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<document id="A69411E943CE0C9CB262973A592580ED" ID-DOI="10.1649/0010-065X(2005)059[0091:LDOESO]2.0.CO;2" ID-Zenodo-Dep="10104898" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630171519151" checkinUser="tatiana" docAuthor="Ratcliffe, Brett C. &amp; Morón, Miguel-Angel" docDate="2005" docId="03ED726E8A183B0BF2FCFA32BB6BFD4E" docLanguage="en" docName="0010-065X_2005_059_0091_LDOESO_2.0.CO_2.pdf.imf" docOrigin="The Coleopterists Bulletin 59 (1)" docSource="http://dx.doi.org/10.1649/0010-065x(2005)059[0091:ldoeso]2.0.co;2" docStyle="DocumentStyle:8F272F586C426DAD54565EB2271A6C17.2:TheColeopBull.2000-2002.journal_article.1cover" docStyleId="8F272F586C426DAD54565EB2271A6C17" docStyleName="TheColeopBull.2000-2002.journal_article.1cover" docStyleVersion="2" docTitle="Megasoma thersites LeConte" docType="treatment" docVersion="3" lastPageNumber="104" masterDocId="FFD40A168A103B06F37CFF9DB947FF9E" masterDocTitle="Larval Descriptions Of Eight Species Of Megasoma Kirby (Coleoptera: Scarabaeidae: Dynastinae) With A Key For Identification And Notes On Biology" masterLastPageNumber="126" masterPageNumber="91" pageNumber="99" updateTime="1699631449810" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8288E2DB73114C0FE0575096C284A329">Larval Descriptions Of Eight Species Of Megasoma Kirby (Coleoptera: Scarabaeidae: Dynastinae) With A Key For Identification And Notes On Biology</mods:title>
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<mods:namePart id="2AF1459FF3926D7E44E616F92F531AF8">Ratcliffe, Brett C.</mods:namePart>
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<mods:namePart id="535013ADC6F9BFF9C1711822E509C183">Morón, Miguel-Angel</mods:namePart>
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<mods:title id="D6F5C0CC23E23832BDB0B82DF6CB050D">The Coleopterists Bulletin</mods:title>
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<mods:date id="C14F29CDD4122A4B130A891FE8B11BE1">2005</mods:date>
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<treatment id="03ED726E8A183B0BF2FCFA32BB6BFD4E" LSID="urn:lsid:plazi:treatment:03ED726E8A183B0BF2FCFA32BB6BFD4E" httpUri="http://treatment.plazi.org/id/03ED726E8A183B0BF2FCFA32BB6BFD4E" lastPageId="13" lastPageNumber="104" pageId="8" pageNumber="99">
<subSubSection id="C35E90F38A183B0EF2FCFA32BBC8FA40" pageId="8" pageNumber="99" type="nomenclature">
<paragraph id="8BFBC3788A183B0EF2FCFA32BBC8FA40" blockId="8.[384,796,1455,1502]" pageId="8" pageNumber="99">
<emphasis id="B9301F6A8A183B0EF2FCFA32BA5BFA5A" bold="true" box="[384,796,1455,1476]" pageId="8" pageNumber="99">
<taxonomicName id="4C44B8FB8A183B0EF2FCFA32BBE0FA5A" authority="LeConte" authorityName="LeConte" box="[384,679,1455,1476]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="99" phylum="Arthropoda" rank="species" species="thersites">
<emphasis id="B9301F6A8A183B0EF2FCFA32BB01FA5A" bold="true" box="[384,582,1455,1476]" italics="true" pageId="8" pageNumber="99">Megasoma thersites</emphasis>
LeConte
</taxonomicName>
, 3rd instar
</emphasis>
(
<figureCitation id="137FDFFD8A183B0EF168FA54BBCEFA40" box="[532,649,1481,1502]" captionStart-0="Figs" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="9.[169,208,1274,1293]" captionStartId-1="10.[197,228,1429,1448]" captionStartId-2="11.[169,200,1199,1218]" captionTargetBox-0="[149,980,194,1252]" captionTargetBox-1="[178,1001,194,1408]" captionTargetBox-2="[151,974,194,1178]" captionTargetId-0="figure-58@9.[148,980,194,1253]" captionTargetId-1="figure-1@10.[178,1002,194,1408]" captionTargetId-2="figure-166@11.[151,975,194,1178]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="Figs. 2940. Megasoma thersites, third instar. 29) Head, frontal view; 30) epipharynx; 3132) mandibles, ventral view, 31) right and 32) left; 33) distal part of right maxillary mala; 34) detail of maxillary stridulatory area; 35) right maxilla and hypopharynx, dorsal view; 3637) distal segment of antenna, 36) dorsal view and 37) ventral view; 38) left posterior tarsal claw, external view; 39) second right abdominal spiracle; 40) microstructure of respiratory plate (5003). Figs. 2932, 35, bar ¼ 1 mm. Figs. 3334, 3639, bar ¼ 0.5 mm." captionText-1="Fig. 41. Megasoma thersites on palo verde, Cercidium microphyllum Rose &amp; I. M. Johnst. Photo by Barney Streit." captionText-2="Fig. 42. Gumming wound made by adult Megasoma thersites on palo verde tree. Photo by Barney Streit." figureDoi-0="http://doi.org/10.5281/zenodo.10104910" figureDoi-1="http://doi.org/10.5281/zenodo.10104912" figureDoi-2="http://doi.org/10.5281/zenodo.10104916" httpUri-0="https://zenodo.org/record/10104910/files/figure.png" httpUri-1="https://zenodo.org/record/10104912/files/figure.png" httpUri-2="https://zenodo.org/record/10104916/files/figure.png" pageId="8" pageNumber="99">Figs. 2942</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35E90F38A183B0BF3BBFA6CBA80FE0F" lastPageId="13" lastPageNumber="104" pageId="8" pageNumber="99" type="description">
<paragraph id="8BFBC3788A183B0EF3BBFA6CBAAAF9BF" blockId="8.[175,1005,1521,1569]" pageId="8" pageNumber="99">
Description based on one third instar larva collected in
<collectingCountry id="F35383E88A183B0EF1BFFA6CBA49F998" box="[707,782,1521,1542]" name="Mexico" pageId="8" pageNumber="99">México</collectingCountry>
:
<collectingRegion id="49800D9A8A183B0EF06AFA6CBA8BF998" box="[790,972,1521,1542]" country="Mexico" name="Baja California Sur" pageId="8" pageNumber="99">Baja California Sur</collectingRegion>
,
<quantity id="4CBC6E9D8A183B0EF0A8FA6CB98FF9BF" metricMagnitude="4" metricUnit="m" metricValue="1.7702784" pageId="8" pageNumber="99" unit="mi" value="11.0">11 mi</quantity>
N Cabo San Lucas,
<date id="FFFAE5B88A183B0EF2F5F991B8BCF9BF" box="[393,507,1548,1569]" pageId="8" pageNumber="99" value="1985-04-26">26-IV-1985</date>
, from soil dug for post hole, R.L. Westcott (UNSM).
</paragraph>
<paragraph id="8BFBC3788A193B0FF3E8FF1DBA4DFF0D" blockId="9.[148,180,128,147]" box="[148,778,128,147]" lastBlockId="9.[347,778,128,147]" pageId="9" pageNumber="100">100 THE COLEOPTERISTS BULLETIN 59(1), 2005</paragraph>
<caption id="DF3B93F08A193B0FF3D5FB67BA53FA1B" ID-DOI="http://doi.org/10.5281/zenodo.10104910" ID-Zenodo-Dep="10104910" httpUri="https://zenodo.org/record/10104910/files/figure.png" pageId="9" pageNumber="100" startId="9.[169,208,1274,1293]" targetBox="[149,980,194,1252]" targetPageId="9">
<paragraph id="8BFBC3788A193B0FF3D5FB67BA53FA1B" blockId="9.[148,978,1274,1413]" pageId="9" pageNumber="100">
<emphasis id="B9301F6A8A193B0FF3D5FB67B85DFA93" bold="true" box="[169,282,1274,1293]" pageId="9" pageNumber="100">Figs. 2940.</emphasis>
<taxonomicName id="4C44B8FB8A193B0FF253FB67B899FA93" box="[303,478,1274,1293]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="100" phylum="Arthropoda" rank="species" species="thersites">
<emphasis id="B9301F6A8A193B0FF253FB67B899FA93" box="[303,478,1274,1293]" italics="true" pageId="9" pageNumber="100">Megasoma thersites</emphasis>
</taxonomicName>
, third instar.
<emphasis id="B9301F6A8A193B0FF11EFB67BB39FA93" bold="true" box="[610,638,1274,1293]" pageId="9" pageNumber="100">29)</emphasis>
Head, frontal view;
<emphasis id="B9301F6A8A193B0FF039FB67BA26FA93" bold="true" box="[837,865,1274,1293]" pageId="9" pageNumber="100">30)</emphasis>
epipharynx;
<emphasis id="B9301F6A8A193B0FF3E8FA8FB997FABB" bold="true" box="[148,208,1298,1317]" pageId="9" pageNumber="100">3132)</emphasis>
mandibles, ventral view,
<emphasis id="B9301F6A8A193B0FF2C9FA8FB896FABB" bold="true" box="[437,465,1298,1317]" pageId="9" pageNumber="100">31)</emphasis>
right and
<emphasis id="B9301F6A8A193B0FF14CFA8FBB0BFABB" bold="true" box="[560,588,1298,1317]" pageId="9" pageNumber="100">32)</emphasis>
left;
<emphasis id="B9301F6A8A193B0FF102FA8FBBDDFABB" bold="true" box="[638,666,1298,1317]" pageId="9" pageNumber="100">33)</emphasis>
distal part of right maxillary mala;
<emphasis id="B9301F6A8A193B0FF3E8FAB4B9F7FAA2" bold="true" box="[148,176,1321,1340]" pageId="9" pageNumber="100">34)</emphasis>
detail of maxillary stridulatory area;
<emphasis id="B9301F6A8A193B0FF28CFAB4BB4BFAA2" bold="true" box="[496,524,1321,1340]" pageId="9" pageNumber="100">35)</emphasis>
right maxilla and hypopharynx, dorsal view;
<emphasis id="B9301F6A8A193B0FF0E9FAB4BA95FAA2" bold="true" box="[917,978,1321,1340]" pageId="9" pageNumber="100">3637)</emphasis>
distal segment of antenna,
<emphasis id="B9301F6A8A193B0FF2FCFADCB8DBFACA" bold="true" box="[384,412,1345,1364]" pageId="9" pageNumber="100">36)</emphasis>
dorsal view and
<emphasis id="B9301F6A8A193B0FF14BFADCBB13FACA" bold="true" box="[567,596,1345,1364]" pageId="9" pageNumber="100">37)</emphasis>
ventral view;
<emphasis id="B9301F6A8A193B0FF1A9FADCBBB6FACA" bold="true" box="[725,753,1345,1364]" pageId="9" pageNumber="100">38)</emphasis>
left posterior tarsal claw, external view;
<emphasis id="B9301F6A8A193B0FF262FAC4B87DFAF2" bold="true" box="[286,314,1369,1388]" pageId="9" pageNumber="100">39)</emphasis>
second right abdominal spiracle;
<emphasis id="B9301F6A8A193B0FF10BFAC4BBD4FAF2" bold="true" box="[631,659,1369,1388]" pageId="9" pageNumber="100">40)</emphasis>
microstructure of respiratory plate (500
<emphasis id="B9301F6A8A193B0FF3C4FAECB980FA1A" bold="true" box="[184,199,1393,1412]" pageId="9" pageNumber="100">3</emphasis>
). Figs. 2932, 35, bar
<emphasis id="B9301F6A8A193B0FF2F2FAEDB8D9FA1B" bold="true" box="[398,414,1392,1413]" italics="true" pageId="9" pageNumber="100">¼</emphasis>
1 mm. Figs. 3334, 3639, bar
<emphasis id="B9301F6A8A193B0FF1C9FAEDBB82FA1B" bold="true" box="[693,709,1392,1413]" italics="true" pageId="9" pageNumber="100">¼</emphasis>
0.5 mm.
</paragraph>
</caption>
<paragraph id="8BFBC3788A193B0FF3D5FA5ABA96F9BC" blockId="9.[148,977,1479,1570]" pageId="9" pageNumber="100">
<emphasis id="B9301F6A8A193B0FF3D5FA5AB850FA44" bold="true" box="[169,279,1479,1498]" pageId="9" pageNumber="100">Description.</emphasis>
<emphasis id="B9301F6A8A193B0FF25EFA5AB811FA44" box="[290,342,1479,1498]" italics="true" pageId="9" pageNumber="100">Head.</emphasis>
Maximum width of head capsule
<quantity id="4CBC6E9D8A193B0FF1F9FA5ABB8CFA44" box="[645,715,1479,1498]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="9" pageNumber="100" unit="mm" value="7.4">7.4 mm</quantity>
. Surface of cranium (
<figureCitation id="137FDFFD8A193B0FF0F7FA5ABA8CFA44" box="[907,971,1479,1498]" captionStart="Figs" captionStartId="9.[169,208,1274,1293]" captionTargetBox="[149,980,194,1252]" captionTargetId="figure-58@9.[148,980,194,1253]" captionTargetPageId="9" captionText="Figs. 2940. Megasoma thersites, third instar. 29) Head, frontal view; 30) epipharynx; 3132) mandibles, ventral view, 31) right and 32) left; 33) distal part of right maxillary mala; 34) detail of maxillary stridulatory area; 35) right maxilla and hypopharynx, dorsal view; 3637) distal segment of antenna, 36) dorsal view and 37) ventral view; 38) left posterior tarsal claw, external view; 39) second right abdominal spiracle; 40) microstructure of respiratory plate (5003). Figs. 2932, 35, bar ¼ 1 mm. Figs. 3334, 3639, bar ¼ 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10104910" httpUri="https://zenodo.org/record/10104910/files/figure.png" pageId="9" pageNumber="100">Fig. 29</figureCitation>
) dark brown, punctate, punctures large, deep. Frons moderately punctate, each side with 3 posterior frontal setae, anterior frontal setae lacking, 1 exterior frontal setae, and 6 anterior angle frontal setae. Remaining cranial surface with 2 dorsoepicranial, short setae, 46 epicranial setae, and 3
</paragraph>
<caption id="DF3B93F08A1A3B0CF3B9FA08B83BFA5E" ID-DOI="http://doi.org/10.5281/zenodo.10104912" ID-Zenodo-Dep="10104912" httpUri="https://zenodo.org/record/10104912/files/figure.png" pageId="10" pageNumber="101" startId="10.[197,228,1429,1448]" targetBox="[178,1001,194,1408]" targetPageId="10">
<paragraph id="8BFBC3788A1A3B0CF3B9FA08BAA9FA36" blockId="10.[175,1006,1429,1472]" box="[197,1006,1429,1448]" pageId="10" pageNumber="101">
<emphasis id="B9301F6A8A1A3B0CF3B9FA08B84DFA36" bold="true" box="[197,266,1429,1448]" pageId="10" pageNumber="101">Fig. 41.</emphasis>
<taxonomicName id="4C44B8FB8A1A3B0CF263FA08B88DFA36" box="[287,458,1429,1448]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="thersites">
<emphasis id="B9301F6A8A1A3B0CF263FA08B88DFA36" box="[287,458,1429,1448]" italics="true" pageId="10" pageNumber="101">Megasoma thersites</emphasis>
</taxonomicName>
on palo verde,
<taxonomicName id="4C44B8FB8A1A3B0CF12AFA08BAA9FA36" authority="Rose &amp; I. M. Johnst." box="[598,1006,1429,1448]" class="Magnoliopsida" family="Fabaceae" genus="Cercidium" kingdom="Plantae" order="Fabales" pageId="10" pageNumber="101" phylum="Tracheophyta" rank="species" species="microphyllum">
<emphasis id="B9301F6A8A1A3B0CF12AFA08BA6BFA36" box="[598,812,1429,1448]" italics="true" pageId="10" pageNumber="101">Cercidium microphyllum</emphasis>
Rose &amp; I. M. Johnst.
</taxonomicName>
</paragraph>
<paragraph id="8BFBC3788A1A3B0CF3D3FA30B83BFA5E" blockId="10.[175,1006,1429,1472]" box="[175,380,1453,1472]" pageId="10" pageNumber="101">Photo by Barney Streit.</paragraph>
</caption>
<paragraph id="8BFBC3788A1B3B0DF3E8FF1DBA4DFF0D" blockId="11.[148,180,128,147]" box="[148,778,128,147]" lastBlockId="11.[347,778,128,147]" pageId="11" pageNumber="102">102 THE COLEOPTERISTS BULLETIN 59(1), 2005</paragraph>
<caption id="DF3B93F08A1B3B0DF3D5FB32B84CFB45" ID-DOI="http://doi.org/10.5281/zenodo.10104916" ID-Zenodo-Dep="10104916" httpUri="https://zenodo.org/record/10104916/files/figure.png" pageId="11" pageNumber="102" startId="11.[169,200,1199,1218]" targetBox="[151,974,194,1178]" targetPageId="11">
<paragraph id="8BFBC3788A1B3B0DF3D5FB32BA97FB5D" blockId="11.[148,976,1199,1243]" box="[169,976,1199,1219]" pageId="11" pageNumber="102">
<emphasis id="B9301F6A8A1B3B0DF3D5FB32B9ABFB5C" bold="true" box="[169,236,1199,1218]" pageId="11" pageNumber="102">Fig. 42.</emphasis>
Gumming wound made by adult
<taxonomicName id="4C44B8FB8A1B3B0DF14EFB2DBB9CFB5D" box="[562,731,1200,1219]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="102" phylum="Arthropoda" rank="species" species="thersites">
<emphasis id="B9301F6A8A1B3B0DF14EFB2DBB9CFB5D" box="[562,731,1200,1219]" italics="true" pageId="11" pageNumber="102">Megasoma thersites</emphasis>
</taxonomicName>
on palo verde tree. Photo by
</paragraph>
<paragraph id="8BFBC3788A1B3B0DF3E8FB55B84CFB45" blockId="11.[148,976,1199,1243]" box="[148,267,1224,1243]" pageId="11" pageNumber="102">Barney Streit.</paragraph>
</caption>
<paragraph id="8BFBC3788A1B3B0AF3E8FA9BBAABFF73" blockId="11.[148,978,1286,1568]" lastBlockId="12.[175,1005,194,237]" lastPageId="12" lastPageNumber="103" pageId="11" pageNumber="102">
paraocellar setae on each side. Clypeus trapezoidal, with 2 lateral setae on each side and 4 posterior clypeal setae. Labrum slightly asymmetrical, subrectangular, with row of 15 anterior labral setae, widely interrupted at middle, 7 posterior setae, central setae absent, and 56 lateral setae. Ocelli not apparent. Epipharynx (
<figureCitation id="137FDFFD8A1B3B0DF299FAD3BB64FAFF" box="[485,547,1358,1377]" captionStart="Figs" captionStartId="9.[169,208,1274,1293]" captionTargetBox="[149,980,194,1252]" captionTargetId="figure-58@9.[148,980,194,1253]" captionTargetPageId="9" captionText="Figs. 2940. Megasoma thersites, third instar. 29) Head, frontal view; 30) epipharynx; 3132) mandibles, ventral view, 31) right and 32) left; 33) distal part of right maxillary mala; 34) detail of maxillary stridulatory area; 35) right maxilla and hypopharynx, dorsal view; 3637) distal segment of antenna, 36) dorsal view and 37) ventral view; 38) left posterior tarsal claw, external view; 39) second right abdominal spiracle; 40) microstructure of respiratory plate (5003). Figs. 2932, 35, bar ¼ 1 mm. Figs. 3334, 3639, bar ¼ 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10104910" httpUri="https://zenodo.org/record/10104910/files/figure.png" pageId="11" pageNumber="102">Fig. 30</figureCitation>
) with fused zygum and epizygum forming weakly raised, subconical, sclerotized tubercle on haptomerum; heli and plegmatia absent; right chaetoparia with 50 spine-like setae and 45 small setae and sensillae toward gymnoparia; left chaetoparia with 35 spine-like setae and 40 small setae and sensillae; acroparia with 9 straight, long, thick setae toward right side and 10 straight, long, thick setae toward left side; right acanthoparia with 12 short, curved, spine-like setae or insertion pits; left acanthoparia with 13 short, curved, spine-like setae or insertion pits; pedium wide and epitorma not defined; dexiotorma long, sinuate, with rounded inner side; laeotorma short, with rounded inner side and subquadrate pternotorma; dexiophoba sparse, formed by 11 setae; laeophoba between haptolachus and inner side of laeotorma formed by 32 setae; sclerotized plate of right nesium large, subpyriform; sense cone of left nesium represented by large, wide, longitudinal, well-sclerotized
</paragraph>
<paragraph id="8BFBC3788A1C3B0AF3D3FF6FBA5AFA97" blockId="12.[175,1006,242,1289]" pageId="12" pageNumber="103">
plate with rounded ends and row of 5 sensillae near base; crepis not defined. Scissorial area of right mandible (
<figureCitation id="137FDFFD8A1C3B0AF244FE97B830FE83" box="[312,375,266,285]" captionStart="Figs" captionStartId="9.[169,208,1274,1293]" captionTargetBox="[149,980,194,1252]" captionTargetId="figure-58@9.[148,980,194,1253]" captionTargetPageId="9" captionText="Figs. 2940. Megasoma thersites, third instar. 29) Head, frontal view; 30) epipharynx; 3132) mandibles, ventral view, 31) right and 32) left; 33) distal part of right maxillary mala; 34) detail of maxillary stridulatory area; 35) right maxilla and hypopharynx, dorsal view; 3637) distal segment of antenna, 36) dorsal view and 37) ventral view; 38) left posterior tarsal claw, external view; 39) second right abdominal spiracle; 40) microstructure of respiratory plate (5003). Figs. 2932, 35, bar ¼ 1 mm. Figs. 3334, 3639, bar ¼ 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10104910" httpUri="https://zenodo.org/record/10104910/files/figure.png" pageId="12" pageNumber="103">Fig. 31</figureCitation>
) widely abraded, without evidence of teeth or notches; molar area with 3 lobes, calx prominent, brustia sparse. Scissorial area of left mandible (
<figureCitation id="137FDFFD8A1C3B0AF06BFEBFBA10FEAB" box="[791,855,290,309]" captionStart="Figs" captionStartId="9.[169,208,1274,1293]" captionTargetBox="[149,980,194,1252]" captionTargetId="figure-58@9.[148,980,194,1253]" captionTargetPageId="9" captionText="Figs. 2940. Megasoma thersites, third instar. 29) Head, frontal view; 30) epipharynx; 3132) mandibles, ventral view, 31) right and 32) left; 33) distal part of right maxillary mala; 34) detail of maxillary stridulatory area; 35) right maxilla and hypopharynx, dorsal view; 3637) distal segment of antenna, 36) dorsal view and 37) ventral view; 38) left posterior tarsal claw, external view; 39) second right abdominal spiracle; 40) microstructure of respiratory plate (5003). Figs. 2932, 35, bar ¼ 1 mm. Figs. 3334, 3639, bar ¼ 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10104910" httpUri="https://zenodo.org/record/10104910/files/figure.png" pageId="12" pageNumber="103">Fig. 32</figureCitation>
) deeply abraded, without evidence of teeth or notches; molar area with 2 wide lobes, brustia sparse, long acia with rounded apex. Stridulatory area of each mandible well-marked by 2224 fine striae. Mala (
<figureCitation id="137FDFFD8A1C3B0AF0B6FECFB982FEE3" captionStart="Figs" captionStartId="9.[169,208,1274,1293]" captionTargetBox="[149,980,194,1252]" captionTargetId="figure-58@9.[148,980,194,1253]" captionTargetPageId="9" captionText="Figs. 2940. Megasoma thersites, third instar. 29) Head, frontal view; 30) epipharynx; 3132) mandibles, ventral view, 31) right and 32) left; 33) distal part of right maxillary mala; 34) detail of maxillary stridulatory area; 35) right maxilla and hypopharynx, dorsal view; 3637) distal segment of antenna, 36) dorsal view and 37) ventral view; 38) left posterior tarsal claw, external view; 39) second right abdominal spiracle; 40) microstructure of respiratory plate (5003). Figs. 2932, 35, bar ¼ 1 mm. Figs. 3334, 3639, bar ¼ 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10104910" httpUri="https://zenodo.org/record/10104910/files/figure.png" pageId="12" pageNumber="103">Fig. 33</figureCitation>
) with apical uncus of galea well-developed and 3 conical unci fused at their bases at apex of lacinia (2 of these abraded); maxillary stridulatory area (
<figureCitation id="137FDFFD8A1C3B0AF1DEFE1FBBA5FE0B" box="[674,738,386,405]" captionStart="Figs" captionStartId="9.[169,208,1274,1293]" captionTargetBox="[149,980,194,1252]" captionTargetId="figure-58@9.[148,980,194,1253]" captionTargetPageId="9" captionText="Figs. 2940. Megasoma thersites, third instar. 29) Head, frontal view; 30) epipharynx; 3132) mandibles, ventral view, 31) right and 32) left; 33) distal part of right maxillary mala; 34) detail of maxillary stridulatory area; 35) right maxilla and hypopharynx, dorsal view; 3637) distal segment of antenna, 36) dorsal view and 37) ventral view; 38) left posterior tarsal claw, external view; 39) second right abdominal spiracle; 40) microstructure of respiratory plate (5003). Figs. 2932, 35, bar ¼ 1 mm. Figs. 3334, 3639, bar ¼ 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10104910" httpUri="https://zenodo.org/record/10104910/files/figure.png" pageId="12" pageNumber="103">Fig. 34</figureCitation>
) with 7 small, subtrapezoidal, rounded teeth and wide, truncate anterior process. Labium (
<figureCitation id="137FDFFD8A1C3B0AF1CFFE07BBB5FE33" box="[691,754,410,429]" captionStart="Figs" captionStartId="9.[169,208,1274,1293]" captionTargetBox="[149,980,194,1252]" captionTargetId="figure-58@9.[148,980,194,1253]" captionTargetPageId="9" captionText="Figs. 2940. Megasoma thersites, third instar. 29) Head, frontal view; 30) epipharynx; 3132) mandibles, ventral view, 31) right and 32) left; 33) distal part of right maxillary mala; 34) detail of maxillary stridulatory area; 35) right maxilla and hypopharynx, dorsal view; 3637) distal segment of antenna, 36) dorsal view and 37) ventral view; 38) left posterior tarsal claw, external view; 39) second right abdominal spiracle; 40) microstructure of respiratory plate (5003). Figs. 2932, 35, bar ¼ 1 mm. Figs. 3334, 3639, bar ¼ 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10104910" httpUri="https://zenodo.org/record/10104910/files/figure.png" pageId="12" pageNumber="103">Fig. 35</figureCitation>
) with numerous long, strong setae located on sides and short, spine-like setae at middle and proximal border; hypopharynx with right anterior lateral process of hypopharyngeal sclerome strongly developed, raised and curved, heavily sclerotized; left hypopharyngeal lateral lobe with patch of 20 posterolateral short setae and a line of 9 minute, thin setae. Last antennal segment on dorsal surface (
<figureCitation id="137FDFFD8A1C3B0AF016FE64BAEEFD92" box="[874,937,505,524]" captionStart="Figs" captionStartId="9.[169,208,1274,1293]" captionTargetBox="[149,980,194,1252]" captionTargetId="figure-58@9.[148,980,194,1253]" captionTargetPageId="9" captionText="Figs. 2940. Megasoma thersites, third instar. 29) Head, frontal view; 30) epipharynx; 3132) mandibles, ventral view, 31) right and 32) left; 33) distal part of right maxillary mala; 34) detail of maxillary stridulatory area; 35) right maxilla and hypopharynx, dorsal view; 3637) distal segment of antenna, 36) dorsal view and 37) ventral view; 38) left posterior tarsal claw, external view; 39) second right abdominal spiracle; 40) microstructure of respiratory plate (5003). Figs. 2932, 35, bar ¼ 1 mm. Figs. 3334, 3639, bar ¼ 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10104910" httpUri="https://zenodo.org/record/10104910/files/figure.png" pageId="12" pageNumber="103">Fig. 36</figureCitation>
) with 7 sensory spots; ventral surface with 6 spots (
<figureCitation id="137FDFFD8A1C3B0AF146FD8CBB39FDBA" box="[570,638,529,548]" captionStart="Figs" captionStartId="9.[169,208,1274,1293]" captionTargetBox="[149,980,194,1252]" captionTargetId="figure-58@9.[148,980,194,1253]" captionTargetPageId="9" captionText="Figs. 2940. Megasoma thersites, third instar. 29) Head, frontal view; 30) epipharynx; 3132) mandibles, ventral view, 31) right and 32) left; 33) distal part of right maxillary mala; 34) detail of maxillary stridulatory area; 35) right maxilla and hypopharynx, dorsal view; 3637) distal segment of antenna, 36) dorsal view and 37) ventral view; 38) left posterior tarsal claw, external view; 39) second right abdominal spiracle; 40) microstructure of respiratory plate (5003). Figs. 2932, 35, bar ¼ 1 mm. Figs. 3334, 3639, bar ¼ 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10104910" httpUri="https://zenodo.org/record/10104910/files/figure.png" pageId="12" pageNumber="103">Fig. 37</figureCitation>
).
<emphasis id="B9301F6A8A1C3B0AF1EDFD8CBB89FDBA" box="[657,718,529,548]" italics="true" pageId="12" pageNumber="103">Thorax</emphasis>
. Pronotum with wide, irregular, reddish brown, lateral scleromes and 140 long, slender setae near anterior border and 44 medium sized, slender setae near posterior border, setae not arranged in definite rows. Mesonotum with 3 transverse groups of 2090 long, slender setae and 4 groups of 1848 medium sized, slender setae; metanotum with 4 transverse groups of 3580 long, slender setae, 3 groups of 1022 medium sized, slender setae and 1 group of 20 short, spine-like setae, setae not arranged in rows. Thoracic spiracles
<quantity id="4CBC6E9D8A1C3B0AF233FD3CB8DCFD2A" box="[335,411,673,692]" metricMagnitude="-4" metricUnit="m" metricValue="6.6" pageId="12" pageNumber="103" unit="mm" value="0.66">0.66 mm</quantity>
long and
<quantity id="4CBC6E9D8A1C3B0AF28DFD3CBB7AFD2A" box="[497,573,673,692]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" pageId="12" pageNumber="103" unit="mm" value="0.81">0.81 mm</quantity>
wide; lobes of respiratory plate nearly contiguous, directed posteriorly. Spiracular bulla rounded, convex, irregular, scarcely prominent. Tarsal claws (
<figureCitation id="137FDFFD8A1C3B0AF3CAFD4DB9B4FD7D" box="[182,243,720,739]" captionStart="Figs" captionStartId="9.[169,208,1274,1293]" captionTargetBox="[149,980,194,1252]" captionTargetId="figure-58@9.[148,980,194,1253]" captionTargetPageId="9" captionText="Figs. 2940. Megasoma thersites, third instar. 29) Head, frontal view; 30) epipharynx; 3132) mandibles, ventral view, 31) right and 32) left; 33) distal part of right maxillary mala; 34) detail of maxillary stridulatory area; 35) right maxilla and hypopharynx, dorsal view; 3637) distal segment of antenna, 36) dorsal view and 37) ventral view; 38) left posterior tarsal claw, external view; 39) second right abdominal spiracle; 40) microstructure of respiratory plate (5003). Figs. 2932, 35, bar ¼ 1 mm. Figs. 3334, 3639, bar ¼ 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10104910" httpUri="https://zenodo.org/record/10104910/files/figure.png" pageId="12" pageNumber="103">Fig. 38</figureCitation>
) similar on all legs, with acute apex, each bearing 2 internal, long, thick, basal setae and 2 external, long, thick, preapical setae.
<emphasis id="B9301F6A8A1C3B0AF297FD75BB7CFD65" box="[491,571,744,763]" italics="true" pageId="12" pageNumber="103">Abdomen</emphasis>
. Dorsum of segment I with 2 rows of 3034 short, spine-like setae, 4 rows of 2046 medium sized, slender setae, and 2 rows of 3638 long, slender setae, and more than 120 long setae on each side. Dorsa of segments IIVI each with 85100 long setae mainly on sides and 90120 spine-like, short setae mixed with long setae; at middle third these setae form more or less regular row of 2024 long setae. Dorsum of segment VII with more than 200 long, slender setae, more than 100 short, spine-like setae on each side, and transverse, anterior row of 36 slender, long setae. Dorsum of segment VIII with 1 row of 26 long, slender setae, more than 100 short, spine-like setae, and 80 long setae on each side. Dorsum of segment IX with 2 rows of 2030 long, slender setae and 50 long slender setae and more than 80 short, spine-like setae on each side. Dorsum of segment X with approximately 200 short spine-like setae and 40 long, slender setae on each side. Venter of abdominal segments III each with 5 rows of 4045 long, slender setae and 2 rows of 3035 medium sized, slender setae; venter of segments IIIV each with 1 row of 20 long, slender setae, and 2 rows of 2530 medium sized, slender setae; venter of segments VIVIII each with 80 long and medium sized, slender setae on each side; venter of segment IX with 70 long and medium sized, slender setae on each side. Raster without pallidia and septula, with 20 long, slender setae on campus, teges formed by 10 medium sized, slender setae; cuticle slightly rugose. Barbula formed by 2025 long, slender setae. Lower anal lip with 140 long and medium sized mixed, slender setae. Upper anal lip with more than 100 short, spine-like setae and long, slender setae mixed, mainly towards sides. Spiracles of segments IVIII progressively shorter (0.7
<quantity id="4CBC6E9D8A1C3B0AF2E9FB32B89FFB5C" box="[405,472,1199,1218]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="103" unit="mm" value="0.6">0.6 mm</quantity>
long; 1.0
<quantity id="4CBC6E9D8A1C3B0AF14BFB32BB3DFB5C" box="[567,634,1199,1218]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="12" pageNumber="103" unit="mm" value="0.8">0.8 mm</quantity>
wide); lobes of respiratory plate separated by distance less than 1/3 diameter of convexity of bulla (
<figureCitation id="137FDFFD8A1C3B0AF1E1FB5BBB9AFB47" box="[669,733,1222,1241]" captionStart="Figs" captionStartId="9.[169,208,1274,1293]" captionTargetBox="[149,980,194,1252]" captionTargetId="figure-58@9.[148,980,194,1253]" captionTargetPageId="9" captionText="Figs. 2940. Megasoma thersites, third instar. 29) Head, frontal view; 30) epipharynx; 3132) mandibles, ventral view, 31) right and 32) left; 33) distal part of right maxillary mala; 34) detail of maxillary stridulatory area; 35) right maxilla and hypopharynx, dorsal view; 3637) distal segment of antenna, 36) dorsal view and 37) ventral view; 38) left posterior tarsal claw, external view; 39) second right abdominal spiracle; 40) microstructure of respiratory plate (5003). Figs. 2932, 35, bar ¼ 1 mm. Figs. 3334, 3639, bar ¼ 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10104910" httpUri="https://zenodo.org/record/10104910/files/figure.png" pageId="12" pageNumber="103">Fig. 39</figureCitation>
), directed anteriorly. Spiracular bulla convex, rounded, weakly prominent; respiratory plate with about 28 irregularly shaped holes (
<figureCitation id="137FDFFD8A1C3B0AF3CAFB6BB9B2FA97" box="[182,245,1270,1289]" captionStart="Figs" captionStartId="9.[169,208,1274,1293]" captionTargetBox="[149,980,194,1252]" captionTargetId="figure-58@9.[148,980,194,1253]" captionTargetPageId="9" captionText="Figs. 2940. Megasoma thersites, third instar. 29) Head, frontal view; 30) epipharynx; 3132) mandibles, ventral view, 31) right and 32) left; 33) distal part of right maxillary mala; 34) detail of maxillary stridulatory area; 35) right maxilla and hypopharynx, dorsal view; 3637) distal segment of antenna, 36) dorsal view and 37) ventral view; 38) left posterior tarsal claw, external view; 39) second right abdominal spiracle; 40) microstructure of respiratory plate (5003). Figs. 2932, 35, bar ¼ 1 mm. Figs. 3334, 3639, bar ¼ 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10104910" httpUri="https://zenodo.org/record/10104910/files/figure.png" pageId="12" pageNumber="103">Fig. 40</figureCitation>
) across any diameter. Approximate dorsal body length
<quantity id="4CBC6E9D8A1C3B0AF1A5FB6BBA5FFA97" box="[729,792,1270,1289]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" pageId="12" pageNumber="103" unit="mm" value="85.0">85 mm</quantity>
.
</paragraph>
<paragraph id="8BFBC3788A1C3B0AF3BBFA83B89DFAF7" blockId="12.[175,1005,1310,1570]" pageId="12" pageNumber="103">
<emphasis id="B9301F6A8A1C3B0AF3BBFA83B8D9FAAD" bold="true" box="[199,414,1310,1331]" pageId="12" pageNumber="103">Second-instar Larva.</emphasis>
<materialsCitation id="3B2CC9258A1C3B0AF2D6FA83B891FAF7" collectingDate="1999-10-02" collectingDateMax="2000-04-05" collectingDateMin="1999-10-02" collectionCode="IEXA" collectorName="G. Nogueira" country="Mexico" location="San Jose del Cabo" municipality="Based" pageId="12" pageNumber="103" specimenCount="1" stateProvince="Baja California Sur">
<collectingMunicipality id="6B9F59028A1C3B0AF2D6FA83B8A3FAAD" box="[426,484,1310,1331]" pageId="12" pageNumber="103">Based</collectingMunicipality>
on one third instar larva reared from egg laid by female collected in
<collectingCountry id="F35383E88A1C3B0AF25FFAA4B828FAD0" box="[291,367,1337,1358]" name="Mexico" pageId="12" pageNumber="103">México</collectingCountry>
:
<collectingRegion id="49800D9A8A1C3B0AF20BFAA4BB76FAD0" box="[375,561,1337,1358]" country="Mexico" name="Baja California Sur" pageId="12" pageNumber="103">Baja California Sur</collectingRegion>
,
<location id="8E9B95A38A1C3B0AF146FAA4BBADFAD0" LSID="urn:lsid:plazi:treatment:03ED726E8A183B0BF2FCFA32BB6BFD4E:8E9B95A38A1C3B0AF146FAA4BBADFAD0" box="[570,746,1337,1358]" country="Mexico" municipality="Based" name="San Jose del Cabo" pageId="12" pageNumber="103" stateProvince="Baja California Sur">San José del Cabo</location>
,
<date id="FFFAE5B88A1C3B0AF18FFAA4BA15FAD0" box="[755,850,1337,1358]" pageId="12" pageNumber="103" value="1999-10-02">
<collectingDate id="EFBE1C508A1C3B0AF18FFAA4BA15FAD0" box="[755,850,1337,1358]" pageId="12" pageNumber="103" value="1999-10-02">
2-X-
<quantity id="4CBC6E9D8A1C3B0AF05CFAA4BA15FAD0" box="[800,850,1337,1358]" metricMagnitude="6" metricUnit="m" metricValue="1.99958" pageId="12" pageNumber="103" unit="m" value="1999580.0">1999</quantity>
</collectingDate>
</date>
, 580 m, fixed
<date id="FFFAE5B88A1C3B0AF0A4FAA4B845FAF7" pageId="12" pageNumber="103" value="2000-04-05">
<collectingDate id="EFBE1C508A1C3B0AF0A4FAA4B845FAF7" pageId="12" pageNumber="103" value="2000-04-05">5- IV-2000</collectingDate>
</date>
,
<collectorName id="26B1A6AE8A1C3B0AF271FAC9B8C4FAF7" box="[269,387,1364,1385]" pageId="12" pageNumber="103">G. Nogueira</collectorName>
(
<collectionCode id="ED555BBD8A1C3B0AF2EDFAC9B888FAF7" box="[401,463,1364,1385]" pageId="12" pageNumber="103">IEXA</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BFBC3788A1C3B0AF3BBFAF3B9BDF9BC" blockId="12.[175,1005,1310,1570]" pageId="12" pageNumber="103">
<emphasis id="B9301F6A8A1C3B0AF3BBFAF3B804FA1D" bold="true" box="[199,323,1390,1411]" pageId="12" pageNumber="103">Description.</emphasis>
Similar to third-instar larva except as follows: maximum width of head capsule 5.0 mm. Right chaetoparia with 38 spine-like setae and 34 small setae and sensillae; left chaetoparia with 24 spine-like setae and 32 small setae and sensillae. Scissorial area of both mandibles with 3 teeth clearly separated by notches. Prothoracic spiracles
<quantity id="4CBC6E9D8A1C3B0AF207FA45B897FA73" box="[379,464,1496,1517]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="12" pageNumber="103" unit="mm" value="0.23">0.23 mm</quantity>
long,
<quantity id="4CBC6E9D8A1C3B0AF171FA45BB25FA73" box="[525,610,1496,1517]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="12" pageNumber="103" unit="mm" value="0.35">0.35 mm</quantity>
wide. Abdominal spiracles progressively smaller (0.16
<quantity id="4CBC6E9D8A1C3B0AF247FA6EB8D4F996" box="[315,403,1523,1544]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="103" unit="mm" value="0.13">0.13 mm</quantity>
long, 0.34
<quantity id="4CBC6E9D8A1C3B0AF177FA6EBB25F996" box="[523,610,1523,1544]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="12" pageNumber="103" unit="mm" value="0.29">0.29 mm</quantity>
wide). Approximate dorsal body length
<quantity id="4CBC6E9D8A1C3B0AF3D3F990B9B1F9BC" box="[175,246,1549,1570]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="12" pageNumber="103" unit="mm" value="26.0">26 mm</quantity>
.
</paragraph>
<paragraph id="8BFBC3788A1D3B0BF3E8FF1DBA4DFF0D" blockId="13.[148,180,128,147]" box="[148,778,128,147]" lastBlockId="13.[347,778,128,147]" pageId="13" pageNumber="104">104 THE COLEOPTERISTS BULLETIN 59(1), 2005</paragraph>
<paragraph id="8BFBC3788A1D3B0BF3E8FF40B884FE93" blockId="13.[148,978,221,720]" pageId="13" pageNumber="104">
<materialsCitation id="3B2CC9258A1D3B0BF3E8FF40B887FE93" collectingDate="1999-10-02" collectingDateMax="1999-12-15" collectingDateMin="1999-10-02" collectionCode="IEXA" collectorName="G. Nogueira" country="Mexico" location="San Jose del Cabo" pageId="13" pageNumber="104" specimenCount="1" stateProvince="Baja California Sur">
collected in
<collectingCountry id="F35383E88A1D3B0BF279FF40B816FF6C" box="[261,337,221,242]" name="Mexico" pageId="13" pageNumber="104">México</collectingCountry>
:
<collectingRegion id="49800D9A8A1D3B0BF225FF40BB56FF6C" box="[345,529,221,242]" country="Mexico" name="Baja California Sur" pageId="13" pageNumber="104">Baja California Sur</collectingRegion>
,
<location id="8E9B95A38A1D3B0BF166FF40BB80FF6C" LSID="urn:lsid:plazi:treatment:03ED726E8A183B0BF2FCFA32BB6BFD4E:8E9B95A38A1D3B0BF166FF40BB80FF6C" box="[538,711,221,242]" country="Mexico" name="San Jose del Cabo" pageId="13" pageNumber="104" stateProvince="Baja California Sur">San José del Cabo</location>
,
<date id="FFFAE5B88A1D3B0BF1B3FF40BA69FF6C" box="[719,814,221,242]" pageId="13" pageNumber="104" value="1999-10-02">
<collectingDate id="EFBE1C508A1D3B0BF1B3FF40BA69FF6C" box="[719,814,221,242]" pageId="13" pageNumber="104" value="1999-10-02">
2-X-
<quantity id="4CBC6E9D8A1D3B0BF180FF40BA69FF6C" box="[764,814,221,242]" metricMagnitude="6" metricUnit="m" metricValue="1.99958" pageId="13" pageNumber="104" unit="m" value="1999580.0">1999</quantity>
</collectingDate>
</date>
, 580 m, fixed
<date id="FFFAE5B88A1D3B0BF0CDFF40B9A9FE93" pageId="13" pageNumber="104" value="1999-12-15">
<collectingDate id="EFBE1C508A1D3B0BF0CDFF40B9A9FE93" pageId="13" pageNumber="104" value="1999-12-15">15- XII-1999</collectingDate>
</date>
,
<collectorName id="26B1A6AE8A1D3B0BF384FF65B829FE93" box="[248,366,248,269]" pageId="13" pageNumber="104">G. Nogueira</collectorName>
(
<collectionCode id="ED555BBD8A1D3B0BF207FF65B8FEFE93" box="[379,441,248,269]" pageId="13" pageNumber="104">IEXA</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BFBC3788A1D3B0BF3D0FE8FBA80FE0F" blockId="13.[148,978,221,720]" pageId="13" pageNumber="104">
<emphasis id="B9301F6A8A1D3B0BF3D0FE8FB86FFEB9" bold="true" box="[172,296,274,295]" pageId="13" pageNumber="104">Description.</emphasis>
Similar to second-instar larva except as follows: maximum width of head capsule 2.73.0 mm. Sclerotized plates on sides of pronotum absent. Eclosion spine on each side of metanotum small, conical, surrounded by 3 short, spine-like setae. Thoracic spiracles
<quantity id="4CBC6E9D8A1D3B0BF23DFEFFB8D3FEE9" box="[321,404,354,375]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="13" pageNumber="104" unit="mm" value="0.13">0.13 mm</quantity>
wide,
<quantity id="4CBC6E9D8A1D3B0BF2B3FEFFBB65FEE9" box="[463,546,354,375]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="13" pageNumber="104" unit="mm" value="0.04">0.04 mm</quantity>
long. Abdominal spiracles similar in size,
<quantity id="4CBC6E9D8A1D3B0BF0DBFEFFB9FEFE0F" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="13" pageNumber="104" unit="mm" value="0.14">0.14 mm</quantity>
wide,
<quantity id="4CBC6E9D8A1D3B0BF385FEE1B80AFE0F" box="[249,333,380,401]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="104" unit="mm" value="0.02">0.02 mm</quantity>
long, without bulla. Approximate dorsal body length 4.56.0 mm.
</paragraph>
</subSubSection>
<subSubSection id="C35E90F38A1D3B0BF3D0FE0ABBEDFD88" pageId="13" pageNumber="104" type="discussion">
<paragraph id="8BFBC3788A1D3B0BF3D0FE0ABBEDFD88" blockId="13.[148,978,221,720]" pageId="13" pageNumber="104">
<emphasis id="B9301F6A8A1D3B0BF3D0FE0AB849FE32" bold="true" box="[172,270,407,428]" pageId="13" pageNumber="104">Remarks.</emphasis>
Larvae of
<taxonomicName id="4C44B8FB8A1D3B0BF200FE0AB8ACFE32" box="[380,491,407,428]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="104" phylum="Arthropoda" rank="species" species="thersites">
<emphasis id="B9301F6A8A1D3B0BF200FE0AB8ACFE32" box="[380,491,407,428]" italics="true" pageId="13" pageNumber="104">M. thersites</emphasis>
</taxonomicName>
are similar to those of
<taxonomicName id="4C44B8FB8A1D3B0BF1B9FE0ABA53FE32" authorityName="Cartwright" authorityYear="1963" box="[709,788,407,428]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="104" phylum="Arthropoda" rank="species" species="vogti">
<emphasis id="B9301F6A8A1D3B0BF1B9FE0ABA53FE32" box="[709,788,407,428]" italics="true" pageId="13" pageNumber="104">M. vogti</emphasis>
</taxonomicName>
and
<taxonomicName id="4C44B8FB8A1D3B0BF03EFE0ABA96FE32" box="[834,977,407,428]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="104" phylum="Arthropoda" rank="species" species="punctulatus">
<emphasis id="B9301F6A8A1D3B0BF03EFE0ABA96FE32" box="[834,977,407,428]" italics="true" pageId="13" pageNumber="104">M. punctulatus</emphasis>
</taxonomicName>
but may be distinguished by 3 postfrontal setae on each side, 6 anterior angle frontal setae, 7 setae near the posterior border of the labrum, pternotorma subquadrate, 7 dorsal sensory spots on the last antennal segment, acanthoparia with 1213 setae, and lobes of the respiratory plate of the spiracles narrowly separated.
</paragraph>
</subSubSection>
<subSubSection id="C35E90F38A1D3B0BF3D0FD86BB6BFD4E" pageId="13" pageNumber="104" type="distribution">
<paragraph id="8BFBC3788A1D3B0BF3D0FD86BB6BFD4E" blockId="13.[148,978,221,720]" pageId="13" pageNumber="104">
<emphasis id="B9301F6A8A1D3B0BF3D0FD86B8F4FDAE" bold="true" box="[172,435,539,560]" pageId="13" pageNumber="104">Distribution and Habits.</emphasis>
Adults of
<taxonomicName id="4C44B8FB8A1D3B0BF156FD81BBD8FDAF" box="[554,671,540,561]" class="Insecta" family="Dynastidae" genus="Megasoma" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="104" phylum="Arthropoda" rank="species" species="thersites">
<emphasis id="B9301F6A8A1D3B0BF156FD81BBD8FDAF" box="[554,671,540,561]" italics="true" pageId="13" pageNumber="104">M. thersites</emphasis>
</taxonomicName>
have been collected in many localities in the state of
<collectingRegion id="49800D9A8A1D3B0BF20CFDABBB6FFDD5" box="[368,552,566,587]" country="Mexico" name="Baja California Sur" pageId="13" pageNumber="104">Baja California Sur</collectingRegion>
,
<collectingCountry id="F35383E88A1D3B0BF14DFDABBB3DFDD5" box="[561,634,566,587]" name="Mexico" pageId="13" pageNumber="104">México</collectingCountry>
such as Todos Santos, La Paz, Cabo San Lucas, and San José del Cabo at elevations from sea level to 600 meters (
<bibRefCitation id="EFD5BE898A1D3B0BF0E8FDCCB980FD1E" author="Hardy" pageId="13" pageNumber="104" pagination="765 - 777" refId="ref19705" refString="Hardy, A. R. 1972. A brief revision of the North and Central American species of Megasoma (Coleoptera: Scarabaeidae). Canadian Entomologist 104: 765 - 777." type="journal article" year="1972">Hardy 1972</bibRefCitation>
). Males and females are attracted to lights during September and October, but they are also commonly found in the soil or on branches of palo verde, (
<taxonomicName id="4C44B8FB8A1D3B0BF011FD1BB807FD28" authority="Rose" authorityName="Rose" class="Magnoliopsida" family="Fabaceae" genus="Cercidium" kingdom="Plantae" order="Fabales" pageId="13" pageNumber="104" phylum="Tracheophyta" rank="species" species="peninsulare">
<emphasis id="B9301F6A8A1D3B0BF011FD1BB843FD28" italics="true" pageId="13" pageNumber="104">Cercidium peninsulare</emphasis>
Rose
</taxonomicName>
, Caesalpinaceae) where adults feed on the cambium (
<figureCitation id="137FDFFD8A1D3B0BF022FD3CBAEFFD28" box="[862,936,673,694]" captionStart="Fig" captionStartId="10.[197,228,1429,1448]" captionTargetBox="[178,1001,194,1408]" captionTargetId="figure-1@10.[178,1002,194,1408]" captionTargetPageId="10" captionText="Fig. 41. Megasoma thersites on palo verde, Cercidium microphyllum Rose &amp; I. M. Johnst. Photo by Barney Streit." figureDoi="http://doi.org/10.5281/zenodo.10104912" httpUri="https://zenodo.org/record/10104912/files/figure.png" pageId="13" pageNumber="104">Fig. 41</figureCitation>
) by gumming away the bark layer (
<figureCitation id="137FDFFD8A1D3B0BF2A5FD26BB66FD4E" box="[473,545,699,720]" captionStart="Fig" captionStartId="11.[169,200,1199,1218]" captionTargetBox="[151,974,194,1178]" captionTargetId="figure-166@11.[151,975,194,1178]" captionTargetPageId="11" captionText="Fig. 42. Gumming wound made by adult Megasoma thersites on palo verde tree. Photo by Barney Streit." figureDoi="http://doi.org/10.5281/zenodo.10104916" httpUri="https://zenodo.org/record/10104916/files/figure.png" pageId="13" pageNumber="104">Fig. 42</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>