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<document id="CC7AD76F18346D905E347483BD28DFC9" ID-DOI="10.21248/contrib.entomol.69.1.033-070" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742766" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638269617685" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2019" docId="03EE87A5FFC13B45FC95F9F474CF9B11" docLanguage="en" docName="ContribEntomol.69.1.33-70.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 69 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) reductus Assing 2019, spec. nov." docType="treatment" docUuid="15761BF9-31AA-4B92-A7A5-8C3535DF876C" docUuidSource="ZooBank" docVersion="3" lastPageNumber="67" masterDocId="FFD7FFDDFFE03B67FFC3FFDE77509866" masterDocTitle="On the Lomechusini fauna of the Palaearctic and Oriental regions. XXVI. New species, a new synonymy, and additional records (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="70" masterPageNumber="33" pageNumber="66" updateTime="1699437235520" updateUser="plazi">
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<mods:title id="0E877A1BF99A6113E77D4C6B766549AB">On the Lomechusini fauna of the Palaearctic and Oriental regions. XXVI. New species, a new synonymy, and additional records (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="0F272C9ECBF1545167F978842C6DAB86">Assing, Volker</mods:namePart>
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<mods:typeOfResource id="21942F3C07CBCCCA336A795FE84C905F">text</mods:typeOfResource>
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<mods:title id="B50B527917362E935A7E26E1C5ED1E54">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:part id="B7810F6F57D9115331A38D5164041ECE">
<mods:date id="7B31C5553A8499714B44AAAE4C704330">2019</mods:date>
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<mods:number id="77EA56C69BCADF3A9835875A89970EBA">2019-06-24</mods:number>
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<mods:number id="AA4F69697AAD59FA52272C2DA74F8A63">69</mods:number>
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<mods:classification id="BDABB5EB4AC3EA798473D5EF9EFF8D09">journal article</mods:classification>
<mods:identifier id="91BC9AA191D03C8A819901374F74B61B" type="DOI">10.21248/contrib.entomol.69.1.033-070</mods:identifier>
<mods:identifier id="9BF3A6CD291976A4017040452B08C1FE" type="ISSN">0005-805X</mods:identifier>
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<treatment id="03EE87A5FFC13B45FC95F9F474CF9B11" LSID="urn:lsid:zoobank.org:act:15761BF9-31AA-4B92-A7A5-8C3535DF876C" httpUri="http://treatment.plazi.org/id/03EE87A5FFC13B45FC95F9F474CF9B11" lastPageId="34" lastPageNumber="67" pageId="33" pageNumber="66">
<subSubSection id="C35D6538FFC13B46FC95F9F472309E03" pageId="33" pageNumber="66" type="nomenclature">
<paragraph id="8BF836B3FFC13B46FC95F9F473F79E20" blockId="33.[854,1376,1578,1669]" box="[854,1191,1578,1606]" pageId="33" pageNumber="66">
<heading id="D0B081DFFFC13B46FC95F9F473F79E20" box="[854,1191,1578,1606]" fontSize="12" level="2" pageId="33" pageNumber="66" reason="2">
<taxonomicName id="4C474D30FFC13B46FC95F9F473679E20" ID-CoL="8YCH2" authority="Assing, 2019" authorityName="Assing" authorityYear="2019" box="[854,1079,1578,1606]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="66" phylum="Arthropoda" rank="species" species="reductus" status="sp. nov." subGenus="Zyras">
<emphasis id="B933EAA1FFC13B46FC95F9F474DC9E20" box="[854,908,1578,1606]" italics="true" pageId="33" pageNumber="66">Zyras</emphasis>
(
<emphasis id="B933EAA1FFC13B46FC59F9F474809E20" box="[922,976,1578,1606]" italics="true" pageId="33" pageNumber="66">Zyras</emphasis>
)
<emphasis id="B933EAA1FFC13B46FC1DF9F473679E20" box="[990,1079,1578,1606]" italics="true" pageId="33" pageNumber="66">reductus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20057DAFFC13B46FBFEF9F473F79E20" box="[1085,1191,1578,1606]" pageId="33" pageNumber="66" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BF836B3FFC13B46FC95F98F72309E03" blockId="33.[854,1376,1578,1669]" box="[854,1376,1617,1637]" pageId="33" pageNumber="66">
<uri id="FFD63AB1FFC13B46FC95F98F72309E03" box="[854,1376,1617,1637]" pageId="33" pageNumber="66">
urn:lsid:zoobank.org:act:
<uuid id="FFE10C66FFC13B46FBCBF98F72309E03" box="[1032,1376,1617,1637]" pageId="33" pageNumber="66">15761BF9-31AA-4B92-A7A5-8C3535DF876C</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C35D6538FFC13B46FC95F9AF74AF9EE3" box="[854,1023,1649,1669]" pageId="33" pageNumber="66" type="description">
<paragraph id="8BF836B3FFC13B46FC95F9AF74AF9EE3" blockId="33.[854,1376,1578,1669]" box="[854,1023,1649,1669]" pageId="33" pageNumber="66">(Figs 7174,160162)</paragraph>
</subSubSection>
<subSubSection id="C35D6538FFC13B46FCF3F97E73C89F5E" pageId="33" pageNumber="66" type="materials_examined">
<paragraph id="8BF836B3FFC13B46FCF3F97E73C89F5E" blockId="33.[816,1427,1695,1848]" pageId="33" pageNumber="66">
<emphasis id="B933EAA1FFC13B46FCF3F97E74999ED1" bold="true" box="[816,969,1695,1720]" pageId="33" pageNumber="66">Type material</emphasis>
:
<materialsCitation id="3B2F3CEEFFC13B46FC1BF97E73C89F5E" collectingDate="2018-03-24" collectingDateMax="2018-03-26" collectingDateMin="2018-03-24" collectionCode="V" collectorName="Shavrin" country="Philippines" county="Kawa-kawa River" elevation="300" latitude="6.7752776" location="Mindanao" longLatPrecision="21" longitude="126.14472" municipality="Sitio Bitaugan" pageId="33" pageNumber="66" specimenCount="2" typeStatus="holotype">
<typeStatus id="54FC8811FFC13B46FC1BF97E73119EDE" box="[984,1089,1696,1720]" pageId="33" pageNumber="66" type="holotype">Holotype</typeStatus>
: “
<collectingCountry id="F3507623FFC13B46FBB3F97E725E9EDE" box="[1136,1294,1696,1720]" name="Philippines" pageId="33" pageNumber="66">PHILIPPINES</collectingCountry>
:
<location id="8E986068FFC13B46FADAF97E72DF9EDE" LSID="urn:lsid:plazi:treatment:03EE87A5FFC13B45FC95F9F474CF9B11:8E986068FFC13B46FADAF97E72DF9EDE" box="[1305,1423,1696,1720]" country="Philippines" county="Kawa-kawa River" latitude="6.7752776" longLatPrecision="21" longitude="126.14472" municipality="Sitio Bitaugan" name="Mindanao" pageId="33" pageNumber="66">Mindanao</location>
,
<collectingMunicipality id="6B9CACC9FFC13B46FCF3F91E74839EBE" box="[816,979,1728,1752]" pageId="33" pageNumber="66">Sitio Bitaugan</collectingMunicipality>
,
<collectingCounty id="62994E3FFFC13B46FC2BF91E73FE9EBE" box="[1000,1198,1728,1752]" pageId="33" pageNumber="66">Kawa-kawa River</collectingCounty>
,
<quantity id="4CBF9B56FFC13B46FB00F91E72589EB1" box="[1219,1288,1728,1752]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="33" pageNumber="66" unit="m" value="300.0">
<elevation id="006AD180FFC13B46FB00F91E72589EB1" box="[1219,1288,1728,1752]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="33" pageNumber="66" unit="m" value="300.0">300 m</elevation>
</quantity>
,
<geoCoordinate id="EE735074FFC13B46FADEF91E72D99EBE" box="[1309,1417,1728,1752]" degrees="6" direction="north" minutes="46" orientation="latitude" pageId="33" pageNumber="66" precision="15" seconds="31" value="6.7752776">6°46'31&quot;N</geoCoordinate>
,
<geoCoordinate id="EE735074FFC13B46FCF3F93E74E19E9E" box="[816,945,1760,1784]" degrees="126" direction="east" minutes="08" orientation="longitude" pageId="33" pageNumber="66" precision="15" seconds="41" value="126.14472">126°08'41&quot;E</geoCoordinate>
, stream bank, wet litter, debris, dead wood,
<date id="FFF91073FFC13B46FCF3F8DE749E9F7E" box="[816,974,1792,1816]" pageId="33" pageNumber="66" value="2018-03-24" valueMax="2018-03-26" valueMin="2018-03-24">
<collectingDate id="EFBDE99BFFC13B46FCF3F8DE749E9F7E" box="[816,974,1792,1816]" pageId="33" pageNumber="66" value="2018-03-24" valueMax="2018-03-26" valueMin="2018-03-24">2426.III.2018</collectingDate>
</date>
, leg.
<collectorName id="26B25365FFC13B46FBC2F8DE73039F7E" box="[1025,1107,1792,1816]" pageId="33" pageNumber="66">Shavrin</collectorName>
/
<typeStatus id="54FC8811FFC13B46FBA4F8DE738C9F7E" box="[1127,1244,1792,1816]" pageId="33" pageNumber="66" type="holotype">Holotypus</typeStatus>
<taxonomicName id="4C474D30FFC13B46FB36F8DE72C39F7E" authority="Assing, 2019" authorityName="Assing" authorityYear="2019" box="[1269,1427,1792,1816]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="66" phylum="Arthropoda" rank="species" species="reductus" status="sp. nov.">
<emphasis id="B933EAA1FFC13B46FB36F8DE72C39F7E" box="[1269,1427,1792,1816]" italics="true" pageId="33" pageNumber="66">Zyras reductus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20057DAFFC13B46FCF3F8FE74399F51" box="[816,873,1824,1847]" pageId="33" pageNumber="66" rank="species">sp. n.</taxonomicNameLabel>
, det.
<collectionCode id="ED56AE76FFC13B46FC61F8FE74E49F51" box="[930,948,1824,1847]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="33" pageNumber="66" type="Museum">V</collectionCode>
. Assing 2018” (cAss).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35D6538FFC13B46FCF3F8BE73599FDE" pageId="33" pageNumber="66" type="etymology">
<paragraph id="8BF836B3FFC13B46FCF3F8BE73599FDE" blockId="33.[816,1427,1888,1976]" pageId="33" pageNumber="66">
<emphasis id="B933EAA1FFC13B46FCF3F8BE74F79F1E" bold="true" box="[816,935,1888,1912]" pageId="33" pageNumber="66">Etymology</emphasis>
: The specific epithet is the past participle of the Latin verb reducere and alludes to the completely reduced hind wings.
</paragraph>
</subSubSection>
<subSubSection id="C35D6538FFC13B45FCF3F80174F39AFE" lastPageId="34" lastPageNumber="67" pageId="33" pageNumber="66" type="description">
<paragraph id="8BF836B3FFC13B45FCF3F80175A399FE" blockId="33.[816,1428,2015,2104]" lastBlockId="34.[212,823,256,1880]" lastPageId="34" lastPageNumber="67" pageId="33" pageNumber="66">
<emphasis id="B933EAA1FFC13B46FCF3F80174E29F91" bold="true" box="[816,946,2015,2039]" pageId="33" pageNumber="66">Description</emphasis>
: Body length
<quantity id="4CBF9B56FFC13B46FB9AF83E73FC9F91" box="[1113,1196,2016,2040]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="33" pageNumber="66" unit="mm" value="6.2">6.2 mm</quantity>
; length of forebody
<quantity id="4CBF9B56FFC13B46FCF3F7DE74D49071" box="[816,900,2048,2072]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="33" pageNumber="66" unit="mm" value="2.7">2.7 mm</quantity>
.Habitus as in Fig. 71. Coloration:forebody brown; abdomen dark-brown with the anterior margins and the narrow posterior margins of the segments reddish-yellow and with the postero-median portions of tergites V and VI blackish; legs yellowish; antennae pale reddish-brown with antennomere XI dark-yellowish and the basal three antennomeres pale-reddish; maxillary palpi yellow.
</paragraph>
<paragraph id="8BF836B3FFC23B45FF17FE7E764A9AD1" blockId="34.[212,823,256,1880]" pageId="34" pageNumber="67">
Head (Fig. 72) 1.1 times as broad as long; punctation sparse and very shallow, median portion extensively impunctate. Eyes strongly convex, longer than postocular region in dorsal view. Antenna (Fig. 73)
<quantity id="4CBF9B56FFC23B45FD1CFDDE74619A71" box="[735,817,512,536]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="34" pageNumber="67" unit="mm" value="2.2">2.2 mm</quantity>
long and rather slender; antennomeres IVV weakly oblong, VI approximately as long as broad, VIIX weakly oblong, X much less than 1.5 times as broad as long, and XI approximately as long as the combined length of IX and X.
</paragraph>
<paragraph id="8BF836B3FFC23B45FF17FD1E76839BFE" blockId="34.[212,823,256,1880]" pageId="34" pageNumber="67">Pronotum (Fig. 72) approximately as broad as long and 1.07 times as broad as head, broadest at anterior angles, rather strongly convex in crosssection; lateral margins not sinuate in dorsal view; posterior angles obtusely marked; punctation rather coarse, moderately sparse, and somewhat irregularly distributed; impunctate median band moderately broad.</paragraph>
<paragraph id="8BF836B3FFC23B45FF17FC7E75949C31" blockId="34.[212,823,256,1880]" pageId="34" pageNumber="67">Elytra (Fig. 72) conspicuously short, 0.55 times as long as pronotum, distinctly dilated posteriad; humeral angles completely obsolete; punctation moderately dense, very coarse, and sharply defined. Hind wings completely reduced. Legs long and slender; metatarsomere I slender, nearly as long as the combined length of IIIV.</paragraph>
<paragraph id="8BF836B3FFC23B45FF17FBBE76859E11" blockId="34.[212,823,256,1880]" pageId="34" pageNumber="67">Abdomen (Fig. 74) distinctly broader than elytra, with rather deep anterior impressions on tergites IIIV; anterior impressions of tergites IIIV each with a transverse row of distinct non-setiferous punctures, with few lateral setiferous punctures on either side, and with six (tergites III and IV) or eight (tergite V) setiferous punctures bearing long black setae at or near posterior margin; tergite VI with a transverse row of non-setiferous punctures anteriorly, with few lateral setiferous punctures on either side and with ten setiferous punctures at or near posterior margin; tergite VII with a transverse band of rather sparse non-setiferous punctures anteriorly and with two highly irregular transverse series of setiferous punctures posteriorly, posterior margin with an indistinct rudiment of a palisade fringe; tergite VIII with setiferous punctation only in posterior third, posterior margin strongly convex.</paragraph>
<paragraph id="8BF836B3FFC23B45FF2AF95E76779F51" blockId="34.[212,823,256,1880]" pageId="34" pageNumber="67">
: sternite VIII strongly produced posteriorly, posterior margin truncate in the middle; median lobe of aedeagus approximately
<quantity id="4CBF9B56FFC23B45FDC9F91E753A9EB1" box="[522,618,1728,1752]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="34" pageNumber="67" unit="mm" value="0.75">0.75 mm</quantity>
long and shaped as in
<figureCitation id="137C2A36FFC23B45FED7F93E76F79E9E" box="[276,423,1760,1784]" captionStart="Figs 145165" captionStartId="13.[174,214,2001,2023]" captionTargetBox="[176,1415,260,1989]" captionTargetId="figure-118@13.[173,1421,255,1995]" captionTargetPageId="13" captionText="Figs 145165:Drusilla spiniventris(145147), D. breviuter(148152), Pheidologitonetes biplicatus (153154), Zyras conlectus (155 157), Z. hetzeli (158159), Z. reductus(160162), andZ. shavrini(163165): male tergite VIII (145, 148); median lobe of aedeagus in lateral and in ventral view (146147, 149150, 153156, 158161, 163164); female tergite VIII (151); spermatheca (152); paramere (157, 162, 165). Scale bars: 145151, 153165: 0.2 mm; 152: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742777" httpUri="https://zenodo.org/record/5742777/files/figure.png" pageId="34" pageNumber="67">Figs 160161</figureCitation>
; paramere (
<figureCitation id="137C2A36FFC23B45FDECF93E75DA9E9E" box="[559,650,1760,1784]" captionStart="Figs 145165" captionStartId="13.[174,214,2001,2023]" captionTargetBox="[176,1415,260,1989]" captionTargetId="figure-118@13.[173,1421,255,1995]" captionTargetPageId="13" captionText="Figs 145165:Drusilla spiniventris(145147), D. breviuter(148152), Pheidologitonetes biplicatus (153154), Zyras conlectus (155 157), Z. hetzeli (158159), Z. reductus(160162), andZ. shavrini(163165): male tergite VIII (145, 148); median lobe of aedeagus in lateral and in ventral view (146147, 149150, 153156, 158161, 163164); female tergite VIII (151); spermatheca (152); paramere (157, 162, 165). Scale bars: 145151, 153165: 0.2 mm; 152: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742777" httpUri="https://zenodo.org/record/5742777/files/figure.png" pageId="34" pageNumber="67">Fig. 162</figureCitation>
)
<quantity id="4CBF9B56FFC23B45FD5FF93E75BF9E91" box="[668,751,1760,1784]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="34" pageNumber="67" unit="mm" value="0.9">0.9 mm</quantity>
long, apical lobe rather short, flattened and apically broadly convex.
</paragraph>
<paragraph id="8BF836B3FFC23B45FF2BF89E76329F3E" blockId="34.[212,823,256,1880]" box="[232,354,1856,1880]" pageId="34" pageNumber="67">: unknown.</paragraph>
<paragraph id="8BF836B3FFC23B45FF17F8A174F39AFE" blockId="34.[212,823,1919,2104]" lastBlockId="34.[854,1465,256,664]" pageId="34" pageNumber="67">
<emphasis id="B933EAA1FFC23B45FF17F8A176F89FFE" bold="true" box="[212,424,1919,1944]" pageId="34" pageNumber="67">Comparative notes</emphasis>
: Based on the slender habitus, the slender antennae, the strongly convex pronotum (crosssection), the punctation pattern of the abdomen, and the morphology of the aedeagus (shape of median lobe and of apical lobe of paramere), this conspicuous species is closely related to
<taxonomicName id="4C474D30FFC23B45FE51F7FE75FC905E" authority="CAMERON, 1944" authorityName="CAMERON" authorityYear="1944" box="[402,684,2080,2104]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="67" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="B933EAA1FFC23B45FE51F7FE76BC9051" box="[402,492,2080,2103]" italics="true" pageId="34" pageNumber="67">Z. nitens</emphasis>
CAMERON, 1944
</taxonomicName>
,
<emphasis id="B933EAA1FFC23B45FD71F7FE743D997E" box="[690,877,256,2104]" italics="true" pageId="34" pageNumber="67">Z. glabricol- lis</emphasis>
SCHEERPELTZ, 1965,
<taxonomicName id="4C474D30FFC23B45FB9DFEDE722C997E" authority="ASSING, 2017" authorityName="ASSING" authorityYear="2017" box="[1118,1404,256,280]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="67" phylum="Arthropoda" rank="species" species="truncatus">
<emphasis id="B933EAA1FFC23B45FB9DFEDE738E9971" box="[1118,1246,256,279]" italics="true" pageId="34" pageNumber="67">Z. truncatus</emphasis>
ASSING, 2017
</taxonomicName>
, and allied species. It is readily distinguished from all other species of
<taxonomicName id="4C474D30FFC23B45FC0CFE9E73599931" authorityName="STEPHENS" authorityYear="1835" box="[975,1033,320,343]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="B933EAA1FFC23B45FC0CFE9E73599931" box="[975,1033,320,343]" italics="true" pageId="34" pageNumber="67">Zyras</emphasis>
</taxonomicName>
sensu strictu by much shorter elytra alone. The only other species that is at least close to this condition is
<taxonomicName id="4C474D30FFC23B45FC25FE5E724199FE" authority="Pace, 2008" authorityName="Pace" authorityYear="2008" box="[998,1297,384,408]" class="Insecta" family="Staphylinidae" genus="Amaurodera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="67" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B933EAA1FFC23B45FC25FE5E73DF99FE" box="[998,1167,384,408]" italics="true" pageId="34" pageNumber="67">Z. kinabaluensis</emphasis>
<bibRefCitation id="EFD64B42FFC23B45FB5FFE5E724199FE" author="PACE, R." box="[1180,1297,384,408]" pageId="34" pageNumber="67" pagination="107 - 156" refId="ref30451" refString="PACE, R. 2008: Generi e specie della tribu Lomechusini del Borneo (Coleoptera, Staphylinidae). - Revue Suisse de Zoologie 115 (1): 107 - 156." type="journal article" year="2008">Pace, 2008</bibRefCitation>
</taxonomicName>
from Borneo, from which
<taxonomicName id="4C474D30FFC23B45FC1EFE7E730299DE" authorityName="Assing" authorityYear="2019" box="[989,1106,416,440]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="67" phylum="Arthropoda" rank="species" species="reductus">
<emphasis id="B933EAA1FFC23B45FC1EFE7E730299DE" box="[989,1106,416,440]" italics="true" pageId="34" pageNumber="67">Z. reductus</emphasis>
</taxonomicName>
is distinguished by even shorter elytra (
<taxonomicName id="4C474D30FFC23B45FC6FFE1E730599BE" authorityName="Pace" authorityYear="2008" box="[940,1109,448,472]" class="Insecta" family="Staphylinidae" genus="Amaurodera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="67" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B933EAA1FFC23B45FC6FFE1E730599BE" box="[940,1109,448,472]" italics="true" pageId="34" pageNumber="67">Z. kinabaluensis</emphasis>
</taxonomicName>
: elytra 0.65 times as long as pronotum), larger body size (
<taxonomicName id="4C474D30FFC23B45FB5AFE3E7212999E" authorityName="Pace" authorityYear="2008" box="[1177,1346,480,504]" class="Insecta" family="Staphylinidae" genus="Amaurodera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="67" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B933EAA1FFC23B45FB5AFE3E7212999E" box="[1177,1346,480,504]" italics="true" pageId="34" pageNumber="67">Z. kinabaluensis</emphasis>
</taxonomicName>
: length of forebody
<quantity id="4CBF9B56FFC23B45FC03FDDE73459A7E" box="[960,1045,512,536]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="34" pageNumber="67" unit="mm" value="2.4">2.4 mm</quantity>
), less slender antennae with a longer antennomere XI, larger and more convex eyes, a broader abdomen, and by the shape of the aedeagus. For illustrations of
<taxonomicName id="4C474D30FFC23B45FC75FDBE730F9A1E" authorityName="Pace" authorityYear="2008" box="[950,1119,608,632]" class="Insecta" family="Staphylinidae" genus="Amaurodera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="67" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B933EAA1FFC23B45FC75FDBE730F9A1E" box="[950,1119,608,632]" italics="true" pageId="34" pageNumber="67">Z. kinabaluensis</emphasis>
</taxonomicName>
see
<bibRefCitation id="EFD64B42FFC23B45FB56FDBE726F9A1E" author="ASSING, V." box="[1173,1343,608,632]" pageId="34" pageNumber="67" pagination="213 - 246" refId="ref29923" refString="ASSING, V. 2017 c: On the Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (2): 213 - 246. - https: // www. contributions-toentomology. org / article / view / 1939 / 1939." type="journal article" year="2017">ASSING (2017c)</bibRefCitation>
and
<bibRefCitation id="EFD64B42FFC23B45FABEFDBE74CF9AFE" author="PACE, R." pageId="34" pageNumber="67" pagination="107 - 156" refId="ref30451" refString="PACE, R. 2008: Generi e specie della tribu Lomechusini del Borneo (Coleoptera, Staphylinidae). - Revue Suisse de Zoologie 115 (1): 107 - 156." type="journal article" year="2008">PACE (2008)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35D6538FFC23B45FC95FD6174CF9B11" pageId="34" pageNumber="67" type="distribution">
<paragraph id="8BF836B3FFC23B45FC95FD6174CF9B11" blockId="34.[854,1465,703,888]" pageId="34" pageNumber="67">
<emphasis id="B933EAA1FFC23B45FC95FD61748F9AB1" bold="true" box="[854,991,703,727]" pageId="34" pageNumber="67">Distribution</emphasis>
: The type locality is situated in
<collectingRegion id="4983F851FFC23B45FAADFD1E73479A9E" country="Philippines" name="Davao Oriental" pageId="34" pageNumber="67">Davao Oriental Province</collectingRegion>
in the southeast of Mindanao, South
<collectingCountry id="F3507623FFC23B45FC95FCDE74819B7E" box="[854,977,768,792]" name="Philippines" pageId="34" pageNumber="67">Philippines</collectingCountry>
. The
<typeStatus id="54FC8811FFC23B45FBE6FCDE73D39B7E" box="[1061,1155,768,792]" pageId="34" pageNumber="67" type="holotype">holotype</typeStatus>
was collected by sifting wet litter between stones, wet debris, and dead wood accumulated near the Kawa-kawa river at an altitude of
<specimenCount id="9D41FD3AFFC23B45FC95FCBE74CF9B11" box="[854,927,864,888]" pageId="34" pageNumber="67" type="generic">
<quantity id="4CBF9B56FFC23B45FC95FCBE74CB9B11" box="[854,923,864,888]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="34" pageNumber="67" unit="m" value="300.0">300 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
</treatment>
</document>