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<mods:title id="8485D09707E982E05DF3A145FCACF71D">Late Neogene Lophophaenidae (Nassellaria, Radiolaria) from the eastern equatorial Pacific</mods:title>
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<taxonomicName id="4C60A5C5FFDCFFB775DFE0DEFE42C7BF" authority="Trubovitz, Renaudie, Lazarus &amp; Noble, 2022" box="[151,389,1193,1219]" family="Lophophaenidae" genus="Pelagomanes" higherTaxonomySource="GBIF" kingdom="Protozoa" order="Nassellaria" pageId="79" pageNumber="80" phylum="Sarcomastigophora" rank="species" species="ibburi" status="sp. nov.">
<emphasis id="B9140254FFDCFFB775DFE0DEFE42C7BF" box="[151,389,1193,1219]" italics="true" pageId="79" pageNumber="80">Pelagomanes ibburi</emphasis>
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<paragraph id="8BDFDE46FFDCFFB775DFE0BAFE50C79B" blockId="79.[151,462,1193,1255]" box="[151,407,1229,1255]" pageId="79" pageNumber="80">
Plate 41,
<figureCitation id="135BC2C3FFDCFFB77448E0BAFE98C79B" box="[256,351,1229,1255]" captionStart="FIGURE 1" captionStartId="6.[151,250,1340,1364]" captionTargetBox="[151,1430,743,1316]" captionTargetId="figure-284@6.[151,1436,741,1317]" captionTargetPageId="6" captionText="FIGURE 1. Diagram illustrating the general skeletal morphology of Lophophaenidae, and terminology used in this manuscript. Dashed lines indicate spines that are not visible from the angle shown, as they extend into the skeleton. An example specimen of Lophophaena casperi n. sp. is included for comparison to the diagrams." figureDoi="http://doi.org/10.5281/zenodo.10544060" httpUri="https://zenodo.org/record/10544060/files/figure.png" pageId="79" pageNumber="80">Figs. 1A</figureCitation>
<figureCitation id="135BC2C3FFDCFFB77438E0BAFE54C79B" box="[368,403,1229,1255]" captionStart="FIGURE 6" captionStartId="15.[151,250,1463,1487]" captionTargetBox="[153,1436,737,1439]" captionTargetId="figure-139@15.[151,1436,736,1440]" captionTargetPageId="15" captionText="FIGURE 6. Internal skeletal structure of Arachnocorallium calvata (Haeckel, 1887) Petrushevskaya, 1971." figureDoi="http://doi.org/10.5281/zenodo.10544070" httpUri="https://zenodo.org/record/10544070/files/figure.png" pageId="79" pageNumber="80">6B</figureCitation>
.
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<paragraph id="8BDFDE46FFDCFFB775DFE147FCACC634" blockId="79.[151,880,1298,1353]" box="[151,875,1328,1353]" pageId="79" pageNumber="80">
<taxonomicName id="4C60A5C5FFDCFFB775DFE147FE93C634" box="[151,340,1328,1352]" class="Polycystina" family="Plagiacanthidae" genus="Lithomelissa" kingdom="Chromista" order="Nassellaria" pageId="79" pageNumber="80" phylum="Radiozoa" rank="species" species="kozoi">
<emphasis id="B9140254FFDCFFB775DFE147FE93C634" box="[151,340,1328,1352]" italics="true" pageId="79" pageNumber="80">Lithomelissa kozoi</emphasis>
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cf sp 1,
<bibRefCitation id="EFF1A3B7FFDCFFB774E3E147FD45C635" author="Trubovitz, S. &amp; Lazarus, D. &amp; Renaudie, J. &amp; Noble, P. J." box="[427,642,1328,1353]" pageId="79" pageNumber="80" pagination="1 - 10" refId="ref96304" refString="Trubovitz, S., Lazarus, D., Renaudie, J. &amp; Noble, P. J. (2020) Marine plankton show threshold extinction response to Neogene climate change. Nature Communications, 11 (5069), 1 - 10. https: // doi. org / 10.1038 / s 41467 - 020 - 18879 - 7" type="journal article" year="2020">
Trubovitz
<emphasis id="B9140254FFDCFFB7775CE146FD80C634" box="[532,583,1328,1352]" italics="true" pageId="79" pageNumber="80">et al.</emphasis>
2020
</bibRefCitation>
, supplementary data 7.
</paragraph>
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<subSubSection id="C37A8DCDFFDCFFB775DFE101FDF5C6A5" pageId="79" pageNumber="80" type="diagnosis">
<paragraph id="8BDFDE46FFDCFFB775DFE101FDF5C6A5" blockId="79.[151,1437,1398,2037]" pageId="79" pageNumber="80">
<emphasis id="B9140254FFDCFFB775DFE101FED5C6EC" bold="true" box="[151,274,1398,1424]" pageId="79" pageNumber="80">Diagnosis.</emphasis>
<taxonomicName id="4C60A5C5FFDCFFB77451E100FE77C6ED" box="[281,432,1399,1425]" family="Lophophaenidae" genus="Pelagomanes" higherTaxonomySource="GBIF" kingdom="Protozoa" order="Nassellaria" pageId="79" pageNumber="80" phylum="Sarcomastigophora" rank="genus">
<emphasis id="B9140254FFDCFFB77451E100FE77C6ED" box="[281,432,1399,1425]" italics="true" pageId="79" pageNumber="80">Pelagomanes</emphasis>
</taxonomicName>
with a cephalis that ranges from having few, small pores to being entirely hyaline, has a relatively weak hooklike apical horn, and an elongated thorax with very strong shoulders at the top and three feet in addition to small teeth at the base.
</paragraph>
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<subSubSection id="C37A8DCDFFDCFFB7758FE195FC28C5A9" pageId="79" pageNumber="80" type="description">
<paragraph id="8BDFDE46FFDCFFB7758FE195FC28C5A9" blockId="79.[151,1437,1398,2037]" pageId="79" pageNumber="80">
<emphasis id="B9140254FFDCFFB7758FE195FE9EC680" bold="true" box="[199,345,1506,1532]" pageId="79" pageNumber="80">Description.</emphasis>
The cephalis of this species has small pores and is often hyaline (i.e., figs. 1 and 2). The apical horn extends from the dorsal side of the cephalis before reaching the apex, and is slightly curved at its base. The thorax is approximately twice the length of the cephalis, and slightly wider than it. Pores on the thorax get larger toward the base, and have the tendency to become more elongated in shape. Prominent shoulders exhibit at the top of the thorax, and the furrows between them extend downwards as ribs along the thorax and finally as bladed feet at the base of the skeleton. The base of the thorax may be enclosed in some specimens (i.e., fig. 3), and fully-preserved individuals show small teeth at the base of the thorax between the three feet.
</paragraph>
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<subSubSection id="C37A8DCDFFDCFFB7758FE2A8FB70C4D1" pageId="79" pageNumber="80" type="discussion">
<paragraph id="8BDFDE46FFDCFFB7758FE2A8FB70C4D1" blockId="79.[151,1437,1398,2037]" pageId="79" pageNumber="80">
<emphasis id="B9140254FFDCFFB7758FE2A8FEFCC585" bold="true" box="[199,315,1759,1785]" pageId="79" pageNumber="80">Remarks.</emphasis>
This species differs from
<taxonomicName id="4C60A5C5FFDCFFB77713E2A8FB45C585" authority="Renaudie and Lazarus, 2013 a" authorityName="Renaudie and Lazarus" authorityYear="2013" box="[603,1154,1759,1785]" family="Lophophaenidae" genus="Pelagomanes" higherTaxonomySource="GBIF" kingdom="Protozoa" order="Nassellaria" pageId="79" pageNumber="80" phylum="Sarcomastigophora" rank="species" species="kozoi">
<emphasis id="B9140254FFDCFFB77713E2A8FCF4C585" box="[603,819,1759,1785]" italics="true" pageId="79" pageNumber="80">Pelagomanes kozoi</emphasis>
<bibRefCitation id="EFF1A3B7FFDCFFB77672E2A8FB45C585" author="Renaudie, J. &amp; Lazarus, D. B." box="[826,1154,1759,1785]" pageId="79" pageNumber="80" pagination="59 - 86" refId="ref94302" refString="Renaudie, J. &amp; Lazarus, D. B. (2013 a) New species of Neogene radiolarians from the Southern Ocean - part II. Journal of Micropalaeontology, 32 (1), 59 - 86." type="journal article" year="2013">Renaudie and Lazarus, 2013a</bibRefCitation>
</taxonomicName>
(Pl. 39,
<figureCitation id="135BC2C3FFDCFFB77197E2A8FA80C585" box="[1247,1351,1759,1785]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="6.[151,250,1340,1364]" captionStartId-1="7.[151,250,1538,1562]" captionStartId-2="9.[151,250,1885,1909]" captionStartId-3="10.[151,250,1579,1603]" captionStartId-4="12.[151,250,1242,1266]" captionStartId-5="15.[151,250,1463,1487]" captionTargetBox-0="[151,1430,743,1316]" captionTargetBox-1="[157,1428,745,1510]" captionTargetBox-2="[189,1398,1065,1862]" captionTargetBox-3="[164,1423,849,1556]" captionTargetBox-4="[151,1436,520,1219]" captionTargetBox-5="[153,1436,737,1439]" captionTargetId-0="figure-284@6.[151,1436,741,1317]" captionTargetId-1="figure-251@7.[151,1436,741,1514]" captionTargetId-2="figure-391@9.[189,1398,1065,1862]" captionTargetId-3="figure-347@10.[164,1423,849,1556]" captionTargetId-4="figure-153@12.[151,1436,520,1219]" captionTargetId-5="figure-139@15.[151,1436,736,1440]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="9" captionTargetPageId-3="10" captionTargetPageId-4="12" captionTargetPageId-5="15" captionText-0="FIGURE 1. Diagram illustrating the general skeletal morphology of Lophophaenidae, and terminology used in this manuscript. Dashed lines indicate spines that are not visible from the angle shown, as they extend into the skeleton. An example specimen of Lophophaena casperi n. sp. is included for comparison to the diagrams." captionText-1="FIGURE 2. Relative abundance of specimens and taxa belonging to Plagiacanthoidea and Lophophaenidae per sample, from IODP Site U1337 in the EEP over the last ~10 Ma. The relative percentages of total taxa are represented by the blue lines and the red lines represent the percentages of total specimens (triangles = Plagiacanthoidea; circles = Lophophaenidae). The Plagiacanthoidea was among the most abundant and speciose of radiolarian groups encountered in Trubovitz et al. (2020), with at least 10% of the specimens and 20% of the taxonomic richness in every sample. Within the Plagiacanthoidea, lophophaenids consistently made up the majority of specimens (~67%) and taxa (~61%). Data are from Trubovitz et al. (2020)." captionText-2="FIGURE 3. Locality map of IODP Site U1337, where all samples for this study were collected. Map from Google EarthTM." captionText-3="FIGURE 4. Examples of specimen measurements discussed in the Systematic Paleontology section of this study." captionText-4="FIGURE 5. Internal skeletal structure of Amphiplecta acrostoma Haeckel, 1887." captionText-5="FIGURE 6. Internal skeletal structure of Arachnocorallium calvata (Haeckel, 1887) Petrushevskaya, 1971." figureDoi-0="http://doi.org/10.5281/zenodo.10544060" figureDoi-1="http://doi.org/10.5281/zenodo.10544062" figureDoi-2="http://doi.org/10.5281/zenodo.10544064" figureDoi-3="http://doi.org/10.5281/zenodo.10544066" figureDoi-4="http://doi.org/10.5281/zenodo.10544068" figureDoi-5="http://doi.org/10.5281/zenodo.10544070" httpUri-0="https://zenodo.org/record/10544060/files/figure.png" httpUri-1="https://zenodo.org/record/10544062/files/figure.png" httpUri-2="https://zenodo.org/record/10544064/files/figure.png" httpUri-3="https://zenodo.org/record/10544066/files/figure.png" httpUri-4="https://zenodo.org/record/10544068/files/figure.png" httpUri-5="https://zenodo.org/record/10544070/files/figure.png" pageId="79" pageNumber="80">Figs. 16</figureCitation>
) in that the shoulders are usually stronger, and the teeth at the base of the thorax are shorter and have more contrast with the three feet. In addition, the thorax of this species tends to be longer than in
<taxonomicName id="4C60A5C5FFDCFFB776ABE350FBFDC43D" authorityName="Renaudie and Lazarus" authorityYear="2013" box="[995,1082,1831,1857]" family="Lophophaenidae" genus="Pelagomanes" higherTaxonomySource="GBIF" kingdom="Protozoa" pageId="79" pageNumber="80" phylum="Sarcomastigophora" rank="species" species="kozoi">
<emphasis id="B9140254FFDCFFB776ABE350FBFDC43D" box="[995,1082,1831,1857]" italics="true" pageId="79" pageNumber="80">P. kozoi</emphasis>
</taxonomicName>
, and the cephalis is often more hyaline. This species differs from
<taxonomicName id="4C60A5C5FFDCFFB77758E33CFCA9C419" authorityName="Caulet" authorityYear="1979" box="[528,878,1867,1893]" class="Polycystina" family="Plagiacanthidae" genus="Pseudodictyophimus" kingdom="Chromista" order="Nassellaria" pageId="79" pageNumber="80" phylum="Radiozoa" rank="species" species="gracilipes">
<emphasis id="B9140254FFDCFFB77758E33CFCA9C419" box="[528,878,1867,1893]" italics="true" pageId="79" pageNumber="80">Pseudodictyophimus gracilipes</emphasis>
</taxonomicName>
in that the thorax is narrower and more elongated. The apical and ventral horns show variable development, but are never as strong as those of
<taxonomicName id="4C60A5C5FFDCFFB771F9E318FA85C4F5" baseAuthorityName="Caulet" baseAuthorityYear="1991" box="[1201,1346,1903,1929]" family="Lophophaenidae" genus="Pelagomanes" higherTaxonomySource="GBIF" kingdom="Protozoa" pageId="79" pageNumber="80" phylum="Sarcomastigophora" rank="species" species="thaumasia">
<emphasis id="B9140254FFDCFFB771F9E318FA85C4F5" box="[1201,1346,1903,1929]" italics="true" pageId="79" pageNumber="80">P. thaumasia</emphasis>
</taxonomicName>
(Pl. 39,
<figureCitation id="135BC2C3FFDCFFB775DFE3E4FEC0C4D1" box="[151,263,1939,1965]" captionStart="FIGURE 10" captionStartId="40.[151,250,1263,1287]" captionTargetBox="[164,1423,418,1240]" captionTargetId="figure-130@40.[164,1423,417,1240]" captionTargetPageId="40" captionText="FIGURE 10. Internal skeletal structure of Lithomelissa alkonost n. sp." figureDoi="http://doi.org/10.5281/zenodo.10544078" httpUri="https://zenodo.org/record/10544078/files/figure.png" pageId="79" pageNumber="80">Figs. 10A</figureCitation>
11),
<taxonomicName id="4C60A5C5FFDCFFB7740EE3E4FDC4C4D0" baseAuthorityName="Funakawa" baseAuthorityYear="1995" box="[326,515,1939,1964]" family="Lophophaenidae" genus="Pelagomanes" higherTaxonomySource="GBIF" kingdom="Protozoa" pageId="79" pageNumber="80" phylum="Sarcomastigophora" rank="species" species="morawanensis">
<emphasis id="B9140254FFDCFFB7740EE3E4FDC4C4D0" box="[326,515,1939,1964]" italics="true" pageId="79" pageNumber="80">P. morawanensis</emphasis>
</taxonomicName>
(Pl. 40,
<figureCitation id="135BC2C3FFDCFFB7772BE3E4FD03C4D1" box="[611,708,1939,1965]" captionStart="FIGURE 4" captionStartId="10.[151,250,1579,1603]" captionTargetBox="[164,1423,849,1556]" captionTargetId="figure-347@10.[164,1423,849,1556]" captionTargetPageId="10" captionText="FIGURE 4. Examples of specimen measurements discussed in the Systematic Paleontology section of this study." figureDoi="http://doi.org/10.5281/zenodo.10544066" httpUri="https://zenodo.org/record/10544066/files/figure.png" pageId="79" pageNumber="80">Figs. 4A</figureCitation>
<figureCitation id="135BC2C3FFDCFFB7779DE3E4FD25C4D1" box="[725,738,1939,1965]" captionStart="FIGURE 6" captionStartId="15.[151,250,1463,1487]" captionTargetBox="[153,1436,737,1439]" captionTargetId="figure-139@15.[151,1436,736,1440]" captionTargetPageId="15" captionText="FIGURE 6. Internal skeletal structure of Arachnocorallium calvata (Haeckel, 1887) Petrushevskaya, 1971." figureDoi="http://doi.org/10.5281/zenodo.10544070" httpUri="https://zenodo.org/record/10544070/files/figure.png" pageId="79" pageNumber="80">6</figureCitation>
,
<figureCitation id="135BC2C3FFDCFFB777A5E3E4FD3AC4D1" box="[749,765,1939,1965]" captionStart="FIGURE 9" captionStartId="30.[151,250,1556,1580]" captionTargetBox="[152,1436,742,1532]" captionTargetId="figure-252@30.[151,1436,741,1532]" captionTargetPageId="30" captionText="FIGURE 9. Internal skeletal structure of Ceratocyrtis sp. C." figureDoi="http://doi.org/10.5281/zenodo.10544076" httpUri="https://zenodo.org/record/10544076/files/figure.png" pageId="79" pageNumber="80">9</figureCitation>
, 1112), or
<taxonomicName id="4C60A5C5FFDCFFB776CCE3E4FC3FC4D1" baseAuthorityName="O'Connor" baseAuthorityYear="1997" box="[900,1016,1939,1965]" family="Lophophaenidae" genus="Pelagomanes" higherTaxonomySource="GBIF" kingdom="Protozoa" pageId="79" pageNumber="80" phylum="Sarcomastigophora" rank="species" species="tekopua">
<emphasis id="B9140254FFDCFFB776CCE3E4FC3FC4D1" box="[900,1016,1939,1965]" italics="true" pageId="79" pageNumber="80">P. tekopua</emphasis>
</taxonomicName>
(Pl. 40,
<figureCitation id="135BC2C3FFDCFFB77110E3E4FB6DC4D1" box="[1112,1194,1939,1965]" captionStart="FIGURE 13" captionStartId="56.[151,250,1336,1360]" captionTargetBox="[169,1418,495,1309]" captionTargetId="figure-127@56.[162,1426,489,1312]" captionTargetPageId="56" captionText="FIGURE 13. Chart illustrating the differences between Lophophaena gozui n. sp. (orange diamonds) and L. ushionii n. sp. (green squares), in terms of cephalis height and the ratio between thorax and cephalis width. L. gozui has a shorter average cephalis height (40 microns) compared to L. ushionii (average cephalis height = 54 microns). In addition, the thorax is proportionally wider compared to the cephalis in L. gozui (average ratio of thorax:cephalis width = 2.5). L. ushionii has a proportionally narrower thorax (average ratio of thorax: cephalis width = 1.5)." figureDoi="http://doi.org/10.5281/zenodo.10544082" httpUri="https://zenodo.org/record/10544082/files/figure.png" pageId="79" pageNumber="80">Fig. 13</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37A8DCDFFDCFFA87586E3C1FC30C165" lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80" type="materials_examined">
<paragraph id="8BDFDE46FFDCFFA87586E3C1FB72C219" blockId="79.[151,1437,1398,2037]" lastBlockId="80.[151,1437,151,717]" lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80">
<emphasis id="B9140254FFDCFFB77586E3C1FE70C4AC" bold="true" box="[206,439,1974,2000]" pageId="79" pageNumber="80">Material examined.</emphasis>
78 specimens observed from samples 321-1337A-35X-1, 106108cm (Middle Miocene), 321-1337A-31X-6, 46cm (Middle Miocene), 321-1337D-30H-3, 103104cm (Middle Miocene), 321-1337D-26H-
<quantity id="4C9873A3FFC3FFA8759DE4E0FEB6C3CD" box="[213,369,151,177]" metricMagnitude="-1" metricUnit="m" metricValue="7.3571" metricValueMax="14.4" metricValueMin="0.3142" pageId="80" pageNumber="81" unit="cm" value="73.571" valueMax="144.0" valueMin="3.142">3, 142144cm</quantity>
(Late Miocene), 321-1337D-23H-
<quantity id="4C9873A3FFC3FFA877B0E4E0FC52C3CD" box="[760,917,151,177]" metricMagnitude="-1" metricUnit="m" metricValue="7.1567" metricValueMax="13.7" metricValueMin="0.6134" pageId="80" pageNumber="81" unit="cm" value="71.567" valueMax="137.0" valueMin="6.134">6, 134137cm</quantity>
(Late Miocene), 321-1337A-18H-6,
<quantity id="4C9873A3FFC3FFA8707CE4E0FA5BC3CD" box="[1332,1436,151,177]" metricMagnitude="-1" metricUnit="m" metricValue="7.85" metricValueMax="8.0" metricValueMin="7.7" pageId="80" pageNumber="81" unit="cm" value="78.5" valueMax="80.0" valueMin="77.0">7780cm</quantity>
(Late Miocene), 321-1337A-16H-
<quantity id="4C9873A3FFC3FFA87768E4CCFD07C3A9" box="[544,704,187,213]" metricMagnitude="-1" metricUnit="m" metricValue="6.50605" metricValueMax="12.4" metricValueMin="0.6121" pageId="80" pageNumber="81" unit="cm" value="65.0605" valueMax="124.0" valueMin="6.121">6, 121124cm</quantity>
(Late Miocene), 321-1337A-14H-7,
<quantity id="4C9873A3FFC3FFA87139E4CCFB1EC3A9" box="[1137,1241,187,213]" metricMagnitude="-1" metricUnit="m" metricValue="4.05" metricValueMax="4.2" metricValueMin="3.9" pageId="80" pageNumber="81" unit="cm" value="40.5" valueMax="42.0" valueMin="39.0">3942cm</quantity>
(Late Miocene), 321-1337A-12H-5,
<quantity id="4C9873A3FFC3FFA87432E4A8FE25C385" box="[378,482,223,249]" metricMagnitude="-1" metricUnit="m" metricValue="2.45" metricValueMax="2.6" metricValueMin="2.3" pageId="80" pageNumber="81" unit="cm" value="24.5" valueMax="26.0" valueMin="23.0">2326cm</quantity>
(Late Miocene), 321-1337A-10H-2,
<quantity id="4C9873A3FFC3FFA876DBE4A8FC3CC385" box="[915,1019,223,249]" metricMagnitude="-1" metricUnit="m" metricValue="9.25" metricValueMax="9.4" metricValueMin="9.1" pageId="80" pageNumber="81" unit="cm" value="92.5" valueMax="94.0" valueMin="91.0">9194cm</quantity>
(Early Pliocene), 321-1337A-7H-
<quantity id="4C9873A3FFC3FFA870C2E4A8FEDBC261" metricMagnitude="-1" metricUnit="m" metricValue="5.6552" metricValueMax="10.7" metricValueMin="0.6104" pageId="80" pageNumber="81" unit="cm" value="56.552" valueMax="107.0" valueMin="6.104">6, 104107cm</quantity>
(Early Pliocene), 321-1337A-6H-3,
<quantity id="4C9873A3FFC3FFA8778CE574FCEBC261" box="[708,812,259,285]" metricMagnitude="-1" metricUnit="m" metricValue="3.05" metricValueMax="3.2" metricValueMin="2.9" pageId="80" pageNumber="81" unit="cm" value="30.5" valueMax="32.0" valueMin="29.0">2932cm</quantity>
(Late Pliocene), 321-1337A-4H-
<quantity id="4C9873A3FFC3FFA871E6E574FA8AC261" box="[1198,1357,258,285]" metricMagnitude="-1" metricUnit="m" metricValue="6.20575" metricValueMax="11.8" metricValueMin="0.6115" pageId="80" pageNumber="81" unit="cm" value="62.0575" valueMax="118.0" valueMin="6.115">6, 115118cm</quantity>
(Early Pleistocene), 321-1337A-4H-2,
<quantity id="4C9873A3FFC3FFA8774FE550FDA8C23D" box="[519,623,295,321]" metricMagnitude="-1" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="80" pageNumber="81" unit="cm" value="17.5" valueMax="19.0" valueMin="16.0">1619cm</quantity>
(Middle Pleistocene), 321-1337A-3H-
<quantity id="4C9873A3FFC3FFA8717BE550FB12C23D" box="[1075,1237,295,321]" metricMagnitude="-1" metricUnit="m" metricValue="5.40515" metricValueMax="10.6" metricValueMin="0.2103" pageId="80" pageNumber="81" unit="cm" value="54.0515" valueMax="106.0" valueMin="2.103">2, 103106cm</quantity>
(Middle Pleistocene), 321-1337A-2H-3,
<quantity id="4C9873A3FFC3FFA874FAE53CFDDCC219" box="[434,539,331,357]" metricMagnitude="-1" metricUnit="m" metricValue="7.75" metricValueMax="7.9" metricValueMin="7.6" pageId="80" pageNumber="81" unit="cm" value="77.5" valueMax="79.0" valueMin="76.0">7679cm</quantity>
(Late Pleistocene), and 321-1337D-1H-1,
<quantity id="4C9873A3FFC3FFA876B2E53CFB80C219" box="[1018,1095,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="3.0" metricValueMin="0.0" pageId="80" pageNumber="81" unit="cm" value="1.5" valueMax="3.0" valueMin="0.0">03cm</quantity>
(Recent).
</paragraph>
<paragraph id="8BDFDE46FFC3FFA8758FE518FB68C2F5" blockId="80.[151,1437,151,717]" box="[199,1199,367,393]" pageId="80" pageNumber="81">
<materialsCitation id="3B08D41BFFC3FFA8758FE518FB68C2F5" box="[199,1199,367,393]" pageId="80" pageNumber="81" specimenCode="321-1337A-16H-6" specimenCount="1" typeStatus="holotype">
<emphasis id="B9140254FFC3FFA8758FE518FEFCC2F5" bold="true" box="[199,315,367,393]" pageId="80" pageNumber="81">
<typeStatus id="54DB60E4FFC3FFA8758FE518FEF1C2F5" box="[199,310,367,393]" pageId="80" pageNumber="81" type="holotype">Holotype</typeStatus>
.
</emphasis>
Pl. 41,
<figureCitation id="135BC2C3FFC3FFA874D9E518FDD1C2F5" box="[401,534,367,393]" captionStart="FIGURE 2" captionStartId="7.[151,250,1538,1562]" captionTargetBox="[157,1428,745,1510]" captionTargetId="figure-251@7.[151,1436,741,1514]" captionTargetPageId="7" captionText="FIGURE 2. Relative abundance of specimens and taxa belonging to Plagiacanthoidea and Lophophaenidae per sample, from IODP Site U1337 in the EEP over the last ~10 Ma. The relative percentages of total taxa are represented by the blue lines and the red lines represent the percentages of total specimens (triangles = Plagiacanthoidea; circles = Lophophaenidae). The Plagiacanthoidea was among the most abundant and speciose of radiolarian groups encountered in Trubovitz et al. (2020), with at least 10% of the specimens and 20% of the taxonomic richness in every sample. Within the Plagiacanthoidea, lophophaenids consistently made up the majority of specimens (~67%) and taxa (~61%). Data are from Trubovitz et al. (2020)." figureDoi="http://doi.org/10.5281/zenodo.10544062" httpUri="https://zenodo.org/record/10544062/files/figure.png" pageId="80" pageNumber="81">Figs. 2AB</figureCitation>
; sample
<specimenCode id="DBC6763DFFC3FFA87730E518FC8CC2F5" box="[632,843,367,393]" pageId="80" pageNumber="81">
321-1337A-16H-
<quantity id="4C9873A3FFC3FFA87675E518FC19C2F5" box="[829,990,367,393]" metricMagnitude="-1" metricUnit="m" metricValue="6.50605" metricValueMax="12.4" metricValueMin="0.6121" pageId="80" pageNumber="81" unit="cm" value="65.0605" valueMax="124.0" valueMin="6.121">
6, 121
<quantity id="4C9873A3FFC3FFA876C6E518FC19C2F5" box="[910,990,367,393]" metricMagnitude="0" metricUnit="m" metricValue="1.24" pageId="80" pageNumber="81" unit="cm" value="124.0">124cm</quantity>
</quantity>
</specimenCode>
; ECO-141; Z12-4.
</materialsCitation>
</paragraph>
<paragraph id="8BDFDE46FFC3FFA8758FE5E4FC30C165" blockId="80.[151,1437,151,717]" pageId="80" pageNumber="81">
<materialsCitation id="3B08D41BFFC3FFA8758FE5E4FC34C165" collectionCode="V" pageId="80" pageNumber="81" specimenCode="321-1337A-16H-6, 321-1337D-23H-6, 321-1337A-14H-7" specimenCount="1" typeStatus="paratype">
<emphasis id="B9140254FFC3FFA8758FE5E4FE81C2D1" bold="true" box="[199,326,403,429]" pageId="80" pageNumber="81">
<typeStatus id="54DB60E4FFC3FFA8758FE5E4FE86C2D1" box="[199,321,403,429]" pageId="80" pageNumber="81" type="paratype">Paratypes</typeStatus>
.
</emphasis>
(1) Pl. 41, figs. 1AB; sample
<specimenCode id="DBC6763DFFC3FFA877E4E5E4FCB8C2D1" box="[684,895,403,429]" pageId="80" pageNumber="81">
321-1337A-16H-
<quantity id="4C9873A3FFC3FFA87639E5E4FBD4C2D1" box="[881,1043,403,429]" metricMagnitude="-1" metricUnit="m" metricValue="6.50605" metricValueMax="12.4" metricValueMin="0.6121" pageId="80" pageNumber="81" unit="cm" value="65.0605" valueMax="124.0" valueMin="6.121">6, 121124cm</quantity>
</specimenCode>
; ECO-143; K12-3. (2) Pl. 41, figs. 4AB; sample
<specimenCode id="DBC6763DFFC3FFA87408E5C0FDD4C2AD" box="[320,531,439,465]" pageId="80" pageNumber="81">
321-1337D-23H-
<quantity id="4C9873A3FFC3FFA8774DE5C0FD61C2AD" box="[517,678,439,465]" metricMagnitude="-1" metricUnit="m" metricValue="7.1567" metricValueMax="13.7" metricValueMin="0.6134" pageId="80" pageNumber="81" unit="cm" value="71.567" valueMax="137.0" valueMin="6.134">6, 134137cm</quantity>
</specimenCode>
; ECO-145; N15-1. (3) Pl. 41, figs. 3AB; sample
<specimenCode id="DBC6763DFFC3FFA87192E5C0FF60C289" pageId="80" pageNumber="81">
321-1337D-23H-
<quantity id="4C9873A3FFC3FFA875DFE5ACFEFCC289" box="[151,315,475,501]" metricMagnitude="-1" metricUnit="m" metricValue="7.1567" metricValueMax="13.7" metricValueMin="0.6134" pageId="80" pageNumber="81" unit="cm" value="71.567" valueMax="137.0" valueMin="6.134">6, 134137cm</quantity>
</specimenCode>
; ECO-145; G17-2. (4) Pl. 41, figs. 6AB; sample
<specimenCode id="DBC6763DFFC3FFA8762FE5ACFBFCC289" box="[871,1083,475,501]" pageId="80" pageNumber="81">321-1337A-14H-7</specimenCode>
,
<quantity id="4C9873A3FFC3FFA8710DE5ACFB76C289" box="[1093,1201,475,501]" metricMagnitude="-1" metricUnit="m" metricValue="4.05" metricValueMax="4.2" metricValueMin="3.9" pageId="80" pageNumber="81" unit="cm" value="40.5" valueMax="42.0" valueMin="39.0">3942cm</quantity>
; ECO-138;
<date id="FFDEF886FFC3FFA8707DE5ACFAB2C289" box="[1333,1397,475,501]" pageId="80" pageNumber="81">V9-2.</date>
(5) Pl. 41, figs. 5AB; sample
<specimenCode id="DBC6763DFFC3FFA8748FE588FD5DC165" box="[455,666,511,537]" pageId="80" pageNumber="81">
321-1337A-16H-
<quantity id="4C9873A3FFC3FFA877C4E588FCEBC165" box="[652,812,511,537]" metricMagnitude="-1" metricUnit="m" metricValue="6.50605" metricValueMax="12.4" metricValueMin="0.6121" pageId="80" pageNumber="81" unit="cm" value="65.0605" valueMax="124.0" valueMin="6.121">6, 121124cm</quantity>
</specimenCode>
; ECO-143; J18-2
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37A8DCDFFC3FFA87586E654FBC5C11D" pageId="80" pageNumber="81" type="description">
<paragraph id="8BDFDE46FFC3FFA87586E654FBC5C11D" blockId="80.[151,1437,151,717]" pageId="80" pageNumber="81">
<emphasis id="B9140254FFC3FFA87586E654FE42C141" bold="true" box="[206,389,547,573]" pageId="80" pageNumber="81">Measurements.</emphasis>
Cephalis height 3037 (34)μm, cephalis width 2631 (29)μm, thorax length (not including feet) 4559 (49)μm, width at shoulders 3847 (42)μm. Based on
<specimenCount id="9D6615CFFFC3FFA87639E630FC39C11D" box="[881,1022,583,609]" pageId="80" pageNumber="81" type="generic">8 specimens</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="C37A8DCDFFC3FFA87586E61CFF30C1D5" pageId="80" pageNumber="81" type="etymology">
<paragraph id="8BDFDE46FFC3FFA87586E61CFF30C1D5" blockId="80.[151,1437,151,717]" pageId="80" pageNumber="81">
<emphasis id="B9140254FFC3FFA87586E61CFE92C1F9" bold="true" box="[206,341,619,645]" pageId="80" pageNumber="81">Etymology.</emphasis>
Named for ibbur in
<collectingRegion id="49A410A4FFC3FFA87770E61CFD43C1F9" box="[568,644,619,645]" country="Russia" name="Yevrey" pageId="80" pageNumber="81">Jewish</collectingRegion>
folklore, a
<typeStatus id="54DB60E4FFC3FFA8764BE61CFCF4C1F9" box="[771,819,619,645]" pageId="80" pageNumber="81">type</typeStatus>
of benevolent spirit possession intended to help people succeed.
</paragraph>
</subSubSection>
<subSubSection id="C37A8DCDFFC3FFA8758FE6C4FD30C1B1" box="[199,759,691,717]" pageId="80" pageNumber="81" type="biology_ecology">
<paragraph id="8BDFDE46FFC3FFA8758FE6C4FD30C1B1" blockId="80.[151,1437,151,717]" box="[199,759,691,717]" pageId="80" pageNumber="81">
<emphasis id="B9140254FFC3FFA8758FE6C4FEDDC1B1" bold="true" box="[199,282,691,717]" pageId="80" pageNumber="81">Range.</emphasis>
Middle Miocene—Recent, EEP (
<tableCitation id="C6E2EBFDFFC3FFA877DAE6C4FD21C1B1" box="[658,742,691,717]" captionStart="TABLE 1" captionStartId="84.[151,248,834,860]" captionTargetPageId="84" captionText="TABLE 1. Occurrence chart of Late NeogeneRecent lophophaenids in the EEP." httpUri="http://table.plazi.org/id/DF1F8ECEFFC7FFAC75DFE735FB46C021" pageId="80" pageNumber="81" tableUuid="DF1F8ECEFFC7FFAC75DFE735FB46C021">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>