorus boehmei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2D257E22A5CFF88CFA1FA7AC3AEB0A5" box="[652,746,1485,1511]" pageId="11" pageNumber="64" rank="species">
<emphasis id="E9E1EA992A5CFF88CFA1FA7AC3AEB0A5" bold="true" box="[652,746,1485,1511]" pageId="11" pageNumber="64">sp. nov.</emphasis>
</taxonomicNameLabel>
is currently known only from Hoang Lien Mountain in Sa Pa and Van Ban districts, 
<collectingCountry id="A382761B2A5CFF88CCE9FA45C0B6B34E" box="[452,498,1522,1548]" name="Laos" pageId="11" pageNumber="64">Lao</collectingCountry>
Cai Province, 
<collectingCountry id="A382761B2A5CFF88CFB2FA45C246B34E" box="[671,770,1522,1548]" name="Vietnam" pageId="11" pageNumber="64">Vietnam</collectingCountry>
(
<figureCitation id="43AE2A0E2A5CFF88CE3FFA45C21FB34E" box="[786,859,1522,1548]" captionStart="FIGURE 4" captionStartId="8.[151,255,1972,1996]" captionTargetBox="[204,1380,186,1951]" captionTargetId="figure@8.[204,1386,186,1955]" captionTargetPageId="8" captionText="FIGURE 4. Map showing the distribution (red circle) of Tropidophorus boehmei sp. nov. in Lao Cai Province, northern Vietnam." httpUri="https://zenodo.org/record/194929/files/figure.png" pageId="11" pageNumber="64">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="DB2A368B2A5CFF88CDE9F9A0C258B385" blockId="11.[151,1437,152,2034]" pageId="11" pageNumber="64">
<emphasis id="E9E1EA992A5CFF88CDE9F9A0C0CFB373" bold="true" box="[196,395,1559,1585]" pageId="11" pageNumber="64">Natural history.</emphasis>
Specimens were collected in the late afternoon and in the evening (1600–2100 h) in and nearby rocky forest streams. When disturbed they disappeared in the water or in rock holes. The stomach content of the female (IEBR A.0923) contained remains of a spider (Arachnida). Stomachs were empty in VNMN 820, 909, and ZFMK 89137. The 
<typeStatus id="042E88292A5CFF88CF86F93FC25DB3E0" box="[683,793,1672,1698]" pageId="11" pageNumber="64" type="holotype">holotype</typeStatus>
had white testes (
<quantity id="1C6D9B6E2A5CFF88C92CF93FC527B3E0" box="[1025,1123,1672,1698]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="11" pageNumber="64" unit="mm" value="7.1">7.1 mm</quantity>
in length and 
<quantity id="1C6D9B6E2A5CFF88C836F93FC438B3E0" box="[1307,1404,1672,1698]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="11" pageNumber="64" unit="mm" value="3.4">3.4 mm</quantity>
in diameter). The females contained small white follicles.
</paragraph>
</subSubSection>
<subSubSection id="938F65002A5CFF8FCDE9F965C055B595" lastPageId="12" lastPageNumber="65" pageId="11" pageNumber="64" type="etymology">
<paragraph id="DB2A368B2A5CFF88CDE9F965C20AB2C0" blockId="11.[151,1437,152,2034]" pageId="11" pageNumber="64">
<emphasis id="E9E1EA992A5CFF88CDE9F965C01CB3AE" bold="true" box="[196,344,1746,1772]" pageId="11" pageNumber="64">Etymology.</emphasis>
We name this new species in honor of Prof. Dr. Wolfgang Böhme, vice director of the Zoological Research Museum Alexander Koenig in Bonn, head of the Vertebrate Section and curator for herpetology, in recognition of his outstanding contributions towards lizard systematics and training of many herpetologist generations. For the common names we suggest Boehme’s Water Skink (English), Thằn lằn tai boê-me (Vietnamese), and Böhmes Wasserskink (
<collectingCountry id="A382761B2A5CFF88CFF0F8DFC206B2C0" box="[733,834,1896,1922]" name="Germany" pageId="11" pageNumber="64">German</collectingCountry>
).
</paragraph>
<paragraph id="DB2A368B2A5CFF8FCDE9F83AC055B595" blockId="11.[151,1437,152,2034]" lastBlockId="12.[151,1437,151,290]" lastPageId="12" lastPageNumber="65" pageId="11" pageNumber="64">
<emphasis id="E9E1EA992A5CFF88CDE9F83AC030B2E5" bold="true" box="[196,372,1933,1959]" pageId="11" pageNumber="64">Comparisons.</emphasis>
Based on data from the literature (e.g., 
<bibRefCitation id="BF044B7A2A5CFF88CE7BF83AC29BB2E5" author="Blyth" box="[854,991,1933,1959]" pageId="11" pageNumber="67" refString="Blyth, E. (1853) Notices and descriptions of various reptiles, new or little known. The Journal of the Asiatic Society of Bengal, Calcutta, 22, 639 - 655." type="journal article" year="1853">Blyth 1853</bibRefCitation>
; 
<bibRefCitation id="BF044B7A2A5CFF88CEC0F83AC5F4B2E5" author="Boulenger" box="[1005,1200,1933,1959]" pageId="11" pageNumber="67" refString="Boulenger, G. A. (1887) Catologue of the lizards in the British Museum (Natural History). Volume III, Trustees, London, xii + 575 pp. + XL pls." type="book" year="1887">Boulenger 1887</bibRefCitation>
; 
<bibRefCitation id="BF044B7A2A5CFF88C993F83AC40FB2E5" author="Smith" box="[1214,1355,1933,1959]" pageId="11" pageNumber="67" refString="Smith, M. A. (1919) The lizards of the genus Tropidophorus in Siam, with description of two new species. Journal of Natural History Society of Siam, 3, 223 - 228." type="journal article" year="1919">Smith 1919</bibRefCitation>
, 
<bibRefCitation id="BF044B7A2A5CFF88C874F83AC4DCB2E5" author="Smith" box="[1369,1432,1933,1959]" pageId="11" pageNumber="67" refString="Smith, M. A. (1923) A review of the lizards of the genus Tropidophorus on the Asiatic mainland. Proceedings of the Zoological Society of London, 1923, 775 - 781." type="proceedings paper" year="1923">1923</bibRefCitation>
, 
<bibRefCitation id="BF044B7A2A5CFF88CDBAF805C191B28E" author="Smith" box="[151,213,1970,1996]" pageId="11" pageNumber="67" refString="Smith, M. A. (1935) The fauna of British India, including Ceylon and Burma: Reptilia and Amphibia. Vol. 2 - Sauria. Taylor and Francis, London, xii + 440 pp." type="book" year="1935">1935</bibRefCitation>
; 
<bibRefCitation id="BF044B7A2A5CFF88CDCFF805C030B28E" author="Taylor" box="[226,372,1970,1996]" pageId="11" pageNumber="67" refString="Taylor, E. H. (1963) The lizards of Thailand. The University of Kansas Science Bulletin, 44, 687 - 1077." type="journal article" year="1963">Taylor 1963</bibRefCitation>
; 
<bibRefCitation id="BF044B7A2A5CFF88CCACF805C0B8B28E" author="Wen" box="[385,508,1970,1996]" pageId="11" pageNumber="68" refString="Wen, Y. - T. (1992) A new species of the genus Tropidophorus (Reptilia: Lacertilia) from Guangxi Zhang Autonomous Region, China. Asiatic Herpetological Research, 4, 18 - 22." type="journal article" year="1992">Wen 1992</bibRefCitation>
; 
<bibRefCitation id="BF044B7A2A5CFF88CF24F805C381B28E" author="Ngo" box="[521,709,1970,1996]" pageId="11" pageNumber="67" refString="Ngo, A., Murphy, R. W., Orlov, N. L., Darevsky, I. S. &amp; Nguyen, V. S. (2000) A redescription of the Ba Vi water skink Tropidophorus baviensis Bourret, 1939. Russian Journal of Herpetology, 7, 155 - 158." type="journal article" year="2000">
Ngo 
<emphasis id="E9E1EA992A5CFF88CF6EF804C333B28E" box="[579,631,1971,1996]" italics="true" pageId="11" pageNumber="64">et al</emphasis>
. 2000
</bibRefCitation>
; 
<bibRefCitation id="BF044B7A2A5CFF88CFFFF805C2EFB28E" author="Hikida" box="[722,939,1970,1996]" pageId="11" pageNumber="67" refString="Hikida, T., Orlov, N. L., Nabhitabhata, J. &amp; Ota, H. (2002) Three new depressed-bodied water skinks of the genus Tropidophorus (Lacertilia: Scincidae) from Thailand and Vietnam. Current Herpetology, 21, 9 - 23." type="journal article" year="2002">
Hikida 
<emphasis id="E9E1EA992A5CFF88CE04F804C21AB28E" box="[809,862,1971,1996]" italics="true" pageId="11" pageNumber="64">et al</emphasis>
. 2002
</bibRefCitation>
; 
<bibRefCitation id="BF044B7A2A5CFF88CE95F805C5F8B28E" author="Greer" box="[952,1212,1970,1996]" pageId="11" pageNumber="67" refString="Greer, A. E. &amp; Biswas, S. (2004) A generic diagnosis for the Southeast Asian scincid lizard genus Tropidophorus Dumeril &amp; Bibron, 1839 with some additional comments on its morphology and distribution. Journal of Herpetology, 38, 426 - 430." type="journal article" year="2004">Greer &amp; Biswas 2004</bibRefCitation>
; 
<bibRefCitation id="BF044B7A2A5CFF88C9E4F805C192B2B0" author="Chuaynkern" pageId="11" pageNumber="67" refString="Chuaynkern, Y., Nabhitabhata, J., Inthara, C., Kamsook, M. &amp; Somsri, K. (2005) A new species of the water skink Tropidophorus (Reptilia: Squamata: Scincidae) from northeastern Thailand. The Thailand Natural History Museum Journal, 1, 165 - 176." type="journal article" year="2005">
Chuaynkern 
<emphasis id="E9E1EA992A5CFF88C84DF804C4D0B28E" box="[1376,1428,1971,1996]" italics="true" pageId="11" pageNumber="64">et al</emphasis>
. 2005
</bibRefCitation>
; 
<bibRefCitation id="BF044B7A2A5CFF88CDCFF86FC085B2B0" author="Ziegler" box="[226,449,2008,2034]" pageId="11" pageNumber="68" refString="Ziegler, T., Vu, N. T. &amp; Bui, N. T. (2005) A new water skink of the genus Tropidophorus from the Phong Nha - Ke Bang National Park, central Vietnam (Squamata: Sauria: Scincidae). Salamandra, 41, 137 - 146." type="journal article" year="2005">
Ziegler 
<emphasis id="E9E1EA992A5CFF88CC12F86FC030B2B3" box="[319,372,2008,2033]" italics="true" pageId="11" pageNumber="64">et al</emphasis>
. 2005
</bibRefCitation>
, 
<bibRefCitation id="BF044B7A2A5CFF88CCE3F86FC349B2B0" author="Ziegler" box="[462,525,2008,2034]" pageId="11" pageNumber="68" refString="Ziegler, T., Heidrich, A., Schmitz, A. &amp; Bohme, W. (2007) Phylogenetic position, morphology and natural history of the Vietnamese water skink Tropidophorus noggei Ziegler, Vu &amp; Bui, 2005 (Sauria: Scincidae). Revue suisse de Zoologie, 114, 599 - 614." type="journal article" year="2007">2007</bibRefCitation>
; 
<bibRefCitation id="BF044B7A2A5CFF88CF34F86FC3FEB2B0" author="Bourret" box="[537,698,2008,2034]" pageId="11" pageNumber="67" refString="Bourret, R. (2009) Les lezards de l'Indochine. Edition Chimaira, Frankfurt am Main, 346 pp." type="book" year="2009">Bourret 2009</bibRefCitation>
; 
<bibRefCitation id="BF044B7A2A5CFF88CFEBF86FC2E8B2B0" author="Nguyen" box="[710,940,2008,2034]" pageId="11" pageNumber="67" refString="Nguyen, Q. T., Nguyen, V. S., Nikolai, O., Hoang, N. T., Bohme, W. &amp; Ziegler, T. (2010) A review of the genus Tropidophorus (Squamata: Scincidae) from Vietnam with new species records and additional data on natural history. Zoosystematics and Evolution, 86 (1), 5 - 19." type="journal article" year="2010">
Nguyen 
<emphasis id="E9E1EA992A5CFF88CE01F86FC224B2B3" box="[812,864,2008,2033]" italics="true" pageId="11" pageNumber="64">et al</emphasis>
. 2010
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) and specimen-based research (Appendix 1), we compare 
<taxonomicName id="1C954D082A5BFF8FCC4DFF2FC3CCB5F3" box="[352,648,152,177]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="boehmei" status="sp. nov.">
<emphasis id="E9E1EA992A5BFF8FCC4DFF2FC3CCB5F3" box="[352,648,152,177]" italics="true" pageId="12" pageNumber="65">Tropidophorus boehmei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2D257E22A5BFF8FCFBCFF20C3B0B5F3" box="[657,756,151,177]" pageId="12" pageNumber="65" rank="species">
<emphasis id="E9E1EA992A5BFF8FCFBCFF20C3B0B5F3" bold="true" box="[657,756,151,177]" pageId="12" pageNumber="65">sp. nov.</emphasis>
</taxonomicNameLabel>
in the following with congeners from the Indochinese Peninsula:
</paragraph>
</subSubSection>
<subSubSection id="938F65002A5BFF8ECDE9FF55C529B5BE" lastPageId="13" lastPageNumber="66" pageId="12" pageNumber="65" type="description">
<paragraph id="DB2A368B2A5BFF8FCDE9FF55C41CB460" blockId="12.[151,1437,151,290]" pageId="12" pageNumber="65">
<taxonomicName id="1C954D082A5BFF8FCDE9FF55C09FB5B9" box="[196,475,226,251]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="boehmei" status="sp. nov.">
<emphasis id="E9E1EA992A5BFF8FCDE9FF55C09FB5B9" box="[196,475,226,251]" italics="true" pageId="12" pageNumber="65">Tropidophorus boehmei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2D257E22A5BFF8FCCCFFF55C37BB5BE" box="[482,575,226,252]" pageId="12" pageNumber="65" rank="species">
<emphasis id="E9E1EA992A5BFF8FCCCFFF55C37BB5BE" bold="true" box="[482,575,226,252]" pageId="12" pageNumber="65">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from 
<taxonomicName id="1C954D082A5BFF8FCFF7FF55C2D1B5B9" box="[730,917,226,251]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="cocincinensis">
<emphasis id="E9E1EA992A5BFF8FCFF7FF55C2D1B5B9" box="[730,917,226,251]" italics="true" pageId="12" pageNumber="65">T. cocincinensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="1C954D082A5BFF8FCE8EFF55C56FB5B9" box="[931,1067,226,251]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="hainanus">
<emphasis id="E9E1EA992A5BFF8FCE8EFF55C56FB5B9" box="[931,1067,226,251]" italics="true" pageId="12" pageNumber="65">T. hainanus</emphasis>
</taxonomicName>
, 
<taxonomicName id="1C954D082A5BFF8FC914FF55C5ABB5B9" box="[1081,1263,226,251]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="guangxiensis">
<emphasis id="E9E1EA992A5BFF8FC914FF55C5ABB5B9" box="[1081,1263,226,251]" italics="true" pageId="12" pageNumber="65">T. guangxiensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="1C954D082A5BFF8FC9D0FF55C4D0B5B9" box="[1277,1428,226,251]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="microlepis">
<emphasis id="E9E1EA992A5BFF8FC9D0FF55C4D0B5B9" box="[1277,1428,226,251]" italics="true" pageId="12" pageNumber="65">T. microlepis</emphasis>
</taxonomicName>
, 
<taxonomicName id="1C954D082A5BFF8FCDBAFEBFC051B463" box="[151,277,264,289]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="robisoni">
<emphasis id="E9E1EA992A5BFF8FCDBAFEBFC051B463" box="[151,277,264,289]" italics="true" pageId="12" pageNumber="65">T. robisoni</emphasis>
</taxonomicName>
, 
<taxonomicName id="1C954D082A5BFF8FCC0EFEBFC0CBB463" box="[291,399,264,289]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="sinicus">
<emphasis id="E9E1EA992A5BFF8FCC0EFEBFC0CBB463" box="[291,399,264,289]" italics="true" pageId="12" pageNumber="65">T. sinicus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="1C954D082A5BFF8FCCFCFEBFC35FB463" box="[465,539,264,289]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="thai">
<emphasis id="E9E1EA992A5BFF8FCCFCFEBFC35FB463" box="[465,539,264,289]" italics="true" pageId="12" pageNumber="65">T. thai</emphasis>
</taxonomicName>
by having smooth dorsal head scales (versus striated in latter species).
</paragraph>
<caption id="8FEA66032A5BFF8FCDBAFEE5C33CB4CB" pageId="12" pageNumber="65">
<paragraph id="DB2A368B2A5BFF8FCDBAFEE5C33CB4CB" blockId="12.[151,1435,338,580]" pageId="12" pageNumber="65">
<emphasis id="E9E1EA992A5BFF8FCDBAFEE5C04CB428" bold="true" box="[151,264,338,362]" pageId="12" pageNumber="65">TABLE 4.</emphasis>
Comparisons of diagnostic characters in 
<taxonomicName id="1C954D082A5BFF8FCFEEFEE5C225B42B" box="[707,865,338,361]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="genus">
<emphasis id="E9E1EA992A5BFF8FCFEEFEE5C225B42B" box="[707,865,338,361]" italics="true" pageId="12" pageNumber="65">Tropidophorus</emphasis>
</taxonomicName>
with smooth dorsal head scales from the Indochinese Peninsula (* = data obtained from literature).
</paragraph>
</caption>
<paragraph id="DB2A368B2A5BFF8FCCA2FE12C4C4B4F8" blockId="12.[151,1435,338,580]" box="[399,1408,421,442]" pageId="12" pageNumber="65">
<emphasis id="E9E1EA992A5BFF8FCCA2FE12C4C4B4F8" box="[399,1408,421,442]" italics="true" pageId="12" pageNumber="65">
<taxonomicName id="1C954D082A5BFF8FCCA2FE12C0ACB4F8" box="[399,488,421,442]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="baviensis">baviensis</taxonomicName>
<taxonomicName id="1C954D082A5BFF8FCF2AFE12C31CB4F8" box="[519,600,421,442]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="boehmei">boehmei</taxonomicName>
<taxonomicName id="1C954D082A5BFF8FCF52FE12C3A5B4F8" box="[639,737,421,442]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="berdmorei">berdmorei</taxonomicName>
<taxonomicName id="1C954D082A5BFF8FCFD9FE12C209B4F8" box="[756,845,421,442]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="hangnam">hangnam</taxonomicName>
<taxonomicName id="1C954D082A5BFF8FCE46FE12C2E1B4F8" box="[875,933,421,442]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="laotus">laotus</taxonomicName>
<taxonomicName id="1C954D082A5BFF8FCEF9FE12C507B4F8" box="[980,1091,421,442]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="latiscutatus">latiscutatus</taxonomicName>
<taxonomicName id="1C954D082A5BFF8FC974FE12C5E4B4F8" box="[1113,1184,421,442]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="matsuii">matsuii</taxonomicName>
<taxonomicName id="1C954D082A5BFF8FC9E1FE12C458B4F8" box="[1228,1308,421,442]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="murphyi">murphyi</taxonomicName>
<taxonomicName id="1C954D082A5BFF8FC812FE12C4C4B4F8" box="[1343,1408,421,442]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="noggei">noggei</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="DB2A368B2A5BFF8FCDB2FE7AC07FB4BD" blockId="12.[151,1435,338,580]" pageId="12" pageNumber="65">Frontonasal divided (2) 1 1 1 1 2 1 2 1 1 or undivided (1)</paragraph>
<paragraph id="DB2A368B2A5BFF8FCDB2FDA5C013B706" blockId="12.[151,1435,338,580]" pageId="12" pageNumber="65">Prefontals in contact 1, rarely 0 0 0 or 1 1 0 or 1 0 1 1 1or 0 (1) or separated (0)</paragraph>
<paragraph id="DB2A368B2A5BFF8FCDB2FDE0C427B690" pageId="12" pageNumber="65">
<table id="A995C42B2A5B007CCDBAFDE0C4DFB690" box="[151,1435,599,978]" gridcols="7" gridrows="7" pageId="12" pageNumber="65">
<tr id="65A534C92A5B007CCDBAFDE0C4DFB7E5" box="[151,1435,599,679]" gridrow="0" pageId="12" pageNumber="65">
<th id="26745DB52A5B007CCDBAFDE0C0F0B7E5" box="[151,436,599,679]" gridcol="0" gridrow="0" pageId="12" pageNumber="65">Parietals in contact (1) 0 or separated (0) posteriorly</th>
<th id="26745DB52A5B007CCF2AFDE0C25CB7E5" box="[519,792,599,679]" gridcol="1" gridrow="0" pageId="12" pageNumber="65">1, rarely 0 1, rarely 0 0</th>
<th id="26745DB52A5B007CCE46FDE0C2D4B7E5" box="[875,912,599,679]" gridcol="2" gridrow="0" pageId="12" pageNumber="65">1</th>
<th id="26745DB52A5B007CCEF9FDE0C2BCB7E5" box="[980,1016,599,679]" gridcol="3" gridrow="0" pageId="12" pageNumber="65">0</th>
<th id="26745DB52A5B007CC974FDE0C53DB7E5" box="[1113,1145,599,679]" gridcol="4" gridrow="0" pageId="12" pageNumber="65">0</th>
<th id="26745DB52A5B007CC9E1FDE0C5B4B7E5" box="[1228,1264,599,679]" gridcol="5" gridrow="0" pageId="12" pageNumber="65">0</th>
<th id="26745DB52A5B007CC812FDE0C4DFB7E5" box="[1343,1435,599,679]" gridcol="6" gridrow="0" pageId="12" pageNumber="65">0</th>
</tr>
<tr id="65A534C92A5B007CCDBAFD0DC4DFB78D" box="[151,1435,698,719]" gridrow="1" pageId="12" pageNumber="65">
<th id="26745DB52A5B007CCDBAFD0DC0F0B78D" box="[151,436,698,719]" gridcol="0" gridrow="1" pageId="12" pageNumber="65">Loreals 2</th>
<td id="26745DB52A5B007CCF2AFD0DC25CB78D" box="[519,792,698,719]" gridcol="1" gridrow="1" pageId="12" pageNumber="65">2 2 1</td>
<td id="26745DB52A5B007CCE46FD0DC2D4B78D" box="[875,912,698,719]" gridcol="2" gridrow="1" pageId="12" pageNumber="65">2</td>
<td id="26745DB52A5B007CCEF9FD0DC2BCB78D" box="[980,1016,698,719]" gridcol="3" gridrow="1" pageId="12" pageNumber="65">2</td>
<td id="26745DB52A5B007CC974FD0DC53DB78D" box="[1113,1145,698,719]" gridcol="4" gridrow="1" pageId="12" pageNumber="65">2</td>
<td id="26745DB52A5B007CC9E1FD0DC5B4B78D" box="[1228,1264,698,719]" gridcol="5" gridrow="1" pageId="12" pageNumber="65">2</td>
<td id="26745DB52A5B007CC812FD0DC4DFB78D" box="[1343,1435,698,719]" gridcol="6" gridrow="1" pageId="12" pageNumber="65">2</td>
</tr>
<tr id="65A534C92A5B007CCDBAFD55C4DFB7B5" box="[151,1435,738,759]" gridrow="2" pageId="12" pageNumber="65">
<th id="26745DB52A5B007CCDBAFD55C0F0B7B5" box="[151,436,738,759]" gridcol="0" gridrow="2" pageId="12" pageNumber="65">Anterior loreal divided yes</th>
<td id="26745DB52A5B007CCF2AFD55C25CB7B5" box="[519,792,738,759]" gridcol="1" gridrow="2" pageId="12" pageNumber="65">no yes, no no</td>
<td id="26745DB52A5B007CCE46FD55C2D4B7B5" box="[875,912,738,759]" gridcol="2" gridrow="2" pageId="12" pageNumber="65">yes</td>
<td id="26745DB52A5B007CCEF9FD55C2BCB7B5" box="[980,1016,738,759]" gridcol="3" gridrow="2" pageId="12" pageNumber="65">yes</td>
<td id="26745DB52A5B007CC974FD55C53DB7B5" box="[1113,1145,738,759]" gridcol="4" gridrow="2" pageId="12" pageNumber="65">yes</td>
<td id="26745DB52A5B007CC9E1FD55C5B4B7B5" box="[1228,1264,738,759]" gridcol="5" gridrow="2" pageId="12" pageNumber="65">yes</td>
<td id="26745DB52A5B007CC812FD55C4DFB7B5" box="[1343,1435,738,759]" gridcol="6" gridrow="2" pageId="12" pageNumber="65">no</td>
</tr>
<tr id="65A534C92A5B007CCDBAFCBDC4DFB65D" box="[151,1435,778,799]" gridrow="3" pageId="12" pageNumber="65">
<th id="26745DB52A5B007CCDBAFCBDC0F0B65D" box="[151,436,778,799]" gridcol="0" gridrow="3" pageId="12" pageNumber="65">Supraciliaries 5–7</th>
<td id="26745DB52A5B007CCF2AFCBDC25CB65D" box="[519,792,778,799]" gridcol="1" gridrow="3" pageId="12" pageNumber="65">7–8 8 4</td>
<td id="26745DB52A5B007CCE46FCBDC2D4B65D" box="[875,912,778,799]" gridcol="2" gridrow="3" pageId="12" pageNumber="65">8–9</td>
<td id="26745DB52A5B007CCEF9FCBDC2BCB65D" box="[980,1016,778,799]" gridcol="3" gridrow="3" pageId="12" pageNumber="65">6–7</td>
<td id="26745DB52A5B007CC974FCBDC53DB65D" box="[1113,1145,778,799]" gridcol="4" gridrow="3" pageId="12" pageNumber="65">8</td>
<td id="26745DB52A5B007CC9E1FCBDC5B4B65D" box="[1228,1264,778,799]" gridcol="5" gridrow="3" pageId="12" pageNumber="65">5–8</td>
<td id="26745DB52A5B007CC812FCBDC4DFB65D" box="[1343,1435,778,799]" gridcol="6" gridrow="3" pageId="12" pageNumber="65">2–5</td>
</tr>
<tr id="65A534C92A5B007CCDBAFC85C4DFB6C0" box="[151,1435,818,898]" gridrow="4" pageId="12" pageNumber="65">
<th id="26745DB52A5B007CCDBAFC85C0F0B6C0" box="[151,436,818,898]" gridcol="0" gridrow="4" pageId="12" pageNumber="65">Supraciliary row no interrupted by fourth supraocular</th>
<td id="26745DB52A5B007CCF2AFC85C25CB6C0" box="[519,792,818,898]" gridcol="1" gridrow="4" pageId="12" pageNumber="65">no yes yes</td>
<td id="26745DB52A5B007CCE46FC85C2D4B6C0" box="[875,912,818,898]" gridcol="2" gridrow="4" pageId="12" pageNumber="65">yes</td>
<td id="26745DB52A5B007CCEF9FC85C2BCB6C0" box="[980,1016,818,898]" gridcol="3" gridrow="4" pageId="12" pageNumber="65">no</td>
<td id="26745DB52A5B007CC974FC85C53DB6C0" box="[1113,1145,818,898]" gridcol="4" gridrow="4" pageId="12" pageNumber="65">no</td>
<td id="26745DB52A5B007CC9E1FC85C5B4B6C0" box="[1228,1264,818,898]" gridcol="5" gridrow="4" pageId="12" pageNumber="65">no</td>
<td id="26745DB52A5B007CC812FC85C4DFB6C0" box="[1343,1435,818,898]" gridcol="6" gridrow="4" pageId="12" pageNumber="65">no</td>
</tr>
<tr id="65A534C92A5B007CCDBAFC22C4DFB6E8" box="[151,1435,917,938]" gridrow="5" pageId="12" pageNumber="65">
<th id="26745DB52A5B007CCDBAFC22C0F0B6E8" box="[151,436,917,938]" gridcol="0" gridrow="5" pageId="12" pageNumber="65">Supralabials 6</th>
<td id="26745DB52A5B007CCF2AFC22C25CB6E8" box="[519,792,917,938]" gridcol="1" gridrow="5" pageId="12" pageNumber="65">6 6 7–8</td>
<td id="26745DB52A5B007CCE46FC22C2D4B6E8" box="[875,912,917,938]" gridcol="2" gridrow="5" pageId="12" pageNumber="65">6–7</td>
<td id="26745DB52A5B007CCEF9FC22C2BCB6E8" box="[980,1016,917,938]" gridcol="3" gridrow="5" pageId="12" pageNumber="65">6</td>
<td id="26745DB52A5B007CC974FC22C53DB6E8" box="[1113,1145,917,938]" gridcol="4" gridrow="5" pageId="12" pageNumber="65">6</td>
<td id="26745DB52A5B007CC9E1FC22C5B4B6E8" box="[1228,1264,917,938]" gridcol="5" gridrow="5" pageId="12" pageNumber="65">6</td>
<td id="26745DB52A5B007CC812FC22C4DFB6E8" box="[1343,1435,917,938]" gridcol="6" gridrow="5" pageId="12" pageNumber="65">6</td>
</tr>
<tr id="65A534C92A5B007CCDBAFC0AC4DFB690" box="[151,1435,957,978]" gridrow="6" pageId="12" pageNumber="65">
<th id="26745DB52A5B007CCDBAFC0AC0F0B690" box="[151,436,957,978]" gridcol="0" gridrow="6" pageId="12" pageNumber="65">Infralabials 5–6</th>
<td id="26745DB52A5B007CCF2AFC0AC25CB690" box="[519,792,957,978]" gridcol="1" gridrow="6" pageId="12" pageNumber="65">5 5–7 5-6</td>
<td id="26745DB52A5B007CCE46FC0AC2D4B690" box="[875,912,957,978]" gridcol="2" gridrow="6" pageId="12" pageNumber="65">5–6</td>
<td id="26745DB52A5B007CCEF9FC0AC2BCB690" box="[980,1016,957,978]" gridcol="3" gridrow="6" pageId="12" pageNumber="65">4–6</td>
<td id="26745DB52A5B007CC974FC0AC53DB690" box="[1113,1145,957,978]" gridcol="4" gridrow="6" pageId="12" pageNumber="65">6</td>
<td id="26745DB52A5B007CC9E1FC0AC5B4B690" box="[1228,1264,957,978]" gridcol="5" gridrow="6" pageId="12" pageNumber="65">5–6</td>
<td id="26745DB52A5B007CC812FC0AC4DFB690" box="[1343,1435,957,978]" gridcol="6" gridrow="6" pageId="12" pageNumber="65">4–5</td>
</tr>
</table>
</paragraph>
<paragraph id="DB2A368B2A5BFF8FCDB2FC52C40FB108" pageId="12" pageNumber="65">
<table id="A995C42B2A5B007CCDB2FC52C43FB125" box="[159,1403,997,1127]" gridcols="8" gridrows="3" pageId="12" pageNumber="65">
<tr id="65A534C92A5B007CCDB2FC52C43FB6B8" box="[159,1403,997,1018]" gridrow="0" pageId="12" pageNumber="65">
<th id="26745DB52A5B007CCDB2FC52C08FB6B8" box="[159,459,997,1018]" gridcol="0" gridrow="0" pageId="12" pageNumber="65">Midbody scale rows 28–30</th>
<th id="26745DB52A5B007CCF2AFC52C307B6B8" box="[519,579,997,1018]" gridcol="1" gridrow="0" pageId="12" pageNumber="65">30–32</th>
<th id="26745DB52A5B007CCF52FC52C3FEB6B8" box="[639,698,997,1018]" gridcol="2" gridrow="0" pageId="12" pageNumber="65">32–40</th>
<th id="26745DB52A5B007CCFD9FC52C274B6B8" box="[756,816,997,1018]" gridcol="3" gridrow="0" pageId="12" pageNumber="65">28–31</th>
<th id="26745DB52A5B007CCE46FC52C554B6B8" box="[875,1040,997,1018]" gridcol="4" gridrow="0" pageId="12" pageNumber="65">30–36 28–30</th>
<th id="26745DB52A5B007CC974FC52C535B6B8" box="[1113,1137,997,1018]" gridcol="5" gridrow="0" pageId="12" pageNumber="65">34</th>
<th id="26745DB52A5B007CC9E1FC52C44CB6B8" box="[1228,1288,997,1018]" gridcol="6" gridrow="0" pageId="12" pageNumber="65">30–32</th>
<th id="26745DB52A5B007CC812FC52C43FB6B8" box="[1343,1403,997,1018]" gridcol="7" gridrow="0" pageId="12" pageNumber="65">22–24</th>
</tr>
<tr id="65A534C92A5B007CCDB2FBBAC43FB160" box="[159,1403,1037,1058]" gridrow="1" pageId="12" pageNumber="65">
<th id="26745DB52A5B007CCDB2FBBAC08FB160" box="[159,459,1037,1058]" gridcol="0" gridrow="1" pageId="12" pageNumber="65">Paravertebrals 47–53</th>
<td id="26745DB52A5B007CCF2AFBBAC307B160" box="[519,579,1037,1058]" gridcol="1" gridrow="1" pageId="12" pageNumber="65">60–69</td>
<td id="26745DB52A5B007CCF52FBBAC3FEB160" box="[639,698,1037,1058]" gridcol="2" gridrow="1" pageId="12" pageNumber="65">64</td>
<td id="26745DB52A5B007CCFD9FBBAC274B160" box="[756,816,1037,1058]" gridcol="3" gridrow="1" pageId="12" pageNumber="65">52</td>
<td id="26745DB52A5B007CCE46FBBAC554B160" box="[875,1040,1037,1058]" gridcol="4" gridrow="1" pageId="12" pageNumber="65">– 58–63</td>
<td id="26745DB52A5B007CC974FBBAC535B160" box="[1113,1137,1037,1058]" gridcol="5" gridrow="1" pageId="12" pageNumber="65">65</td>
<td id="26745DB52A5B007CC9E1FBBAC44CB160" box="[1228,1288,1037,1058]" gridcol="6" gridrow="1" pageId="12" pageNumber="65">55–67</td>
<td id="26745DB52A5B007CC812FBBAC43FB160" box="[1343,1403,1037,1058]" gridcol="7" gridrow="1" pageId="12" pageNumber="65">43–49</td>
</tr>
<tr id="65A534C92A5B007CCDB2FB82C43FB125" box="[159,1403,1077,1127]" gridrow="2" pageId="12" pageNumber="65">
<th id="26745DB52A5B007CCDB2FB82C08FB125" box="[159,459,1077,1127]" gridcol="0" gridrow="2" pageId="12" pageNumber="65">Paravertebral scales 0 widened (1) or not (0)</th>
<td id="26745DB52A5B007CCF2AFB82C307B125" box="[519,579,1077,1127]" gridcol="1" gridrow="2" pageId="12" pageNumber="65">0</td>
<td id="26745DB52A5B007CCF52FB82C3FEB125" box="[639,698,1077,1127]" gridcol="2" gridrow="2" pageId="12" pageNumber="65">–</td>
<td id="26745DB52A5B007CCFD9FB82C274B125" box="[756,816,1077,1127]" gridcol="3" gridrow="2" pageId="12" pageNumber="65">0</td>
<td id="26745DB52A5B007CCE46FB82C554B125" box="[875,1040,1077,1127]" gridcol="4" gridrow="2" pageId="12" pageNumber="65">– 1</td>
<td id="26745DB52A5B007CC974FB82C535B125" box="[1113,1137,1077,1127]" gridcol="5" gridrow="2" pageId="12" pageNumber="65">0</td>
<td id="26745DB52A5B007CC9E1FB82C44CB125" box="[1228,1288,1077,1127]" gridcol="6" gridrow="2" pageId="12" pageNumber="65">0</td>
<td id="26745DB52A5B007CC812FB82C43FB125" box="[1343,1403,1077,1127]" gridcol="7" gridrow="2" pageId="12" pageNumber="65">0</td>
</tr>
</table>
</paragraph>
<paragraph id="DB2A368B2A5BFF8FCDB2FBCDC434B36D" pageId="12" pageNumber="65">
<table id="A995C42B2A5B007CCDBAFBCDC4DFB36D" box="[151,1435,1146,1583]" gridcols="7" gridrows="7" pageId="12" pageNumber="65">
<tr id="65A534C92A5B007CCDBAFBCDC4DFB188" box="[151,1435,1146,1226]" gridrow="0" pageId="12" pageNumber="65">
<th id="26745DB52A5B007CCDBAFBCDC09DB188" box="[151,473,1146,1226]" gridcol="0" gridrow="0" pageId="12" pageNumber="65">Dorsal body scales kl smooth (sm) or keeled (kl)</th>
<th id="26745DB52A5B007CCF2AFBCDC308B188" box="[519,588,1146,1226]" gridcol="1" gridrow="0" pageId="12" pageNumber="65">sm</th>
<th id="26745DB52A5B007CCF52FBCDC276B188" box="[639,818,1146,1226]" gridcol="2" gridrow="0" pageId="12" pageNumber="65">sm (kl in kl juveniles)</th>
<th id="26745DB52A5B007CCE46FBCDC556B188" box="[875,1042,1146,1226]" gridcol="3" gridrow="0" pageId="12" pageNumber="65">sm kl</th>
<th id="26745DB52A5B007CC974FBCDC5D2B188" box="[1113,1174,1146,1226]" gridcol="4" gridrow="0" pageId="12" pageNumber="65">kl</th>
<th id="26745DB52A5B007CC9E1FBCDC451B188" box="[1228,1301,1146,1226]" gridcol="5" gridrow="0" pageId="12" pageNumber="65">kl</th>
<th id="26745DB52A5B007CC812FBCDC4DFB188" box="[1343,1435,1146,1226]" gridcol="6" gridrow="0" pageId="12" pageNumber="65">kl</th>
</tr>
<tr id="65A534C92A5B007CCDBAFB6AC4DFB1B0" box="[151,1435,1245,1266]" gridrow="1" pageId="12" pageNumber="65">
<th id="26745DB52A5B007CCDBAFB6AC09DB1B0" box="[151,473,1245,1266]" gridcol="0" gridrow="1" pageId="12" pageNumber="65">Ventral scales 44–52</th>
<td id="26745DB52A5B007CCF2AFB6AC308B1B0" box="[519,588,1245,1266]" gridcol="1" gridrow="1" pageId="12" pageNumber="65">56–66</td>
<td id="26745DB52A5B007CCF52FB6AC276B1B0" box="[639,818,1245,1266]" gridcol="2" gridrow="1" pageId="12" pageNumber="65">53 –</td>
<td id="26745DB52A5B007CCE46FB6AC556B1B0" box="[875,1042,1245,1266]" gridcol="3" gridrow="1" pageId="12" pageNumber="65">– –</td>
<td id="26745DB52A5B007CC974FB6AC5D2B1B0" box="[1113,1174,1245,1266]" gridcol="4" gridrow="1" pageId="12" pageNumber="65">–</td>
<td id="26745DB52A5B007CC9E1FB6AC451B1B0" box="[1228,1301,1245,1266]" gridcol="5" gridrow="1" pageId="12" pageNumber="65">–</td>
<td id="26745DB52A5B007CC812FB6AC4DFB1B0" box="[1343,1435,1245,1266]" gridcol="6" gridrow="1" pageId="12" pageNumber="65">40–44</td>
</tr>
<tr id="65A534C92A5B007CCDBAFAB2C4DFB075" box="[151,1435,1285,1335]" gridrow="2" pageId="12" pageNumber="65">
<th id="26745DB52A5B007CCDBAFAB2C09DB075" box="[151,473,1285,1335]" gridcol="0" gridrow="2" pageId="12" pageNumber="65">Scale rows at position 11 of tenth subcaudal</th>
<td id="26745DB52A5B007CCF2AFAB2C308B075" box="[519,588,1285,1335]" gridcol="1" gridrow="2" pageId="12" pageNumber="65">17–18</td>
<td id="26745DB52A5B007CCF52FAB2C276B075" box="[639,818,1285,1335]" gridcol="2" gridrow="2" pageId="12" pageNumber="65">– –</td>
<td id="26745DB52A5B007CCE46FAB2C556B075" box="[875,1042,1285,1335]" gridcol="3" gridrow="2" pageId="12" pageNumber="65">– 13–15</td>
<td id="26745DB52A5B007CC974FAB2C5D2B075" box="[1113,1174,1285,1335]" gridcol="4" gridrow="2" pageId="12" pageNumber="65">15</td>
<td id="26745DB52A5B007CC9E1FAB2C451B075" box="[1228,1301,1285,1335]" gridcol="5" gridrow="2" pageId="12" pageNumber="65">13</td>
<td id="26745DB52A5B007CC812FAB2C4DFB075" box="[1343,1435,1285,1335]" gridcol="6" gridrow="2" pageId="12" pageNumber="65">9–10</td>
</tr>
<tr id="65A534C92A5B007CCDBAFAFDC4DFB0D8" box="[151,1435,1354,1434]" gridrow="3" pageId="12" pageNumber="65">
<th id="26745DB52A5B007CCDBAFAFDC09DB0D8" box="[151,473,1354,1434]" gridcol="0" gridrow="3" pageId="12" pageNumber="65">Subcaudals divided on 1 anterior part of tail (1) or on whole tail (2)</th>
<td id="26745DB52A5B007CCF2AFAFDC308B0D8" box="[519,588,1354,1434]" gridcol="1" gridrow="3" pageId="12" pageNumber="65">1</td>
<td id="26745DB52A5B007CCF52FAFDC276B0D8" box="[639,818,1354,1434]" gridcol="2" gridrow="3" pageId="12" pageNumber="65">1 2</td>
<td id="26745DB52A5B007CCE46FAFDC556B0D8" box="[875,1042,1354,1434]" gridcol="3" gridrow="3" pageId="12" pageNumber="65">1 1</td>
<td id="26745DB52A5B007CC974FAFDC5D2B0D8" box="[1113,1174,1354,1434]" gridcol="4" gridrow="3" pageId="12" pageNumber="65">1</td>
<td id="26745DB52A5B007CC9E1FAFDC451B0D8" box="[1228,1301,1354,1434]" gridcol="5" gridrow="3" pageId="12" pageNumber="65">1</td>
<td id="26745DB52A5B007CC812FAFDC4DFB0D8" box="[1343,1435,1354,1434]" gridcol="6" gridrow="3" pageId="12" pageNumber="65">1</td>
</tr>
<tr id="65A534C92A5B007CCDBAFA1AC4DFB09D" box="[151,1435,1453,1503]" gridrow="4" pageId="12" pageNumber="65">
<th id="26745DB52A5B007CCDBAFA1AC09DB09D" box="[151,473,1453,1503]" gridcol="0" gridrow="4" pageId="12" pageNumber="65">Lamellae under fourth 18–22 toe</th>
<td id="26745DB52A5B007CCF2AFA1AC308B09D" box="[519,588,1453,1503]" gridcol="1" gridrow="4" pageId="12" pageNumber="65">16–19</td>
<td id="26745DB52A5B007CCF52FA1AC276B09D" box="[639,818,1453,1503]" gridcol="2" gridrow="4" pageId="12" pageNumber="65">22 18–19</td>
<td id="26745DB52A5B007CCE46FA1AC556B09D" box="[875,1042,1453,1503]" gridcol="3" gridrow="4" pageId="12" pageNumber="65">18–22 18–22</td>
<td id="26745DB52A5B007CC974FA1AC5D2B09D" box="[1113,1174,1453,1503]" gridcol="4" gridrow="4" pageId="12" pageNumber="65">22–23</td>
<td id="26745DB52A5B007CC9E1FA1AC451B09D" box="[1228,1301,1453,1503]" gridcol="5" gridrow="4" pageId="12" pageNumber="65">20–25</td>
<td id="26745DB52A5B007CC812FA1AC4DFB09D" box="[1343,1435,1453,1503]" gridcol="6" gridrow="4" pageId="12" pageNumber="65">18–20</td>
</tr>
<tr id="65A534C92A5B007CCDBAFA45C4DFB345" box="[151,1435,1522,1543]" gridrow="5" pageId="12" pageNumber="65">
<th id="26745DB52A5B007CCDBAFA45C09DB345" box="[151,473,1522,1543]" gridcol="0" gridrow="5" pageId="12" pageNumber="65">White spots on dorsum absent</th>
<td id="26745DB52A5B007CCF2AFA45C308B345" box="[519,588,1522,1543]" gridcol="1" gridrow="5" pageId="12" pageNumber="65">present</td>
<td id="26745DB52A5B007CCF52FA45C276B345" box="[639,818,1522,1543]" gridcol="2" gridrow="5" pageId="12" pageNumber="65">present absent</td>
<td id="26745DB52A5B007CCE46FA45C556B345" box="[875,1042,1522,1543]" gridcol="3" gridrow="5" pageId="12" pageNumber="65">present absent</td>
<td id="26745DB52A5B007CC974FA45C5D2B345" box="[1113,1174,1522,1543]" gridcol="4" gridrow="5" pageId="12" pageNumber="65">absent</td>
<td id="26745DB52A5B007CC9E1FA45C451B345" box="[1228,1301,1522,1543]" gridcol="5" gridrow="5" pageId="12" pageNumber="65">absent</td>
<td id="26745DB52A5B007CC812FA45C4DFB345" box="[1343,1435,1522,1543]" gridcol="6" gridrow="5" pageId="12" pageNumber="65">absent</td>
</tr>
<tr id="65A534C92A5B007CCDBAF9ADC4DFB36D" box="[151,1435,1562,1583]" gridrow="6" pageId="12" pageNumber="65">
<th id="26745DB52A5B007CCDBAF9ADC09DB36D" box="[151,473,1562,1583]" gridcol="0" gridrow="6" pageId="12" pageNumber="65">Sample size (n) n = 5, *</th>
<td id="26745DB52A5B007CCF2AF9ADC308B36D" box="[519,588,1562,1583]" gridcol="1" gridrow="6" pageId="12" pageNumber="65">n = 10</td>
<td id="26745DB52A5B007CCF52F9ADC276B36D" box="[639,818,1562,1583]" gridcol="2" gridrow="6" pageId="12" pageNumber="65">* *</td>
<td id="26745DB52A5B007CCE46F9ADC556B36D" box="[875,1042,1562,1583]" gridcol="3" gridrow="6" pageId="12" pageNumber="65">* *</td>
<td id="26745DB52A5B007CC974F9ADC5D2B36D" box="[1113,1174,1562,1583]" gridcol="4" gridrow="6" pageId="12" pageNumber="65">*</td>
<td id="26745DB52A5B007CC9E1F9ADC451B36D" box="[1228,1301,1562,1583]" gridcol="5" gridrow="6" pageId="12" pageNumber="65">n = 2, *</td>
<td id="26745DB52A5B007CC812F9ADC4DFB36D" box="[1343,1435,1562,1583]" gridcol="6" gridrow="6" pageId="12" pageNumber="65">n = 8</td>
</tr>
</table>
</paragraph>
<paragraph id="DB2A368B2A5BFF8ECDE9F9C2C529B5BE" blockId="12.[151,1437,1653,2015]" lastBlockId="13.[151,1436,151,252]" lastPageId="13" lastPageNumber="66" pageId="12" pageNumber="65">
<taxonomicName id="1C954D082A5BFF8FCDE9F9C2C09FB3CC" box="[196,475,1653,1678]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="boehmei" status="sp. nov.">
<emphasis id="E9E1EA992A5BFF8FCDE9F9C2C09FB3CC" box="[196,475,1653,1678]" italics="true" pageId="12" pageNumber="65">Tropidophorus boehmei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2D257E22A5BFF8FCCCEF9C2C305B3CD" box="[483,577,1653,1679]" pageId="12" pageNumber="65" rank="species">
<emphasis id="E9E1EA992A5BFF8FCCCEF9C2C305B3CD" bold="true" box="[483,577,1653,1679]" pageId="12" pageNumber="65">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from remaining water skink species, that also have smooth dorsal head scales (
<tableCitation id="961703302A5BFF8FCC06F92DC0C2B3F6" box="[299,390,1690,1716]" captionStart="TABLE 4" captionStartId="12.[151,239,338,362]" captionText="TABLE 4. Comparisons of diagnostic characters in Tropidophorus with smooth dorsal head scales from the Indochinese Peninsula (* = data obtained from literature)." pageId="12" pageNumber="65">Table 4</tableCitation>
) as follows: from 
<emphasis id="E9E1EA992A5BFF8FCF73F92CC571B3F6" box="[606,1077,1691,1716]" italics="true" pageId="12" pageNumber="65">
<taxonomicName id="1C954D082A5BFF8FCF73F92CC3AFB3F6" box="[606,747,1691,1716]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="baviensis">T. baviensis</taxonomicName>
, 
<taxonomicName id="1C954D082A5BFF8FCFDAF92CC2C0B3F6" box="[759,900,1691,1716]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="hangnam">T. hangnam</taxonomicName>
, 
<taxonomicName id="1C954D082A5BFF8FCEBDF92CC571B3F6" box="[912,1077,1691,1716]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="latiscutatus">T. latiscutatus</taxonomicName>
</emphasis>
, 
<taxonomicName id="1C954D082A5BFF8FC969F92CC5F1B3F6" box="[1092,1205,1691,1716]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="matsuii">
<emphasis id="E9E1EA992A5BFF8FC969F92CC5F1B3F6" box="[1092,1205,1691,1716]" italics="true" pageId="12" pageNumber="65">T. matsuii</emphasis>
</taxonomicName>
, 
<taxonomicName id="1C954D082A5BFF8FC9EAF92CC402B3F6" box="[1223,1350,1691,1716]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="murphyi">
<emphasis id="E9E1EA992A5BFF8FC9EAF92CC402B3F6" box="[1223,1350,1691,1716]" italics="true" pageId="12" pageNumber="65">T. murphyi</emphasis>
</taxonomicName>
, and 
<taxonomicName id="1C954D082A5BFF8FC8ABF92CC1AEB39B" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="noggei">
<emphasis id="E9E1EA992A5BFF8FC8ABF92CC1AEB39B" italics="true" pageId="12" pageNumber="65">T. noggei</emphasis>
</taxonomicName>
in having smooth dorsal body scales (versus being keeled in the latter species). The new species further differs from 
<taxonomicName id="1C954D082A5BFF8FCCA3F952C35AB3BC" box="[398,542,1765,1790]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="baviensis">
<emphasis id="E9E1EA992A5BFF8FCCA3F952C35AB3BC" box="[398,542,1765,1790]" italics="true" pageId="12" pageNumber="65">T. baviensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="1C954D082A5BFF8FCF03F952C39EB3BC" box="[558,730,1765,1790]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="latiscutatus">
<emphasis id="E9E1EA992A5BFF8FCF03F952C39EB3BC" box="[558,730,1765,1790]" italics="true" pageId="12" pageNumber="65">T. latiscutatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="1C954D082A5BFF8FCFC6F952C225B3BC" box="[747,865,1765,1790]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="matsuii">
<emphasis id="E9E1EA992A5BFF8FCFC6F952C225B3BC" box="[747,865,1765,1790]" italics="true" pageId="12" pageNumber="65">T. matsuii</emphasis>
</taxonomicName>
, 
<taxonomicName id="1C954D082A5BFF8FCE59F952C2B1B3BC" box="[884,1013,1765,1790]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="murphyi">
<emphasis id="E9E1EA992A5BFF8FCE59F952C2B1B3BC" box="[884,1013,1765,1790]" italics="true" pageId="12" pageNumber="65">T. murphyi</emphasis>
</taxonomicName>
, and 
<taxonomicName id="1C954D082A5BFF8FC911F952C5EAB3BC" box="[1084,1198,1765,1790]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="noggei">
<emphasis id="E9E1EA992A5BFF8FC911F952C5EAB3BC" box="[1084,1198,1765,1790]" italics="true" pageId="12" pageNumber="65">T. noggei</emphasis>
</taxonomicName>
by having a higher number of scale rows at position of tenth subcaudal (17–18 versus 9–15). 
<taxonomicName id="1C954D082A5BFF8FC92FF8BCC45EB266" box="[1026,1306,1803,1828]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="boehmei" status="sp. nov.">
<emphasis id="E9E1EA992A5BFF8FC92FF8BCC45EB266" box="[1026,1306,1803,1828]" italics="true" pageId="12" pageNumber="65">Tropidophorus boehmei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2D257E22A5BFF8FC80FF8BDC4C4B266" box="[1314,1408,1802,1828]" pageId="12" pageNumber="65" rank="species">
<emphasis id="E9E1EA992A5BFF8FC80FF8BDC4C4B266" bold="true" box="[1314,1408,1802,1828]" pageId="12" pageNumber="65">sp. nov.</emphasis>
</taxonomicNameLabel>
is most similar to 
<taxonomicName id="1C954D082A5BFF8FCC7CF887C33AB20B" box="[337,638,1840,1865]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="berdmorei">
<emphasis id="E9E1EA992A5BFF8FCC7CF887C33AB20B" box="[337,638,1840,1865]" italics="true" pageId="12" pageNumber="65">Tropidophorus berdmorei</emphasis>
</taxonomicName>
and 
<taxonomicName id="1C954D082A5BFF8FCF94F887C25AB20B" box="[697,798,1840,1865]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="laotus">
<emphasis id="E9E1EA992A5BFF8FCF94F887C25AB20B" box="[697,798,1840,1865]" italics="true" pageId="12" pageNumber="65">T. laotus</emphasis>
</taxonomicName>
due to the presence of smooth body scales. However, 
<taxonomicName id="1C954D082A5BFF8FCDBAF8E2C0F2B22C" box="[151,438,1877,1902]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="boehmei" status="sp. nov.">
<emphasis id="E9E1EA992A5BFF8FCDBAF8E2C0F2B22C" box="[151,438,1877,1902]" italics="true" pageId="12" pageNumber="65">Tropidophorus boehmei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2D257E22A5BFF8FCC93F8E2C35BB22D" box="[446,543,1877,1903]" pageId="12" pageNumber="65" rank="species">
<emphasis id="E9E1EA992A5BFF8FCC93F8E2C35BB22D" bold="true" box="[446,543,1877,1903]" pageId="12" pageNumber="65">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from both latter species by having the supraciliary row completed along the entire length of the lateral edge of the supraoculars. The new species is further distinguished from 
<taxonomicName id="1C954D082A5BFF8FC8ABF8CCC04AB2FB" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="berdmorei">
<emphasis id="E9E1EA992A5BFF8FC8ABF8CCC04AB2FB" italics="true" pageId="12" pageNumber="65">T. berdmorei</emphasis>
</taxonomicName>
by having lower counts of midbody scale rows (30–32 versus 
<quantity id="1C6D9B6E2A5BFF8FCEC6F817C516B2F8" box="[1003,1106,1952,1978]" metricMagnitude="-1" metricUnit="m" metricValue="9.144" metricValueMax="10.16" metricValueMin="8.128" pageId="12" pageNumber="65" unit="in" value="36.0" valueMax="40.0" valueMin="32.0">32–40 in</quantity>
<taxonomicName id="1C954D082A5BFF8FC974F817C5A8B2FB" box="[1113,1260,1952,1977]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="berdmorei">
<emphasis id="E9E1EA992A5BFF8FC974F817C5A8B2FB" box="[1113,1260,1952,1977]" italics="true" pageId="12" pageNumber="65">T. berdmorei</emphasis>
</taxonomicName>
), lower counts of lamellae under the fourth toe (16–19 versus 
<quantity id="1C6D9B6E2A5BFF8FCF97F872C3B2B29D" box="[698,758,1989,2015]" metricMagnitude="-1" metricUnit="m" metricValue="5.588" pageId="12" pageNumber="65" unit="in" value="22.0">22 in</quantity>
<taxonomicName id="1C954D082A5BFF8FCFD0F872C2CBB29C" box="[765,911,1989,2014]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="65" phylum="Chordata" rank="species" species="berdmorei">
<emphasis id="E9E1EA992A5BFF8FCFD0F872C2CBB29C" box="[765,911,1989,2014]" italics="true" pageId="12" pageNumber="65">T. berdmorei</emphasis>
</taxonomicName>
), and a higher number of ventral scales (56– 66 versus 
<quantity id="1C6D9B6E2A5AFF8ECC3DFF2FC008B5F0" box="[272,332,152,178]" metricMagnitude="0" metricUnit="m" metricValue="1.3462" pageId="13" pageNumber="66" unit="in" value="53.0">53 in</quantity>
<taxonomicName id="1C954D082A5AFF8ECC79FF2FC0ADB5F3" box="[340,489,152,177]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="66" phylum="Chordata" rank="species" species="berdmorei">
<emphasis id="E9E1EA992A5AFF8ECC79FF2FC0ADB5F3" box="[340,489,152,177]" italics="true" pageId="13" pageNumber="66">T. berdmorei</emphasis>
</taxonomicName>
). 
<taxonomicName id="1C954D082A5AFF8ECF29FF2FC25AB5F3" box="[516,798,152,177]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="66" phylum="Chordata" rank="species" species="boehmei" status="sp. nov.">
<emphasis id="E9E1EA992A5AFF8ECF29FF2FC25AB5F3" box="[516,798,152,177]" italics="true" pageId="13" pageNumber="66">Tropidophorus boehmei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2D257E22A5AFF8ECE0BFF20C2C0B5F3" box="[806,900,151,177]" pageId="13" pageNumber="66" rank="species">
<emphasis id="E9E1EA992A5AFF8ECE0BFF20C2C0B5F3" bold="true" box="[806,900,151,177]" pageId="13" pageNumber="66">sp. nov.</emphasis>
</taxonomicNameLabel>
furthermore differs from 
<taxonomicName id="1C954D082A5AFF8EC995FF2FC45AB5F3" box="[1208,1310,152,177]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="66" phylum="Chordata" rank="species" species="laotus">
<emphasis id="E9E1EA992A5AFF8EC995FF2FC45AB5F3" box="[1208,1310,152,177]" italics="true" pageId="13" pageNumber="66">T. laotus</emphasis>
</taxonomicName>
by having the tympanum diameter (
<quantity id="1C6D9B6E2A5AFF8ECC91FF0AC30FB595" box="[444,587,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.9" metricValueMin="1.7" pageId="13" pageNumber="66" unit="mm" value="2.3" valueMax="2.9" valueMin="1.7">1.7–2.9 mm</quantity>
) smaller than the eye length (
<quantity id="1C6D9B6E2A5AFF8ECE89FF0AC570B595" box="[932,1076,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" metricValueMax="5.2" metricValueMin="3.6" pageId="13" pageNumber="66" unit="mm" value="4.4" valueMax="5.2" valueMin="3.6">3.6–5.2 mm</quantity>
) (versus tympanum as large as eye in 
<taxonomicName id="1C954D082A5AFF8ECDC8FF55C003B5B9" box="[229,327,226,251]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="66" phylum="Chordata" rank="species" species="laotus">
<emphasis id="E9E1EA992A5AFF8ECDC8FF55C003B5B9" box="[229,327,226,251]" italics="true" pageId="13" pageNumber="66">T. laotus</emphasis>
</taxonomicName>
) and frontonasal undivided (versus frontonasal divided in 
<taxonomicName id="1C954D082A5AFF8ECED5FF55C51EB5B9" box="[1016,1114,226,251]" class="Reptilia" family="Scincidae" genus="Tropidophorus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="66" phylum="Chordata" rank="species" species="laotus">
<emphasis id="E9E1EA992A5AFF8ECED5FF55C51EB5B9" box="[1016,1114,226,251]" italics="true" pageId="13" pageNumber="66">T. laotus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>