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<document id="96E11F36AD682D79219D596A94F906C5" ID-CLB-Dataset="5684" ID-DOI="10.11646/zootaxa.4985.4.5" ID-GBIF-Dataset="9db1f64f-288e-430e-82fa-0e811af468a7" ID-ISSN="1175-5326" ID-Zenodo-Dep="4964249" ID-ZooBank="BBBAFB98-1377-42FF-91A0-79F28A046CA3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623832345623" checkinUser="plazi" docAuthor="Tan, Ming Kai, Rahmadi, Cahyo &amp; Robillard, Tony" docDate="2021" docId="03EE87E7FFC6FFBB88EC7CE8FE7AC714" docLanguage="en" docName="zootaxa.4985.4.5.pdf" docOrigin="Zootaxa 4985 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phaloria lobulata Tan &amp; Rahmadi &amp; Robillard 2021, sp. nov." docType="treatment" docVersion="6" lastPageNumber="528" masterDocId="FFD7FF9FFFCBFFB4887B7C7FFFC4C321" masterDocTitle="New species of Phaloria (Orthoptera: Phalangopsidae: Phaloriinae) from West Papua (Indonesia)" masterLastPageNumber="530" masterPageNumber="513" pageNumber="526" updateTime="1699017048324" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="30F2174A9FA71E2D4B851A52AC3F2951">
<mods:title id="2B854A696F0B858EA82C871183EBFD50">New species of Phaloria (Orthoptera: Phalangopsidae: Phaloriinae) from West Papua (Indonesia)</mods:title>
</mods:titleInfo>
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<mods:namePart id="5A13A110B3B37290FC3ACE0B17F7AEC1">Tan, Ming Kai</mods:namePart>
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<mods:affiliation id="F3DA63B59754D64B34463024EE9BE31D">Institut de Systématique, Evolution et Biodiversité (ISYEB), Muséum national dHistoire naturelle, CNRS, SU, EPHE, UA, 57 rue Cuvier, CP 50, 75231 Paris Cedex 05, France. &amp; orthoptera. mingkai @ gmail. com; https: // orcid. org / 0000 - 0002 - 4324 - 6305</mods:affiliation>
<mods:nameIdentifier id="1DD5286DB6388DCB665DA8E48D553C1D" type="email">orthoptera.mingkai@gmail.com</mods:nameIdentifier>
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<mods:namePart id="5B25179FB0EF505655BC233C2CDD23D4">Rahmadi, Cahyo</mods:namePart>
<mods:affiliation id="0ED90B91E6B9C0E70A453222291703FF">Museum Zoologicum Bogoriense, Research Center for Biology, Cibinong Science Center-LIPI, Jl. Raya Jakarta- Bogor Km. 46, Cibinong, 16911 Indonesia.</mods:affiliation>
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<mods:namePart id="28507029E6AA059823D711E2BE03AF75">Robillard, Tony</mods:namePart>
<mods:nameIdentifier id="8B2260CF44EE99667190B400CEE5341F" type="ORCID">0000-0002-2177-9549</mods:nameIdentifier>
<mods:affiliation id="080A9AF51ED9BF577F3ADAD134FE5F7F">Institut de Systématique, Evolution et Biodiversité (ISYEB), Muséum national dHistoire naturelle, CNRS, SU, EPHE, UA, 57 rue Cuvier, CP 50, 75231 Paris Cedex 05, France. &amp; tony. robillard @ mnhn. fr; https: // orcid. org / 0000 - 0002 - 2177 - 9549</mods:affiliation>
<mods:nameIdentifier id="2C5B6D58F40CE41114C85C0EC2F33AE0" type="email">tony.robillard@mnhn.fr</mods:nameIdentifier>
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<heading id="D0B0819DFFC6FFB988EC7CE8FD6AC390" bold="true" box="[151,686,151,178]" fontSize="11" level="1" pageId="13" pageNumber="526" reason="1">
<emphasis id="B933EAE3FFC6FFB988EC7CE8FD6AC390" bold="true" box="[151,686,151,178]" pageId="13" pageNumber="526">
<taxonomicName id="4C474D72FFC6FFB988EC7CE8FD80C390" authority="Tan &amp; Robillard" authorityName="Tan &amp; Rahmadi &amp; Robillard" authorityYear="2021" box="[151,580,151,178]" class="Insecta" family="Haglotettigoniidae" genus="Phaloria" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="526" phylum="Arthropoda" rank="species" species="lobulata" status="sp. nov.">
<emphasis id="B933EAE3FFC6FFB988EC7CE8FEAFC393" bold="true" box="[151,363,151,178]" italics="true" pageId="13" pageNumber="526">Phaloria lobulata</emphasis>
Tan &amp; Robillard
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,
<taxonomicNameLabel id="A2005798FFC6FFB98A2A7CE8FD6AC390" box="[593,686,151,177]" pageId="13" pageNumber="526" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
</paragraph>
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<subSubSection id="C35D657AFFC6FFB988EC7CC4FE2EC3F7" box="[151,490,187,214]" pageId="13" pageNumber="526" type="description">
<paragraph id="8BF836F1FFC6FFB988EC7CC4FE2EC3F7" blockId="13.[151,686,151,214]" box="[151,490,187,214]" pageId="13" pageNumber="526">
(
<figureCitation id="137C2A74FFC6FFB988E47CC4FEC5C3F4" box="[159,257,187,213]" captionStart="FIGURE 1" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[195,1408,189,1918]" captionTargetId="figure-26@8.[179,1408,181,1920]" captionTargetPageId="8" captionText="FIGURE 1. Habitus of Phaloria males (AD, FI) and females (E, J, K) from Lobo and Kumawa in dorsal (AH, J) and lateral (I, K) views: P. dischidia sp. nov. (A), P. paradischidia sp. nov. (B), P. tukul sp. nov. (C), P. brevis sp. nov. (D, E), P. berbeda sp. nov. (F), P. lobulata sp. nov. (G), P. nr. aspersa (HK). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4964251" httpUri="https://zenodo.org/record/4964251/files/figure.png" pageId="13" pageNumber="526">Figs. 1G</figureCitation>
,
<figureCitation id="137C2A74FFC6FFB989747CC4FEE9C3F4" box="[271,301,187,213]" captionStart="FIGURE 2" captionStartId="9.[151,250,1814,1838]" captionTargetBox="[160,1417,196,1778]" captionTargetId="figure-19@9.[151,1436,181,1790]" captionTargetPageId="9" captionText="FIGURE 2. Head and pronotum of P. dischidia sp. nov. (A), P. paradischidia sp. nov. (B), P. tukul sp. nov. (C), P. brevis sp. nov. (D), P. berbeda sp. nov. (E), P. lobulata sp. nov. (F), P. nr. aspersa (G) in dorsal view. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4964253" httpUri="https://zenodo.org/record/4964253/files/figure.png" pageId="13" pageNumber="526">2F</figureCitation>
,
<figureCitation id="137C2A74FFC6FFB989437CC4FE92C3F4" box="[312,342,187,213]" captionStart="FIGURE 3" captionStartId="10.[151,250,1696,1720]" captionTargetBox="[197,1378,236,1639]" captionTargetId="figure-26@10.[183,1404,181,1672]" captionTargetPageId="10" captionText="FIGURE 3. Faces of P. dischidia sp. nov. (A), P. paradischidia sp. nov. (B), P. tukul sp. nov. (C), P. brevis sp. nov. (D), P. berbeda sp. nov. (E), P. lobulata sp. nov. (F), P. nr. aspersa (G) in anterior views. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4964255" httpUri="https://zenodo.org/record/4964255/files/figure.png" pageId="13" pageNumber="526">3F</figureCitation>
,
<figureCitation id="137C2A74FFC6FFB989197CC4FE40C3F4" box="[354,388,187,213]" captionStart="FIGURE 4" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[195,1397,192,1918]" captionTargetId="figure-19@11.[183,1404,181,1920]" captionTargetPageId="11" captionText="FIGURE 4. Male FWs of P. dischidia sp. nov. (A), P. paradischidia sp. nov. (B), P. tukul sp. nov. (C), P. brevis sp. nov. (D), P. berbeda sp. nov. (F), P. lobulata sp. nov. (G), P. nr. aspersa (H) and female FWs of P. brevis sp. nov. (E) and P. nr. aspersa (I) in dorsal views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4964258" httpUri="https://zenodo.org/record/4964258/files/figure.png" pageId="13" pageNumber="526">4G</figureCitation>
,
<figureCitation id="137C2A74FFC6FFB989E97CC4FE25C3F7" box="[402,481,187,214]" captionStart="FIGURE 5" captionStartId="12.[151,250,1841,1865]" captionTargetBox="[244,1363,190,1783]" captionTargetId="figure-26@12.[215,1372,181,1818]" captionTargetPageId="12" captionText="FIGURE 5. Male genitalia of Phaloria in dorsal (A, C, E, G, I, K, M) and ventral (B, D, F, H, J, L, N) views: P. dischidia sp. nov. (A, B), P. paradischidia sp. nov. (C, D), P. tukul sp. nov. (E, F), P. brevis sp. nov. (G, H), P. berbeda sp. nov. (I, J), P. lobulata sp. nov. (K, L), P. nr. aspersa (M, N). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4964262" httpUri="https://zenodo.org/record/4964262/files/figure.png" pageId="13" pageNumber="526">5K, 5L</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35D657AFFC6FFB988EC7D7CFB9DC2AB" pageId="13" pageNumber="526" type="materials_examined">
<paragraph id="8BF836F1FFC6FFB988EC7D7CFC39C247" blockId="13.[151,1437,259,934]" pageId="13" pageNumber="526">
<emphasis id="B933EAE3FFC6FFB988EC7D7CFE4EC23C" bold="true" box="[151,394,259,285]" pageId="13" pageNumber="526">Material examined.</emphasis>
<materialsCitation id="3B2F3CACFFC6FFB989E17D7CFC39C247" ID-GBIF-Occurrence="3314109304" collectingDate="2014-11-11" collectingDateMax="2014-11-17" collectingDateMin="2014-11-11" collectionCode="T, MZB" country="Indonesia" elevation="27" latitude="-4.0688334" location="Kumawa" longLatPrecision="1" longitude="133.03644" municipality="New Guinea" pageId="13" pageNumber="526" specimenCode="LEN2014, KUM2" specimenCount="27" stateProvince="Papua Barat" typeStatus="holotype">
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<typeStatus id="54FC8853FFC6FFB989E17D7CFDC3C23C" box="[410,519,259,285]" pageId="13" pageNumber="526" type="holotype">Holotype</typeStatus>
</emphasis>
(male,
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-TR542),
<collectingMunicipality id="6B9CAC8BFFC6FFB98B2F7D7BFC29C23C" box="[852,1005,259,286]" pageId="13" pageNumber="526">New Guinea</collectingMunicipality>
,
<emphasis id="B933EAE3FFC6FFB98C7A7D7CFBB2C23C" bold="true" box="[1025,1142,259,285]" pageId="13" pageNumber="526">
<collectingCountry id="F3507661FFC6FFB98C7A7D7CFBB2C23C" box="[1025,1142,259,285]" name="Indonesia" pageId="13" pageNumber="526">Indonesia</collectingCountry>
</emphasis>
,
<collectingRegion id="4983F813FFC6FFB98CF67D7BFADFC23F" box="[1165,1307,260,286]" country="Indonesia" name="Papua Barat" pageId="13" pageNumber="526">West Papua</collectingRegion>
,
<location id="8E98602AFFC6FFB98D4B7D7BFA5CC23F" LSID="urn:lsid:plazi:treatment:03EE87E7FFC6FFBB88EC7CE8FE7AC714:8E98602AFFC6FFB98D4B7D7BFA5CC23F" box="[1328,1432,260,286]" country="Indonesia" latitude="-4.0688334" longLatPrecision="1" longitude="133.03644" municipality="New Guinea" name="Kumawa" pageId="13" pageNumber="526" stateProvince="Papua Barat">Kumawa</location>
,
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,
<geoCoordinate id="EE735036FFC6FFB989557D58FE00C260" box="[302,452,295,321]" degrees="133.036444" direction="east" orientation="longitude" pageId="13" pageNumber="526" precision="1" value="133.03644">E133.036444</geoCoordinate>
(
<specimenCode id="DBE19E8AFFC6FFB989AE7D58FDEEC260" box="[469,554,295,321]" collectionCode="KUM" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="13" pageNumber="526" type="Herbarium">KUM2</specimenCode>
),
<specimenCount id="9D41FD78FFC6FFB98A447D58FD41C260" box="[575,645,295,321]" pageId="13" pageNumber="526" type="generic">
<quantity id="4CBF9B14FFC6FFB98A447D58FD45C260" box="[575,641,295,321]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="13" pageNumber="526" unit="m" value="27.0">
<elevation id="006AD1C2FFC6FFB98A447D58FD45C260" box="[575,641,295,321]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="13" pageNumber="526" unit="m" value="27.0">27 m</elevation>
</quantity>
.
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a.s.l., forêt littorale karstique sur pente [karst coastal forest on slope],
<date id="FFF91031FFC6FFB988EC7D34FEFEC244" box="[151,314,331,357]" pageId="13" pageNumber="526" value="2014-11-11" valueMax="2014-11-17" valueMin="2014-11-11">
<collectingDate id="EFBDE9D9FFC6FFB988EC7D34FEFEC244" box="[151,314,331,357]" pageId="13" pageNumber="526" value="2014-11-11" valueMax="2014-11-17" valueMin="2014-11-11">1117.xi.2014</collectingDate>
</date>
, jour [day], litière [leaf litter],
<collectionCode id="ED56AE34FFC6FFB98AEA7D33FD67C247" box="[657,675,332,358]" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="526">T</collectionCode>
. Robillard (
<collectionCode id="ED56AE34FFC6FFB98B5C7D34FCA1C244" box="[807,869,331,357]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="13" pageNumber="526" type="Museum">MZB</collectionCode>
.Orth.21844).
</materialsCitation>
</paragraph>
<paragraph id="8BF836F1FFC6FFB988BC7D10FB9DC2AB" blockId="13.[151,1437,259,934]" box="[199,1113,367,394]" pageId="13" pageNumber="526">
<materialsCitation id="3B2F3CACFFC6FFB988BC7D10FB9DC2AB" ID-GBIF-Occurrence="3314109305" box="[199,1113,367,394]" country="Indonesia" county="New Guinea" location="Kumawa" municipality="Kaimana Regency" pageId="13" pageNumber="526" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
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<typeStatus id="54FC8853FFC6FFB988BC7D10FEC6C2A8" box="[199,258,367,393]" pageId="13" pageNumber="526">Type</typeStatus>
locality.
</emphasis>
<collectingCounty id="62994E7DFFC6FFB989177D0FFE38C2A8" box="[364,508,367,394]" pageId="13" pageNumber="526">New Guinea</collectingCounty>
,
<emphasis id="B933EAE3FFC6FFB98A7C7D10FDB8C2A8" bold="true" box="[519,636,367,393]" pageId="13" pageNumber="526">
<collectingCountry id="F3507661FFC6FFB98A7C7D10FDB8C2A8" box="[519,636,367,393]" name="Indonesia" pageId="13" pageNumber="526">Indonesia</collectingCountry>
</emphasis>
,
<collectingRegion id="4983F813FFC6FFB98AF27D0FFCCBC2AB" box="[649,783,368,394]" country="Indonesia" name="Papua Barat" pageId="13" pageNumber="526">West Papua</collectingRegion>
,
<collectingMunicipality id="6B9CAC8BFFC6FFB98B617D10FC2DC2AB" box="[794,1001,367,394]" pageId="13" pageNumber="526">Kaimana Regency</collectingMunicipality>
,
<location id="8E98602AFFC6FFB98B8F7D0FFB9DC2AB" LSID="urn:lsid:plazi:treatment:03EE87E7FFC6FFBB88EC7CE8FE7AC714:8E98602AFFC6FFB98B8F7D0FFB9DC2AB" box="[1012,1113,368,394]" country="Indonesia" county="New Guinea" municipality="Kaimana Regency" name="Kumawa" pageId="13" pageNumber="526" stateProvince="Papua Barat">Kumawa</location>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35D657AFFC6FFB988BC7DECFE1AC2D4" pageId="13" pageNumber="526" type="etymology">
<paragraph id="8BF836F1FFC6FFB988BC7DECFE1AC2D4" blockId="13.[151,1437,259,934]" pageId="13" pageNumber="526">
<emphasis id="B933EAE3FFC6FFB988BC7DECFE8AC28C" bold="true" box="[199,334,403,429]" pageId="13" pageNumber="526">Etymology.</emphasis>
The species name refers to the median lobe present in the ectophallic fold and the lobule at the apical third external margin of the pseudepiphallic lobe that distinguish this species from other similar congeners; lobulata = lobulated in Latin.
</paragraph>
</subSubSection>
<subSubSection id="C35D657AFFC6FFBA88BC7D80FB0EC5F2" lastPageId="14" lastPageNumber="527" pageId="13" pageNumber="526" type="diagnosis">
<paragraph id="8BF836F1FFC6FFB988BC7D80FA44C143" blockId="13.[151,1437,259,934]" pageId="13" pageNumber="526">
<emphasis id="B933EAE3FFC6FFB988BC7D80FE86C138" bold="true" box="[199,322,511,537]" pageId="13" pageNumber="526">Diagnosis.</emphasis>
This new species is unique among congeners by pseudepiphallic lobes forming pincer-like paired processes, dorsal process bends inward, with setose lobule at apical third along external margin; ectophallic fold with lateral arms pointing postero-externally and with a narrow median lobe tapering into a narrow obtuse apex.
</paragraph>
<paragraph id="8BF836F1FFC6FFB988BC7E14FE3EC087" blockId="13.[151,1437,259,934]" pageId="13" pageNumber="526">
This species is most similar to
<taxonomicName id="4C474D72FFC6FFB98A657E13FCA0C1A4" authority="Gorochov, 1999" authorityName="Gorochov" authorityYear="1999" box="[542,868,619,645]" class="Insecta" family="Haglotettigoniidae" genus="Phaloria" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="526" phylum="Arthropoda" rank="species" species="pareximia">
<emphasis id="B933EAE3FFC6FFB98A657E13FD6EC1A4" box="[542,682,620,645]" italics="true" pageId="13" pageNumber="526">P. pareximia</emphasis>
<bibRefCitation id="EFD64B00FFC6FFB98ACB7E14FCA0C1A4" author="Gorochov, A. V." box="[688,868,619,645]" pageId="13" pageNumber="526" pagination="27 - 60" refId="ref9986" refString="Gorochov, A. V. (1999) New and little known Phaloriinae (Orthoptera: Gryllidae). Zoosystematica Rossica, 8 (1), 27 - 60." type="journal article" year="1999">Gorochov, 1999</bibRefCitation>
</taxonomicName>
from Cyclops Mountains (north-eastern West Papua) and then
<taxonomicName id="4C474D72FFC6FFB989527EEFFD88C188" authority="Gorochov, 1996" authorityName="Gorochov" authorityYear="1996" box="[297,588,655,681]" class="Insecta" family="Haglotettigoniidae" genus="Phaloria" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="526" phylum="Arthropoda" rank="species" species="eximia">
<emphasis id="B933EAE3FFC6FFB989527EEFFE54C188" box="[297,400,656,681]" italics="true" pageId="13" pageNumber="526">P. eximia</emphasis>
<bibRefCitation id="EFD64B00FFC6FFB989EC7EF0FD88C188" author="Gorochov, A. V." box="[407,588,655,681]" pageId="13" pageNumber="526" pagination="29 - 90" refId="ref9941" refString="Gorochov, A. V. (1996) New and little-known crickets from the collection of the Humboldt University and some other collections (Orthoptera: Grylloidea). Part 2. Zoosystematica Rossica, 5 (1), 29 - 90." type="journal article" year="1996">Gorochov, 1996</bibRefCitation>
</taxonomicName>
from New
<collectingCountry id="F3507661FFC6FFB98AB67EF0FCDAC188" box="[717,798,655,681]" name="Guinea-Bissau" pageId="13" pageNumber="526">Guinea</collectingCountry>
in pseudepiphallic lobes, pseudepiphallic parameres and endophallic sclerite; but differs by pseudepiphallic lobes with setose nodule at apical third, shape of pseudepiphallic parameres and ectophallic fold with median lobe and with lateral arms pointing slightly externally. It also differs from
<taxonomicName id="4C474D72FFC6FFB988AD7E83FEA0C034" authorityName="Gorochov" authorityYear="1999" box="[214,356,764,789]" class="Insecta" family="Haglotettigoniidae" genus="Phaloria" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="526" phylum="Arthropoda" rank="species" species="pareximia">
<emphasis id="B933EAE3FFC6FFB988AD7E83FEA0C034" box="[214,356,764,789]" italics="true" pageId="13" pageNumber="526">P. pareximia</emphasis>
</taxonomicName>
by basal process of pseudepiphallic lobes pointing interno-anteriorly (instead of interno-posteiorly). It differs from
<taxonomicName id="4C474D72FFC6FFB989F27F5FFE34C018" authorityName="Gorochov" authorityYear="1996" box="[393,496,800,825]" class="Insecta" family="Haglotettigoniidae" genus="Phaloria" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="526" phylum="Arthropoda" rank="species" species="eximia">
<emphasis id="B933EAE3FFC6FFB989F27F5FFE34C018" box="[393,496,800,825]" italics="true" pageId="13" pageNumber="526">P. eximia</emphasis>
</taxonomicName>
by basal lobe of pseudepiphallic lobes acute (rather than stout) and pseudepiphallic parameres without long acute process. This species is also vaguely similar in pseudepiphallic lobes as
<taxonomicName id="4C474D72FFC6FFB98D647F3BFE8FC0A3" authority="Gorochov, 2005" authorityName="Gorochov" authorityYear="2005" class="Insecta" family="Haglotettigoniidae" genus="Phaloria" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="526" phylum="Arthropoda" rank="species" species="aleximia">
<emphasis id="B933EAE3FFC6FFB98D647F3BFA58C07C" box="[1311,1436,835,861]" italics="true" pageId="13" pageNumber="526">P. aleximia</emphasis>
<bibRefCitation id="EFD64B00FFC6FFB988EC7F18FE8FC0A3" author="Gorochov, A. V." box="[151,331,871,898]" pageId="13" pageNumber="526" pagination="25 - 46" refId="ref10018" refString="Gorochov, A. V. (2005) New and little known crickets of the subfamilies Phaloriinae, Phalangopsinae and Landrevinae (Orthoptera, Gryllidae) from Indonesia and South Africa. Trudy Russkago Entomologicheskago Obshchestva [Horae Societatis Entomologicae Rossicae], 76, 25 - 46." type="journal article" year="2005">Gorochov, 2005</bibRefCitation>
</taxonomicName>
from Supiori Island (north of West Papua) but differs by shape of pseudepiphallic parameres without long acute external process.
</paragraph>
<caption id="DF386679FFC6FFB988EC7BE3FCE1C4F9" ID-DOI="http://doi.org/10.5281/zenodo.4964264" ID-Zenodo-Dep="4964264" httpUri="https://zenodo.org/record/4964264/files/figure.png" pageId="13" pageNumber="526" startId="13.[151,250,1948,1972]" targetBox="[151,1436,957,1924]" targetPageId="13">
<paragraph id="8BF836F1FFC6FFB988EC7BE3FCE1C4F9" blockId="13.[151,1436,1948,2008]" pageId="13" pageNumber="526">
<emphasis id="B933EAE3FFC6FFB988EC7BE3FED7C495" bold="true" box="[151,275,1948,1972]" pageId="13" pageNumber="526">FIGURE 6.</emphasis>
<taxonomicName id="4C474D72FFC6FFB989617BE3FEB0C495" authorityName="Stal" authorityYear="1877" box="[282,372,1948,1972]" class="Insecta" family="Haglotettigoniidae" genus="Phaloria" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="526" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAE3FFC6FFB989617BE3FEB0C495" box="[282,372,1948,1972]" italics="true" pageId="13" pageNumber="526">Phaloria</emphasis>
</taxonomicName>
in the natural environment: probably
<taxonomicName id="4C474D72FFC6FFB98A837BE2FCAAC495" authority="Tan &amp; Rahmadi &amp; Robillard, 2021" authorityName="Tan &amp; Rahmadi &amp; Robillard" authorityYear="2021" box="[760,878,1948,1972]" class="Insecta" family="Haglotettigoniidae" genus="Phaloria" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="526" phylum="Arthropoda" rank="species" species="dischidia" status="sp. nov.">
<emphasis id="B933EAE3FFC6FFB98A837BE2FCAAC495" box="[760,878,1948,1972]" italics="true" pageId="13" pageNumber="526">P. dischidia</emphasis>
</taxonomicName>
<emphasis id="B933EAE3FFC6FFB98B0E7BE3FC02C495" bold="true" box="[885,966,1948,1972]" pageId="13" pageNumber="526">
<taxonomicNameLabel id="A2005798FFC6FFB98B0E7BE3FC02C495" box="[885,966,1948,1972]" pageId="13" pageNumber="526" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male (A) and female (B) in Lobo;
<taxonomicName id="4C474D72FFC6FFB98D567BE2FF12C4F9" authority="Tan &amp; Rahmadi &amp; Robillard, 2021" authorityName="Tan &amp; Rahmadi &amp; Robillard" authorityYear="2021" class="Insecta" family="Haglotettigoniidae" genus="Phaloria" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="526" phylum="Arthropoda" rank="species" species="paradischidia" status="sp. nov.">
<emphasis id="B933EAE3FFC6FFB98D567BE2FF12C4F9" italics="true" pageId="13" pageNumber="526">P. paradischidia</emphasis>
</taxonomicName>
<emphasis id="B933EAE3FFC6FFB988A67BBFFEE9C4F9" bold="true" box="[221,301,1984,2008]" pageId="13" pageNumber="526">
<taxonomicNameLabel id="A2005798FFC6FFB988A67BBFFEE9C4F9" box="[221,301,1984,2008]" pageId="13" pageNumber="526" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from Kumawa (C);
<taxonomicName id="4C474D72FFC6FFB989867BBEFD8DC4F9" authority="Tan &amp; Rahmadi &amp; Robillard, 2021" authorityName="Tan &amp; Rahmadi &amp; Robillard" authorityYear="2021" box="[509,585,1984,2008]" class="Insecta" family="Haglotettigoniidae" genus="Phaloria" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="526" phylum="Arthropoda" rank="species" species="tukul" status="sp. nov.">
<emphasis id="B933EAE3FFC6FFB989867BBEFD8DC4F9" box="[509,585,1984,2008]" italics="true" pageId="13" pageNumber="526">P. tukul</emphasis>
</taxonomicName>
<emphasis id="B933EAE3FFC6FFB98A347BBFFD5BC4F9" bold="true" box="[591,671,1984,2008]" pageId="13" pageNumber="526">
<taxonomicNameLabel id="A2005798FFC6FFB98A347BBFFD5BC4F9" box="[591,671,1984,2008]" pageId="13" pageNumber="526" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in Lobo (D).
</paragraph>
</caption>
<caption id="DF386679FFC5FFBA88EC7A54FD05C546" ID-DOI="http://doi.org/10.5281/zenodo.4964266" ID-Zenodo-Dep="4964266" httpUri="https://zenodo.org/record/4964266/files/figure.png" pageId="14" pageNumber="527" startId="14.[151,250,1579,1603]" targetBox="[214,1369,210,1536]" targetPageId="14">
<paragraph id="8BF836F1FFC5FFBA88EC7A54FD05C546" blockId="14.[151,1436,1579,1639]" pageId="14" pageNumber="527">
<emphasis id="B933EAE3FFC5FFBA88EC7A54FED6C562" bold="true" box="[151,274,1579,1603]" pageId="14" pageNumber="527">FIGURE 7.</emphasis>
Calling song of
<taxonomicName id="4C474D72FFC5FFBA89C37A54FD61C562" authority="Tan &amp; Rahmadi &amp; Robillard, 2021" authorityName="Tan &amp; Rahmadi &amp; Robillard" authorityYear="2021" box="[440,677,1579,1603]" class="Insecta" family="Haglotettigoniidae" genus="Phaloria" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="527" phylum="Arthropoda" rank="species" species="paradischidia" status="sp. nov.">
<emphasis id="B933EAE3FFC5FFBA89C37A54FD61C562" box="[440,677,1579,1603]" italics="true" pageId="14" pageNumber="527">Phaloria paradischidia</emphasis>
</taxonomicName>
<emphasis id="B933EAE3FFC5FFBA8AD17A54FD3EC562" bold="true" box="[682,762,1579,1603]" pageId="14" pageNumber="527">
<taxonomicNameLabel id="A2005798FFC5FFBA8AD17A54FD3EC562" box="[682,762,1579,1603]" pageId="14" pageNumber="527" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: a sequence of five calling songs (A); a single calling song consisting of two syllables (B); spectral plot of a syllable (C).
</paragraph>
</caption>
<paragraph id="8BF836F1FFC5FFBA88BC7AEAFB0EC5F2" blockId="14.[151,1437,1685,2036]" pageId="14" pageNumber="527">
<emphasis id="B933EAE3FFC5FFBA88BC7AEAFE65C58E" bold="true" box="[199,417,1685,1711]" pageId="14" pageNumber="527">Subgeneric status.</emphasis>
This species should belong to the subgenus
<taxonomicName id="4C474D72FFC5FFBA8BEA7AEAFBC7C58E" authorityName="Gorochov" authorityYear="1996" box="[913,1027,1685,1711]" class="Insecta" family="Haglotettigoniidae" genus="Phaloria" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="527" phylum="Arthropoda" rank="subGenus" subGenus="Papuloria">
<emphasis id="B933EAE3FFC5FFBA8BEA7AEAFBC7C58E" box="[913,1027,1685,1711]" italics="true" pageId="14" pageNumber="527">Papuloria</emphasis>
</taxonomicName>
based on the pseudepiphallus with a pair of lateral lophi and endophallic sclerite without large unpaired apodeme directed anteriorly.
</paragraph>
</subSubSection>
<subSubSection id="C35D657AFFC5FFBB88BC7AA2FE7AC714" lastPageId="15" lastPageNumber="528" pageId="14" pageNumber="527" type="description">
<paragraph id="8BF836F1FFC5FFBB88BC7AA2FD5DC2A8" blockId="14.[151,1437,1685,2036]" lastBlockId="15.[151,1437,151,1077]" lastPageId="15" lastPageNumber="528" pageId="14" pageNumber="527">
<emphasis id="B933EAE3FFC5FFBA88BC7AA2FE9DC5D6" bold="true" box="[199,345,1757,1783]" pageId="14" pageNumber="527">Description.</emphasis>
Similar to
<taxonomicName id="4C474D72FFC5FFBA89A07AA2FC96C5D6" authority="Tan &amp; Robillard" authorityName="Tan &amp; Rahmadi &amp; Robillard" authorityYear="2021" box="[475,850,1757,1784]" class="Insecta" family="Haglotettigoniidae" genus="Phaloria" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="527" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
<emphasis id="B933EAE3FFC5FFBA89A07AA2FD4FC5D6" box="[475,651,1757,1783]" italics="true" pageId="14" pageNumber="527">Phaloria brevis</emphasis>
Tan &amp; Robillard
</taxonomicName>
,
<emphasis id="B933EAE3FFC5FFBA8B257AA2FC7EC5D6" bold="true" box="[862,954,1757,1783]" pageId="14" pageNumber="527">
<taxonomicNameLabel id="A2005798FFC5FFBA8B257AA2FC7EC5D6" box="[862,954,1757,1783]" pageId="14" pageNumber="527" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in habitus, size and colour patterns (more contrasting but could be due to artefact of preservation) (
<figureCitation id="137C2A74FFC5FFBA8B757B7EFCA7C43A" box="[782,867,1793,1819]" captionStart="FIGURE 1" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[195,1408,189,1918]" captionTargetId="figure-26@8.[179,1408,181,1920]" captionTargetPageId="8" captionText="FIGURE 1. Habitus of Phaloria males (AD, FI) and females (E, J, K) from Lobo and Kumawa in dorsal (AH, J) and lateral (I, K) views: P. dischidia sp. nov. (A), P. paradischidia sp. nov. (B), P. tukul sp. nov. (C), P. brevis sp. nov. (D, E), P. berbeda sp. nov. (F), P. lobulata sp. nov. (G), P. nr. aspersa (HK). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4964251" httpUri="https://zenodo.org/record/4964251/files/figure.png" pageId="14" pageNumber="527">Fig. 1G</figureCitation>
). Head dorsum yellow brown with irregular brown bands at posterior end (
<figureCitation id="137C2A74FFC5FFBA89DF7B5AFE38C41E" box="[420,508,1829,1855]" captionStart="FIGURE 2" captionStartId="9.[151,250,1814,1838]" captionTargetBox="[160,1417,196,1778]" captionTargetId="figure-19@9.[151,1436,181,1790]" captionTargetPageId="9" captionText="FIGURE 2. Head and pronotum of P. dischidia sp. nov. (A), P. paradischidia sp. nov. (B), P. tukul sp. nov. (C), P. brevis sp. nov. (D), P. berbeda sp. nov. (E), P. lobulata sp. nov. (F), P. nr. aspersa (G) in dorsal view. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4964253" httpUri="https://zenodo.org/record/4964253/files/figure.png" pageId="14" pageNumber="527">Fig. 2F</figureCitation>
). Fastigium darker brown. Scapes yellow brown, ventrally dark brown. Antennae yellow brown. Fastigium verticis brown. Frons, clypeus and mouthparts pale brown; frons with a horizontal dark brown stripe ventral to eye; clypeus with another horizontal dark brown stripe (
<figureCitation id="137C2A74FFC5FFBA8C647B12FBB1C4A6" box="[1055,1141,1901,1927]" captionStart="FIGURE 3" captionStartId="10.[151,250,1696,1720]" captionTargetBox="[197,1378,236,1639]" captionTargetId="figure-26@10.[183,1404,181,1672]" captionTargetPageId="10" captionText="FIGURE 3. Faces of P. dischidia sp. nov. (A), P. paradischidia sp. nov. (B), P. tukul sp. nov. (C), P. brevis sp. nov. (D), P. berbeda sp. nov. (E), P. lobulata sp. nov. (F), P. nr. aspersa (G) in anterior views. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4964255" httpUri="https://zenodo.org/record/4964255/files/figure.png" pageId="14" pageNumber="527">Fig. 3F</figureCitation>
). Maxillary palpi pale coloured, apices of segments faintly brownish. Lateral part of head, including genae, pale yellow brown; gena with a vertical brown band ventral to eye; with a brown horizontal band at posterior of eye; posterior margin of head dark brown. Pronotal disk generally dark brown with median carina pale brown and two pairs of lateral oval pale brown spots; posterior margin slightly sinuous (
<figureCitation id="137C2A74FFC4FFBB8A1C7CE8FD7AC390" box="[615,702,151,177]" captionStart="FIGURE 2" captionStartId="9.[151,250,1814,1838]" captionTargetBox="[160,1417,196,1778]" captionTargetId="figure-19@9.[151,1436,181,1790]" captionTargetPageId="9" captionText="FIGURE 2. Head and pronotum of P. dischidia sp. nov. (A), P. paradischidia sp. nov. (B), P. tukul sp. nov. (C), P. brevis sp. nov. (D), P. berbeda sp. nov. (E), P. lobulata sp. nov. (F), P. nr. aspersa (G) in dorsal view. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4964253" httpUri="https://zenodo.org/record/4964253/files/figure.png" pageId="15" pageNumber="528">Fig. 2F</figureCitation>
). Lateral lobes also dark brown, with two pale brown spots near ventro-anterior angle. Legs pale yellow brown with brown patterns and spots. Inner tympanum oblong; outer tympanum smaller and oval. FIs and FIIs with two brown oblong spots on inner and outer surfaces at the base (sometimes forming an incomplete ring), followed by brown rings near knees. TIs with three brown rings (one around tympana, one in middle and one at apex). FIIIs yellow brown basally with some brown spots and mottled patterns on the dorsal surface, distal third with a brown ring and knee area brown. TIIIs yellow brown with brown rings, particularly around bases of spurs. Tergites yellow brown.
</paragraph>
<paragraph id="8BF836F1FFC4FFBB88BC7DECFAC2C11C" blockId="15.[151,1437,151,1077]" pageId="15" pageNumber="528">
Male. FWs slender, about 2.9 times longer than wide, surpassing abdominal apex, mostly yellow brown and some parts hyaline (
<figureCitation id="137C2A74FFC4FFBB89017DC8FE15C2F0" box="[378,465,439,465]" captionStart="FIGURE 4" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[195,1397,192,1918]" captionTargetId="figure-19@11.[183,1404,181,1920]" captionTargetPageId="11" captionText="FIGURE 4. Male FWs of P. dischidia sp. nov. (A), P. paradischidia sp. nov. (B), P. tukul sp. nov. (C), P. brevis sp. nov. (D), P. berbeda sp. nov. (F), P. lobulata sp. nov. (G), P. nr. aspersa (H) and female FWs of P. brevis sp. nov. (E) and P. nr. aspersa (I) in dorsal views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4964258" httpUri="https://zenodo.org/record/4964258/files/figure.png" pageId="15" pageNumber="528">Fig. 4G</figureCitation>
). FW venation typical of genus, 8 fairly straight veins in harp; mirror as long as wide, very large and separated by two dividing veins: basal one mostly straight, distal one angularly bent. Apical field long, 1.1 times longer than length of mirror. Lateral field with R and M diverging before converging strongly at apical third, with 6 cross veins between R and M. Hind wings reaching apex of FW, not exceeding FWs.
</paragraph>
<paragraph id="8BF836F1FFC4FFBB88BC7E38FC2AC0CC" blockId="15.[151,1437,151,1077]" pageId="15" pageNumber="528">
Male genitalia (
<figureCitation id="137C2A74FFC4FFBB89027E38FDCEC140" box="[377,522,582,609]" captionStart="FIGURE 5" captionStartId="12.[151,250,1841,1865]" captionTargetBox="[244,1363,190,1783]" captionTargetId="figure-26@12.[215,1372,181,1818]" captionTargetPageId="12" captionText="FIGURE 5. Male genitalia of Phaloria in dorsal (A, C, E, G, I, K, M) and ventral (B, D, F, H, J, L, N) views: P. dischidia sp. nov. (A, B), P. paradischidia sp. nov. (C, D), P. tukul sp. nov. (E, F), P. brevis sp. nov. (G, H), P. berbeda sp. nov. (I, J), P. lobulata sp. nov. (K, L), P. nr. aspersa (M, N). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4964262" httpUri="https://zenodo.org/record/4964262/files/figure.png" pageId="15" pageNumber="528">Figs. 5K, 5L</figureCitation>
). Pseudepiphallus with lateral margins slightly converging posterior; with posterior margin deeply, roundly and broadly incised in the middle, in the middle with a pair of small stout triangular lobules. Pseudepiphallic lobe forming pincer-like paired processes; both processes pointing internally, slender and with acute apex; dorsal process anterior of ventral process; dorsal process bends inward, with setose lobule at apical third along external margin; ventral process more gently curved. Rami long, longer than the pseudepiphallus length, curved inwards apically, but not connected. Pseudepiphallic parameres strongly sclerotized, obliquely transverse and curved, with two posterior lobes, one dorsal and one ventral; dorsal lobe basal of ventral lobe, triangular; ventral lobe tongue-shaped, pointing interno-posteriorly. Ectophallic apodemes fork-shaped, with basal end straight and fairly long; inner apical arm pointing interno-posteriorly, apex rounded; outer apical arm longer, pointing posteriorly, apex enlarged. Ectophallic fold broad, laterally with lateral arms pointing postero-externally; posterior margin concave with a narrow median lobe tapering into a narrow obtuse apex. Endophallic sclerite small with two lateral plate-like structures, star-shaped; with a pair of stout and obtuse basal lobes.
</paragraph>
<paragraph id="8BF836F1FFC4FFBB88BC7F88FE7AC714" blockId="15.[151,1437,151,1077]" pageId="15" pageNumber="528">
Measurements (in mm). Male
<typeStatus id="54FC8853FFC4FFBB8A5F7F88FD42C730" box="[548,646,1015,1041]" pageId="15" pageNumber="528" type="holotype">holotype</typeStatus>
PronL = 2.4, PronW = 4.3, FWL = 17.3, FWW = 5.9, FIIIL = 10.9, FIIIW = 2.1, TIIIL = 10.0.
</paragraph>
</subSubSection>
</treatment>
</document>