167 lines
20 KiB
XML
167 lines
20 KiB
XML
<document id="2D5171710690D11E1544FAC18EB2B008" ID-CLB-Dataset="31536" ID-DOI="10.11646/zootaxa.4349.1.1" ID-GBIF-Dataset="3e5cd781-3051-4527-bace-d222e83d8398" ID-ISSN="1175-5326" ID-Zenodo-Dep="1049190" ID-ZooBank="DDFDC23A-FB21-41E2-B38B-A0FD19F5BFAE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510732256065" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2017" docId="03EE87F7FFA4DE61FF25D660FD23A826" docLanguage="en" docName="zootaxa.4349.1.1.pdf.imf" docOrigin="Zootaxa 4349 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Cephennomicrus acupunctatus Jałoszyński 2017, sp. n." docType="treatment" docVersion="7" lastPageNumber="9" masterDocId="FFD7FF8FFFA3DE69FFB2D111FFADAD5E" masterDocTitle="Ant-like stone beetles on the roof of the world. Cephenniini of Nepal and Bhutan (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="120" masterPageNumber="1" pageNumber="8" updateTime="1698484713136" updateUser="ExternalLinkService">
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<mods:title id="D4D4BD7725D8CCC7C9CE5E4A80C3DD19">Ant-like stone beetles on the roof of the world. Cephenniini of Nepal and Bhutan (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:name id="0E8A96498C25FE0297025EF22A39FAFF" type="personal">
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<mods:namePart id="4D6DDC30E86663D5C764D43814B8F719">Jałoszyński, Paweł</mods:namePart>
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<mods:typeOfResource id="09F5D5F3AC42165762949BBB11150755">text</mods:typeOfResource>
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<mods:title id="5560A89A8776217DA9ABA2406FE23642">Zootaxa</mods:title>
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<mods:part id="5AA75467672F46CB67B8D9B8F6438D0E">
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<mods:date id="276A06F3D7F112C10316FDD89509D2EA">2017</mods:date>
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<mods:number id="DC226C334C190128C50C13B4D9C2AD18">2017-11-15</mods:number>
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<mods:number id="58BFF00265EB6985766536A06A7C0FA6">4349</mods:number>
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<mods:identifier id="3E3EEE033E052BD2C4B2F3675308B644" type="DOI">10.11646/zootaxa.4349.1.1</mods:identifier>
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<treatment id="03EE87F7FFA4DE61FF25D660FD23A826" ID-DOI="http://doi.org/10.5281/zenodo.6026666" ID-GBIF-Taxon="137306712" ID-Zenodo-Dep="6026666" LSID="urn:lsid:plazi:treatment:03EE87F7FFA4DE61FF25D660FD23A826" httpUri="http://treatment.plazi.org/id/03EE87F7FFA4DE61FF25D660FD23A826" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
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<subSubSection id="C35D656AFFA4DE6EFF25D660FE25AAF2" pageId="7" pageNumber="8" type="nomenclature">
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<paragraph id="8BF836E1FFA4DE6EFF25D660FDF4AAD5" blockId="7.[151,601,1905,1964]" box="[151,601,1905,1931]" pageId="7" pageNumber="8">
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<heading id="D0B0818DFFA4DE6EFF25D660FDF4AAD5" bold="true" box="[151,601,1905,1931]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
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<emphasis id="B933EAF3FFA4DE6EFF25D660FDF4AAD5" bold="true" box="[151,601,1905,1931]" pageId="7" pageNumber="8">
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<taxonomicName id="4C474D62FFA4DE6EFF25D660FDA2AAD5" ID-CoL="SFMB" authority="Jałoszyński, 2017" authorityName="Jałoszyński" authorityYear="2017" box="[151,527,1905,1931]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="acupunctatus" status="sp. nov.">
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<emphasis id="B933EAF3FFA4DE6EFF25D660FDA2AAD5" bold="true" box="[151,527,1905,1931]" italics="true" pageId="7" pageNumber="8">Cephennomicrus acupunctatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2005788FFA4DE6EFDA5D660FDF4AAD5" box="[535,601,1905,1931]" pageId="7" pageNumber="8" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF836E1FFA4DE6EFF25D682FE25AAF2" blockId="7.[151,601,1905,1964]" box="[151,392,1939,1964]" pageId="7" pageNumber="8">
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(
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<figureCitation id="137C2A64FFA4DE6EFF2DD682FF4AAAF2" box="[159,231,1939,1964]" captionStart="FIGURES 2 – 4" captionStartId="7.[151,264,1776,1799]" captionTargetBox="[173,1420,815,1738]" captionTargetId="figure@7.[151,1436,807,1755]" captionTargetPageId="7" captionText="FIGURES 2 – 4. Dorsal habitus of Cephennomicrus arunensis sp. n. (2), Cm. acupunctatus sp. n. (3), and Cm. taplejungensis sp. n. (4)." httpUri="https://zenodo.org/record/1049194/files/figure.png" pageId="7" pageNumber="8">Figs 3</figureCitation>
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,
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<figureCitation id="137C2A64FFA4DE6EFF41D682FEFAAAF2" box="[243,343,1939,1964]" captionStart="FIGURES 5 – 13" captionStartId="9.[151,264,1952,1975]" captionTargetBox="[156,1428,872,1902]" captionTargetId="figure@9.[154,1439,860,1921]" captionTargetPageId="9" captionText="FIGURES 5 – 13. Body outline (5 – 7) and aedeagus in ventral (8, 10, 12) and lateral (9, 11, 13) views. Cephennomicrus arunensis sp. n. (5, 8 – 9), Cm. acupunctatus sp. n. (6, 10 – 11), and Cm. taplejungensis sp. n. (7, 12 – 13)." httpUri="https://zenodo.org/record/1049196/files/figure.png" pageId="7" pageNumber="8">6, 10–11</figureCitation>
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,
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<figureCitation id="137C2A64FFA4DE6EFED0D685FED2AAF2" box="[354,383,1940,1964]" captionStart="FIGURES 17 – 18" captionStartId="12.[151,264,1363,1386]" captionTargetBox="[151,1436,193,1336]" captionTargetId="figure@12.[151,1436,193,1342]" captionTargetPageId="12" captionText="FIGURES 17 – 18. Distribution of Cephennomicrus arunensis sp. n. (17, circle), Cm. acupunctatus sp. n. (17, circle), Cm. taplejungensis sp. n. (17, triangle) and Hlavaciellus primitivus sp. n. (18, circle)." httpUri="https://zenodo.org/record/1049200/files/figure.png" pageId="7" pageNumber="8">17</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C35D656AFFA4DE61FF25D6CDFF53AD8A" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="materials_examined">
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<paragraph id="8BF836E1FFA4DE61FF25D6CDFF53AD8A" blockId="7.[151,1436,2012,2037]" lastBlockId="8.[151,1437,151,1400]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
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<emphasis id="B933EAF3FFA4DE6EFF25D6CDFCC1AAAB" bold="true" box="[151,876,2012,2037]" pageId="7" pageNumber="8">
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<typeStatus id="54FC8843FFA4DE6EFF25D6CDFF7FAAAB" box="[151,210,2012,2037]" pageId="7" pageNumber="8">Type</typeStatus>
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material.
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<typeStatus id="54FC8843FFA4DE6EFEFDD6CDFE6DAAAB" box="[335,448,2012,2037]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
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:
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<collectingCountry id="F3507671FFA4DE6EFE7FD6CDFD87AAAB" box="[461,554,2012,2037]" name="Nepal" pageId="7" pageNumber="8">NEPAL</collectingCountry>
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(Sankhuwasabha District)
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</emphasis>
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:
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<specimenCount id="9D41FD68FFA4DE6EFCCED6CCFC3EAAAA" box="[892,915,2013,2036]" pageId="7" pageNumber="8" type="male">♂</specimenCount>
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, "
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<collectingCountry id="F3507671FFA4DE6EFC18D6CDFBA5AAAA" box="[938,1032,2012,2036]" name="Nepal" pageId="7" pageNumber="8">NEPAL</collectingCountry>
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, Sanhua Sabha Distr. / Arun Valley between Mure and Hurure /
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<quantity id="4CBF9B04FFABDE61FE6BD186FDD8ADF1" box="[473,629,151,176]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.15" metricValueMin="2.05" pageId="8" pageNumber="9" unit="m" value="2100.0" valueMax="2150.0" valueMin="2050.0">2050–2150 m</quantity>
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alt.,
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<date id="FFF91021FFABDE61FD1DD189FCFCADEE" box="[687,849,152,176]" pageId="8" pageNumber="9" value="1988-06-09" valueMax="1988-06-11" valueMin="1988-06-09">9.–11.VI.1988</date>
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/ leg. J. Martens & W. Schawaller" [white, printed] (
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<collectionCode id="ED56AE24FFABDE61FF2DD1ADFF5CAD8A" box="[159,241,188,212]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="8" pageNumber="9">SMNS</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C35D656AFFABDE61FF75D1CEFD18AC1E" pageId="8" pageNumber="9" type="diagnosis">
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<paragraph id="8BF836E1FFABDE61FF75D1CEFD18AC1E" blockId="8.[151,1437,151,1400]" pageId="8" pageNumber="9">
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<emphasis id="B933EAF3FFABDE61FF75D1CEFEECADA6" bold="true" box="[199,321,223,248]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
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BL ~
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<quantity id="4CBF9B04FFABDE61FE3DD1F1FE47ADA9" box="[399,490,224,248]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="9" unit="mm" value="0.8">0.8 mm</quantity>
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; head and pronotum covered with small but deep and dense punctures; elytra with fine but distinct punctures; aedeagus with lightly sclerotized bell-shaped internal armature, in lateral view subapical margin of dorsal wall strongly curved.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35D656AFFABDE61FF75D05DFE33A853" pageId="8" pageNumber="9" type="description">
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<paragraph id="8BF836E1FFABDE61FF75D05DFB82ACD9" blockId="8.[151,1437,151,1400]" pageId="8" pageNumber="9">
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<emphasis id="B933EAF3FFABDE61FF75D05DFEF4AC3B" bold="true" box="[199,345,332,357]" pageId="8" pageNumber="9">Description.</emphasis>
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Body of male (
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<figureCitation id="137C2A64FFABDE61FDA9D05DFDC5AC3A" box="[539,616,332,357]" captionStart="FIGURES 2 – 4" captionStartId="7.[151,264,1776,1799]" captionTargetBox="[173,1420,815,1738]" captionTargetId="figure@7.[151,1436,807,1755]" captionTargetPageId="7" captionText="FIGURES 2 – 4. Dorsal habitus of Cephennomicrus arunensis sp. n. (2), Cm. acupunctatus sp. n. (3), and Cm. taplejungensis sp. n. (4)." httpUri="https://zenodo.org/record/1049194/files/figure.png" pageId="8" pageNumber="9">Figs 3</figureCitation>
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,
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<figureCitation id="137C2A64FFABDE61FDC9D05DFD27AC3B" box="[635,650,332,357]" captionStart="FIGURES 5 – 13" captionStartId="9.[151,264,1952,1975]" captionTargetBox="[156,1428,872,1902]" captionTargetId="figure@9.[154,1439,860,1921]" captionTargetPageId="9" captionText="FIGURES 5 – 13. Body outline (5 – 7) and aedeagus in ventral (8, 10, 12) and lateral (9, 11, 13) views. Cephennomicrus arunensis sp. n. (5, 8 – 9), Cm. acupunctatus sp. n. (6, 10 – 11), and Cm. taplejungensis sp. n. (7, 12 – 13)." httpUri="https://zenodo.org/record/1049196/files/figure.png" pageId="8" pageNumber="9">6</figureCitation>
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) elongate, with distinct constriction between pronotum and elytra, strongly convex, dark brown, covered with slightly lighter vestiture; BL
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<quantity id="4CBF9B04FFABDE61FC76D061FB86ACD9" box="[964,1067,368,392]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="8" pageNumber="9" unit="mm" value="0.83">0.83 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BF836E1FFABDE61FF75D085FD78AF93" blockId="8.[151,1437,151,1400]" pageId="8" pageNumber="9">
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Head subtrapezoidal, HL
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<quantity id="4CBF9B04FFABDE61FE5ED085FDF9ACF2" box="[492,596,404,428]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="9" unit="mm" value="0.1">0.10 mm</quantity>
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, HW
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<quantity id="4CBF9B04FFABDE61FD2BD085FCAFACF2" box="[665,770,404,428]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="9" unit="mm" value="0.2">0.20 mm</quantity>
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; vertex and frons confluent, evenly convex; supraantennal tubercles barely marked; eyes large, strongly convex, coarsely faceted. Punctures on vertex and frons moderately large and deep, densely and unevenly distributed, separated by spaces 0.5–1 × as wide as diameters of punctures; setae short, sparse and nearly recumbent. Antennae short, compact, slender, with trimerous and strongly thickened club, AnL
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<quantity id="4CBF9B04FFABDE61FEA2D335FEDAAF62" box="[272,375,548,572]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="8" pageNumber="9" unit="mm" value="0.33">0.33 mm</quantity>
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; antennomere I nearly twice as long as broad; II distinctly shorter and slightly narrower than I, 1.7 × as long as broad; III much narrower and shorter than II, slightly broader than long; IV–VIII subequal in length and width, each as broad as III but slightly longer, 1.1–1.2 × as long as broad; IX distinctly broader and slightly longer than VIII, as long as broad; X much larger than IX, distinctly transverse; XI as broad as X but slightly longer, with distinct subapical constriction.
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</paragraph>
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<paragraph id="8BF836E1FFABDE61FF75D3C9FCDFAEFA" blockId="8.[151,1437,151,1400]" pageId="8" pageNumber="9">
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Pronotum in dorsal view nearly subquadrate, equally broad between anterior fourth and base, PL
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<quantity id="4CBF9B04FFABDE61FAB3D3C6FACAAFB1" box="[1281,1383,727,752]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="9" unit="mm" value="0.25">0.25 mm</quantity>
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, PW
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<quantity id="4CBF9B04FFABDE61FF25D3EDFEADAE4A" box="[151,256,764,788]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="8" pageNumber="9" unit="mm" value="0.33">0.33 mm</quantity>
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; anterior margin strongly arcuate; lateral margins indistinctly microserrate, nearly straight and parallel; hind corners nearly right-angled; posterior margin slightly arcuate; base of pronotum with two pairs of small but distinct lateral antebasal pits, obscured by very dense, moderately large and very deep punctures covering pronotal disc, distances between punctures distinctly shorter than diameters of punctures. Setae short and sparse, slightly suberect, additionally each hind corner bears one long, erect seta.
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</paragraph>
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<paragraph id="8BF836E1FFABDE61FF75D2BEFD20A906" blockId="8.[151,1437,151,1400]" pageId="8" pageNumber="9">
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Elytra oval, moderately stout, broadest near anterior third, EL
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<quantity id="4CBF9B04FFABDE61FC35D2A1FC43AE99" box="[903,1006,944,968]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="8" pageNumber="9" unit="mm" value="0.48">0.48 mm</quantity>
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, EW
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<quantity id="4CBF9B04FFABDE61FB9CD2A1FB39AE99" box="[1070,1172,944,968]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="9" unit="mm" value="0.4">0.40 mm</quantity>
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, EI 1.19. Humeral calli small but well marked, mesally delimited by oblique, deep, relatively narrow basal impression; apices of elytra separately rounded. Punctures very small but relatively deep and sharply marked, dense; setae very short, but well visible under magnification 40 ×, dense and nearly recumbent, additionally each elytron bears a single long seta located in subhumeral area of lateral margin.
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</paragraph>
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<paragraph id="8BF836E1FFABDE61FF75D575FE7EA923" blockId="8.[151,1437,151,1400]" box="[199,467,1124,1149]" pageId="8" pageNumber="9">Hind wings not studied.</paragraph>
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<paragraph id="8BF836E1FFABDE61FF75D596FE8DA9B6" blockId="8.[151,1437,151,1400]" pageId="8" pageNumber="9">
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Aedeagus (
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<figureCitation id="137C2A64FFABDE61FEF4D596FE68A9FE" box="[326,453,1159,1184]" captionStart="FIGURES 5 – 13" captionStartId="9.[151,264,1952,1975]" captionTargetBox="[156,1428,872,1902]" captionTargetId="figure@9.[154,1439,860,1921]" captionTargetPageId="9" captionText="FIGURES 5 – 13. Body outline (5 – 7) and aedeagus in ventral (8, 10, 12) and lateral (9, 11, 13) views. Cephennomicrus arunensis sp. n. (5, 8 – 9), Cm. acupunctatus sp. n. (6, 10 – 11), and Cm. taplejungensis sp. n. (7, 12 – 13)." httpUri="https://zenodo.org/record/1049196/files/figure.png" pageId="8" pageNumber="9">Figs 10–11</figureCitation>
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) thin-walled, AeL
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<quantity id="4CBF9B04FFABDE61FD12D599FCA5A9FE" box="[672,776,1160,1184]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="8" pageNumber="9" unit="mm" value="0.13">0.13 mm</quantity>
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, with subtrapezoidal apex; internal armature composed of lightly sclerotized, bell-shaped structures located in middle of median lobe; parameres very slender, each with three apical setae.
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</paragraph>
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<paragraph id="8BF836E1FFABDE61FF75D5E5FE33A853" blockId="8.[151,1437,151,1400]" box="[199,414,1268,1293]" pageId="8" pageNumber="9">Female. Unknown.</paragraph>
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</subSubSection>
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<subSubSection id="C35D656AFFABDE61FF75D406FDD8A86E" box="[199,629,1303,1328]" pageId="8" pageNumber="9" type="distribution">
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<paragraph id="8BF836E1FFABDE61FF75D406FDD8A86E" blockId="8.[151,1437,151,1400]" box="[199,629,1303,1328]" pageId="8" pageNumber="9">
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||
<emphasis id="B933EAF3FFABDE61FF75D406FEF4A86E" bold="true" box="[199,345,1303,1328]" pageId="8" pageNumber="9">Distribution</emphasis>
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||
(
|
||
<figureCitation id="137C2A64FFABDE61FEDAD406FE11A86E" box="[360,444,1303,1328]" captionStart="FIGURES 17 – 18" captionStartId="12.[151,264,1363,1386]" captionTargetBox="[151,1436,193,1336]" captionTargetId="figure@12.[151,1436,193,1342]" captionTargetPageId="12" captionText="FIGURES 17 – 18. Distribution of Cephennomicrus arunensis sp. n. (17, circle), Cm. acupunctatus sp. n. (17, circle), Cm. taplejungensis sp. n. (17, triangle) and Hlavaciellus primitivus sp. n. (18, circle)." httpUri="https://zenodo.org/record/1049200/files/figure.png" pageId="8" pageNumber="9">Fig. 17</figureCitation>
|
||
).
|
||
<collectingRegion id="4983F803FFABDE61FE62D408FD89A86E" box="[464,548,1305,1328]" country="Sri Lanka" name="Eastern" pageId="8" pageNumber="9">Eastern</collectingRegion>
|
||
<collectingCountry id="F3507671FFABDE61FD99D406FDDFA86E" box="[555,626,1303,1328]" name="Nepal" pageId="8" pageNumber="9">Nepal</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35D656AFFABDE61FF75D42DFAEBA80A" box="[199,1350,1340,1365]" pageId="8" pageNumber="9" type="etymology">
|
||
<paragraph id="8BF836E1FFABDE61FF75D42DFAEBA80A" blockId="8.[151,1437,151,1400]" box="[199,1350,1340,1365]" pageId="8" pageNumber="9">
|
||
<emphasis id="B933EAF3FFABDE61FF75D42DFEE3A80B" bold="true" box="[199,334,1340,1365]" pageId="8" pageNumber="9">Etymology.</emphasis>
|
||
The name refers to particularly sharply marked, distinct punctures covering the pronotum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35D656AFFABDE61FF75D44EFD23A826" box="[199,654,1375,1400]" pageId="8" pageNumber="9" type="discussion">
|
||
<paragraph id="8BF836E1FFABDE61FF75D44EFD23A826" blockId="8.[151,1437,151,1400]" box="[199,654,1375,1400]" pageId="8" pageNumber="9">
|
||
<emphasis id="B933EAF3FFABDE61FF75D44EFE96A826" bold="true" box="[199,315,1375,1400]" pageId="8" pageNumber="9">Remarks.</emphasis>
|
||
See remarks for
|
||
<taxonomicName id="4C474D62FFABDE61FE48D471FD25A829" authorityName="Jałoszyński" authorityYear="2017" box="[506,648,1376,1399]" class="Insecta" family="Staphylinidae" genus="Cephennomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="arunensis">
|
||
<emphasis id="B933EAF3FFABDE61FE48D471FD25A829" box="[506,648,1376,1399]" italics="true" pageId="8" pageNumber="9">C. arunensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |