273 lines
33 KiB
XML
273 lines
33 KiB
XML
<document ID-CLB-Dataset="24300" ID-DOI="http://dx.doi.org/10.3897/zookeys.905.39592" ID-GBIF-Dataset="ee5a0817-7cfc-4c21-ba7d-3beabb06cccd" ID-PMC="PMC7004242" ID-Pensoft-Pub="1313-2970-905-1" ID-Pensoft-UUID="4C7E0CB428DF56BDB15B7AC8D6707961" ID-PubMed="32063726" ID-ZooBank="F4C766E4633A41039FFDE952718F41FB" ModsDocID="1313-2970-905-1" checkinTime="1579539902851" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="48EAF341D85A208D580E62E905FCC1DE" docLanguage="en" docName="ZooKeys 905: 1-235" docOrigin="ZooKeys 905" docSource="http://dx.doi.org/10.3897/zookeys.905.39592" docTitle="Pheidole vatovavensis Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="93F87A9F-0EC1-4FF5-80BA-A0FFD0E38D60" docUuidSource="ZooBank" docVersion="7" id="4C7E0CB428DF56BDB15B7AC8D6707961" lastPageNumber="1" masterDocId="4C7E0CB428DF56BDB15B7AC8D6707961" masterDocTitle="Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups" masterLastPageNumber="235" masterPageNumber="1" pageNumber="1" updateTime="1732649601836" updateUser="ExternalLinkService">
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<mods:titleInfo id="C91FFE9729A541A993C2C54A6BB2383C">
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<mods:title id="7F4F0566C7BE60C87CFF18B9510D2E00">Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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<mods:name id="66251E591DDA89F97ACA35FAFEEE5DC7" type="personal">
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<mods:namePart id="488E9E37D2041B462AE6E68801FD5225">Salata, Sebastian</mods:namePart>
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<mods:name id="A895DF3F7D84CAFF6F87276DF8D84F88" type="personal">
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<mods:roleTerm id="EF4CECD3A29F2F89705A25571E97AF61">Author</mods:roleTerm>
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<mods:namePart id="CA48087F22388522FA62BDC92F5BAE54">Fisher, Brian L.</mods:namePart>
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<mods:typeOfResource id="4FFCEEA8416DF5167AA5478FE1A1F07F">text</mods:typeOfResource>
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<mods:title id="7B9A041D960B66AD7C9641F90FE7253E">ZooKeys</mods:title>
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<mods:part id="E545C8E0862DB765DB56B8D0E7AA2F0F">
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<mods:date id="E12F09D231BBE3B89CB6E1D15BE0DE10">2020</mods:date>
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<mods:detail id="FDF0E42CB7148B2DC6FF475C9FEAD52B" type="volume">
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<mods:number id="652EF3A5DD74D93ABCDED8D6B68D7F44">905</mods:number>
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<mods:start id="5A7193A5DCDD83EBDA53210F363891C7">1</mods:start>
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<mods:end id="08395AD9E09546E079049E3C944C454F">235</mods:end>
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<mods:url id="C7F3802291AAEB711586297B91173543">http://dx.doi.org/10.3897/zookeys.905.39592</mods:url>
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<mods:classification id="AAD072BE260D432206CED75B1C8F33F3">journal article</mods:classification>
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<mods:identifier id="AF420067301D412E0D9DB66047865FF2" type="DOI">http://dx.doi.org/10.3897/zookeys.905.39592</mods:identifier>
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<mods:identifier id="537E1700837A9A2DA1CD306207D785BA" type="Pensoft-Pub">1313-2970-905-1</mods:identifier>
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<treatment id="48EAF341D85A208D580E62E905FCC1DE" ID-GBIF-Taxon="161777010" LSID="urn:lsid:zoobank.org:act:93F87A9F-0EC1-4FF5-80BA-A0FFD0E38D60" httpUri="http://treatment.plazi.org/id/48EAF341D85A208D580E62E905FCC1DE" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
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<subSubSection id="142922866DC5DF619A6AB35D7A7A67E5" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="D14CBFCAF058BE8EC779C154FBD5D532" pageId="0" pageNumber="1">
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<taxonomicName id="6E5D60C0EBA703B9225C7A160FD316F7" ID-CoL="CMM3C" LSID="http://zoobank.org/93F87A9F-0EC1-4FF5-80BA-A0FFD0E38D60" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole vatovavensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vatovavensis" status="sp. nov.">Pheidole vatovavensis</taxonomicName>
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<taxonomicNameLabel id="A4D9B289F3075E51D5393749A7FBEDEE" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation id="A13744BCC39D6A0F66FD9DF4375B55B2" captionStart="Figure 72" captionStartId="F72" captionText="Figure 72. Pheidole vatovavensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923179) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0060282)." figureDoi="10.3897/zookeys.905.39592.figure72" httpUri="https://binary.pensoft.net/fig/373119" pageId="0" pageNumber="1">Figs 72A-F</figureCitation>
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<figureCitation id="91E6E9F3B524DB0EFF5702450E62EAA3" captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">, 85Y</figureCitation>
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<figureCitation id="85B9AAC0C0602ABCEE66A85FC45EBA11" captionStart="Figure 88" captionStartId="F88" captionText="Figure 88. Distribution. Pheidole petax Forel (A). P. podargea sp. nov. (B). P. praegrandis sp. nov. (C). P. ranohirensis sp. nov. (D). P. rugocephala sp. nov. (E). P. rugofitarata sp. nov. (F). P. scabrata Forel (G). P. typhlos sp. nov. (H). P. vatovavensis sp. nov. (I). P. voasara sp. nov. (J). P. vohemarensis sp. nov. (K). P. zavamanira sp. nov. (L)." figureDoi="10.3897/zookeys.905.39592.figure88" httpUri="https://binary.pensoft.net/fig/373135" pageId="0" pageNumber="1">, 88I</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="8295F19EEA07DBECEDDD8337827126FE" pageId="0" pageNumber="1" type="type material">
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<paragraph id="1ECF8B967A771F592C00DD93233B30C0" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="519962A961318DFB5593FD9D43A165A4" pageId="0" pageNumber="1">
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<emphasis id="07509379BA5051D1E6A7D1203ADDA797" bold="true" pageId="0" pageNumber="1">
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<emphasis id="0EB782C44E47EAB65683A7A8B678D251" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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.
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</emphasis>
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Madagascar. •1 major worker; Fianarantsoa; 7.6 km 122° Kianjavato,
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<normalizedToken id="B91E6F42F951F3BD3D50DAA2C8EEE079" originalValue="Forêt">Foret</normalizedToken>
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<normalizedToken id="45C41A198A97512F52060B3FEA061DB2" originalValue="Classée">Classee</normalizedToken>
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Vatovavy; -21.4, 47.94; alt. 175 m; 6 Jun 2005; B.L. Fisher et al. leg.; BLF12315, CASENT0060282 (CASC).
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<emphasis id="7AAEEB0384612756BC65AE95EF72FEB5" bold="true" pageId="0" pageNumber="1">
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<emphasis id="F98A02B81FDB358885C02A7863D048BC" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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.
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</emphasis>
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Madagascar. •2w.; same data as for holotype; CASENT0060283, CASENT0923179 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection id="A074BC9C5A38E5406C84342F110F1DAF" pageId="0" pageNumber="1" type="materials_examined">
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<paragraph id="9CA693D9257E0A797346C6F58D6D1874" pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph id="64A6C89C3838907F6A3BFA8C8E0AB9D4" pageId="0" pageNumber="1">
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Madagascar. -
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<emphasis id="2722583CDD2DCCE76326A3E2390BBF7F" bold="true" italics="true" pageId="0" pageNumber="1">Fianarantsoa</emphasis>
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: •7w., 10s., 1q.; 7.6 km 122° Kianjavato,
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<normalizedToken id="ACB1912C780DBEF4EFB672DEAA28E2D4" originalValue="Forêt">Foret</normalizedToken>
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<normalizedToken id="968356CC659A43307EEDBB6965C4A5AA" originalValue="Classée">Classee</normalizedToken>
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Vatovavy; -21.4, 47.94; alt. 175 m; 6 Jun 2005; B.L. Fisher et al. leg.; CASENT0060466, CASENT0060467CASENT0060470-CASENT0060472, CASENT0061080, CASENT0061081, CASENT0061146, CASENT0061179, CASENT0061183, CASENT0061257, CASENT0061259, CASENT0061351, CASENT0061352, CASENT0061435 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection id="BBDA3E33489175EEB248251B885A191E" pageId="0" pageNumber="1" type="diagnosis">
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<paragraph id="6A04654E9DBB6774E6B8D320BC99A150" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="8F680BD8C16798FCF0B8A34E5E14557F" pageId="0" pageNumber="1">
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<emphasis id="3EE27F88E941B08BD2777BEF84885626" bold="true" pageId="0" pageNumber="1">
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<emphasis id="6D4AA01720C2CAA79B71812491571DE7" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
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.
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</emphasis>
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Head in full-face view sub-rectangular, with lateral sides relatively straight, only posterior-most part slightly convex; sides of the head with moderately dense, long, erect pilosity; occipital lobes shiny, with thick, irregular rugae, interspaces with indistinct, irregular rugulae fading posteriorly; frons with moderately dense, thick, and longitudinal rugae, interspaces with dense but fine and irregular rugulae; inner hypostomal teeth distinct, small, and low, closely spaced, triangular, with rounded apex and wide base; outer hypostomal teeth bigger and wider than inner hypostomal teeth, triangular, with tops directed inward, triangular; inner and outer teeth closely spaced and connected by concavity; gaster smooth; body reddish brown.
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<emphasis id="7F62E4667BA665EB0BEDFC81FC3E4164" bold="true" pageId="0" pageNumber="1">
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<emphasis id="63D6E4347AD9905858B4812A5DFA7022" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
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.
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</emphasis>
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Head foveolate; genae with fading sculpture; promesonotum low, convex, moderately long, with posterior declivity smoothly declining towards propodeum; propodeal spines relatively long, triangular; mesosoma foveolate, katepisternum smooth.
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</paragraph>
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</subSubSection>
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<subSubSection id="E2A22A2B4A77D55BBB6A2506E7B8EB1D" pageId="0" pageNumber="1" type="description">
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<paragraph id="E372E58EF1BE90BBD8466ED7CC38DED9" pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph id="4CC50DD8081AD7F6A60A0834FDF93013" pageId="0" pageNumber="1">
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<emphasis id="C5F0AE0404AB20C0718259B60BE35368" bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Measurements (
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<emphasis id="16495867C7E72898C9E8F6EAD433B992" italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 8): HL: 1.22-1.37 (1.3); HW: 1.19-1.31 (1.24); SL: 0.53-0.59 (0.56); EL: 0.13-0.16 (0.15); WL: 0.92-1.01 (0.97); PSL: 0.19-0.23 (0.21); MTL: 0.52-0.56 (0.54); PNW: 0.58-0.65 (0.61); PTW: 0.12-0.15 (0.13); PPW: 0.45-0.51 (0.47); CI: 94.2-99.4 (95.4); SI: 42.3-46.7 (44.9); PSLI: 14.7-16.8 (15.9); PPI: 26.1-33.3 (28.0); PNI: 46.6-51.7 (49.2); MTI: 41.4-45.2 (43.3).
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<emphasis id="141C3744913766BE9FEC26BE3BB73309" bold="true" pageId="0" pageNumber="1">
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<emphasis id="1622C97C2CB47EECA54FE48615886F2F" italics="true" pageId="0" pageNumber="1">Head</emphasis>
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.
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</emphasis>
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In full-face view longer than wide, anterior of eyes straight, posterior of eyes convex (Fig.
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<figureCitation id="A2A41E1AC76BC0A607416CDFEDD46456" captionStart="Figure 72" captionStartId="F72" captionText="Figure 72. Pheidole vatovavensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923179) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0060282)." figureDoi="10.3897/zookeys.905.39592.figure72" httpUri="https://binary.pensoft.net/fig/373119" pageId="0" pageNumber="1">72B</figureCitation>
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). In lateral view sub-rectangular; ventral and dorsal faces finely convex; dorsal face finely depressed posteriorly, forming shallow transverse depression between frons and occipital lobes; inner hypostomal teeth invisible. Sides of the head with moderately dense, long, erect pilosity; whole head with dense, long, suberect to erect pilosity. Antennal scrobes weakly impressed and indistinctly delimited ventrally and posteriorly by carinulae; scrobe surface shiny, foveolate with thick, longitudinal, and long rugae. Occipital lobes shiny, with thick, irregular rugae, interspaces with indistinct, irregular rugulae fading posteriorly; frons with moderately dense, thick, and longitudinal rugae, interspaces with dense but fine and irregular rugulae; genae shiny, with dense and thin, irregular rugulae, central part with reduced sculpture; malar area with thin, longitudinal, dense rugoreticulation. Centre of clypeus shiny and smooth, lateral sides with longitudinal rugulae; median notch present, moderately wide and shallow; median longitudinal carina present; lateral longitudinal carinae present. Scape, when laid back, slightly surpassing the midlength of head; pilosity suberect (Fig.
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<figureCitation id="A7964C8782BABC03F1F4C8EA7876FF47" captionStart="Figure 72" captionStartId="F72" captionText="Figure 72. Pheidole vatovavensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923179) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0060282)." figureDoi="10.3897/zookeys.905.39592.figure72" httpUri="https://binary.pensoft.net/fig/373119" pageId="0" pageNumber="1">72B, D</figureCitation>
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). Inner hypostomal teeth distinct, small, and low, closely spaced, triangular, with rounded apex and wide base; outer hypostomal teeth bigger and wider than inner hypostomal teeth, triangular, with tops directed inward, triangular; inner and outer teeth closely spaced and connected by concavity (Fig.
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<figureCitation id="DF30FAAD91BE5A42E817D1C7FA57F215" captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">85Y</figureCitation>
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).
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<emphasis id="24E6FD5C65E2B587DD2ED53922E6DB61" bold="true" pageId="0" pageNumber="1">
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<emphasis id="66CA82AC31F620982DF45BD7711D4866" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
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.
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</emphasis>
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In lateral view, promesonotum relatively low and arched, posterior mesonotum steep, without tubercle-like projections; promesonotal groove absent; metanotal groove absent; propodeal spines moderately long, triangular, with rounded apex and wide base; humeral area laterally well produced (Fig.
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<figureCitation id="8FACC13CD5796C7E063A90486DDD4560" captionStart="Figure 72" captionStartId="F72" captionText="Figure 72. Pheidole vatovavensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923179) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0060282)." figureDoi="10.3897/zookeys.905.39592.figure72" httpUri="https://binary.pensoft.net/fig/373119" pageId="0" pageNumber="1">72D</figureCitation>
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). Surface shiny, with fine and sparse foveolae and additional rugoreticulation; dorsal surface of promesonotum with fading foveolae and a few thick, transverse rugae; katepisternum smooth. Pilosity moderately sparse, long, and erect (Fig.
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<figureCitation id="3E87DA7813E97F507FF66E4EBBA44939" captionStart="Figure 72" captionStartId="F72" captionText="Figure 72. Pheidole vatovavensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923179) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0060282)." figureDoi="10.3897/zookeys.905.39592.figure72" httpUri="https://binary.pensoft.net/fig/373119" pageId="0" pageNumber="1">72D, F</figureCitation>
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).
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<emphasis id="600E7958D644BCA0CB7A2FC239D24A82" bold="true" pageId="0" pageNumber="1">
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<emphasis id="C771BA391721953BFCF464A74015B520" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
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.
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</emphasis>
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Shiny and with fine foveolae; peduncle short, with indistinct horizontal lobes on its basal part; node relatively high, triangular, with rounded apex, in rear view node slightly convex; pilosity moderately sparse and erect (Fig.
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<figureCitation id="B3A513C203F6D146AFC0C30244A8C940" captionStart="Figure 72" captionStartId="F72" captionText="Figure 72. Pheidole vatovavensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923179) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0060282)." figureDoi="10.3897/zookeys.905.39592.figure72" httpUri="https://binary.pensoft.net/fig/373119" pageId="0" pageNumber="1">72D, F</figureCitation>
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).
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<emphasis id="C6BC4BF1D070CCE48C2902291672A56F" bold="true" pageId="0" pageNumber="1">
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<emphasis id="5535FFD5B310ECE7E099049C324DDEF3" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
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.
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</emphasis>
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Shiny, finely shagreened; in dorsal view sides with moderately long, triangular projections; pilosity long, moderately sparse, and erect (Fig.
|
|
<figureCitation id="1B36E77884540640EB622AB1A0E214B5" captionStart="Figure 72" captionStartId="F72" captionText="Figure 72. Pheidole vatovavensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923179) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0060282)." figureDoi="10.3897/zookeys.905.39592.figure72" httpUri="https://binary.pensoft.net/fig/373119" pageId="0" pageNumber="1">72D, F</figureCitation>
|
|
).
|
|
<emphasis id="5622CB0F8423502E269AE7A038FA5A5E" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="F9021173DB54BE65E3B099AC1826B285" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
Shiny and smooth; pilosity moderately sparse, long, and erect (Fig.
|
|
<figureCitation id="32A9EB6589734DA0C8B8358DA21C0FFA" captionStart="Figure 72" captionStartId="F72" captionText="Figure 72. Pheidole vatovavensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923179) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0060282)." figureDoi="10.3897/zookeys.905.39592.figure72" httpUri="https://binary.pensoft.net/fig/373119" pageId="0" pageNumber="1">72D, F</figureCitation>
|
|
).
|
|
<emphasis id="C494623E912FE2624D9B8C425D357F97" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="B9E1FBFD6000D6D92B9AD1F990DC2D24" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
|
.
|
|
</emphasis>
|
|
Head reddish brown; mesosoma and legs yellowish brown; gaster brown (Fig.
|
|
<figureCitation id="E5F039CA9FFD3A20CDDE93BBA0F1EF38" captionStart="Figure 72" captionStartId="F72" captionText="Figure 72. Pheidole vatovavensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923179) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0060282)." figureDoi="10.3897/zookeys.905.39592.figure72" httpUri="https://binary.pensoft.net/fig/373119" pageId="0" pageNumber="1">72D, F</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption id="2D34656BA82B985E73A50514B30F8B14" doi="10.3897/zookeys.905.39592.figure72" httpUri="https://binary.pensoft.net/fig/373119" pageId="0" pageNumber="1" start="Figure 72" startId="F72">
|
|
<paragraph id="327BF171DCF3105A35FA78F1BF4EE719" pageId="0" pageNumber="1">
|
|
<emphasis id="754B0A6F3073B8F1C3020D02234222C6" bold="true" pageId="0" pageNumber="1">Figure 72.</emphasis>
|
|
<taxonomicName id="F69081E39ED271ABABAFCA06ABBBF26B" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole vatovavensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vatovavensis">
|
|
<emphasis id="A1581CA2965F3CC1A14CBACC0916564E" italics="true" pageId="0" pageNumber="1">Pheidole vatovavensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., full-face view (
|
|
<emphasis id="0716D3F03119C9D0C3CAAAF98F2A9CB0" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
|
), profile (
|
|
<emphasis id="55B843D1FE6E7EE9CA80C632A9856BFD" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
|
), and dorsal view (
|
|
<emphasis id="E1C39E233F7AC5C9535868DE4C81F401" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
|
) of paratype minor worker (CASENT0923179) and full-face view (
|
|
<emphasis id="BB8FECABED9814A3B279EA3670F83D78" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
|
), profile (
|
|
<emphasis id="50E3B25B849AA1268B5305020F588485" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
|
), and dorsal view (
|
|
<emphasis id="19B82F2512A440E69DA37753B89B1624" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
|
) of holotype major worker (CASENT0060282).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="E8D82BD85EB4A18A77718B5B467ED5C5" pageId="0" pageNumber="1">
|
|
<emphasis id="B3BCF3F5069D09CC828263EBABC87190" bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
|
Measurements (
|
|
<emphasis id="FD2736A82742396D7FF9EE5F83A2C54A" italics="true" pageId="0" pageNumber="1">N</emphasis>
|
|
= 2): HL: 0.56-0.52; HW: 0.48-0.47; SL: 0.5-0.48; EL: 0.1-0.1; WL: 0.64-0.6; PSL: 0.11-0.1; MTL: 0.38-0.37; PNW: 0.33-0.33; PTW: 0.06-0.06; PPW: 0.14-0.16; CI: 86.4-90.5; SI: 103.7-100.2; PSLI: 19.8-19.3; PPI: 42.2-47.2; PNI: 67.1-69.6; MTI: 77.7-77.2.
|
|
<emphasis id="2E64C349DC32CBA469120B031D3F4CE0" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="13227919A61DB4C0D81ED43C6752D0FE" italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
|
.
|
|
</emphasis>
|
|
Occipital margin straight or indistinctly concave; occipital carina absent (Fig.
|
|
<figureCitation id="5C5E4D1691A410F6914FEBF94D20A942" captionStart="Figure 72" captionStartId="F72" captionText="Figure 72. Pheidole vatovavensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923179) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0060282)." figureDoi="10.3897/zookeys.905.39592.figure72" httpUri="https://binary.pensoft.net/fig/373119" pageId="0" pageNumber="1">72A</figureCitation>
|
|
). Pilosity moderately sparse, long, suberect to erect. Head foveolate; genae with fading sculpture. Clypeus foveolate; median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by two-fifths of its length; pilosity suberect (Fig.
|
|
<figureCitation id="86CCFC20CA817F957EE05E42CD2108ED" captionStart="Figure 72" captionStartId="F72" captionText="Figure 72. Pheidole vatovavensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923179) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0060282)." figureDoi="10.3897/zookeys.905.39592.figure72" httpUri="https://binary.pensoft.net/fig/373119" pageId="0" pageNumber="1">72A, C</figureCitation>
|
|
).
|
|
<emphasis id="C1ACDAD1211CB31978B4F6E44F7FCC2D" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="7B0A37AEEAF0B3DF7B73DF8588ED254E" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
|
.
|
|
</emphasis>
|
|
In lateral view, promesonotum low, convex, moderately long, with posterior declivity smoothly declining toward propodeum; promesonotal groove absent; metanotal groove distinct; propodeal spines relatively long, triangular, with acute apex (Fig.
|
|
<figureCitation id="06CCEFA79065906187C4FAB2849B5A7D" captionStart="Figure 72" captionStartId="F72" captionText="Figure 72. Pheidole vatovavensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923179) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0060282)." figureDoi="10.3897/zookeys.905.39592.figure72" httpUri="https://binary.pensoft.net/fig/373119" pageId="0" pageNumber="1">72C</figureCitation>
|
|
). Sculpture foveolate; katepisternum smooth. Pilosity moderately sparse, short, and erect (Fig.
|
|
<figureCitation id="6E0E28010D00DB4E3E29B7A8464B7DE8" captionStart="Figure 72" captionStartId="F72" captionText="Figure 72. Pheidole vatovavensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923179) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0060282)." figureDoi="10.3897/zookeys.905.39592.figure72" httpUri="https://binary.pensoft.net/fig/373119" pageId="0" pageNumber="1">72C, E</figureCitation>
|
|
).
|
|
<emphasis id="D4B09C23A10864DF709609ECD3C1AB60" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="0B1390E2F60912364EDCC6EB5B1ADFCC" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
Peduncle moderately short and thin with ventral face slightly convex; with few long, erect setae (Fig.
|
|
<figureCitation id="D2724A4701CD92B38763EB4099C8F016" captionStart="Figure 72" captionStartId="F72" captionText="Figure 72. Pheidole vatovavensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923179) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0060282)." figureDoi="10.3897/zookeys.905.39592.figure72" httpUri="https://binary.pensoft.net/fig/373119" pageId="0" pageNumber="1">72C, E</figureCitation>
|
|
).
|
|
<emphasis id="124FD3E005B9F888B1E5305F9E337E5B" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="BDD1072122A3E6F59E1BEAAD7C3E0D85" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
Short, low, and convex; with few short, erect setae (Fig.
|
|
<figureCitation id="7031FE711B8D1DEEAC8E5F5893437CA5" captionStart="Figure 72" captionStartId="F72" captionText="Figure 72. Pheidole vatovavensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923179) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0060282)." figureDoi="10.3897/zookeys.905.39592.figure72" httpUri="https://binary.pensoft.net/fig/373119" pageId="0" pageNumber="1">72C, E</figureCitation>
|
|
).
|
|
<emphasis id="D76B8EC80BE4A8302BFCA070922C44A3" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="DCE5F5170825D5E6D20B850F7CC8CC21" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
With sparse, erect pilosity (Fig.
|
|
<figureCitation id="1BC59FD7DA067B73CA2E829F938706B2" captionStart="Figure 72" captionStartId="F72" captionText="Figure 72. Pheidole vatovavensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923179) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0060282)." figureDoi="10.3897/zookeys.905.39592.figure72" httpUri="https://binary.pensoft.net/fig/373119" pageId="0" pageNumber="1">72C, E</figureCitation>
|
|
).
|
|
<emphasis id="9D16E06E4B8BA10581FB037825305DAF" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="020C5B57BEA9ECF64E61567800449253" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
|
.
|
|
</emphasis>
|
|
Yellow (Fig.
|
|
<figureCitation id="3CF441312152983C41667A4A701DE1F3" captionStart="Figure 72" captionStartId="F72" captionText="Figure 72. Pheidole vatovavensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923179) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0060282)." figureDoi="10.3897/zookeys.905.39592.figure72" httpUri="https://binary.pensoft.net/fig/373119" pageId="0" pageNumber="1">72C, E</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="B9A5B620A812A94C0C72D66A70073667" pageId="0" pageNumber="1" type="etymology">
|
|
<paragraph id="5E8650341B9E874297446B4104CD9607" pageId="0" pageNumber="1">Etymology.</paragraph>
|
|
<paragraph id="1F7840AAEFFAB437081421C67B8757CB" pageId="0" pageNumber="1">From the type locality.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="F4378F50CE0EA3065588AA8EA2A7D97A" pageId="0" pageNumber="1" type="biology_ecology">
|
|
<paragraph id="E404CA55B2D72A001F77CB32F08496E5" pageId="0" pageNumber="1">Biology.</paragraph>
|
|
<paragraph id="80ABB048498E093777A9107EF0E98AAA" pageId="0" pageNumber="1">The species was collected at 175 m in elevation, in rainforest. Nests were located in rotten logs.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="9B34A524B072587177FE9AC6416BB1FF" pageId="0" pageNumber="1" type="comments">
|
|
<paragraph id="34D836996A0575E6FFBB2155DA7F666D" pageId="0" pageNumber="1">Comments.</paragraph>
|
|
<paragraph id="AEFD8EF0D6F33C59B5EC58DF61A00187" pageId="0" pageNumber="1">
|
|
<taxonomicName id="6DB78505D94AB0AAC81F36AD7F62AB0B" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole vatovavensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vatovavensis">
|
|
<emphasis id="0562794836F2DC347AFEA05AC6F9E80F" italics="true" pageId="0" pageNumber="1">Pheidole vatovavensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. is most similar to
|
|
<taxonomicName id="BBFD8F8E3341AAABF43ECD43C5DFFE0C" lsidName="P. ankerana" pageId="0" pageNumber="1" rank="species" species="ankerana">
|
|
<emphasis id="EB93C3D5DDD5E7447019F9E8DFD96CC3" italics="true" pageId="0" pageNumber="1">P. ankerana</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.
|
|
<emphasis id="077E0F7ABE52FD29684839AEF59008FE" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="A8162131B6788A35D19FCBB12456CFFE" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
|
.
|
|
</emphasis>
|
|
<taxonomicName id="691E372634F8F5D52B836BE216B4B5E4" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole vatovavensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vatovavensis">
|
|
<emphasis id="092695BCD6D4F86FC76E61F369F27ED9" italics="true" pageId="0" pageNumber="1">Pheidole vatovavensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName id="62C6335D983A93F20AA53BA536C0E529" lsidName="P. ankerana" pageId="0" pageNumber="1" rank="species" species="ankerana">
|
|
<emphasis id="F91549D3D7757A6D726E436B7AEC5C35" italics="true" pageId="0" pageNumber="1">P. ankerana</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. in sparser and shorter pilosity on the sides of head, frons with interspaces never smooth, foveolate sculpture on scrobes, higher promesonotum, and triangular shape of inner hypostomal teeth.
|
|
<emphasis id="3D4AA174685D0E6DE8EB83C96A646AE1" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="8D1574D6ECB8BA76602F43734D5B064C" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
|
.
|
|
</emphasis>
|
|
<taxonomicName id="C05D7449A94F752361B2508FA386DB13" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole vatovavensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vatovavensis">
|
|
<emphasis id="0323F96E4FFF586E4199739DEE5F99EC" italics="true" pageId="0" pageNumber="1">Pheidole vatovavensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName id="7A34D2196A8A05009BA2A6C4B4F73D9F" lsidName="P. ankerana" pageId="0" pageNumber="1" rank="species" species="ankerana">
|
|
<emphasis id="36BBD94F86BAC4752CF1A25B3480C229" italics="true" pageId="0" pageNumber="1">P. ankerana</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. in lack of additional thin, sparse rugae on the head and mesosoma and presence of relatively long propodeal spines.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |