318 lines
34 KiB
XML
318 lines
34 KiB
XML
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<mods:title id="0DC3346F6AC192598C8A93A96137B127">Annotated checklist and distribution of the lizards of Iran</mods:title>
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<mods:namePart id="38A3804C6FF2C0170313E60452954368">Šmíd, Jiří</mods:namePart>
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<mods:namePart id="65B4DB38359E9900434DFC25330715C6">Moravec, Jiří</mods:namePart>
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<mods:namePart id="792DA0FEC1159161835909F0B6E64ECB">Kodym, Petr</mods:namePart>
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<mods:namePart id="7EA6167D7D8DF32C2245F0240B0B1F1F">Kratochvíl, Lukáš</mods:namePart>
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<mods:namePart id="BC1F80E96F37FC0EA1B345C1DF8A6EB6">Yousefkhani, Seyyed Saeed Hosseinian</mods:namePart>
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<mods:namePart id="120F060EFB5D6C80BC83BA66576C0B06">Rastegar-Pouyani, Eskandar</mods:namePart>
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<mods:nameIdentifier id="610AD34C5CDB9542C0437F512AFF8319" type="email">jirismd@gmail.com.</mods:nameIdentifier>
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<mods:namePart id="A2F967D377161D7D425A019FB0089D22">Frynta, Daniel</mods:namePart>
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<mods:title id="D9B682CD7E05D231AD968F8CA248F1E6">Zootaxa</mods:title>
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<mods:date id="F491531854CD1AB1AE14285677B1E1A5">2014</mods:date>
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<mods:number id="BE488D73B5C8F8D729B251E03A22FB79">3855</mods:number>
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<mods:identifier id="9731430F04026F1A5ED1D85E0A8F233E" type="DOI">10.11646/zootaxa.3855.1.1</mods:identifier>
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<treatment id="03C387F2FF85FFB0FF5B4DC85750F9EF" ID-DOI="http://doi.org/10.5281/zenodo.4929112" ID-GBIF-Taxon="185468319" ID-Zenodo-Dep="4929112" LSID="urn:lsid:plazi:treatment:03C387F2FF85FFB0FF5B4DC85750F9EF" httpUri="http://treatment.plazi.org/id/03C387F2FF85FFB0FF5B4DC85750F9EF" lastPageNumber="10" pageId="9" pageNumber="10">
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<subSubSection id="C370656FFF85FFB0FF5B4DC85557FC8A" pageId="9" pageNumber="10" type="nomenclature">
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<paragraph id="8BD536E4FF85FFB0FF5B4DC854F1FCAF" blockId="9.[151,1436,746,1603]" box="[151,649,746,771]" pageId="9" pageNumber="10">
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<heading id="D09D8188FF85FFB0FF5B4DC854F1FCAF" bold="true" box="[151,649,746,771]" fontSize="10" level="2" pageId="9" pageNumber="10" reason="2">
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<taxonomicName id="4C6A4D67FF85FFB0FF5B4DC854F1FCAF" authority="(Pallas, 1771)" baseAuthorityName="Pallas" baseAuthorityYear="1771" box="[151,649,746,771]" class="Reptilia" family="Agamidae" genus="Phrynocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="helioscopus">
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<emphasis id="B91EEAF6FF85FFB0FF5B4DC854F1FCAF" bold="true" box="[151,649,746,771]" pageId="9" pageNumber="10">
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<emphasis id="B91EEAF6FF85FFB0FF5B4DC85799FCAF" bold="true" box="[151,481,746,771]" italics="true" pageId="9" pageNumber="10">Phrynocephalus helioscopus</emphasis>
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(
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<bibRefCitation id="EFFB4B15FF85FFB0FE3C4DC854F8FCAF" author="Pallas, P. S." box="[496,640,746,771]" pageId="9" pageNumber="10" refId="ref61618" refString="Pallas, P. S. (1771) Reise durch verschiedene Provinzen des Russisches Reichs. Kaiserlichen Akademie der Wissenschaften, St. Petersburg, 504 pp." type="book" year="1771">Pallas, 1771</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BD536E4FF85FFB0FF5B4C2C5557FC8A" blockId="9.[151,1436,746,1603]" box="[151,815,781,806]" pageId="9" pageNumber="10">
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<materialsCitation id="3B023CB9FF85FFB0FF5B4C2C5557FC8A" ID-GBIF-Occurrence="3342274359" box="[151,815,781,806]" collectionCode="ZMB" pageId="9" pageNumber="10" specimenCode="ZMB 781" specimenCount="1" typeStatus="lectotype">
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<typeStatus id="54D18846FF85FFB0FF5B4C2C5763FC88" box="[151,283,782,805]" pageId="9" pageNumber="10" type="lectotype">LECTOTYPE</typeStatus>
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.
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<specimenCode id="DBCC9E9FFF85FFB0FEE44C2C57E1FC8A" box="[296,409,781,806]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="9" pageNumber="10" type="Museum">ZMB 781</specimenCode>
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, designated by
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<bibRefCitation id="EFFB4B15FF85FFB0FD8A4C2C5553FC8A" author="Denzer, W. & Gunther, R. & Manthey, U." box="[582,811,781,806]" pageId="9" pageNumber="10" pagination="309 - 332" refId="ref49420" refString="Denzer, W., Gunther, R. & Manthey, U. (1997) Kommentierter Typenkatalog der Agamen (Reptilia: Squamata: Agamidae) des Museums fur Naturkunde der Humboldt-Universitat zu Berlin (ehemals Zoologisches Museum Berlin). Mitteilung aus dem Zoologischen Museum in Berlin, 73, 309 - 332. http: // dx. doi. org / 10.1002 / mmnz. 4830730209" type="journal article" year="1997">
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Denzer
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<emphasis id="B91EEAF6FF85FFB0FD514C2C54AEFC8A" box="[669,726,781,806]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
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(1997)
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</bibRefCitation>
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.
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C370656FFF85FFB0FF5B4C1257C4FC1C" pageId="9" pageNumber="10" type="materials_examined">
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<paragraph id="8BD536E4FF85FFB0FF5B4C1257C4FC1C" blockId="9.[151,1436,746,1603]" pageId="9" pageNumber="10">
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TYPE LOCALITY. Originally “in deserti australioris collibus ardentissimis” (
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<bibRefCitation id="EFFB4B15FF85FFB0FBE04C0D52C5FCE4" author="Pallas, P. S." box="[1068,1213,815,840]" pageId="9" pageNumber="10" refId="ref61618" refString="Pallas, P. S. (1771) Reise durch verschiedene Provinzen des Russisches Reichs. Kaiserlichen Akademie der Wissenschaften, St. Petersburg, 504 pp." type="book" year="1771">Pallas 1771</bibRefCitation>
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); restricted to the “Inderskija Gory, Gebiet des unteren Uralflusses” [region of lower Ural River, W
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<collectingCountry id="F37D7674FF85FFB0FBAD4C70529FFCC7" box="[1121,1255,850,875]" name="Kazakhstan" pageId="9" pageNumber="10">Kazakhstan</collectingCountry>
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] by
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<bibRefCitation id="EFFB4B15FF85FFB0FAEC4C715743FC22" author="Mertens, R. & Muller, L." pageId="9" pageNumber="10" pagination="1 - 62" refId="ref58855" refString="Mertens, R. & Muller, L. (1928) Liste der Amphibien und Reptilien Europas. Abhandlungen der Senckenbergischen Naturfoschenden Gesellschaft, 41, 1 - 62." type="journal article" year="1928">Mertens & Müller (1928)</bibRefCitation>
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. This restriction was not accompanied by a
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<typeStatus id="54D18846FF85FFB0FCF04C5755DBFC22" box="[828,931,885,910]" pageId="9" pageNumber="10" type="lectotype">lectotype</typeStatus>
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or
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<typeStatus id="54D18846FF85FFB0FC014C54525EFC22" box="[973,1062,886,910]" pageId="9" pageNumber="10" type="neotype">neotype</typeStatus>
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designation and is therefore not valid as such (
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<bibRefCitation id="EFFB4B15FF85FFB0FEFA4CB557C9FC1C" author="Uetz, P." box="[310,433,919,944]" pageId="9" pageNumber="10" refId="ref67280" refString="Uetz, P. (2013) The Reptile database. Available from: http: // www. reptile-database. org / (accessed 8 July 2014)" type="url" year="2013">Uetz 2013</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C370656FFF85FFB0FF5B4C99576BFBF2" pageId="9" pageNumber="10" type="distribution">
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<paragraph id="8BD536E4FF85FFB0FF5B4C9957FDFC5A" blockId="9.[151,1436,746,1603]" pageId="9" pageNumber="10">
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DISTRIBUTION. SE part of European
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<collectingCountry id="F37D7674FF85FFB0FDF54C9854FEFC7F" box="[569,646,954,979]" name="Russia" pageId="9" pageNumber="10">Russia</collectingCountry>
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,
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<collectingCountry id="F37D7674FF85FFB0FD5F4C985561FC7F" box="[659,793,954,979]" name="Kazakhstan" pageId="9" pageNumber="10">Kazakhstan</collectingCountry>
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,
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<collectingCountry id="F37D7674FF85FFB0FCE94C9855BBFC7F" box="[805,963,954,979]" name="Turkmenistan" pageId="9" pageNumber="10">Turkmenistan</collectingCountry>
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,
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<collectingCountry id="F37D7674FF85FFB0FC1D4C98522AFC7F" box="[977,1106,954,979]" name="Uzbekistan" pageId="9" pageNumber="10">Uzbekistan</collectingCountry>
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,
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<collectingCountry id="F37D7674FF85FFB0FB924C99529AFC7F" box="[1118,1250,955,979]" name="Kyrgyzstan" pageId="9" pageNumber="10">Kyrgyzstan</collectingCountry>
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,
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<collectingCountry id="F37D7674FF85FFB0FB234C98531AFC7F" box="[1263,1378,954,979]" name="Tajikistan" pageId="9" pageNumber="10">Tajikistan</collectingCountry>
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, NW
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<collectingCountry id="F37D7674FF85FFB0FF5B4CFF56A2FC5A" box="[151,218,989,1014]" name="China" pageId="9" pageNumber="10">China</collectingCountry>
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and
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<collectingCountry id="F37D7674FF85FFB0FEDC4CFF57F9FC5A" box="[272,385,989,1014]" name="Mongolia" pageId="9" pageNumber="10">Mongolia</collectingCountry>
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.
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</paragraph>
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<paragraph id="8BD536E4FF85FFB0FF5B4B225288FBB4" blockId="9.[151,1436,746,1603]" box="[151,1264,1023,1048]" pageId="9" pageNumber="10">
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DISTRIBUTION IN
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<collectingCountry id="F37D7674FF85FFB0FE914B2257EDFBBB" box="[349,405,1024,1047]" name="Iran" pageId="9" pageNumber="10">IRAN</collectingCountry>
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.
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<figureCitation id="13512A61FF85FFB0FE684CDD578EFBB4" box="[420,502,1023,1048]" captionStart="FIGURES 14–19. 14" captionStartId="74.[151,264,1557,1580]" captionTargetBox="[171,1416,213,1516]" captionTargetId="figure-63@74.[151,1436,193,1536]" captionTargetPageId="74" captionText="FIGURES 14–19. 14. Phrynocephalus helioscopus. 15. Phrynocephalus maculatus. 16. Phrynocephalus mystaceus. 17. Phrynocephalus ornatus. Blue square denotes the type locality for P. o. vindumi. 18. Phrynocephalus persicus. 19. Phrynocephalus scutellatus." figureDoi="http://doi.org/10.5281/zenodo.4929713" httpUri="https://zenodo.org/record/4929713/files/figure.png" pageId="9" pageNumber="10">Fig. 14</figureCitation>
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. Occurs only by the borders with Turkmenistan by the Caspian Sea.
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</paragraph>
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<paragraph id="8BD536E4FF85FFB0FF5B4B01576BFBF2" blockId="9.[151,1436,746,1603]" pageId="9" pageNumber="10">HABITAT. Clay and stony deserts, sometimes sandy areas with high admixture of pebbles, usually in areas of sparse vegetation.</paragraph>
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</subSubSection>
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<subSubSection id="C370656FFF85FFB0FF5B4B4A5694FA52" pageId="9" pageNumber="10" type="discussion">
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<paragraph id="8BD536E4FF85FFB0FF5B4B4A5694FA52" blockId="9.[151,1436,746,1603]" pageId="9" pageNumber="10">
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REMARKS.
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<taxonomicName id="4C6A4D67FF85FFB0FEDB4B45544BFBD3" authorityName="de Filippi" authorityYear="1863" box="[279,563,1127,1152]" class="Reptilia" family="Agamidae" genus="Phrynocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="persicus">
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<emphasis id="B91EEAF6FF85FFB0FEDB4B45544BFBD3" box="[279,563,1127,1152]" italics="true" pageId="9" pageNumber="10">Phrynocephalus persicus</emphasis>
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</taxonomicName>
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was separated from
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<taxonomicName id="4C6A4D67FF85FFB0FCE84B4B55B2FB2C" baseAuthorityName="Pallas" baseAuthorityYear="1771" box="[804,970,1127,1152]" class="Reptilia" family="Agamidae" genus="Phrynocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="helioscopus">
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<emphasis id="B91EEAF6FF85FFB0FCE84B4B55B2FB2C" box="[804,970,1127,1152]" italics="true" pageId="9" pageNumber="10">P. helioscopus</emphasis>
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</taxonomicName>
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on the basis of the analysis of allozyme variation (
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<bibRefCitation id="EFFB4B15FF85FFB0FEC74BA85427FB0F" author="Mezhzherin, S. V. & Golubev, M. L." box="[267,607,1162,1187]" pageId="9" pageNumber="10" pagination="72 - 74" refId="ref59156" refString="Mezhzherin, S. V. & Golubev, M. L. (1989) The genetic divergence of Phrynocephalus Kaup (Reptilia, Agamidae) of the USSR fauna [in Russian]. Reports of Ukraine SSR Academy of Sciences. Series B Geology, Chemistry and Biological Sciences, 12, 72 - 74." type="journal article" year="1989">Mezhzherin & Golubev 1989</bibRefCitation>
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). There is an immense chaos regarding the form
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<taxonomicName id="4C6A4D67FF85FFB0FB5E4BA85336FB0F" authority="Mehely" authorityName="Mehely" box="[1170,1358,1162,1187]" class="Reptilia" family="Agamidae" genus="Phrynocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="horvathi">
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<emphasis id="B91EEAF6FF85FFB0FB5E4BA85289FB0F" box="[1170,1265,1162,1187]" italics="true" pageId="9" pageNumber="10">horvathi</emphasis>
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Méhely
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</taxonomicName>
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which was originally described as a subspecies of
|
||
<taxonomicName id="4C6A4D67FF85FFB0FDB54B8C5564FB6A" baseAuthorityName="Pallas" baseAuthorityYear="1771" box="[633,796,1197,1222]" class="Reptilia" family="Agamidae" genus="Phrynocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="helioscopus">
|
||
<emphasis id="B91EEAF6FF85FFB0FDB54B8C5564FB6A" box="[633,796,1197,1222]" italics="true" pageId="9" pageNumber="10">P. helioscopus</emphasis>
|
||
</taxonomicName>
|
||
from the foothills of
|
||
<collectingRegion id="49AEF806FF85FFB0FBC54B8F522CFB6A" box="[1033,1108,1197,1222]" country="Armenia" name="Ararat" pageId="9" pageNumber="10">Ararat</collectingRegion>
|
||
,
|
||
<collectingCountry id="F37D7674FF85FFB0FB924B8F52B8FB6A" box="[1118,1216,1197,1222]" name="Armenia" pageId="9" pageNumber="10">Armenia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFFB4B15FF85FFB0FB024B8F531DFB6A" author="Mehely, L." box="[1230,1381,1197,1222]" pageId="9" pageNumber="10" pagination="81 - 87" refId="ref58344" refString="Mehely, L. (1894) Beitrage zur Herpetologie Transkaukasiens und Armeniens. Zoologischer Anzeiger, 17, 81 - 87." type="journal article" year="1894">Méhely 1894</bibRefCitation>
|
||
) and which is endemic to E
|
||
<collectingCountry id="F37D7674FF85FFB0FE6F4BED547FFB44" box="[419,519,1231,1256]" name="Turkey" pageId="9" pageNumber="10">Anatolia</collectingCountry>
|
||
, S
|
||
<collectingCountry id="F37D7674FF85FFB0FDE14BED54EAFB44" box="[557,658,1231,1256]" name="Armenia" pageId="9" pageNumber="10">Armenia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F37D7674FF85FFB0FD6C4BED5563FB44" box="[672,795,1231,1256]" name="Azerbaijan" pageId="9" pageNumber="10">Azerbaijan</collectingCountry>
|
||
and NW
|
||
<collectingCountry id="F37D7674FF85FFB0FC424BF255C7FB4B" box="[910,959,1232,1255]" name="Iran" pageId="9" pageNumber="10">Iran</collectingCountry>
|
||
. In
|
||
<collectingCountry id="F37D7674FF85FFB0FC204BF25261FB4B" box="[1004,1049,1232,1255]" name="Iran" pageId="9" pageNumber="10">Iran</collectingCountry>
|
||
it is restricted to the vicinity and north of the Lake Urmia (
|
||
<bibRefCitation id="EFFB4B15FF85FFB0FE7B4BD054D1FAA7" author="Melnikov, D. A. & Ananjeva, N. B. & Agasyan, A. L. & Rajabizadeh, M." box="[439,681,1266,1291]" pageId="9" pageNumber="10" pagination="286 - 297" refId="ref58367" refString="Melnikov, D. A., Ananjeva, N. B., Agasyan, A. L. & Rajabizadeh, M. (2008) Historical background and taxonomic status of the Persian toad head agama, Phrynocephalus persicus de Filippi, 1863 and Horvath's sun watcher toad head agama Phrynocephalus helioscopus horvathi Mehely, 1894. In: Ananjeva, N. B., Danilov, I. G., Dunayev, E. A., Ishchenko, V. G., Lada, G. A., Litvinchuk, S. N., Orlova, V. F., Smirina, E. M., Tuniyev, B. S. & Khalikov, R. G. (Eds.), The problems of herpetology. Proceedings of the 3 th Meeting of the Nikolsky Herpetological Society 9 - 13 October 2006, Putschino, pp. 286 - 297." type="book chapter" year="2008">
|
||
Melnikov
|
||
<emphasis id="B91EEAF6FF85FFB0FDE04BD1541DFAA7" box="[556,613,1266,1291]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
|
||
2008
|
||
</bibRefCitation>
|
||
). Some authors treat this form as it was described—a subspecies of
|
||
<taxonomicName id="4C6A4D67FF85FFB0FF5B4A345738FA82" baseAuthorityName="Pallas" baseAuthorityYear="1771" box="[151,320,1301,1326]" class="Reptilia" family="Agamidae" genus="Phrynocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="helioscopus">
|
||
<emphasis id="B91EEAF6FF85FFB0FF5B4A345738FA82" box="[151,320,1301,1326]" italics="true" pageId="9" pageNumber="10">P. helioscopus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFFB4B15FF85FFB0FE994A3754CAFA82" author="Rastegar-Pouyani, N. & Kami, H. G. & Rajabzadeh, M. & Shafiei, S. & Anderson, S. C." box="[341,690,1301,1326]" pageId="9" pageNumber="10" pagination="43 - 66" refId="ref63314" refString="Rastegar-Pouyani, N., Kami, H. G., Rajabzadeh, M., Shafiei, S. & Anderson, S. C. (2008) Annotated checklist of Amphibians and Reptiles of Iran. Iranian Journal of Animal Biosystematics, 4, 43 - 66." type="journal article" year="2008">
|
||
Rastegar-Pouyani
|
||
<emphasis id="B91EEAF6FF85FFB0FDE74A345412FA82" box="[555,618,1301,1326]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
|
||
2008
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFB4B15FF85FFB0FD084A3755A8FA82" author="Tosunoglu, M. & Gul, C. & Topyildiz, H. & Uysal, I." box="[708,976,1301,1326]" pageId="9" pageNumber="10" pagination="247 - 252" refId="ref67094" refString="Tosunoglu, M., Gul, C., Topyildiz, H. & Uysal, I. (2011) Notes on Distribution, Ecology, and Morphological Characters of Phrynocephalus helioscopus horvathi Mehely, 1894 from Northeast Anatolia. Russian Journal of Herpetology, 18, 247 - 252." type="journal article" year="2011">
|
||
Tosunoğlu
|
||
<emphasis id="B91EEAF6FF85FFB0FC844A3455FFFA82" box="[840,903,1301,1326]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
), some as a subspecies of
|
||
<taxonomicName id="4C6A4D67FF85FFB0FAD54A3453E4FA81" authorityName="de Filippi" authorityYear="1863" box="[1305,1436,1302,1325]" class="Reptilia" family="Agamidae" genus="Phrynocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="persicus">
|
||
<emphasis id="B91EEAF6FF85FFB0FAD54A3453E4FA81" box="[1305,1436,1302,1325]" italics="true" pageId="9" pageNumber="10">P. persicus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFFB4B15FF85FFB0FF524A155789FAFC" author="Barabanov, A. V. & Ananjeva, N. B." box="[158,497,1335,1360]" pageId="9" pageNumber="10" pagination="1 - 56" refId="ref46583" refString="Barabanov, A. V. & Ananjeva, N. B. (2007) Catalogue of the available scientific species-group names for lizards of the genus Phrynocephalus Kaup, 1825 (Reptilia, Sauria, Agamidae). Zootaxa, 1399, 1 - 56." type="journal article" year="2007">Barabanov & Ananjeva 2007</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFB4B15FF85FFB0FE334A155484FAFC" author="Arakelyan, M. & Danielyan, F. & Corti, C. & Sindaco, R. & Leviton, A. E." box="[511,764,1335,1360]" pageId="9" pageNumber="10" refId="ref45580" refString="Arakelyan, M., Danielyan, F., Corti, C., Sindaco, R. & Leviton, A. E. (2011) Herpetofauna of Armenia and Nagorno-Karabakh. Society for the Study of Amphibians and Reptiles, Oxford, Ohio, 149 pp." type="book" year="2011">
|
||
Arakelyan
|
||
<emphasis id="B91EEAF6FF85FFB0FDB14A1B54CFFAFC" box="[637,695,1335,1360]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFB4B15FF85FFB0FCC54A15527FFAFC" author="Solovyeva, E. N. & Poyarkov, N. A. & Dunaev, E. A. & Duysebayeva, T. N. & Bannikova, A. A." box="[777,1031,1335,1360]" pageId="9" pageNumber="10" pagination="842 - 856" refId="ref65720" refString="Solovyeva, E. N., Poyarkov, N. A., Dunaev, E. A., Duysebayeva, T. N. & Bannikova, A. A. (2011) Molecular Differentiation and Taxonomy of the Sunwatcher Toad Headed Agama Species Complex Phrynocephalus Superspecies helioscopus (Pallas 1771) (Reptilia: Agamidae). Russian Journal of Genetics, 47, 842 - 856. http: // dx. doi. org / 10.1134 / s 1022795411070155" type="journal article" year="2011">
|
||
Solovyeva
|
||
<emphasis id="B91EEAF6FF85FFB0FC444A1B55BBFAFC" box="[904,963,1335,1360]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFB4B15FF85FFB0FBD94A155340FAFC" author="Milto, K. D. & Barabanov, A. V." box="[1045,1336,1335,1360]" pageId="9" pageNumber="10" pagination="155 - 170" refId="ref59215" refString="Milto, K. D. & Barabanov, A. V. (2012) A catalogueof the agamid chamaeleonid types in the collection of the Zoological institute, Russian Academy of Sciences, St. Petersburg. Russian Journal of Herpetology, 19, 155 - 170." type="journal article" year="2012">Milto & Barabanov 2012</bibRefCitation>
|
||
) and yet others as a full species (
|
||
<bibRefCitation id="EFFB4B15FF85FFB0FE614A785401FADF" author="Cicek, K. & Ayaz, D. & Tok, C. V. & Tayhan, Y." box="[429,633,1370,1395]" pageId="9" pageNumber="10" pagination="69 - 73" refId="ref48566" refString="Cicek, K., Ayaz, D., Tok, C. V. & Tayhan, Y. (2011) Data on food composition of Phrynocephalus horvathi Mehely, 1894 (Reptilia: Agamidae) in Mount Ararat (Northeastern Anatolia, Turkey). Ecologia Balkanica, 3, 69 - 73." type="journal article" year="2011">
|
||
Çiçek
|
||
<emphasis id="B91EEAF6FF85FFB0FE354A79544CFADF" box="[505,564,1370,1395]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
). According to
|
||
<bibRefCitation id="EFFB4B15FF85FFB0FCE24A785589FADF" author="Anderson, S. C." box="[814,1009,1370,1395]" pageId="9" pageNumber="10" refId="ref45318" refString="Anderson, S. C. (1999) The lizards of Iran. Society for the Study of Amphibians and Reptiles, Oxford, Ohio, 442 pp." type="book" year="1999">Anderson (1999)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C6A4D67FF85FFB0FC334A795207FADF" box="[1023,1151,1370,1395]" class="Reptilia" family="Agamidae" genus="Phrynocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="horvathi">
|
||
<emphasis id="B91EEAF6FF85FFB0FC334A795207FADF" box="[1023,1151,1370,1395]" italics="true" pageId="9" pageNumber="10">P. horvathi</emphasis>
|
||
</taxonomicName>
|
||
appears to be a younger synonym of
|
||
<taxonomicName id="4C6A4D67FF85FFB0FEE94A5C57DAFA39" authorityName="de Filippi" authorityYear="1863" box="[293,418,1406,1429]" class="Reptilia" family="Agamidae" genus="Phrynocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="persicus">
|
||
<emphasis id="B91EEAF6FF85FFB0FEE94A5C57DAFA39" box="[293,418,1406,1429]" italics="true" pageId="9" pageNumber="10">P. persicus</emphasis>
|
||
</taxonomicName>
|
||
. This assumption is in accordance with the results of mtDNA analyses of
|
||
<taxonomicName id="4C6A4D67FF85FFB0FB224A5C568CFA1B" class="Reptilia" family="Agamidae" genus="Phrynocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="helioscopuspersicus">
|
||
<emphasis id="B91EEAF6FF85FFB0FB224A5C568CFA1B" italics="true" pageId="9" pageNumber="10">P. helioscopuspersicus</emphasis>
|
||
</taxonomicName>
|
||
complex by
|
||
<bibRefCitation id="EFFB4B15FF85FFB0FE424ABD54E6FA14" author="Solovyeva, E. N. & Poyarkov, N. A. & Dunaev, E. A. & Duysebayeva, T. N. & Bannikova, A. A." box="[398,670,1439,1464]" pageId="9" pageNumber="10" pagination="842 - 856" refId="ref65720" refString="Solovyeva, E. N., Poyarkov, N. A., Dunaev, E. A., Duysebayeva, T. N. & Bannikova, A. A. (2011) Molecular Differentiation and Taxonomy of the Sunwatcher Toad Headed Agama Species Complex Phrynocephalus Superspecies helioscopus (Pallas 1771) (Reptilia: Agamidae). Russian Journal of Genetics, 47, 842 - 856. http: // dx. doi. org / 10.1134 / s 1022795411070155" type="journal article" year="2011">
|
||
Solovyeva
|
||
<emphasis id="B91EEAF6FF85FFB0FDC34A835432FA14" box="[527,586,1439,1464]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
|
||
(2011)
|
||
</bibRefCitation>
|
||
which proved
|
||
<taxonomicName id="4C6A4D67FF85FFB0FC824ABD55D5FA14" box="[846,941,1439,1464]" class="Reptilia" family="Agamidae" genus="Phrynocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="horvathi">
|
||
<emphasis id="B91EEAF6FF85FFB0FC824ABD55D5FA14" box="[846,941,1439,1464]" italics="true" pageId="9" pageNumber="10">horvathi</emphasis>
|
||
</taxonomicName>
|
||
to be an inner lineage of
|
||
<taxonomicName id="4C6A4D67FF85FFB0FB2C4A835327FA1B" authorityName="de Filippi" authorityYear="1863" box="[1248,1375,1440,1464]" class="Reptilia" family="Agamidae" genus="Phrynocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="persicus">
|
||
<emphasis id="B91EEAF6FF85FFB0FB2C4A835327FA1B" box="[1248,1375,1440,1464]" italics="true" pageId="9" pageNumber="10">P. persicus</emphasis>
|
||
</taxonomicName>
|
||
. The name
|
||
<taxonomicName id="4C6A4D67FF85FFB0FF124AE05745FA77" box="[222,317,1474,1499]" class="Reptilia" family="Agamidae" genus="Phrynocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="horvathi">
|
||
<emphasis id="B91EEAF6FF85FFB0FF124AE05745FA77" box="[222,317,1474,1499]" italics="true" pageId="9" pageNumber="10">horvathi</emphasis>
|
||
</taxonomicName>
|
||
was stabilized by
|
||
<bibRefCitation id="EFFB4B15FF85FFB0FDD04AE0555FFA77" author="Melnikov, D. & Melnikova, E. & Nazarov, R. & Rajabizadeh, M." box="[540,807,1474,1499]" pageId="9" pageNumber="10" pagination="34 - 46" refId="ref58533" refString="Melnikov, D., Melnikova, E., Nazarov, R. & Rajabizadeh, M. (2013) Taxonomic revision of Phrynocephalus persicus de Filippi, 1863 complex with description of a new species from Zagros, southern Iran. Modern Herpetology, 2013, 34 - 46." type="journal article" year="2013">
|
||
Melnikov
|
||
<emphasis id="B91EEAF6FF85FFB0FD5F4AE154B7FA77" box="[659,719,1474,1499]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
|
||
(2013)
|
||
</bibRefCitation>
|
||
, who designated
|
||
<typeStatus id="54D18846FF85FFB0FC384AE15235FA77" box="[1012,1101,1475,1499]" pageId="9" pageNumber="10" type="neotype">neotype</typeStatus>
|
||
(see remarks by
|
||
<taxonomicName id="4C6A4D67FF85FFB0FAD04AE153E4FA76" authorityName="de Filippi" authorityYear="1863" box="[1308,1436,1475,1498]" class="Reptilia" family="Agamidae" genus="Phrynocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="persicus">
|
||
<emphasis id="B91EEAF6FF85FFB0FAD04AE153E4FA76" box="[1308,1436,1475,1498]" italics="true" pageId="9" pageNumber="10">P. persicus</emphasis>
|
||
</taxonomicName>
|
||
below).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C370656FFF85FFB0FF5B492A5750F9EF" pageId="9" pageNumber="10" type="reference_group">
|
||
<paragraph id="8BD536E4FF85FFB0FF5B492A5750F9EF" blockId="9.[151,1436,746,1603]" pageId="9" pageNumber="10">
|
||
REFERENCES.
|
||
<bibRefCitation id="EFFB4B15FF85FFB0FEF84925578DF98C" author="Anderson, S. C." box="[308,501,1543,1568]" pageId="9" pageNumber="10" refId="ref45318" refString="Anderson, S. C. (1999) The lizards of Iran. Society for the Study of Amphibians and Reptiles, Oxford, Ohio, 442 pp." type="book" year="1999">Anderson (1999)</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFB4B15FF85FFB0FDCD49255486F98C" author="Ananjeva, N. B. & Orlov, N. & Khalikov, R. G. & Darevskij, I. S. & Ryabov, S. A. & Barabanov, A. V." box="[513,766,1543,1568]" pageId="9" pageNumber="10" refId="ref44840" refString="Ananjeva, N. B., Orlov, N., Khalikov, R. G., Darevskij, I. S., Ryabov, S. A. & Barabanov, A. V. (2006) The reptiles of the Northern Eurasia. Taxonomic diversity, distribution, conservation status. Pensoft Publishers, Sofia, 245 pp." type="book" year="2006">
|
||
Ananjeva
|
||
<emphasis id="B91EEAF6FF85FFB0FDBF492B54D3F98C" box="[627,683,1543,1568]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
|
||
(2006)
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFB4B15FF85FFB0FCC549255270F98C" author="Melnikov, D. A. & Ananjeva, N. B. & Agasyan, A. L. & Rajabizadeh, M." box="[777,1032,1543,1568]" pageId="9" pageNumber="10" pagination="286 - 297" refId="ref58367" refString="Melnikov, D. A., Ananjeva, N. B., Agasyan, A. L. & Rajabizadeh, M. (2008) Historical background and taxonomic status of the Persian toad head agama, Phrynocephalus persicus de Filippi, 1863 and Horvath's sun watcher toad head agama Phrynocephalus helioscopus horvathi Mehely, 1894. In: Ananjeva, N. B., Danilov, I. G., Dunayev, E. A., Ishchenko, V. G., Lada, G. A., Litvinchuk, S. N., Orlova, V. F., Smirina, E. M., Tuniyev, B. S. & Khalikov, R. G. (Eds.), The problems of herpetology. Proceedings of the 3 th Meeting of the Nikolsky Herpetological Society 9 - 13 October 2006, Putschino, pp. 286 - 297." type="book chapter" year="2008">
|
||
Melnikov
|
||
<emphasis id="B91EEAF6FF85FFB0FCB1492B55CDF98C" box="[893,949,1543,1568]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
|
||
(2008)
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFB4B15FF85FFB0FBD849255362F98C" author="Arakelyan, M. & Danielyan, F. & Corti, C. & Sindaco, R. & Leviton, A. E." box="[1044,1306,1543,1568]" pageId="9" pageNumber="10" refId="ref45580" refString="Arakelyan, M., Danielyan, F., Corti, C., Sindaco, R. & Leviton, A. E. (2011) Herpetofauna of Armenia and Nagorno-Karabakh. Society for the Study of Amphibians and Reptiles, Oxford, Ohio, 149 pp." type="book" year="2011">
|
||
Arakelyan
|
||
<emphasis id="B91EEAF6FF85FFB0FB43492B52BFF98C" box="[1167,1223,1543,1568]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
|
||
(2011)
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFB4B15FF85FFB0FAE94925575BF9EF" author="Solovyeva, E. N. & Poyarkov, N. A. & Dunaev, E. A. & Duysebayeva, T. N. & Bannikova, A. A." pageId="9" pageNumber="10" pagination="842 - 856" refId="ref65720" refString="Solovyeva, E. N., Poyarkov, N. A., Dunaev, E. A., Duysebayeva, T. N. & Bannikova, A. A. (2011) Molecular Differentiation and Taxonomy of the Sunwatcher Toad Headed Agama Species Complex Phrynocephalus Superspecies helioscopus (Pallas 1771) (Reptilia: Agamidae). Russian Journal of Genetics, 47, 842 - 856. http: // dx. doi. org / 10.1134 / s 1022795411070155" type="journal article" year="2011">
|
||
Solovyeva
|
||
<emphasis id="B91EEAF6FF85FFB0FF5B490956B7F9EF" box="[151,207,1578,1603]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
|
||
(2011)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |