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<document id="F73D7ED0DD2A79E98935D2DDF6028B61" ID-DOI="10.21248/contrib.entomol.69.1.033-070" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742766" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638269617685" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2019" docId="03EE87A5FFFC3B7AFF3AF934754B9D9E" docLanguage="en" docName="ContribEntomol.69.1.33-70.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 69 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Drusilla breviuter Assing 2019, spec. nov." docType="treatment" docUuid="43EDA558-1DBF-4861-B73F-F292EF6169C5" docUuidSource="ZooBank" docVersion="3" lastPageNumber="62" masterDocId="FFD7FFDDFFE03B67FFC3FFDE77509866" masterDocTitle="On the Lomechusini fauna of the Palaearctic and Oriental regions. XXVI. New species, a new synonymy, and additional records (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="70" masterPageNumber="33" pageNumber="61" updateTime="1699437235520" updateUser="plazi">
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<mods:title id="517E12CB922D0BF4335DAC772E2B4553">On the Lomechusini fauna of the Palaearctic and Oriental regions. XXVI. New species, a new synonymy, and additional records (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:name id="AD5BFC8CA5BA7D0C01B0D2D75305F2CE" type="personal">
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<mods:namePart id="B1C9FF91AE1B10C36D2E344D27D8E025">Assing, Volker</mods:namePart>
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<mods:title id="F744D26628C0EF6137E2DCA4F31213F7">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:part id="08B78B8481E23DFA749BDBAE9A48A325">
<mods:date id="CF700B31C4FB0B790AF406D3D5E02A33">2019</mods:date>
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<mods:number id="EED134BE677D4A2555B98F469021C052">2019-06-24</mods:number>
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<mods:identifier id="492A36FE640ADF612DBB79BBD580E3A9" type="ISSN">0005-805X</mods:identifier>
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<treatment id="03EE87A5FFFC3B7AFF3AF934754B9D9E" LSID="urn:lsid:zoobank.org:act:43EDA558-1DBF-4861-B73F-F292EF6169C5" httpUri="http://treatment.plazi.org/id/03EE87A5FFFC3B7AFF3AF934754B9D9E" lastPageId="29" lastPageNumber="62" pageId="28" pageNumber="61">
<subSubSection id="C35D6538FFFC3B7BFF3AF93475AB9F43" pageId="28" pageNumber="61" type="nomenclature">
<paragraph id="8BF836B3FFFC3B7BFF3AF93475489F60" blockId="28.[249,763,1770,1861]" box="[249,536,1770,1798]" pageId="28" pageNumber="61">
<heading id="D0B081DFFFFC3B7BFF3AF93475489F60" box="[249,536,1770,1798]" fontSize="12" level="2" pageId="28" pageNumber="61" reason="2">
<taxonomicName id="4C474D30FFFC3B7BFF3AF93476F89F60" ID-CoL="8XZV2" authority="Assing, 2019" authorityName="Assing" authorityYear="2019" box="[249,424,1770,1798]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="61" phylum="Arthropoda" rank="species" species="breviuter" status="sp. nov.">
<emphasis id="B933EAA1FFFC3B7BFF3AF93476F89F60" box="[249,424,1770,1798]" italics="true" pageId="28" pageNumber="61">Drusilla breviuter</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20057DAFFFC3B7BFE6CF93475489F60" box="[431,536,1770,1798]" pageId="28" pageNumber="61" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BF836B3FFFC3B7BFF3AF8CF75AB9F43" blockId="28.[249,763,1770,1861]" box="[249,763,1809,1829]" pageId="28" pageNumber="61">
<uri id="FFD63AB1FFFC3B7BFF3AF8CF75AB9F43" box="[249,763,1809,1829]" pageId="28" pageNumber="61">
urn:lsid:zoobank.org: act:
<uuid id="FFE10C66FFFC3B7BFE6BF8CF75AB9F43" box="[424,763,1809,1829]" pageId="28" pageNumber="61">43EDA558-1DBF-4861-B73F-F292EF6169C5</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C35D6538FFFC3B7BFF3AF8EF76F29F23" box="[249,418,1841,1861]" pageId="28" pageNumber="61" type="description">
<paragraph id="8BF836B3FFFC3B7BFF3AF8EF76F29F23" blockId="28.[249,763,1770,1861]" box="[249,418,1841,1861]" pageId="28" pageNumber="61">(Figs 5963, 148152)</paragraph>
</subSubSection>
<subSubSection id="C35D6538FFFC3B7BFF17F8BE76D5907E" pageId="28" pageNumber="61" type="materials_examined">
<paragraph id="8BF836B3FFFC3B7BFF17F8BE76D5907E" blockId="28.[212,823,1887,2072]" pageId="28" pageNumber="61">
<emphasis id="B933EAA1FFFC3B7BFF17F8BE763D9F11" bold="true" box="[212,365,1887,1912]" pageId="28" pageNumber="61">Type material</emphasis>
:
<materialsCitation id="3B2F3CEEFFFC3B7BFEB9F8BE76319F9E" collectingDate="2018-03-28" collectionCode="V" collectorName="Shavrin" country="Philippines" latitude="7.2833333" location="Boss Carreon Function Hall" longLatPrecision="1303" longitude="125.683334" municipality="Mindanao" pageId="28" pageNumber="61" specimenCount="2" stateProvince="Davao" typeStatus="holotype">
<typeStatus id="54FC8811FFFC3B7BFEB9F8BE76B59F1E" box="[378,485,1888,1912]" pageId="28" pageNumber="61" type="holotype">Holotype</typeStatus>
: “
<collectingCountry id="F3507623FFFC3B7BFDD3F8BE75E29F1E" box="[528,690,1888,1912]" name="Philippines" pageId="28" pageNumber="61">PHILIPPINES</collectingCountry>
:
<collectingMunicipality id="6B9CACC9FFFC3B7BFD79F8BE74629F1E" box="[698,818,1888,1912]" pageId="28" pageNumber="61">Mindanao</collectingMunicipality>
,
<collectingRegion id="4983F851FFFC3B7BFF17F85E764C9FF1" box="[212,284,1920,1943]" country="Philippines" name="Davao" pageId="28" pageNumber="61">Davao</collectingRegion>
,
<location id="8E986068FFFC3B7BFEEAF85E75349FFE" LSID="urn:lsid:plazi:treatment:03EE87A5FFFC3B7AFF3AF934754B9D9E:8E986068FFFC3B7BFEEAF85E75349FFE" box="[297,612,1920,1944]" country="Philippines" latitude="7.2833333" longLatPrecision="1303" longitude="125.683334" municipality="Mindanao" name="Boss Carreon Function Hall" pageId="28" pageNumber="61" stateProvince="Davao">Boss Carreon Function Hall</location>
,
<geoCoordinate id="EE735074FFFC3B7BFDB1F85E75ED9FFE" box="[626,701,1920,1944]" degrees="7" direction="north" minutes="17" orientation="latitude" pageId="28" pageNumber="61" precision="925" value="7.2833333">7°17'N</geoCoordinate>
,
<geoCoordinate id="EE735074FFFC3B7BFD08F85E747A9FFE" box="[715,810,1920,1944]" degrees="125" direction="east" minutes="41" orientation="longitude" pageId="28" pageNumber="61" precision="925" value="125.683334">125°41'E</geoCoordinate>
, slope near river, litter sifted,
<date id="FFF91073FFFC3B7BFDE0F87E75CC9FDE" box="[547,668,1952,1976]" pageId="28" pageNumber="61" value="2018-03-28">
<collectingDate id="EFBDE99BFFFC3B7BFDE0F87E75CC9FDE" box="[547,668,1952,1976]" pageId="28" pageNumber="61" value="2018-03-28">28.III.2018</collectingDate>
</date>
, leg.
<collectorName id="26B25365FFFC3B7BFD1EF87E74679FDE" box="[733,823,1952,1976]" pageId="28" pageNumber="61">Shavrin</collectorName>
/
<typeStatus id="54FC8811FFFC3B7BFF24F81E760F9FBE" box="[231,351,1984,2008]" pageId="28" pageNumber="61" type="holotype">Holotypus</typeStatus>
<taxonomicName id="4C474D30FFFC3B7BFE41F81E75149FBE" authority="Assing, 2019" authorityName="Assing" authorityYear="2019" box="[386,580,1984,2008]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="61" phylum="Arthropoda" rank="species" species="breviuter" status="sp. nov.">
<emphasis id="B933EAA1FFFC3B7BFE41F81E75149FBE" box="[386,580,1984,2008]" italics="true" pageId="28" pageNumber="61">Drusilla breviuter</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20057DAFFFC3B7BFD8CF81E75D99FB1" box="[591,649,1984,2007]" pageId="28" pageNumber="61" rank="species">sp. n.</taxonomicNameLabel>
, det.
<collectionCode id="ED56AE76FFFC3B7BFD0FF81E758F9FB1" box="[716,735,1984,2007]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="28" pageNumber="61" type="Museum">V</collectionCode>
. Assing 2018” (cAss)
</materialsCitation>
.
<typeStatus id="54FC8811FFFC3B7BFEA9F83E76869F9E" box="[362,470,2016,2040]" pageId="28" pageNumber="61" type="paratype">Paratypes</typeStatus>
: 1 [teneral], 1: same data as holotype (cAss).
</paragraph>
</subSubSection>
<subSubSection id="C35D6538FFFC3B7BFC95FEDE74B4991E" pageId="28" pageNumber="61" type="etymology">
<paragraph id="8BF836B3FFFC3B7BFC95FEDE74B4991E" blockId="28.[854,1465,256,376]" pageId="28" pageNumber="61">
<emphasis id="B933EAA1FFFC3B7BFC95FEDE749D997E" bold="true" box="[854,973,256,280]" pageId="28" pageNumber="61">Etymology</emphasis>
: The specific epithet is a noun in apposition composed of the Latin adjective brevis (short) and the Latin noun uter (hose). It alludes to the small and short spermatheca.
</paragraph>
</subSubSection>
<subSubSection id="C35D6538FFFC3B7AFC95FE4175E49D5E" lastPageId="29" lastPageNumber="62" pageId="28" pageNumber="61" type="description">
<paragraph id="8BF836B3FFFC3B7BFC95FE41730C9ADE" blockId="28.[854,1466,415,2072]" pageId="28" pageNumber="61">
<emphasis id="B933EAA1FFFC3B7BFC95FE41748899D1" bold="true" box="[854,984,415,439]" pageId="28" pageNumber="61">Description</emphasis>
: Body length 4.0
<quantity id="4CBF9B56FFFC3B7BFB6FFE7E73AF99D1" box="[1196,1279,416,440]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="28" pageNumber="61" unit="mm" value="4.7">4.7 mm</quantity>
; length of forebody 1.82.0 mm. Habitus as in Fig. 59. Coloration: head black; pronotum and elytra black with slight bronze hue; abdomen black with segments IIIIV and the narrow posterior margin of tergite V reddish; legs uniformly yellow; antennae blackish-brown with antennomeres III dark-reddish and the apex of antennomere XI reddishyellow; maxillary palpi yellow to pale-reddish with the apical palpomere yellow.
</paragraph>
<paragraph id="8BF836B3FFFC3B7BFC95FD1E73A19C71" blockId="28.[854,1466,415,2072]" pageId="28" pageNumber="61">
Head (Figs 6061) transverse, approximately 1.15 times as broad as long, without sexual dimorphism; punctation very fine and moderately dense; interstices without microreticulation. Eyes large and strongly convex, approximately twice as long as distance from posterior margin of eye to posterior constriction in dorsal view. Antenna (Fig. 62)
<quantity id="4CBF9B56FFFC3B7BFBDAFC5E73C89BF1" box="[1049,1176,896,920]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="28" pageNumber="61" unit="mm" value="1.55" valueMax="1.6" valueMin="1.5">1.51.6 mm</quantity>
long and distinctly incrassate apically; antennomeres IV weakly transverse, VX of increasing width and increasingly transverse, X approximately 1.5 times as broad as long, and XI slightly longer than the combined length of IX and X.
</paragraph>
<paragraph id="8BF836B3FFFC3B7BFC95FBFE72769D5E" blockId="28.[854,1466,415,2072]" pageId="28" pageNumber="61">Pronotum (Figs 6061) with moderate sexual dimorphism, weakly convex in cross-section, moderately transverse, approximately 1.1 times as broad as long and approximately as broad as head, broadest near anterior angles, moderately tapering posteriad; along midline with very fine sulcus posteriorly terminating in a small impression; lateral margins straight in dorsal view; posterior angles obtusely marked; punctation dense and distinct; interstices without microsculpture.</paragraph>
<paragraph id="8BF836B3FFFC3B7BFC95FA9E74269D91" blockId="28.[854,1466,415,2072]" pageId="28" pageNumber="61">Elytra (Figs 6061) approximately 0.85 times as long as pronotum; punctation similar to that of pronotum, but somewhat denser; interstices without microsculpture. Hind wings fully developed. Metatarsomere I relatively short, barely as long as the combined length of III and IV.</paragraph>
<paragraph id="8BF836B3FFFC3B7BFC95F9DE74819EDE" blockId="28.[854,1466,415,2072]" pageId="28" pageNumber="61">Abdomen (Fig. 63) broad and relatively short, approximately as broad as elytra; tergite IIIVII with few punctures at posterior margin, otherwise nearly impunctate; all tergites with shallow and fine transverse microsculpture; posterior margin of tergite VII with palisade fringe.</paragraph>
<paragraph id="8BF836B3FFFC3B7AFCA8F91E7665995E" blockId="28.[854,1466,415,2072]" lastBlockId="29.[174,785,256,632]" lastPageId="29" lastPageNumber="62" pageId="28" pageNumber="61">
: pronotum (Fig. 60) with rather extensive, but not very deep median impression, posteriorly with a granulose elevation on either side, punctation slightly granulose; tergite VIII (
<figureCitation id="137C2A36FFFC3B7BFBA4F8FE73EF9F5E" box="[1127,1215,1824,1848]" captionStart="Figs 145165" captionStartId="13.[174,214,2001,2023]" captionTargetBox="[176,1415,260,1989]" captionTargetId="figure-118@13.[173,1421,255,1995]" captionTargetPageId="13" captionText="Figs 145165:Drusilla spiniventris(145147), D. breviuter(148152), Pheidologitonetes biplicatus (153154), Zyras conlectus (155 157), Z. hetzeli (158159), Z. reductus(160162), andZ. shavrini(163165): male tergite VIII (145, 148); median lobe of aedeagus in lateral and in ventral view (146147, 149150, 153156, 158161, 163164); female tergite VIII (151); spermatheca (152); paramere (157, 162, 165). Scale bars: 145151, 153165: 0.2 mm; 152: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742777" httpUri="https://zenodo.org/record/5742777/files/figure.png" pageId="28" pageNumber="61">Fig. 148</figureCitation>
) strongly transverse, posterior margin with a pronounced tooth on either side, in the middle between these teeth with a broadly truncate projection, in anterior portion with few scattered gland openings; sternite VIII strongly transverse, longer than tergite VIII, with strongly convex posterior margin, in anterior portion without gland openings; median lobe of aedeagus
<quantity id="4CBF9B56FFFC3B7BFC03F7DE734E9071" box="[960,1054,2048,2072]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="28" pageNumber="61" unit="mm" value="0.43">0.43 mm</quantity>
long and shaped as in
<figureCitation id="137C2A36FFFC3B7BFAE2F7DE72FF907E" box="[1313,1455,2048,2072]" captionStart="Figs 145165" captionStartId="13.[174,214,2001,2023]" captionTargetBox="[176,1415,260,1989]" captionTargetId="figure-118@13.[173,1421,255,1995]" captionTargetPageId="13" captionText="Figs 145165:Drusilla spiniventris(145147), D. breviuter(148152), Pheidologitonetes biplicatus (153154), Zyras conlectus (155 157), Z. hetzeli (158159), Z. reductus(160162), andZ. shavrini(163165): male tergite VIII (145, 148); median lobe of aedeagus in lateral and in ventral view (146147, 149150, 153156, 158161, 163164); female tergite VIII (151); spermatheca (152); paramere (157, 162, 165). Scale bars: 145151, 153165: 0.2 mm; 152: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742777" httpUri="https://zenodo.org/record/5742777/files/figure.png" pageId="28" pageNumber="61">Figs 149150</figureCitation>
; ventral process broad, apically broadly triangular in ventral view.
</paragraph>
<paragraph id="8BF836B3FFFD3B7AFF00FE9E76DE9A1E" blockId="29.[174,785,256,632]" pageId="29" pageNumber="62">
: pronotum (Fig. 61) only with shallow oblong impression along middle in posterior portion, without elevation, andwithnon-granulosepunctation; tergite VIII (
<figureCitation id="137C2A36FFFD3B7AFD68FE5E745399FE" box="[683,771,384,408]" captionStart="Figs 145165" captionStartId="13.[174,214,2001,2023]" captionTargetBox="[176,1415,260,1989]" captionTargetId="figure-118@13.[173,1421,255,1995]" captionTargetPageId="13" captionText="Figs 145165:Drusilla spiniventris(145147), D. breviuter(148152), Pheidologitonetes biplicatus (153154), Zyras conlectus (155 157), Z. hetzeli (158159), Z. reductus(160162), andZ. shavrini(163165): male tergite VIII (145, 148); median lobe of aedeagus in lateral and in ventral view (146147, 149150, 153156, 158161, 163164); female tergite VIII (151); spermatheca (152); paramere (157, 162, 165). Scale bars: 145151, 153165: 0.2 mm; 152: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742777" httpUri="https://zenodo.org/record/5742777/files/figure.png" pageId="29" pageNumber="62">Fig. 151</figureCitation>
) strongly transverse, posterior margin with broad and shallow excision laterally delimited by a triangular projection on either side, anterior half with scattered gland openings; sternite VIII strongly transverse, posterior margin convex, with long and thin marginal setae in the middle; spermatheca (
<figureCitation id="137C2A36FFFD3B7AFE06FD9E754C9A3E" box="[453,540,576,600]" captionStart="Figs 145165" captionStartId="13.[174,214,2001,2023]" captionTargetBox="[176,1415,260,1989]" captionTargetId="figure-118@13.[173,1421,255,1995]" captionTargetPageId="13" captionText="Figs 145165:Drusilla spiniventris(145147), D. breviuter(148152), Pheidologitonetes biplicatus (153154), Zyras conlectus (155 157), Z. hetzeli (158159), Z. reductus(160162), andZ. shavrini(163165): male tergite VIII (145, 148); median lobe of aedeagus in lateral and in ventral view (146147, 149150, 153156, 158161, 163164); female tergite VIII (151); spermatheca (152); paramere (157, 162, 165). Scale bars: 145151, 153165: 0.2 mm; 152: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742777" httpUri="https://zenodo.org/record/5742777/files/figure.png" pageId="29" pageNumber="62">Fig. 152</figureCitation>
) comma-shaped, very small and very short.
</paragraph>
<paragraph id="8BF836B3FFFD3B7AFF6DFD4175E49D5E" blockId="29.[174,785,671,1336]" pageId="29" pageNumber="62">
<emphasis id="B933EAA1FFFD3B7AFF6DFD41762D9ADE" bold="true" box="[174,381,671,696]" pageId="29" pageNumber="62">Comparative notes</emphasis>
: Based on the primary sexual characters,
<taxonomicName id="4C474D30FFFD3B7AFF22FD1E760D9ABE" authorityName="Assing" authorityYear="2019" box="[225,349,704,728]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="62" phylum="Arthropoda" rank="species" species="breviuter">
<emphasis id="B933EAA1FFFD3B7AFF22FD1E760D9ABE" box="[225,349,704,728]" italics="true" pageId="29" pageNumber="62">D. breviuter</emphasis>
</taxonomicName>
is closely allied to
<taxonomicName id="4C474D30FFFD3B7AFDE0FD1E77B19A9E" authority="PACE, 2000" authorityName="PACE" authorityYear="2000" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="62" phylum="Arthropoda" rank="species" species="endopilosella">
<emphasis id="B933EAA1FFFD3B7AFDE0FD1E75999ABE" box="[547,713,704,728]" italics="true" pageId="29" pageNumber="62">D. endopilosella</emphasis>
<bibRefCitation id="EFD64B42FFFD3B7AFD0DFD1E77B19A9E" author="PACE, R." pageId="29" pageNumber="62" pagination="39 - 86" refId="ref30339" refString="PACE, R. 2000: Aleocharinae della Thailandia (Coleoptera, Staphylinidae). - Bollettino del Museo Regionale di Scienze Naturali, Torino 17 (1): 39 - 86." type="journal article" year="2000">PACE, 2000</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F3507623FFFD3B7AFEEDFD3E76DE9A9E" box="[302,398,736,760]" name="Thailand" pageId="29" pageNumber="62">Thailand</collectingCountry>
and
<taxonomicName id="4C474D30FFFD3B7AFE0DFD3E74409A9E" authority="PACE, 2008" authorityName="PACE" authorityYear="2008" box="[462,784,736,760]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="62" phylum="Arthropoda" rank="species" species="semimonticola">
<emphasis id="B933EAA1FFFD3B7AFE0DFD3E75D39A9E" box="[462,643,736,760]" italics="true" pageId="29" pageNumber="62">D. semimonticola</emphasis>
<bibRefCitation id="EFD64B42FFFD3B7AFD4CFD3E74409A9E" author="PACE, R." box="[655,784,736,760]" pageId="29" pageNumber="62" pagination="107 - 156" refId="ref30451" refString="PACE, R. 2008: Generi e specie della tribu Lomechusini del Borneo (Coleoptera, Staphylinidae). - Revue Suisse de Zoologie 115 (1): 107 - 156." type="journal article" year="2008">PACE, 2008</bibRefCitation>
</taxonomicName>
(male unknown) from Borneo. It is readily distinguished from both of them by the coloration of the abdomen (completely reddish in the other species) and by the shape of the spermatheca. For illustrations of
<taxonomicName id="4C474D30FFFD3B7AFD73FCBE76529BFE" authorityName="PACE" authorityYear="2000" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="62" phylum="Arthropoda" rank="species" species="endopilosella">
<emphasis id="B933EAA1FFFD3B7AFD73FCBE76529BFE" italics="true" pageId="29" pageNumber="62">D. endopilosella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C474D30FFFD3B7AFE87FC5E76AA9BFE" authorityName="PACE" authorityYear="2008" box="[324,506,896,920]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="62" phylum="Arthropoda" rank="species" species="semimonticola">
<emphasis id="B933EAA1FFFD3B7AFE87FC5E76AA9BFE" box="[324,506,896,920]" italics="true" pageId="29" pageNumber="62">D. semimonticola</emphasis>
</taxonomicName>
see
<bibRefCitation id="EFD64B42FFFD3B7AFDF6FC5E75E79BFE" author="PACE, R." box="[565,695,896,920]" pageId="29" pageNumber="62" pagination="39 - 86" refId="ref30339" refString="PACE, R. 2000: Aleocharinae della Thailandia (Coleoptera, Staphylinidae). - Bollettino del Museo Regionale di Scienze Naturali, Torino 17 (1): 39 - 86." type="journal article" year="2000">PACE (2000</bibRefCitation>
,
<bibRefCitation id="EFD64B42FFFD3B7AFD0BFC5E74559BFE" author="PACE, R." box="[712,773,896,920]" pageId="29" pageNumber="62" pagination="107 - 156" refId="ref30451" refString="PACE, R. 2008: Generi e specie della tribu Lomechusini del Borneo (Coleoptera, Staphylinidae). - Revue Suisse de Zoologie 115 (1): 107 - 156." type="journal article" year="2008">2008</bibRefCitation>
). It differs from the widespread and common
<taxonomicName id="4C474D30FFFD3B7AFD5DFC7E76DB9BBE" authority="(BERNHAUER, 1916)" baseAuthorityName="BERNHAUER" baseAuthorityYear="1916" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="62" phylum="Arthropoda" rank="species" species="obliqua">
<emphasis id="B933EAA1FFFD3B7AFD5DFC7E745B9BDE" box="[670,779,928,952]" italics="true" pageId="29" pageNumber="62">D. obliqua</emphasis>
(BERNHAUER, 1916)
</taxonomicName>
by numerous characters, e.g., a smaller and more slender body with narrower elytra, the coloration of the abdomen and elytra (
<taxonomicName id="4C474D30FFFD3B7AFD5BFBDE74549C7E" baseAuthorityName="BERNHAUER" baseAuthorityYear="1916" box="[664,772,1024,1048]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="62" phylum="Arthropoda" rank="species" species="obliqua">
<emphasis id="B933EAA1FFFD3B7AFD5BFBDE74549C7E" box="[664,772,1024,1048]" italics="true" pageId="29" pageNumber="62">D. obliqua</emphasis>
</taxonomicName>
: abdomen blackish with the narrow posterior margins of the segments yellow; elytra yellow with the posterolateral portions extensively blackish), much less massive antennae with more transverse antennomeres VX, the shape of the head (depressed or impressed in
<taxonomicName id="4C474D30FFFD3B7AFD4CFB7E75AB9CDE" baseAuthorityName="BERNHAUER" baseAuthorityYear="1916" box="[655,763,1184,1208]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="62" phylum="Arthropoda" rank="species" species="obliqua">
<emphasis id="B933EAA1FFFD3B7AFD4CFB7E75AB9CDE" box="[655,763,1184,1208]" italics="true" pageId="29" pageNumber="62">D. obliqua</emphasis>
</taxonomicName>
), the modifications of the male pronotum, the shapes of the male and female tergite and sternite VIII, and by the completely different primary sexual characters. For illustrations of
<taxonomicName id="4C474D30FFFD3B7AFEE2FAFE76DE9D5E" baseAuthorityName="BERNHAUER" baseAuthorityYear="1916" box="[289,398,1312,1336]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="62" phylum="Arthropoda" rank="species" species="obliqua">
<emphasis id="B933EAA1FFFD3B7AFEE2FAFE76DE9D5E" box="[289,398,1312,1336]" italics="true" pageId="29" pageNumber="62">D. obliqua</emphasis>
</taxonomicName>
see
<bibRefCitation id="EFD64B42FFFD3B7AFE78FAFE750A9D5E" author="ASSING, V." box="[443,602,1312,1336]" pageId="29" pageNumber="62" pagination="243 - 262" refId="ref29450" refString="ASSING, V. 2015 b: New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). - Contributions to Entomology 65 (2): 243 - 262. - https: // www. contributions-toentomology. org / article / view / 1882 / 1881." type="journal article" year="2015">ASSING (2015b</bibRefCitation>
,
<bibRefCitation id="EFD64B42FFFD3B7AFDA6FAFE75F99D5E" author="ASSING, V." box="[613,681,1312,1336]" pageId="29" pageNumber="62" pagination="63 - 106" refId="ref29777" refString="ASSING, V. 2017 a: On the Lomechusini fauna of the East Palaearctic and Oriental regions, with a focus on the genera Orphnebius and Amaurodera (Coleoptera: Staphylinidae: Aleocharinae). - Contributions to Entomology 67 (1): 63 - 106. - https: // www. contributions-to-entomology. org / article / view / 1918 / 1915." type="journal article" year="2017">2017a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35D6538FFFD3B7AFF6DFA81754B9D9E" pageId="29" pageNumber="62" type="distribution">
<paragraph id="8BF836B3FFFD3B7AFF6DFA81754B9D9E" blockId="29.[174,785,1375,1528]" pageId="29" pageNumber="62">
<emphasis id="B933EAA1FFFD3B7AFF6DFA8175709D11" bold="true" box="[174,544,1375,1400]" pageId="29" pageNumber="62">Distribution and natural history</emphasis>
: The type locality is situated to the northeast of
<collectingRegion id="4983F851FFFD3B7AFE18FA5E75719DF1" box="[475,545,1408,1431]" country="Philippines" name="Davao" pageId="29" pageNumber="62">Davao</collectingRegion>
, near the coast of the Gulf of
<collectingRegion id="4983F851FFFD3B7AFF3FFA7E76129DD1" box="[252,322,1440,1463]" country="Philippines" name="Davao" pageId="29" pageNumber="62">Davao</collectingRegion>
, Central Mindanao,
<collectingCountry id="F3507623FFFD3B7AFDDAFA7E75C49DDE" box="[537,660,1440,1464]" name="Philippines" pageId="29" pageNumber="62">Philippines</collectingCountry>
. The specimens were sifted from litter on a slope near a river. The male
<typeStatus id="54FC8811FFFD3B7AFF2BFA3E76159D9E" box="[232,325,1504,1528]" pageId="29" pageNumber="62" type="paratype">paratype</typeStatus>
is distinctly teneral.
</paragraph>
</subSubSection>
</treatment>
</document>