treatments-xml/data/03/EE/87/03EE87AEFFA3FF8FFD33FE678C840062.xml
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<document ID-DOI="10.5281/zenodo.3245327" ID-GBIF-Dataset="9bf3a9f5-cd36-4be6-bba9-49607815bcd0" ID-Zenodo-Dep="3245327" approvalRequired="3" approvalRequired_for_bibRefs="3" checkinTime="1559900965784" checkinUser="jeremy" docAuthor="Holtz, T. R." docDate="2001" docId="03EE87AEFFA3FF8FFD33FE678C840062" docLanguage="en" docName="Holtz2001TyrannosauridaeGS.pdf.imf" docOrigin="Mesozoic Vertebrate Life, Bloomington: Indiana University Press" docStyle="DocumentStyle{}" docTitle="Daspletosaurus torosus" docType="treatment" docVersion="17" lastPageNumber="70" masterDocId="FFD7FFD6FFA5FF89FFE1FFFB8A660428" masterDocTitle="The phylogeny and taxonomy of the Tyrannosauridae" masterLastPageNumber="83" masterPageNumber="64" pageNumber="70" updateTime="1645527105248" updateUser="jeremy">
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<mods:titleInfo>
<mods:title>The phylogeny and taxonomy of the Tyrannosauridae</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Holtz, T. R.</mods:namePart>
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<mods:dateIssued>2001</mods:dateIssued>
<mods:dateOther type="pubDate">2001-12-31</mods:dateOther>
<mods:publisher>Indiana University Press</mods:publisher>
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<mods:roleTerm>Editor</mods:roleTerm>
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<mods:namePart>Tanke D. H.</mods:namePart>
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<mods:roleTerm>Editor</mods:roleTerm>
</mods:role>
<mods:namePart>Carpenter K.</mods:namePart>
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<mods:title>Mesozoic Vertebrate Life</mods:title>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5500186" ID-GBIF-Taxon="157533275" ID-Zenodo-Dep="5500186" LSID="urn:lsid:plazi:treatment:03EE87AEFFA3FF8FFD33FE678C840062" httpUri="http://treatment.plazi.org/id/03EE87AEFFA3FF8FFD33FE678C840062" lastPageNumber="70" pageId="6" pageNumber="70">
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<taxonomicName authority="Russell, 1970" box="[722,1132,412,448]" class="Reptilia" family="Tyrannosauridae" genus="Daspletosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="6" pageNumber="70" phylum="Chordata" rank="species" species="torosus">
<emphasis box="[722,1132,412,448]" italics="true" pageId="6" pageNumber="70">Daspletosaurus torosus</emphasis>
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,
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the Two Medicine tyrannosaurine, and
<taxonomicName authorityName="Osborn" authorityYear="1905" box="[648,896,462,497]" class="Reptilia" family="Tyrannosauridae" genus="Tyrannosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="6" pageNumber="70" phylum="Chordata" rank="genus">
<emphasis box="[648,896,462,497]" italics="true" pageId="6" pageNumber="70">Tyrannosaurus</emphasis>
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share several derived features lacking in other tyrannosaurids. Many of the synapomorphies of this clade suggest a more forcefully built and muscular skull and neck than in other tyrannosaurids, In some of the most parsimonious trees
<taxonomicName box="[1445,1842,614,650]" class="Reptilia" family="Tyrannosauridae" genus="Daspletosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="6" pageNumber="70" phylum="Chordata" rank="species" species="torosus">
<emphasis box="[1445,1842,614,650]" pageId="6" pageNumber="70">Daspletosaurus torosus</emphasis>
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and a presently unnamed tyrannosaurine from the upper Two Medicine Formation were united by a potential synapomorphy: 53.1 Lacrimal horn directly dorsal to descending ramus. However, in other trees
<emphasis box="[646,904,813,848]" italics="true" pageId="6" pageNumber="70">Daspletosaurzs</emphasis>
is closer to
<taxonomicName authorityName="Osborn" authorityYear="1905" box="[1126,1376,814,849]" class="Reptilia" family="Tyrannosauridae" genus="Tyrannosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="6" pageNumber="70" phylum="Chordata" rank="genus">
<emphasis box="[1126,1376,814,849]" italics="true" pageId="6" pageNumber="70">Tyrannosaurus</emphasis>
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than to the Two Medicine form, and in still others the Two Medicine form is closer to
<taxonomicName authorityName="Osborn" authorityYear="1905" class="Reptilia" family="Tyrannosauridae" genus="Tyrannosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="6" pageNumber="70" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="6" pageNumber="70">Tyrannosaurus</emphasis>
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.
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<paragraph blockId="6.[645,1850,159,2654]" box="[721,1391,963,1000]" pageId="6" pageNumber="70">
<taxonomicName authority="Russell 1970" authorityName="Russell" authorityYear="1970" box="[721,1386,963,1000]" class="Reptilia" family="Tyrannosauridae" genus="Daspletosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="6" pageNumber="70" phylum="Chordata" rank="species" species="torosus">
<emphasis box="[721,1131,963,1000]" italics="true" pageId="6" pageNumber="70">Daspletosaurus torosus</emphasis>
<bibRefCitation author="Russell" box="[1150,1386,963,1000]" firstAuthor="Russell" journalOrPublisher="National Musewm Natural Sciences Publications in Palaeontology" pageId="6" pageNumber="70" pagination="1 - - 34" part="1" refId="ref6245" refString="Russell, D. A. 1970. Tyrannosaurs from the Late Cretaceous of western Canada. National Musewm Natural Sciences Publications in Palaeontology 1: 1 - - 34." title="Tyrannosaurs from the Late Cretaceous of western Canada" type="journal volume" year="1970">Russell 1970</bibRefCitation>
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:
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<subSubSection pageId="6" pageNumber="70" type="discussion">
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From the Dinosaur Park Formation of Alberta, this form is more robustly and powerfully constructed than the sympatric
<taxonomicName authorityName="Lambe" authorityYear="1914" box="[1132,1494,1064,1101]" class="Reptilia" family="Tyrannosauridae" genus="Gorgosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="6" pageNumber="83" phylum="Chordata" rank="species" species="libratus">
<emphasis box="[1132,1494,1064,1101]" italics="true" pageId="6" pageNumber="70">Gorgosaurus libratus</emphasis>
</taxonomicName>
(
<bibRefCitation author="Russell" box="[1521,1746,1062,1098]" firstAuthor="Russell" journalOrPublisher="National Musewm Natural Sciences Publications in Palaeontology" pageId="6" pageNumber="70" pagination="1 - - 34" part="1" refId="ref6245" refString="Russell, D. A. 1970. Tyrannosaurs from the Late Cretaceous of western Canada. National Musewm Natural Sciences Publications in Palaeontology 1: 1 - - 34." title="Tyrannosaurs from the Late Cretaceous of western Canada" type="journal volume" year="1970">Russell 1970</bibRefCitation>
).
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