273 lines
34 KiB
XML
273 lines
34 KiB
XML
<document id="53DBAA40D97CC42F92B9C2B91FD57061" ID="10.11646/zootaxa.4193.2.9" ID-DOI="10.11646/zootaxa.4193.2.9" ID-GBIF-Dataset="82b05856-1eeb-41d1-bc8b-c2dcb7883310" ID-ISSN="1175-5326" ID-Zenodo-Dep="166958" ID-ZooBank="52692E2F-8D7D-4C55-8A04-917C7D2A4D3D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1479284196931" checkinUser="plazi" docAuthor="Kompier, Tom" docDate="2016" docId="03EE87DF575F3137FF368E7B4734D566" docLanguage="en" docName="zootaxa.4193.2.9.pdf" docOrigin="Zootaxa 4193 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Protosticta nigra Kompier, 2016, spec. nov." docType="treatment" docVersion="10" lastPageNumber="356" masterDocId="FFD7FFA75758313DFFA18D63427ED47A" masterDocTitle="New species of Protosticta Selys from Vietnam with a key to the males of the P. curiosa group (Odonata: Platystictidae)" masterLastPageNumber="360" masterPageNumber="347" pageNumber="354" updateTime="1698393694562" updateUser="plazi">
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<mods:title id="03F6A2859A56704A74CA2A912E774021">New species of Protosticta Selys from Vietnam with a key to the males of the P. curiosa group (Odonata: Platystictidae)</mods:title>
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<mods:name id="87A6A5EB2BDF4289A68AAC79C7887CA5" type="personal">
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<mods:roleTerm id="8A7F2517BA1C597EB5124C0B48B6ED42">Author</mods:roleTerm>
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<mods:namePart id="692A4928DA46AC263A402A589F50F3AD">Kompier, Tom</mods:namePart>
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<mods:typeOfResource id="8784D1CF474CEA4E901FD3E2088BEB04">text</mods:typeOfResource>
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<mods:title id="372FEBBBF0EA3FE12A252047B568E12B">Zootaxa</mods:title>
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<mods:part id="7F4A72FAE9AD7E7A91D2DDAA72E9E4C6">
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<mods:date id="5DD3CEF26DC9FEC2D53D63EDFB315C5E">2016</mods:date>
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<mods:number id="8CF30BA3EABC3C15002BC843DEF2BE76">4193</mods:number>
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<mods:number id="ACAECFBF353CD6052B69A70545CF3927">2</mods:number>
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<mods:identifier id="E2525D312FBF3B1D9FA7DFCEF65DC335" type="DOI">10.11646/zootaxa.4193.2.9</mods:identifier>
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<treatment id="03EE87DF575F3137FF368E7B4734D566" ID-DOI="http://doi.org/10.5281/zenodo.5692075" ID-GBIF-Taxon="124736668" ID-Zenodo-Dep="5692075" LSID="urn:lsid:plazi:treatment:03EE87DF575F3137FF368E7B4734D566" httpUri="http://treatment.plazi.org/id/03EE87DF575F3137FF368E7B4734D566" lastPageId="10" lastPageNumber="356" pageId="7" pageNumber="354">
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<subSubSection id="C35D6542575F313AFF368E7B435DD72E" pageId="7" pageNumber="354" type="nomenclature">
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<paragraph id="8BF836C9575F313AFF368E7B439ED749" blockId="7.[151,480,792,853]" box="[151,480,792,819]" pageId="7" pageNumber="354">
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<heading id="D0B081A5575F313AFF368E7B439ED749" bold="true" box="[151,480,792,819]" fontSize="11" level="1" pageId="7" pageNumber="354" reason="1">
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<emphasis id="B933EADB575F313AFF368E7B439ED749" bold="true" box="[151,480,792,819]" pageId="7" pageNumber="354">
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<taxonomicName id="4C474D4A575F313AFF368E7B431FD748" ID-CoL="9X45X" box="[151,353,792,818]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="nigra" status="sp. nov.">
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<emphasis id="B933EADB575F313AFF368E7B431FD748" bold="true" box="[151,353,792,818]" italics="true" pageId="7" pageNumber="354">Protosticta nigra</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20057A0575F313AFEC98E7A439ED749" box="[360,480,793,819]" pageId="7" pageNumber="354" rank="species">spec. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF836C9575F313AFF368E5F435DD72E" blockId="7.[151,480,792,853]" box="[151,291,828,853]" pageId="7" pageNumber="354">
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(
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<figureCitation id="137C2A4C575F313AFF3E8E5F4365D72E" box="[159,283,828,853]" captionStart-0="FIGURE 12" captionStart-1="FIGURE 13" captionStart-2="FIGURE 14" captionStartId-0="7.[151,250,1926,1948]" captionStartId-1="8.[151,250,829,851]" captionStartId-2="8.[151,250,1585,1607]" captionTargetBox-0="[151,1436,1013,1905]" captionTargetBox-1="[384,1203,193,807]" captionTargetBox-2="[340,1066,988,1507]" captionTargetId-0="figure@7.[151,1436,1013,1905]" captionTargetId-1="figure@8.[384,1203,193,807]" captionTargetId-2="figure@8.[340,1247,900,1563]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="8" captionText-0="FIGURE 12. Male P. nigra spec. nov, 22. V. 2016, Tam Dao National Park, Vinh Phuc Province." captionText-1="FIGURE 13. Thorax of male P. nigra spec. nov., 22. V. 2016, Tam Dao National Park, Vinh Phuc Province, T. Kompier leg." captionText-2="FIGURE 14. Appendages of male P. nigra spec. nov., 22. V. 2016, Tam Dao National Park, Vinh Phuc Province, T. Kompier leg." httpUri-0="https://zenodo.org/record/266147/files/figure.png" httpUri-1="https://zenodo.org/record/266148/files/figure.png" httpUri-2="https://zenodo.org/record/166971/files/figure.png" pageId="7" pageNumber="354">Fig. 12–14</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C35D6542575F3137FF368EE74734D566" lastPageId="10" lastPageNumber="357" pageId="7" pageNumber="354" type="etymology">
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<paragraph id="8BF836C9575F313AFF368EE74291D7BA" blockId="7.[151,1436,900,960]" pageId="7" pageNumber="354">
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<emphasis id="B933EADB575F313AFF368EE74360D7E7" bold="true" box="[151,286,900,925]" pageId="7" pageNumber="354">Etymology.</emphasis>
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The adjective
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<taxonomicName id="4C474D4A575F313AFE628EE64381D7E6" box="[451,511,901,924]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="nigra">
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<emphasis id="B933EADB575F313AFE628EE64381D7E6" box="[451,511,901,924]" italics="true" pageId="7" pageNumber="354">nigra</emphasis>
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</taxonomicName>
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in the specific name means black and refers to the blackish thorax (
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<figureCitation id="137C2A4C575F313AFB5B8EE74732D7E6" box="[1274,1356,900,925]" captionStart="FIGURE 13" captionStartId="8.[151,250,829,851]" captionTargetBox="[384,1203,193,807]" captionTargetId="figure@8.[384,1203,193,807]" captionTargetPageId="8" captionText="FIGURE 13. Thorax of male P. nigra spec. nov., 22. V. 2016, Tam Dao National Park, Vinh Phuc Province, T. Kompier leg." httpUri="https://zenodo.org/record/266148/files/figure.png" pageId="7" pageNumber="354">Fig. 13</figureCitation>
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) of the species.
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</paragraph>
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<caption id="DF386641575F313AFF368AE5461CD3E6" box="[151,1122,1926,1948]" httpUri="https://zenodo.org/record/266147/files/figure.png" pageId="7" pageNumber="354" targetBox="[151,1436,1013,1905]" targetPageId="7">
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<paragraph id="8BF836C9575F313AFF368AE5461CD3E6" blockId="7.[151,1122,1926,1948]" box="[151,1122,1926,1948]" pageId="7" pageNumber="354">
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<emphasis id="B933EADB575F313AFF368AE54361D3E6" bold="true" box="[151,287,1926,1948]" pageId="7" pageNumber="354">FIGURE 12.</emphasis>
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Male
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<taxonomicName id="4C474D4A575F313AFEC18AE443D1D3E6" box="[352,431,1927,1948]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="nigra" status="sp. nov.">
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<emphasis id="B933EADB575F313AFEC18AE443D1D3E6" box="[352,431,1927,1948]" italics="true" pageId="7" pageNumber="354">P. nigra</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20057A0575F313AFE148AE44068D3E6" box="[437,534,1927,1948]" pageId="7" pageNumber="354" rank="species">spec. nov</taxonomicNameLabel>
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, 22.V.2016, Tam Dao National Park, Vinh Phuc Province.
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</paragraph>
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</caption>
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<caption id="DF38664157503135FF368E5E4708D729" box="[151,1398,829,851]" httpUri="https://zenodo.org/record/266148/files/figure.png" pageId="8" pageNumber="355" targetBox="[384,1203,193,807]" targetPageId="8">
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<paragraph id="8BF836C957503135FF368E5E4708D729" blockId="8.[151,1398,829,851]" box="[151,1398,829,851]" pageId="8" pageNumber="355">
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<emphasis id="B933EADB57503135FF368E5E4361D728" bold="true" box="[151,287,829,851]" pageId="8" pageNumber="355">FIGURE 13.</emphasis>
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Thorax of male
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<taxonomicName id="4C474D4A57503135FE698E5D4069D729" box="[456,535,830,851]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="nigra" status="sp. nov.">
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<emphasis id="B933EADB57503135FE698E5D4069D729" box="[456,535,830,851]" italics="true" pageId="8" pageNumber="355">P. nigra</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20057A057503135FDBF8E5D40FBD729" box="[542,645,830,851]" pageId="8" pageNumber="355" rank="species">spec. nov.</taxonomicNameLabel>
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, 22.V.2016, Tam Dao National Park, Vinh Phuc Province, T. Kompier leg.
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</paragraph>
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</caption>
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<caption id="DF38664157503135FF368B5242C3D21C" httpUri="https://zenodo.org/record/166971/files/figure.png" pageId="8" pageNumber="355" targetBox="[340,1066,988,1507]" targetPageId="8">
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<paragraph id="8BF836C957503135FF368B5242C3D21C" blockId="8.[151,1436,1585,1638]" pageId="8" pageNumber="355">
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<emphasis id="B933EADB57503135FF368B52435FD23C" bold="true" box="[151,289,1585,1607]" pageId="8" pageNumber="355">FIGURE 14.</emphasis>
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Appendages of male
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<taxonomicName id="4C474D4A57503135FDA58B514028D23D" box="[516,598,1586,1607]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="nigra" status="sp. nov.">
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<emphasis id="B933EADB57503135FDA58B514028D23D" box="[516,598,1586,1607]" italics="true" pageId="8" pageNumber="355">P. nigra</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20057A057503135FDFC8B5140B8D23D" box="[605,710,1586,1607]" pageId="8" pageNumber="355" rank="species">spec. nov.</taxonomicNameLabel>
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, 22.V.2016, Tam Dao National Park, Vinh Phuc Province, T. Kompier leg.
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</paragraph>
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</caption>
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<paragraph id="8BF836C957503134FF668BF0407FD53A" blockId="8.[151,1437,1682,2032]" lastBlockId="9.[151,1437,151,861]" lastPageId="9" lastPageNumber="356" pageId="8" pageNumber="355">
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<materialsCitation id="3B2F3C9457503135FF668BF04611D289" ID-GBIF-Occurrence="1325928574" collectorName="Van Tol" location="Vinh Phuc Province" pageId="8" pageNumber="355" specimenCount="1" stateProvince="Vinh Phuc Province">
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<collectorName id="26B2531F57503135FF668BF0435CD2D1" box="[199,290,1682,1707]" pageId="8" pageNumber="355">
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Van
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<bibRefCitation id="EFD64B3857503135FF5C8BF1435CD2D1" author="Tol" box="[253,290,1682,1707]" pageId="8" pageNumber="355" refString="van Tol, J. (2008) Notes on some species of the genus Protosticta from Vietnam (Odonata, Platystictidae). Zoologische Mededelingen Leiden, 82, 217 - 234." type="journal article" year="2008">Tol</bibRefCitation>
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</collectorName>
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(2008) provided thorough descriptions of both light and dark form of
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<emphasis id="B933EADB57503135FBF88BF046D4D2D0" box="[1113,1194,1683,1706]" italics="true" pageId="8" pageNumber="355">P. s a t o i</emphasis>
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. The dark form was described based on specimens from Tam Dao National Park,
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<collectingRegion id="4983F82B57503135FCF38BD54647D2B5" box="[850,1081,1718,1743]" country="Vietnam" name="Vinh Phuc" pageId="8" pageNumber="355">
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Vinh
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<collectingRegion id="4983F82B57503135FC308BD54647D2B5" box="[913,1081,1718,1743]" inferredBySuffix="true" pageId="8" pageNumber="355">Phuc Province</collectingRegion>
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</collectingRegion>
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, whereas the description of the light form (
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<figureCitation id="137C2A4C57503135FEB78BB943E8D289" box="[278,406,1754,1779]" captionStart-0="FIGURE 15" captionStart-1="FIGURE 16" captionStartId-0="9.[151,250,1774,1796]" captionStartId-1="10.[151,250,1608,1630]" captionTargetBox-0="[151,1436,913,1752]" captionTargetBox-1="[151,1436,708,1586]" captionTargetId-0="figure@9.[151,1436,913,1753]" captionTargetId-1="figure@10.[151,1436,708,1587]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURE 15. Male P. satoi, 24. VII. 2016, Cuc Phuong National Park, Ninh Binh Province. Note yellowish white prothorax with black marking on posterior lobe only." captionText-1="FIGURE 16. Male P. satoi, 01. VIII. 2016, Xuan Son National Park, Phu Tho Province. This male with black markings on all lobes of prothorax." httpUri-0="https://zenodo.org/record/266149/files/figure.png" httpUri-1="https://zenodo.org/record/166973/files/figure.png" pageId="8" pageNumber="355">Figs. 15-16</figureCitation>
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) was based on specimens collected in Cuc Phuong National Park
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</materialsCitation>
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,
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<materialsCitation id="3B2F3C9457503135FBDB8BB940E0D36D" ID-GBIF-Occurrence="1325928571" country="Vietnam" location="Ninh Binh Province" pageId="8" pageNumber="355" specimenCount="1" stateProvince="Ninh Binh Province">
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<collectingRegion id="4983F82B57503135FBDB8BB94722D289" box="[1146,1372,1754,1779]" country="Vietnam" name="Ninh Binh" pageId="8" pageNumber="355">
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Ninh
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<collectingRegion id="4983F82B57503135FB198BB94722D289" box="[1208,1372,1754,1779]" inferredBySuffix="true" pageId="8" pageNumber="355">Binh Province</collectingRegion>
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</collectingRegion>
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. Both provinces are located in the north of
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<collectingCountry id="F350765957503135FD9A8B9D40E0D36D" box="[571,670,1790,1815]" name="Vietnam" pageId="8" pageNumber="355">Vietnam</collectingCountry>
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</materialsCitation>
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||
.
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<materialsCitation id="3B2F3C9457503134FD088B9D4300D48D" ID-GBIF-Occurrence="1325928570" collectorName="Asahina & Tam Dao. The & From & This & van Tol & It" country="Vietnam" lastPageId="9" lastPageNumber="356" location="The" municipality="Tam Dao Town" pageId="8" pageNumber="355" specimenCount="1" typeStatus="holotype">
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<location id="8E98601257503135FD088B9D40ABD36D" LSID="urn:lsid:plazi:treatment:03EE87DF575F3137FF368E7B4734D566:8E98601257503135FD088B9D40ABD36D" box="[681,725,1790,1815]" country="Vietnam" municipality="Tam Dao Town" name="The" pageId="8" pageNumber="355">The</location>
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<typeStatus id="54FC886B57503135FD7C8A634173D362" box="[733,781,1792,1816]" pageId="8" pageNumber="355">type</typeStatus>
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specimen (a female) on which
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||
<bibRefCitation id="EFD64B3857503135FBD48B9D475DD36D" author="Asahina" box="[1141,1315,1790,1815]" pageId="8" pageNumber="355" refString="Asahina, S. (1997) Records of the Northern Vietnamese Odonata taken by the expedition members from the National Science Museum, Tokyo. 6. Platystictidae, Megapodagrionidae, Lestidae and Synlestidae. Bulletin of the National Science Museum, Series A (Zoology), 23, 107 - 113." type="journal article" year="1997">
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||
<collectorName id="26B2531F57503135FBD48B9D46AFD36D" box="[1141,1233,1790,1815]" pageId="8" pageNumber="355">Asahina</collectorName>
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||
(1997)
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||
</bibRefCitation>
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||
had based his description of
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<taxonomicName id="4C474D4A57503135FEC38A40400BD340" box="[354,629,1827,1851]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="355" phylum="Arthropoda" rank="subSpecies" species="khaosoidaoensis" subSpecies="satoi">
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<emphasis id="B933EADB57503135FEC38A40400BD340" box="[354,629,1827,1851]" italics="true" pageId="8" pageNumber="355">P. khaosoidaoensis satoi</emphasis>
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</taxonomicName>
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(which currently cannot be located) had been collected from
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<collectorName id="26B2531F57503135FA8C8A4042BDD325" pageId="8" pageNumber="355">Tam Dao. The</collectorName>
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description he provided is rather limited but mentions three characteristics: 1) the prothorax has a large broad black area on the posterior lobe, 2) the black stripe over the metapleural suture is only weakly darkened, 3) S9 is adorned by a large pale bluish spot.
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<collectorName id="26B2531F57503135FD9B8AF34009D3DD" box="[570,631,1936,1959]" pageId="8" pageNumber="355">From</collectorName>
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the accompanying drawings it is clear that the “large pale bluish spot covers about three-fifths of the dorsal side of S9 and does not extend laterally towards the ventral margin.
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<collectorName id="26B2531F57503135FACB8AD147E2D3B1" box="[1386,1436,1970,1995]" pageId="8" pageNumber="355">This</collectorName>
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||
combination of characteristics only fits the female of
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<emphasis id="B933EADB57503135FD488ABB4149D395" box="[745,823,2008,2031]" italics="true" pageId="8" pageNumber="355">P. s a t o i</emphasis>
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light form sensu
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<collectorName id="26B2531F57503135FC5D8ABB462ED395" box="[1020,1104,2006,2031]" pageId="8" pageNumber="355">van Tol</collectorName>
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amongst species known from northern
|
||
<collectingCountry id="F350765957513134FF5E8DF4431CD4CA" box="[255,354,151,176]" name="Vietnam" pageId="9" pageNumber="356">Vietnam</collectingCountry>
|
||
. Tam Dao National Park covers a large area but has been developed rapidly in the area where sampling took place in the nineties.
|
||
<collectorName id="26B2531F57513134FD888DDE4044D4AE" box="[553,570,189,212]" pageId="9" pageNumber="356">It</collectorName>
|
||
is certainly possible that
|
||
<emphasis id="B933EADB57513134FCF68DDE41DBD4AE" box="[855,933,189,212]" italics="true" pageId="9" pageNumber="356">P. s a t o i</emphasis>
|
||
light form sensu
|
||
<collectorName id="26B2531F57513134FBCD8DDE46BED4AF" box="[1132,1216,188,213]" pageId="9" pageNumber="356">van Tol</collectorName>
|
||
occurred in what is now
|
||
<collectingMunicipality id="6B9CACB357513134FF6F8D834300D48D" box="[206,382,224,247]" pageId="9" pageNumber="356">Tam Dao Town</collectingMunicipality>
|
||
</materialsCitation>
|
||
. I am convinced, based on the match between the characteristics listed by Asahina for the type of
|
||
<taxonomicName id="4C474D4A57513134FF148C6643B9D566" box="[181,455,260,284]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="356" phylum="Arthropoda" rank="subSpecies" species="khaosoidaoensis" subSpecies="satoi">
|
||
<emphasis id="B933EADB57513134FF148C6643B9D566" box="[181,455,260,284]" italics="true" pageId="9" pageNumber="356">P. khaosoidaoensis satoi</emphasis>
|
||
</taxonomicName>
|
||
and the species from Cuc Phuong, that van Tol’s identification of his specimens of Cuc Phuong as
|
||
<taxonomicName id="4C474D4A57513134FEB38C4A431DD545" box="[274,355,296,320]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="satoi">
|
||
<emphasis id="B933EADB57513134FEB38C4A431DD545" box="[274,355,296,320]" italics="true" pageId="9" pageNumber="356">P. satoi</emphasis>
|
||
</taxonomicName>
|
||
was justified.
|
||
</paragraph>
|
||
<paragraph id="8BF836C957513134FF668C2F41F1D58F" blockId="9.[151,1437,151,861]" pageId="9" pageNumber="356">
|
||
Both forms occur widely in northern Vietnam (
|
||
<bibRefCitation id="EFD64B3857513134FD768C2F41A5D51F" author="Phan" box="[727,987,332,357]" pageId="9" pageNumber="356" refString="Phan, Q. T. & Kompier, T. (2016) A study of the genus Protosticta Selys, 1855, with descriptions of four new species from Vietnam (Odonata: Platystictidae). Zootaxa, 4098 (3), 529 - 544. http: // dx. doi. org / 10.11646 / zootaxa. 4098.3.6" type="journal article" year="2016">Phan & Kompier 2016</bibRefCitation>
|
||
), but based on many observations it can be concluded that the two forms never occur side by side at the same locations. Most commonly they do not even occur in the same general areas. The dark form, for instance, does not occur in Cuc Phuong, whereas I have not been able to relocate the light form in Tam Dao, where the dark form (
|
||
<figureCitation id="137C2A4C57513134FC068CD44186D5AA" box="[935,1016,439,464]" captionStart="FIGURE 12" captionStartId="7.[151,250,1926,1948]" captionTargetBox="[151,1436,1013,1905]" captionTargetId="figure@7.[151,1436,1013,1905]" captionTargetPageId="7" captionText="FIGURE 12. Male P. nigra spec. nov, 22. V. 2016, Tam Dao National Park, Vinh Phuc Province." httpUri="https://zenodo.org/record/266147/files/figure.png" pageId="9" pageNumber="356">Fig. 12</figureCitation>
|
||
) is common. In Ba Be National Park, Bac Kan Province, both forms occur, but at very different locations.
|
||
</paragraph>
|
||
<paragraph id="8BF836C957513134FF668F6341D3D61A" blockId="9.[151,1437,151,861]" pageId="9" pageNumber="356">
|
||
In their treatment of known
|
||
<taxonomicName id="4C474D4A57513134FD8B8F6340DBD66D" box="[554,677,512,535]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="356" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B933EADB57513134FD8B8F6340DBD66D" box="[554,677,512,535]" italics="true" pageId="9" pageNumber="356">Protosticta</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F350765957513134FD5A8C9C4124D662" box="[763,858,511,536]" name="Vietnam" pageId="9" pageNumber="356">Vietnam</collectingCountry>
|
||
<bibRefCitation id="EFD64B3857513134FCC98C9C46ECD662" author="Phan" box="[872,1170,511,536]" pageId="9" pageNumber="356" refString="Phan, Q. T. & Kompier, T. (2016) A study of the genus Protosticta Selys, 1855, with descriptions of four new species from Vietnam (Odonata: Platystictidae). Zootaxa, 4098 (3), 529 - 544. http: // dx. doi. org / 10.11646 / zootaxa. 4098.3.6" type="journal article" year="2016">Phan & Kompier (2016)</bibRefCitation>
|
||
followed the original interpretation that these forms belonged to one species. But with increasing insight into the high diversity of the
|
||
<taxonomicName id="4C474D4A57513134FA268F464295D625" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="curiosa">
|
||
<emphasis id="B933EADB57513134FA268F464295D625" italics="true" pageId="9" pageNumber="356">P. curiosa</emphasis>
|
||
</taxonomicName>
|
||
group of species in
|
||
<collectingCountry id="F350765957513134FE6C8F244052D61A" box="[461,556,583,608]" name="Vietnam" pageId="9" pageNumber="356">Vietnam</collectingCountry>
|
||
a reassessment seemed necessary.
|
||
</paragraph>
|
||
<paragraph id="8BF836C957513134FF668F0F40B2D727" blockId="9.[151,1437,151,861]" pageId="9" pageNumber="356">
|
||
Although in lateral view differences between the appendages of the two forms seem minimal, close inspection in dorsolateral view does reveal consistent differences (
|
||
<figureCitation id="137C2A4C57513134FCA18FEC4122D6D2" box="[768,860,655,680]" captionStart="FIGURE 14" captionStartId="8.[151,250,1585,1607]" captionTargetBox="[340,1066,988,1507]" captionTargetId="figure@8.[340,1247,900,1563]" captionTargetPageId="8" captionText="FIGURE 14. Appendages of male P. nigra spec. nov., 22. V. 2016, Tam Dao National Park, Vinh Phuc Province, T. Kompier leg." httpUri="https://zenodo.org/record/166971/files/figure.png" pageId="9" pageNumber="356">Figs. 14</figureCitation>
|
||
,
|
||
<figureCitation id="137C2A4C57513134FCC98FF341F9D6D2" box="[872,903,656,680]" captionStart="FIGURE 17" captionStartId="11.[151,250,876,898]" captionTargetBox="[340,1044,324,781]" captionTargetId="figure@11.[340,1247,195,855]" captionTargetPageId="11" captionText="FIGURE 17. Appendages of male P. satoi, 24. VII. 2016, Cuc Phuong National Park, Ninh Binh Province, T. Kompier leg." httpUri="https://zenodo.org/record/166974/files/figure.png" pageId="9" pageNumber="356">17</figureCitation>
|
||
). Having said that, within the light form there is still some variation, both in shape of the cerci (presence of absence of dorsal tooth halfway) and pattern of the prothorax. However, they all share the same thoracic pattern (
|
||
<figureCitation id="137C2A4C57513134FCEA8FB441ADD68A" box="[843,979,727,752]" captionStart-0="FIGURE 18" captionStart-1="FIGURE 19" captionStartId-0="11.[151,250,1662,1684]" captionStartId-1="12.[151,250,1711,1733]" captionTargetBox-0="[371,1218,976,1611]" captionTargetBox-1="[384,1203,1076,1690]" captionTargetId-0="figure@11.[371,1219,976,1612]" captionTargetId-1="figure@12.[384,1203,1076,1690]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="FIGURE 18. Thorax of male P. satoi, 24. VII. 2016, Cuc Phuong National Park, Ninh Binh Province, T. Kompier leg." captionText-1="FIGURE 19. Thorax of male P. satoi, 01. VIII. 2016, Xuan Son National Park, Phu Tho Province, T. Kompier leg." httpUri-0="https://zenodo.org/record/266150/files/figure.png" httpUri-1="https://zenodo.org/record/166976/files/figure.png" pageId="9" pageNumber="356">Figs. 18–19</figureCitation>
|
||
), and the general structure of the cerci is also similar. These differences seem too weak to warrant further division. In
|
||
<tableCitation id="C6C5037257513134FC598F9F4635D76E" box="[1016,1099,764,789]" captionStart="TABLE 1" captionStartId="10.[151,239,332,355]" captionTargetBox="[159,1394,383,631]" captionTargetPageId="10" captionText="TABLE 1. Differences between the cerci of P. satoi and P. nigra spec. nov." httpUri="http://table.plazi.org/id/DF38664157523137FF368C2F41C6D519" pageId="9" pageNumber="356" tableUuid="DF38664157523137FF368C2F41C6D519">Table 1</tableCitation>
|
||
the main differences between the cerci of
|
||
<emphasis id="B933EADB57513134FEBE8E424311D74D" box="[287,367,800,824]" italics="true" pageId="9" pageNumber="356">P. s a t o i</emphasis>
|
||
and
|
||
<emphasis id="B933EADB57513134FE0A8E42407CD742" box="[427,514,800,824]" italics="true" pageId="9" pageNumber="356">P. ni g r a</emphasis>
|
||
spec. nov. are summarized. Obviously the differences in patterning of the thorax are evident and do not warrant further elaboration.
|
||
</paragraph>
|
||
<caption id="DF38664157513134FF368B8D406DD359" httpUri="https://zenodo.org/record/266149/files/figure.png" pageId="9" pageNumber="356" targetBox="[151,1436,913,1752]" targetPageId="9">
|
||
<paragraph id="8BF836C957513134FF368B8D406DD359" blockId="9.[151,1435,1774,1827]" pageId="9" pageNumber="356">
|
||
<emphasis id="B933EADB57513134FF368B8D4361D379" bold="true" box="[151,287,1774,1796]" pageId="9" pageNumber="356">FIGURE 15.</emphasis>
|
||
Male
|
||
<taxonomicName id="4C474D4A57513134FEFF8B8C43D9D37E" box="[350,423,1775,1796]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="satoi">
|
||
<emphasis id="B933EADB57513134FEFF8B8C43D9D37E" box="[350,423,1775,1796]" italics="true" pageId="9" pageNumber="356">P. satoi</emphasis>
|
||
</taxonomicName>
|
||
, 24.VII.2016, Cuc Phuong National Park, Ninh Binh Province. Note yellowish white prothorax with black marking on posterior lobe only.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF836C957513137FF668A284734D566" blockId="9.[151,1437,1866,2035]" lastBlockId="10.[151,1436,151,285]" lastPageId="10" lastPageNumber="357" pageId="9" pageNumber="356">
|
||
Van
|
||
<bibRefCitation id="EFD64B3857513134FF5A8A29430DD319" author="Tol" box="[251,371,1866,1891]" pageId="9" pageNumber="356" refString="van Tol, J. (2008) Notes on some species of the genus Protosticta from Vietnam (Odonata, Platystictidae). Zoologische Mededelingen Leiden, 82, 217 - 234." type="journal article" year="2008">Tol (2008)</bibRefCitation>
|
||
suggested that dark form
|
||
<emphasis id="B933EADB57513134FD3F8A284090D318" box="[670,750,1867,1890]" italics="true" pageId="9" pageNumber="356">P. s a t oi</emphasis>
|
||
(=
|
||
<taxonomicName id="4C474D4A57513134FCBA8A28410AD318" box="[795,884,1867,1890]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="nigra" status="sp. nov.">
|
||
<emphasis id="B933EADB57513134FCBA8A28410AD318" box="[795,884,1867,1890]" italics="true" pageId="9" pageNumber="356">P. nigra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20057A057513134FCDC8A28418ED319" box="[893,1008,1867,1891]" pageId="9" pageNumber="356" rank="species">spec. nov.</taxonomicNameLabel>
|
||
) and dark form
|
||
<taxonomicName id="4C474D4A57513134FB088A2842ACD3FD" authority="Wilson, 1997" authorityName="Wilson" authorityYear="1997" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="beaumonti">
|
||
<emphasis id="B933EADB57513134FB088A284742D318" box="[1193,1340,1866,1890]" italics="true" pageId="9" pageNumber="356">P. beaumonti</emphasis>
|
||
Wilson, 1997
|
||
</taxonomicName>
|
||
, may in fact be the same taxon.
|
||
<bibRefCitation id="EFD64B3857513134FD9A8A0D4133D3FD" author="Phan" box="[571,845,1902,1927]" pageId="9" pageNumber="356" refString="Phan, Q. T. & Kompier, T. (2016) A study of the genus Protosticta Selys, 1855, with descriptions of four new species from Vietnam (Odonata: Platystictidae). Zootaxa, 4098 (3), 529 - 544. http: // dx. doi. org / 10.11646 / zootaxa. 4098.3.6" type="journal article" year="2016">Phan & Kompier (2016)</bibRefCitation>
|
||
published first records of
|
||
<taxonomicName id="4C474D4A57513134FBD48A0C4778D3FC" box="[1141,1286,1902,1926]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="beaumonti">
|
||
<emphasis id="B933EADB57513134FBD48A0C4778D3FC" box="[1141,1286,1902,1926]" italics="true" pageId="9" pageNumber="356">P. beaumonti</emphasis>
|
||
</taxonomicName>
|
||
for
|
||
<collectingCountry id="F350765957513134FA978A0D47E7D3FD" box="[1334,1433,1902,1927]" name="Vietnam" pageId="9" pageNumber="356">Vietnam</collectingCountry>
|
||
. Although morphological differences appear to be slight (prominence of teeth on cerci), they pointed to two differences in patterning between
|
||
<taxonomicName id="4C474D4A57513134FDB58AD4401DD3B4" box="[532,611,1975,1998]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="satoi">
|
||
<emphasis id="B933EADB57513134FDB58AD4401DD3B4" box="[532,611,1975,1998]" italics="true" pageId="9" pageNumber="356">P. satoi</emphasis>
|
||
</taxonomicName>
|
||
dark form (=
|
||
<taxonomicName id="4C474D4A57513134FCA58AD44123D3B4" box="[772,861,1975,1998]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="nigra" status="sp. nov.">
|
||
<emphasis id="B933EADB57513134FCA58AD44123D3B4" box="[772,861,1975,1998]" italics="true" pageId="9" pageNumber="356">P. nigra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20057A057513134FCC58AD441ABD3B5" box="[868,981,1975,1999]" pageId="9" pageNumber="356" rank="species">spec. nov.</taxonomicNameLabel>
|
||
) and
|
||
<taxonomicName id="4C474D4A57513134FBB28AD446DAD3B4" box="[1043,1188,1974,1998]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="beaumonti">
|
||
<emphasis id="B933EADB57513134FBB28AD446DAD3B4" box="[1043,1188,1974,1998]" italics="true" pageId="9" pageNumber="356">P. beaumonti</emphasis>
|
||
</taxonomicName>
|
||
. Of these the thoracic pattern with a large white area on the posterior half of the metepisternum is most characteristic. The difference in patterning on S9 for the females in fact does not provide a good character to separate
|
||
<taxonomicName id="4C474D4A57523137FBC88DFA4685D4CA" box="[1129,1275,152,176]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="357" phylum="Arthropoda" rank="species" species="beaumonti">
|
||
<emphasis id="B933EADB57523137FBC88DFA4685D4CA" box="[1129,1275,152,176]" italics="true" pageId="10" pageNumber="357">P. beaumonti</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C474D4A57523137FAE28DFA47E5D4D5" box="[1347,1435,152,176]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="357" phylum="Arthropoda" rank="species" species="nigra" status="sp. nov.">
|
||
<emphasis id="B933EADB57523137FAE28DFA47E5D4D5" box="[1347,1435,152,176]" italics="true" pageId="10" pageNumber="357">P. nigra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20057A057523137FF368DDE4377D4AE" box="[151,265,189,212]" pageId="10" pageNumber="357" rank="species">spec. nov.</taxonomicNameLabel>
|
||
, which also has dark S9. In view of van Tol’s suggestion regarding the conspecificity of these two dark species, it is advisable to support the slight differences in morphology and thoracic patterning with a detailed molecular study in the future. For now we treat
|
||
<taxonomicName id="4C474D4A57523137FD0F8C664141D566" box="[686,831,260,284]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="357" phylum="Arthropoda" rank="species" species="beaumonti">
|
||
<emphasis id="B933EADB57523137FD0F8C664141D566" box="[686,831,260,284]" italics="true" pageId="10" pageNumber="357">P. beaumonti</emphasis>
|
||
</taxonomicName>
|
||
dark form as separate from
|
||
<emphasis id="B933EADB57523137FBDA8C6646AED566" box="[1147,1232,261,284]" italics="true" pageId="10" pageNumber="357">P. n i gr a</emphasis>
|
||
spec. nov.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |