treatments-xml/data/03/EE/87/03EE8794FFB9FFCBFD02FAB0FD061CB6.xml

<document id="00D7005A58F8AF9DA22B37E264EEC470" ID-CLB-Dataset="9470" ID-DOI="10.5852/ejt.2020.724.1157" ID-GBIF-Dataset="8b7e3a10-8d56-4d68-9516-80e08d0565ef" ID-Zenodo-Dep="4298074" ID-ZooBank="F8E5876F-1438-4055-9DB8-656969643176" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1606737149627" checkinUser="plazi" docAuthor="Rivera-Quiroz, Francisco Andrés, Petcharad, Booppa &amp; Miller, Jeremy A." docDate="2020" docId="03EE8794FFB9FFCBFD02FAB0FD061CB6" docLanguage="en" docName="ejt-724_rivera-quiroz_petcharad_miller.pdf" docOrigin="European Journal of Taxonomy 724" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Hahnia ngai Rivera-Quiroz &amp; Petcharad &amp; Miller 2020, sp. nov." docType="treatment" docVersion="8" lastPageNumber="61" masterDocId="FFD7FFECFFB1FFC1FF9EFF9DFFFA1F2F" masterDocTitle="First records and a new genus of comb-tailed spiders (Araneae: Hahniidae) from Thailand with comments on the six-eyed species of this family" masterLastPageNumber="69" masterPageNumber="51" pageNumber="59" updateTime="1698846946028" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="BDD4A564F9427446784B707C5F30DE02">First records and a new genus of comb-tailed spiders (Araneae: Hahniidae) from Thailand with comments on the six-eyed species of this family</mods:title>
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<mods:namePart id="F45AE2B6D927F7466E9AC0E9A3F7F2CF">Rivera-Quiroz, Francisco Andrés</mods:namePart>
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<mods:affiliation id="07B1D8CBD3BF51CFC13BE7BAAC0BB570">Department of Terrestrial Zoology, Understanding Evolution group, Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, The Netherlands. Institute for Biology Leiden (IBL), Leiden University, Sylviusweg 72, 2333 BE Leiden, The Netherlands. Faculty of Science and Technology, Thammasat University, Rangsit, Pathum Thani, 12121 Thailand.</mods:affiliation>
<mods:nameIdentifier id="A295953DA6DFE3A50A20046E15A421F4" type="email">andres.riveraquiroz@naturalis.nl</mods:nameIdentifier>
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<mods:namePart id="728956368F3F225553A31102ACF64F49">Petcharad, Booppa</mods:namePart>
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<mods:namePart id="BA328FABE47D374472B9AAF99381B20B">Miller, Jeremy A.</mods:namePart>
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<mods:affiliation id="97A16D7350BCC539FDCD960BF01A6969">Department of Terrestrial Zoology, Understanding Evolution group, Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, The Netherlands. Institute for Biology Leiden (IBL), Leiden University, Sylviusweg 72, 2333 BE Leiden, The Netherlands. Faculty of Science and Technology, Thammasat University, Rangsit, Pathum Thani, 12121 Thailand.</mods:affiliation>
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<taxonomicName id="4C474D01FFB9FFC9FD02FAB0FCCC1A68" ID-CoL="98ZG3" authority="Rivera-Quiroz &amp; Petcharad &amp; Miller, 2020" authorityName="Rivera-Quiroz &amp; Petcharad &amp; Miller" authorityYear="2020" box="[668,822,1325,1352]" class="Arachnida" family="Hahniidae" genus="Hahnia" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="ngai" status="sp. nov.">
<emphasis id="B933EA90FFB9FFC9FD02FAB0FCCC1A68" bold="true" box="[668,822,1325,1352]" italics="true" pageId="8" pageNumber="59">Hahnia ngai</emphasis>
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<figureCitation id="137C2A07FFB9FFC9FD4CFAE9FCE71AA1" box="[722,797,1396,1422]" captionStart="Fig" captionStartId="9.[189,232,1920,1946]" captionTargetBox="[219,1370,677,1872]" captionTargetId="figure-89@9.[205,1382,665,1884]" captionTargetPageId="9" captionText="Fig. 4. Hahnia ngai sp. nov., holotype, ♀ (RMNH.ARA.18415). a–c. Habitus. a. Ventral view. b. Lateral view. c. Dorsal view. d. Prosoma, anterior view. e. Chelicerae, posterior view. f–g. Epigynum. f. Dorsal view, cleared. g. Ventral view. Scale bars: a–c = 1.0 mm; d–e, g = 0.25 mm; f = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4298084" httpUri="https://zenodo.org/record/4298084/files/figure.png" pageId="8" pageNumber="59">Figs 4</figureCitation>
, 
<figureCitation id="137C2A07FFB9FFC9FCB4FAE9FCC21AA1" box="[810,824,1396,1422]" captionStart="Fig" captionStartId="12.[189,232,1888,1914]" captionTargetBox="[189,1396,639,1811]" captionTargetId="figure-148@12.[189,1398,630,1824]" captionTargetPageId="12" captionText="Fig. 6. Female spinnerets and genitals. a–c. Hahnia ngai sp. nov., holotype (RMNH.ARA.18415). a. Spinnerets, ventral view. b. Epigynum, cleared, dorsal view. c. Ventral view. d–f. Hahnia saccata Zhang, Li &amp; Zheng, 2011 (RMNH.ARA.18412). d. Spinnerets, ventral view. e. Epigynum, cleared, dorsal view. f. Ventral view. Scale bars: a, d–f = 0.25 mm; b–c = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4298088" httpUri="https://zenodo.org/record/4298088/files/figure.png" pageId="8" pageNumber="59">6</figureCitation>
a–c
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<emphasis id="B933EA90FFB9FFC9FF23FA28FECC1AE0" bold="true" box="[189,310,1461,1487]" pageId="8" pageNumber="59">Diagnosis</emphasis>
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<taxonomicName id="4C474D01FFB9FFC9FF23FA79FEAA1AD1" authority="Rivera-Quiroz &amp; Petcharad &amp; Miller, 2020" authorityName="Rivera-Quiroz &amp; Petcharad &amp; Miller" authorityYear="2020" box="[189,336,1508,1534]" class="Arachnida" family="Hahniidae" genus="Hahnia" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="ngai" status="sp. nov.">
<emphasis id="B933EA90FFB9FFC9FF23FA79FEAA1AD1" box="[189,336,1508,1534]" italics="true" pageId="8" pageNumber="59">Hahnia ngai</emphasis>
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can be easily separated from other members of this genus by the simplified female genitalia. Copulatory ducts show only slightly swollen areas with glandular insertions (
<figureCitation id="137C2A07FFB9FFC9FB54F99AFADD190E" box="[1226,1319,1543,1569]" captionStart="Fig" captionStartId="9.[189,232,1920,1946]" captionTargetBox="[219,1370,677,1872]" captionTargetId="figure-89@9.[205,1382,665,1884]" captionTargetPageId="9" captionText="Fig. 4. Hahnia ngai sp. nov., holotype, ♀ (RMNH.ARA.18415). a–c. Habitus. a. Ventral view. b. Lateral view. c. Dorsal view. d. Prosoma, anterior view. e. Chelicerae, posterior view. f–g. Epigynum. f. Dorsal view, cleared. g. Ventral view. Scale bars: a–c = 1.0 mm; d–e, g = 0.25 mm; f = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4298084" httpUri="https://zenodo.org/record/4298084/files/figure.png" pageId="8" pageNumber="59">
Figs 
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, 
<figureCitation id="137C2A07FFB9FFC9FAAAF99AFAB8190D" box="[1332,1346,1543,1570]" captionStart="Fig" captionStartId="12.[189,232,1888,1914]" captionTargetBox="[189,1396,639,1811]" captionTargetId="figure-148@12.[189,1398,630,1824]" captionTargetPageId="12" captionText="Fig. 6. Female spinnerets and genitals. a–c. Hahnia ngai sp. nov., holotype (RMNH.ARA.18415). a. Spinnerets, ventral view. b. Epigynum, cleared, dorsal view. c. Ventral view. d–f. Hahnia saccata Zhang, Li &amp; Zheng, 2011 (RMNH.ARA.18412). d. Spinnerets, ventral view. e. Epigynum, cleared, dorsal view. f. Ventral view. Scale bars: a, d–f = 0.25 mm; b–c = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4298088" httpUri="https://zenodo.org/record/4298088/files/figure.png" pageId="8" pageNumber="59">6</figureCitation>
b–c) but do not form a receptacle or secondary spermathecae (as seen in 
<figureCitation id="137C2A07FFB9FFC9FC46F9B7FBCB196A" box="[984,1073,1578,1605]" captionStart="Fig" captionStartId="11.[189,232,1920,1946]" captionTargetBox="[202,1386,675,1842]" captionTargetId="figure-79@11.[189,1398,663,1855]" captionTargetPageId="11" captionText="Fig. 5. Hahnia saccata Zhang, Li &amp; Zheng, 2011, ♀ (RMNH.ARA.18412). a–c. Habitus.a. Ventral view. b. Lateral view. c. Dorsal view. d. Prosoma, anterior view. e. Chelicerae, posterior view. f–g. Epigynum. f. Dorsal view, cleared. g. Ventral view. Scale bars: a–c = 1.0 mm; d = 0.50 mm; e–g = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4298086" httpUri="https://zenodo.org/record/4298086/files/figure.png" pageId="8" pageNumber="59">
Figs 
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, 
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e–f).
</paragraph>
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<subSubSection id="C35D6509FFB9FFC9FF23F9F1FAF919F7" pageId="8" pageNumber="59" type="etymology">
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<emphasis id="B933EA90FFB9FFC9FF23F9F1FEBE19A9" bold="true" box="[189,324,1644,1670]" pageId="8" pageNumber="59">Etymology</emphasis>
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The species epithet is a derivation of the Thai 
<taxonomicName id="4C474D01FFB9FFC9FD41F906FCE9199B" authorityName="Rivera-Quiroz &amp; Petcharad &amp; Miller" authorityYear="2020" box="[735,787,1691,1716]" class="Arachnida" family="Hahniidae" genus="Hahnia" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="ngai">
<emphasis id="B933EA90FFB9FFC9FD41F906FCE9199B" box="[735,787,1691,1716]" italics="true" pageId="8" pageNumber="59">ngai</emphasis>
</taxonomicName>
(simple), in reference to the relatively simple vulva without the well-formed secondary spermathecae commonly seen in other species of 
<taxonomicName id="4C474D01FFB9FFC9FB3AF920FB0619F8" authorityName="C. L. Koch" authorityYear="1841" box="[1188,1276,1725,1751]" class="Arachnida" family="Hahniidae" genus="Hahnia" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B933EA90FFB9FFC9FB3AF920FB0619F8" box="[1188,1276,1725,1751]" italics="true" pageId="8" pageNumber="59">Hahnia</emphasis>
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.
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<paragraph id="8BF83682FFB9FFC9FF23F962FE961836" blockId="8.[189,364,1791,1817]" box="[189,364,1791,1817]" pageId="8" pageNumber="59">
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<emphasis id="B933EA90FFB9FFC9FF23F962FE961836" bold="true" box="[189,364,1791,1817]" pageId="8" pageNumber="59">Type material</emphasis>
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</paragraph>
<paragraph id="8BF83682FFB9FFC9FF74F8B0FA83189D" blockId="8.[189,1401,1837,1970]" pageId="8" pageNumber="59">
<materialsCitation id="3B2F3CDFFFB9FFC9FF74F8B0FA83189D" ID-GBIF-Occurrence="2985908305" collectedFrom="montane evergreen forest with pine" collectingDate="2018-07-24" collectingDateMax="2018-07-28" collectingDateMin="2018-07-24" collectingMethod="Winkler extractor" collectionCode="RMNH" collectorName="Booppa Petcharad &amp; Jeremy Miller &amp; F. Andres Rivera-Quiroz" country="Thailand" elevation="1409" latitude="18.808367" location="Doi Suthep National Park" longLatPrecision="1" longitude="98.892136" municipality="Doi Suthep National Park" pageId="8" pageNumber="59" specimenCode="RMNH.ARA.18415" specimenCount="1" specimenCount-female="1" stateProvince="Chiang Mai Province" typeStatus="holotype">
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<typeStatus id="54FC8820FFB9FFC9FF74F8B0FEA61868" box="[234,348,1837,1863]" pageId="8" pageNumber="59" type="holotype">Holotype</typeStatus>
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<collectingCountry id="F3507612FFB9FFC9FF23F8CCFEAF1844" box="[189,341,1873,1899]" name="Thailand" pageId="8" pageNumber="59">THAILAND</collectingCountry>
• 
<specimenCount id="9D41FD0BFFB9FFC9FEF2F8CCFE791844" box="[364,387,1873,1899]" pageId="8" pageNumber="59" type="female">♀</specimenCount>
; 
<collectingRegion id="4983F860FFB9FFC9FE0FF8CDFD771845" box="[401,653,1872,1898]" country="Thailand" name="Chiang Mai" pageId="8" pageNumber="59">Chiang Mai Province</collectingRegion>
, 
<collectingMunicipality id="6B9CACF8FFB9FFC9FD06F8CDFC321845" box="[664,968,1872,1898]" country="Thailand" latitude="18.808367" longLatPrecision="1" longitude="98.892136" name="Doi Suthep National Park" pageId="8" pageNumber="59" stateProvince="Chiang Mai Province">Doi Suthep National Park</collectingMunicipality>
; 
<geoCoordinate id="EE735045FFB9FFC9FC4DF8D2FB951844" box="[979,1135,1871,1899]" degrees="18" direction="north" minutes="48.502" orientation="latitude" pageId="8" pageNumber="59" precision="1" value="18.808367">18º48.502′ N</geoCoordinate>
, 
<geoCoordinate id="EE735045FFB9FFC9FBE4F8D2FAE91844" box="[1146,1299,1871,1899]" degrees="98" direction="east" minutes="53.528" orientation="longitude" pageId="8" pageNumber="59" precision="1" value="98.892136">98º53.528′ E</geoCoordinate>
; 
<elevation id="006AD1B1FFB9FFC9FA80F8CCFF0C18A1" metricMagnitude="3" metricUnit="m" metricValue="1.409" pageId="8" pageNumber="59" unit="m" value="1409.0">
<quantity id="4CBF9B67FFB9FFC9FA80F8CCFA8D1844" box="[1310,1399,1873,1899]" metricMagnitude="3" metricUnit="m" metricValue="1.409" pageId="8" pageNumber="59" unit="m" value="1409.0">1409 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFF91042FFB9FFC9FE9BF8E9FE3318A1" box="[261,457,1908,1934]" pageId="8" pageNumber="59" value="2018-07-24" valueMax="2018-07-28" valueMin="2018-07-24">
<collectingDate id="EFBDE9AAFFB9FFC9FE9BF8E9FE3318A1" box="[261,457,1908,1934]" pageId="8" pageNumber="59" value="2018-07-24" valueMax="2018-07-28" valueMin="2018-07-24">24–28 Jul. 2018</collectingDate>
</date>
; 
<collectorName id="26B25354FFB9FFC9FE46F8E9FD4918A1" box="[472,691,1908,1934]" pageId="8" pageNumber="59">Booppa Petcharad</collectorName>
, 
<collectorName id="26B25354FFB9FFC9FD5FF8E9FC9218A1" box="[705,872,1908,1934]" pageId="8" pageNumber="59">Jeremy Miller</collectorName>
and 
<collectorName id="26B25354FFB9FFC9FC38F8E9FB3118A1" box="[934,1227,1908,1934]" pageId="8" pageNumber="59">F. Andrés Rivera-Quiroz</collectorName>
leg.; 
<collectedFrom id="385B757FFFB9FFC9FA8FF8E9FE0E189E" pageId="8" pageNumber="59">montane evergreen forest with pine</collectedFrom>
; 
<collectingMethod id="52064E95FFB9FFC9FE61F80AFD2A189E" box="[511,720,1943,1969]" pageId="8" pageNumber="59">Winkler extractor</collectingMethod>
; 
<specimenCode id="DBE19EF9FFB9FFC9FD42F80AFC30189D" box="[732,970,1943,1970]" pageId="8" pageNumber="59">RMNH.ARA.18415</specimenCode>
(four legs used for DNA extraction).
</materialsCitation>
</paragraph>
<paragraph id="8BF83682FFB8FFC8FF74FE94FE9D1E0C" blockId="9.[189,1398,265,431]" box="[234,359,265,291]" pageId="9" pageNumber="60">
<emphasis id="B933EA90FFB8FFC8FF74FE94FE9D1E0C" bold="true" box="[234,359,265,291]" pageId="9" pageNumber="60">
<typeStatus id="54FC8820FFB8FFC8FF74FE94FE9D1E0C" box="[234,359,265,291]" pageId="9" pageNumber="60" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BF83682FFB8FFC8FF23FEB1FE0C1E80" blockId="9.[189,1398,265,431]" pageId="9" pageNumber="60">
<materialsCitation id="3B2F3CDFFFB8FFC8FF23FEB1FCDE1EA3" ID-GBIF-Occurrence="2985908302" collectedFrom="cloud forest" collectingDate="2018-07-24" collectingDateMax="2018-07-28" collectingDateMin="2018-07-24" collectingMethod="Winkler extractor" collectionCode="RMNH" collectorName="Booppa Petcharad &amp; Jeremy Miller &amp; F. Andres Rivera-Quiroz" country="Thailand" elevation="2572" latitude="18.5878" location="Doi Inthanon National Park" longLatPrecision="1" longitude="98.487335" pageId="9" pageNumber="60" specimenCode="RMNH.ARA.18414" specimenCount="1" specimenCount-female="1" stateProvince="Chiang Mai Province" typeStatus="paratype">
<collectingCountry id="F3507612FFB8FFC8FF23FEB1FEAF1E69" box="[189,341,300,326]" name="Thailand" pageId="9" pageNumber="60">THAILAND</collectingCountry>
• 
<specimenCount id="9D41FD0BFFB8FFC8FEEDFEB1FE591E69" box="[371,419,300,326]" pageId="9" pageNumber="60" type="female">1 ♀</specimenCount>
; 
<collectingRegion id="4983F860FFB8FFC8FE2AFEB1FD421E69" box="[436,696,300,326]" country="Thailand" name="Chiang Mai" pageId="9" pageNumber="60">Chiang Mai Province</collectingRegion>
, 
<location id="8E986059FFB8FFC8FD58FEB1FBE21E69" LSID="urn:lsid:plazi:treatment:03EE8794FFB9FFCBFD02FAB0FD061CB6:8E986059FFB8FFC8FD58FEB1FBE21E69" box="[710,1048,300,326]" country="Thailand" latitude="18.5878" longLatPrecision="1" longitude="98.487335" name="Doi Inthanon National Park" pageId="9" pageNumber="60" stateProvince="Chiang Mai Province">Doi Inthanon National Park</location>
; 
<geoCoordinate id="EE735045FFB8FFC8FBB8FEB7FB3C1E69" box="[1062,1222,298,326]" degrees="18" direction="north" minutes="35.268" orientation="latitude" pageId="9" pageNumber="60" precision="1" value="18.5878">18º35.268′ N</geoCoordinate>
, 
<geoCoordinate id="EE735045FFB8FFC8FB4BFEB7FA8B1E69" box="[1237,1393,298,326]" degrees="98" direction="east" minutes="29.240" orientation="longitude" pageId="9" pageNumber="60" precision="1" value="98.487335">98º29.240′ E</geoCoordinate>
; 
<elevation id="006AD1B1FFB8FFC8FF23FED2FEA61E46" box="[189,348,335,362]" metricMagnitude="3" metricUnit="m" metricValue="2.572" pageId="9" pageNumber="60" unit="m" value="2572.0">
<quantity id="4CBF9B67FFB8FFC8FF23FED2FEE31E46" box="[189,281,335,362]" metricMagnitude="3" metricUnit="m" metricValue="2.572" pageId="9" pageNumber="60" unit="m" value="2572.0">2572 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFF91042FFB8FFC8FEF2FED2FDCB1E46" box="[364,561,335,361]" pageId="9" pageNumber="60" value="2018-07-24" valueMax="2018-07-28" valueMin="2018-07-24">
<collectingDate id="EFBDE9AAFFB8FFC8FEF2FED2FDCB1E46" box="[364,561,335,361]" pageId="9" pageNumber="60" value="2018-07-24" valueMax="2018-07-28" valueMin="2018-07-24">24–28 Jul. 2018</collectingDate>
</date>
; 
<collectorName id="26B25354FFB8FFC8FDDFFED2FCE71E46" box="[577,797,335,361]" pageId="9" pageNumber="60">Booppa Petcharad</collectorName>
, 
<collectorName id="26B25354FFB8FFC8FCB5FED2FC291E46" box="[811,979,335,361]" pageId="9" pageNumber="60">Jeremy Miller</collectorName>
and 
<collectorName id="26B25354FFB8FFC8FB8DFED2FAC21E46" box="[1043,1336,335,361]" pageId="9" pageNumber="60">F. Andrés Rivera-Quiroz</collectorName>
leg.; 
<collectedFrom id="385B757FFFB8FFC8FF23FEEFFEB71EA3" box="[189,333,370,396]" pageId="9" pageNumber="60">cloud forest</collectedFrom>
; 
<collectingMethod id="52064E95FFB8FFC8FEC6FEEFFDD01EA2" box="[344,554,370,397]" pageId="9" pageNumber="60">Winkler extractor</collectingMethod>
; 
<specimenCode id="DBE19EF9FFB8FFC8FDA8FEEFFCDE1EA3" box="[566,804,370,396]" pageId="9" pageNumber="60">RMNH.ARA.18414</specimenCode>
</materialsCitation>
• 
<materialsCitation id="3B2F3CDFFFB8FFC8FCA3FEEFFE0B1E80" ID-GBIF-Occurrence="2985908303" collectedFrom="cloud forest" collectingDate="2018-07-24" collectingDateMax="2018-07-28" collectingDateMin="2018-07-24" collectingMethod="hand coll" collectionCode="RMNH" collectorName="Booppa Petcharad &amp; Jeremy Miller &amp; F. Andres Rivera-Quiroz &amp; Winkler" country="Thailand" elevation="2572" latitude="18.5878" location="Doi Inthanon National Park" longLatPrecision="1" longitude="98.487335" pageId="9" pageNumber="60" specimenCode="RMNH.ARA.18413" specimenCount="1" specimenCount-female="1" stateProvince="Chiang Mai" typeStatus="paratype">
<specimenCount id="9D41FD0BFFB8FFC8FCA3FEEFFC901EA2" box="[829,874,370,397]" pageId="9" pageNumber="60" type="female">1 ♀</specimenCount>
; same collection data as for preceding; 
<collectingMethod id="52064E95FFB8FFC8FAA3FEEFFF0A1E80" pageId="9" pageNumber="60">hand coll</collectingMethod>
.; 
<specimenCode id="DBE19EF9FFB8FFC8FE9EFE08FE0B1E80" box="[256,497,405,431]" pageId="9" pageNumber="60">RMNH.ARA.18413</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35D6509FFB8FFCBFF23FE4BFCEA1C29" lastPageId="10" lastPageNumber="61" pageId="9" pageNumber="60" type="description">
<paragraph id="8BF83682FFB8FFC8FF23FE4BFEB41EDF" blockId="9.[189,334,470,496]" box="[189,334,470,496]" pageId="9" pageNumber="60">
<heading id="D0B081EEFFB8FFC8FF23FE4BFEB41EDF" bold="true" box="[189,334,470,496]" fontSize="11" level="3" pageId="9" pageNumber="60" reason="3">
<emphasis id="B933EA90FFB8FFC8FF23FE4BFEB41EDF" bold="true" box="[189,334,470,496]" pageId="9" pageNumber="60">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BF83682FFB8FFC8FF74FD98FEBF1D30" blockId="9.[189,1399,517,614]" box="[234,325,517,543]" pageId="9" pageNumber="60">
<emphasis id="B933EA90FFB8FFC8FF74FD98FEBF1D30" bold="true" box="[234,325,517,543]" pageId="9" pageNumber="60">Female</emphasis>
</paragraph>
<paragraph id="8BF83682FFB8FFCBFF23FDB4FD4B1EA2" blockId="9.[189,1399,517,614]" lastBlockId="10.[189,1400,265,397]" lastPageId="10" lastPageNumber="61" pageId="9" pageNumber="60">
Carapace pear-shaped, reddish-brown, slightly darker in cephalic region; texture smooth (
<figureCitation id="137C2A07FFB8FFC8FAD9FDB5FF261D49" captionStart="Fig" captionStartId="9.[189,232,1920,1946]" captionTargetBox="[219,1370,677,1872]" captionTargetId="figure-89@9.[205,1382,665,1884]" captionTargetPageId="9" captionText="Fig. 4. Hahnia ngai sp. nov., holotype, ♀ (RMNH.ARA.18415). a–c. Habitus. a. Ventral view. b. Lateral view. c. Dorsal view. d. Prosoma, anterior view. e. Chelicerae, posterior view. f–g. Epigynum. f. Dorsal view, cleared. g. Ventral view. Scale bars: a–c = 1.0 mm; d–e, g = 0.25 mm; f = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4298084" httpUri="https://zenodo.org/record/4298084/files/figure.png" pageId="9" pageNumber="60">Fig. 4c</figureCitation>
). AME 
<quantity id="4CBF9B67FFB8FFC8FEDFFDD1FE4B1D49" box="[321,433,588,614]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="60" unit="mm" value="0.04">0.04 mm</quantity>
, ALE 
<quantity id="4CBF9B67FFB8FFC8FD9AFDD1FD8F1D49" box="[516,629,588,614]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="60" unit="mm" value="0.06">0.06 mm</quantity>
, PME 
<quantity id="4CBF9B67FFB8FFC8FD53FDD1FCC71D49" box="[717,829,588,614]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="9" pageNumber="60" unit="mm" value="0.07">0.07 mm</quantity>
, PLE 
<quantity id="4CBF9B67FFB8FFC8FC13FDD1FC071D49" box="[909,1021,588,614]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="60" unit="mm" value="0.04">0.04 mm</quantity>
, AME–AME 
<quantity id="4CBF9B67FFB8FFC8FB36FDD1FAE21D49" box="[1192,1304,588,614]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="60" unit="mm" value="0.03">0.03 mm</quantity>
, AMEALE 
<quantity id="4CBF9B67FFBBFFCBFF63FE94FE931E0C" box="[253,361,265,291]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="61" unit="mm" value="0.02">0.02 mm</quantity>
, PME–PME 
<quantity id="4CBF9B67FFBBFFCBFD9DFE94FD951E0C" box="[515,623,265,292]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="61" unit="mm" value="0.05">0.05 mm</quantity>
, PME–PLE 
<quantity id="4CBF9B67FFBBFFCBFC9EFE94FC931E0C" box="[768,873,265,291]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="61" unit="mm" value="0.03">0.03 mm</quantity>
(
<figureCitation id="137C2A07FFBBFFCBFCE7FE94FC2B1E0B" box="[889,977,265,292]" captionStart="Fig" captionStartId="9.[189,232,1920,1946]" captionTargetBox="[219,1370,677,1872]" captionTargetId="figure-89@9.[205,1382,665,1884]" captionTargetPageId="9" captionText="Fig. 4. Hahnia ngai sp. nov., holotype, ♀ (RMNH.ARA.18415). a–c. Habitus. a. Ventral view. b. Lateral view. c. Dorsal view. d. Prosoma, anterior view. e. Chelicerae, posterior view. f–g. Epigynum. f. Dorsal view, cleared. g. Ventral view. Scale bars: a–c = 1.0 mm; d–e, g = 0.25 mm; f = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4298084" httpUri="https://zenodo.org/record/4298084/files/figure.png" pageId="10" pageNumber="61">Fig. 4d</figureCitation>
). Chelicerae with three promarginal and three retromarginal teeth (
<figureCitation id="137C2A07FFBBFFCBFDBAFEB1FD861E69" box="[548,636,300,326]" captionStart="Fig" captionStartId="9.[189,232,1920,1946]" captionTargetBox="[219,1370,677,1872]" captionTargetId="figure-89@9.[205,1382,665,1884]" captionTargetPageId="9" captionText="Fig. 4. Hahnia ngai sp. nov., holotype, ♀ (RMNH.ARA.18415). a–c. Habitus. a. Ventral view. b. Lateral view. c. Dorsal view. d. Prosoma, anterior view. e. Chelicerae, posterior view. f–g. Epigynum. f. Dorsal view, cleared. g. Ventral view. Scale bars: a–c = 1.0 mm; d–e, g = 0.25 mm; f = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4298084" httpUri="https://zenodo.org/record/4298084/files/figure.png" pageId="10" pageNumber="61">Fig. 4e</figureCitation>
). Legs pale brown, slightly darker on distal segments. Abdomen dark grey with light patches forming five to six chevron bands, oval, longer than wide (
<figureCitation id="137C2A07FFBBFFCBFB34FED2FB051E46" box="[1194,1279,335,361]" captionStart="Fig" captionStartId="9.[189,232,1920,1946]" captionTargetBox="[219,1370,677,1872]" captionTargetId="figure-89@9.[205,1382,665,1884]" captionTargetPageId="9" captionText="Fig. 4. Hahnia ngai sp. nov., holotype, ♀ (RMNH.ARA.18415). a–c. Habitus. a. Ventral view. b. Lateral view. c. Dorsal view. d. Prosoma, anterior view. e. Chelicerae, posterior view. f–g. Epigynum. f. Dorsal view, cleared. g. Ventral view. Scale bars: a–c = 1.0 mm; d–e, g = 0.25 mm; f = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4298084" httpUri="https://zenodo.org/record/4298084/files/figure.png" pageId="10" pageNumber="61">Fig. 4c</figureCitation>
). Tracheal spiracle near middle of abdomen (
<figureCitation id="137C2A07FFBBFFCBFDD0FEEFFD5E1EA2" box="[590,676,370,397]" captionStart="Fig" captionStartId="9.[189,232,1920,1946]" captionTargetBox="[219,1370,677,1872]" captionTargetId="figure-89@9.[205,1382,665,1884]" captionTargetPageId="9" captionText="Fig. 4. Hahnia ngai sp. nov., holotype, ♀ (RMNH.ARA.18415). a–c. Habitus. a. Ventral view. b. Lateral view. c. Dorsal view. d. Prosoma, anterior view. e. Chelicerae, posterior view. f–g. Epigynum. f. Dorsal view, cleared. g. Ventral view. Scale bars: a–c = 1.0 mm; d–e, g = 0.25 mm; f = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4298084" httpUri="https://zenodo.org/record/4298084/files/figure.png" pageId="10" pageNumber="61">Fig. 4a</figureCitation>
).
</paragraph>
<caption id="DF38660AFFB8FFC8FF23F81DFB3C18CE" ID-DOI="http://doi.org/10.5281/zenodo.4298084" ID-Zenodo-Dep="4298084" httpUri="https://zenodo.org/record/4298084/files/figure.png" pageId="9" pageNumber="60" startId="9.[189,232,1920,1946]" targetBox="[219,1370,677,1872]" targetPageId="9">
<paragraph id="8BF83682FFB8FFC8FF23F81DFB3C18CE" blockId="9.[188,1398,1920,2018]" pageId="9" pageNumber="60">
<emphasis id="B933EA90FFB8FFC8FF23F81DFEF318B5" bold="true" box="[189,265,1920,1946]" pageId="9" pageNumber="60">Fig. 4.</emphasis>
<taxonomicName id="4C474D01FFB8FFC8FE91F81DFE5B18B5" authority="Rivera-Quiroz &amp; Petcharad &amp; Miller, 2020" authorityName="Rivera-Quiroz &amp; Petcharad &amp; Miller" authorityYear="2020" box="[271,417,1920,1946]" class="Arachnida" family="Hahniidae" genus="Hahnia" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="ngai" status="sp. nov.">
<emphasis id="B933EA90FFB8FFC8FE91F81DFE5B18B5" box="[271,417,1920,1946]" italics="true" pageId="9" pageNumber="60">Hahnia ngai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20057EBFFB8FFC8FE39F81CFDF818B4" box="[423,514,1921,1947]" pageId="9" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♀ (RMNH.ARA.18415). 
<emphasis id="B933EA90FFB8FFC8FC35F81CFC2F18B4" bold="true" box="[939,981,1921,1947]" pageId="9" pageNumber="60">a–c</emphasis>
. Habitus. 
<emphasis id="B933EA90FFB8FFC8FBD4F81CFBA318B4" bold="true" box="[1098,1113,1921,1947]" pageId="9" pageNumber="60">a</emphasis>
. Ventral view. 
<emphasis id="B933EA90FFB8FFC8FA9AF81CFAEE18B4" bold="true" box="[1284,1300,1921,1947]" pageId="9" pageNumber="60">b</emphasis>
. Lateral view. 
<emphasis id="B933EA90FFB8FFC8FE9DF839FEEA1891" bold="true" box="[259,272,1956,1982]" pageId="9" pageNumber="60">c</emphasis>
. Dorsal view. 
<emphasis id="B933EA90FFB8FFC8FE27F839FE331891" bold="true" box="[441,457,1956,1982]" pageId="9" pageNumber="60">d</emphasis>
. Prosoma, anterior view. 
<emphasis id="B933EA90FFB8FFC8FD6BF839FCF81891" bold="true" box="[757,770,1956,1982]" pageId="9" pageNumber="60">e</emphasis>
. Chelicerae, posterior view. 
<emphasis id="B933EA90FFB8FFC8FBCFF83EFB821892" bold="true" box="[1105,1144,1955,1981]" pageId="9" pageNumber="60">f–g</emphasis>
. Epigynum. 
<emphasis id="B933EA90FFB8FFC8FA8EF83EFAE01892" bold="true" box="[1296,1306,1955,1981]" pageId="9" pageNumber="60">f</emphasis>
. Dorsal view, cleared. 
<emphasis id="B933EA90FFB8FFC8FEF8F85AFE8F18CE" bold="true" box="[358,373,1991,2017]" pageId="9" pageNumber="60">g</emphasis>
. Ventral view. Scale bars: a–c = 1.0 mm; d–e, g = 0.25 mm; f = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BF83682FFBBFFCBFF23FE28FEFC1D17" blockId="10.[188,1398,436,568]" pageId="10" pageNumber="61">
VULVA. Epigynal plate semitransparent, spermathecae well visible due to transparency. Copulatory openings close together, forming small semi-circular atrium (
<figureCitation id="137C2A07FFBBFFCBFC16FE4AFC281EDE" box="[904,978,471,497]" captionStart="Fig" captionStartId="9.[189,232,1920,1946]" captionTargetBox="[219,1370,677,1872]" captionTargetId="figure-89@9.[205,1382,665,1884]" captionTargetPageId="9" captionText="Fig. 4. Hahnia ngai sp. nov., holotype, ♀ (RMNH.ARA.18415). a–c. Habitus. a. Ventral view. b. Lateral view. c. Dorsal view. d. Prosoma, anterior view. e. Chelicerae, posterior view. f–g. Epigynum. f. Dorsal view, cleared. g. Ventral view. Scale bars: a–c = 1.0 mm; d–e, g = 0.25 mm; f = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4298084" httpUri="https://zenodo.org/record/4298084/files/figure.png" pageId="10" pageNumber="61">Figs 4</figureCitation>
f–g, 6c). Spermatheca sub-spherical with brownish-red coloration (
<figureCitation id="137C2A07FFBBFFCBFD81FE67FD881D3A" box="[543,626,506,533]" captionStart="Fig" captionStartId="9.[189,232,1920,1946]" captionTargetBox="[219,1370,677,1872]" captionTargetId="figure-89@9.[205,1382,665,1884]" captionTargetPageId="9" captionText="Fig. 4. Hahnia ngai sp. nov., holotype, ♀ (RMNH.ARA.18415). a–c. Habitus. a. Ventral view. b. Lateral view. c. Dorsal view. d. Prosoma, anterior view. e. Chelicerae, posterior view. f–g. Epigynum. f. Dorsal view, cleared. g. Ventral view. Scale bars: a–c = 1.0 mm; d–e, g = 0.25 mm; f = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4298084" httpUri="https://zenodo.org/record/4298084/files/figure.png" pageId="10" pageNumber="61">Fig. 4f</figureCitation>
). Copulatory ducts very simple, slightly swollen centrally (
<figureCitation id="137C2A07FFBBFFCBFA82FE67FA881D3B" box="[1308,1394,506,532]" captionStart="Fig" captionStartId="9.[189,232,1920,1946]" captionTargetBox="[219,1370,677,1872]" captionTargetId="figure-89@9.[205,1382,665,1884]" captionTargetPageId="9" captionText="Fig. 4. Hahnia ngai sp. nov., holotype, ♀ (RMNH.ARA.18415). a–c. Habitus. a. Ventral view. b. Lateral view. c. Dorsal view. d. Prosoma, anterior view. e. Chelicerae, posterior view. f–g. Epigynum. f. Dorsal view, cleared. g. Ventral view. Scale bars: a–c = 1.0 mm; d–e, g = 0.25 mm; f = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4298084" httpUri="https://zenodo.org/record/4298084/files/figure.png" pageId="10" pageNumber="61">Figs 4f</figureCitation>
, 
<figureCitation id="137C2A07FFBBFFCBFF22FD80FF311D17" box="[188,203,541,568]" captionStart="Fig" captionStartId="12.[189,232,1888,1914]" captionTargetBox="[189,1396,639,1811]" captionTargetId="figure-148@12.[189,1398,630,1824]" captionTargetPageId="12" captionText="Fig. 6. Female spinnerets and genitals. a–c. Hahnia ngai sp. nov., holotype (RMNH.ARA.18415). a. Spinnerets, ventral view. b. Epigynum, cleared, dorsal view. c. Ventral view. d–f. Hahnia saccata Zhang, Li &amp; Zheng, 2011 (RMNH.ARA.18412). d. Spinnerets, ventral view. e. Epigynum, cleared, dorsal view. f. Ventral view. Scale bars: a, d–f = 0.25 mm; b–c = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4298088" httpUri="https://zenodo.org/record/4298088/files/figure.png" pageId="10" pageNumber="61">6</figureCitation>
b–c).
</paragraph>
<paragraph id="8BF83682FFBBFFCBFF23FDFDFCEA1C29" blockId="10.[188,1401,607,775]" pageId="10" pageNumber="61">MEASUREMENTS (in mm). Total length 2.8, carapace 1.25 long, 0.91 wide; clypeus 0.09; chelicera 0.45 long, 0.25 wide; leg I: femur 0.95, patella 0.37, tibia 0.71, metatarsus 0.72, tarsus 0.55; leg II: femur 0.94, patella 0.34, tibia 0.72, metatarsus 0.68, tarsus 0.55; leg III: femur 0.89, patella 0.33, tibia 0.63, metatarsus 0.71, tarsus 0.51; leg IV: femur 1.12, patella 0.34, tibia 0.93, metatarsus 1.01, tarsus 0.62; leg formula IV-I-II-III; abdomen 1.65 long, 1.23 wide.</paragraph>
</subSubSection>
<subSubSection id="C35D6509FFBBFFCBFF23FCB0FD061CB6" pageId="10" pageNumber="61" type="distribution">
<paragraph id="8BF83682FFBBFFCBFF23FCB0FEAC1C68" blockId="10.[189,342,813,839]" box="[189,342,813,839]" pageId="10" pageNumber="61">
<heading id="D0B081EEFFBBFFCBFF23FCB0FEAC1C68" bold="true" box="[189,342,813,839]" fontSize="11" level="3" pageId="10" pageNumber="61" reason="3">
<emphasis id="B933EA90FFBBFFCBFF23FCB0FEAC1C68" bold="true" box="[189,342,813,839]" pageId="10" pageNumber="61">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BF83682FFBBFFCBFF23FCC1FD061CB6" blockId="10.[189,1398,860,921]" pageId="10" pageNumber="61">
Known from two localities in 
<collectingRegion id="4983F860FFBBFFCBFDBEFCC1FD541C59" box="[544,686,860,886]" country="Thailand" name="Chiang Mai" pageId="10" pageNumber="61">Chiang Mai</collectingRegion>
, 
<collectingCountry id="F3507612FFBBFFCBFD24FCC1FCD81C59" box="[698,802,860,886]" name="Thailand" pageId="10" pageNumber="61">Thailand</collectingCountry>
(
<figureCitation id="137C2A07FFBBFFCBFCADFCC1FC811C59" box="[819,891,860,886]" captionStart="Fig" captionStartId="14.[189,232,1884,1910]" captionTargetBox="[417,1187,916,1830]" captionTargetId="figure-236@14.[400,1187,904,1845]" captionTargetPageId="14" captionText="Fig. 8. Map of mainland Southeast Asia, showing the collecting sites of Zhang et al. (2011) (Hahnia saccata Zhang, Li &amp; Zheng, 2011 and Hexamatia senaria (Zhang, Li &amp; Zheng, 2011) gen. et comb. nov.), circle; and our new hahniid specimens (Hexamatia seekhaow gen. et sp. nov., Hahnia ngai sp. nov. and Hahnia saccata), square." figureDoi="http://doi.org/10.5281/zenodo.4298092" httpUri="https://zenodo.org/record/4298092/files/figure.png" pageId="10" pageNumber="61">Fig. 8</figureCitation>
): Doi Suthep National Park (
<typeStatus id="54FC8820FFBBFFCBFB4FFCC1FAFF1C59" box="[1233,1285,860,886]" pageId="10" pageNumber="61">type</typeStatus>
locality), and the neighboring Doi Inthanon National Park.
</paragraph>
</subSubSection>
</treatment>
</document>