126 lines
14 KiB
XML
126 lines
14 KiB
XML
<document id="559FE700225E5895D590422E69E59E19" ID-CLB-Dataset="4530" ID-DOI="10.14446/FI.2016.251" ID-GBIF-Dataset="0679bb20-5192-433f-b780-45d5c35ab853" ID-ISSN="2533-4069" ID-Zenodo-Dep="5387908" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630306085997" checkinUser="diego" docAuthor="Benicio, Jose R. W., Pires, Etiene F., Da Rosa, Átila A. S., Spiekermann, Rafael, Uhl, Dieter & Jasper, André" docDate="2016" docId="03C387E6FE32FFF82286FF18AED4FDD3" docLanguage="en" docName="NM_IF_c3_4_16_clanek11_Benicio.pdf" docOrigin="Fossil Imprint 72 (3 - 4)" docSource="http://dx.doi.org/10.14446/fi.2016.251" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.4:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="4" docTitle="Itaquixylon heterogenum BENICIO, PIRES, DA ROSA, SPIEKERMANN, UHL et JASPER 2016, sp. nov." docType="treatment" docVersion="7" lastPageNumber="254" masterDocId="FFFAFF9EFE31FFFB220DFF8FAD76FF9C" masterDocTitle="A New Fossil Fabaceae Wood From The Pleistocene Touro Passo Formation Of Rio Grande Do Sul, Brazil" masterLastPageNumber="264" masterPageNumber="251" pageNumber="254" updateTime="1699240847107" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="996ADC906165F2402DBAC0D53A038BF2">A New Fossil Fabaceae Wood From The Pleistocene Touro Passo Formation Of Rio Grande Do Sul, Brazil</mods:title>
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<mods:namePart id="864EA7B5DC6A286F61E154AFEB6D9819">Benicio, Jose R. W.</mods:namePart>
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<mods:namePart id="76EFB3629E599974675FCF0F5A7CE0AC">Pires, Etiene F.</mods:namePart>
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<mods:roleTerm id="7EECDB5564598D7BA46A826ECAA656C3">Author</mods:roleTerm>
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<mods:namePart id="74B99B3B1342EB16341F9476FCB965AA">Da Rosa, Átila A. S.</mods:namePart>
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<mods:namePart id="B0699530704A65DB8F04D6AB4E199837">Spiekermann, Rafael</mods:namePart>
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<mods:namePart id="904FCF9120BC91B6804D7A73BD2DD7DD">Uhl, Dieter</mods:namePart>
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<mods:namePart id="0791E42F24206FAD9AFD624F98515CDD">Jasper, André</mods:namePart>
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<mods:title id="B66AE06FDD45D0869F339E367BE50F77">Fossil Imprint</mods:title>
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<mods:part id="8C782E2E9890CB7F88D72940182E974C">
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<mods:date id="C12F86A2B0CBF8832B1C5C5096123B92">2016</mods:date>
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<mods:number id="53706268D25A4872F595004BC178CC31">2016-12-30</mods:number>
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<mods:number id="ED5D5878C4EC7A3E627D43DDDCE04B46">72</mods:number>
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<mods:number id="DDD14AFCDDFF3FB383AA969A6FFC9526">3 - 4</mods:number>
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<mods:start id="B9E3304BF051B79B3F63374A80F4820C">251</mods:start>
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<mods:url id="3B748C6E0E679885BB9946D93168CD3E">http://dx.doi.org/10.14446/fi.2016.251</mods:url>
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<mods:classification id="E3C6961172718CA6BE94F6AACCE68437">journal article</mods:classification>
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<mods:identifier id="CEF86110870AA09CB6DFE0FAAEC6B2BF" type="DOI">10.14446/FI.2016.251</mods:identifier>
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<mods:identifier id="C5A905A7F0409A0BDA16802EAD8CEBC1" type="ISSN">2533-4069</mods:identifier>
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<treatment id="03C387E6FE32FFF82286FF18AED4FDD3" ID-DOI="http://doi.org/10.5281/zenodo.5467488" ID-GBIF-Taxon="185367240" ID-Zenodo-Dep="5467488" LSID="urn:lsid:plazi:treatment:03C387E6FE32FFF82286FF18AED4FDD3" httpUri="http://treatment.plazi.org/id/03C387E6FE32FFF82286FF18AED4FDD3" lastPageNumber="254" pageId="3" pageNumber="254">
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<subSubSection id="C370657BFE32FFF82286FF18AC15FEA1" pageId="3" pageNumber="254" type="nomenclature">
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<paragraph id="8BD536F0FE32FFF82286FF18AF0DFF53" blockId="3.[139,712,151,207]" pageId="3" pageNumber="254">
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<heading id="D09D819CFE32FFF82286FF18AF0DFF53" centered="true" fontSize="8" level="2" pageId="3" pageNumber="254" reason="2">
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<emphasis id="B91EEAE2FE32FFF82286FF18AF0DFF53" bold="true" pageId="3" pageNumber="254">
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<taxonomicName id="4C6A4D73FE32FFF82286FF18AF69FF52" authority="BENICIO, PIRES, DA ROSA, SPIEKERMANN, UHL et JASPER" authorityName="BENICIO, PIRES, DA ROSA, SPIEKERMANN, UHL et JASPER" authorityYear="2016" class="Magnoliopsida" family="Fabaceae" genus="Itaquixylon" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="254" phylum="Tracheophyta" rank="species" species="heterogenum" status="sp. nov.">
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<emphasis id="B91EEAE2FE32FFF82286FF18ACD4FF33" bold="true" box="[139,418,151,175]" italics="true" pageId="3" pageNumber="254">Itaquixylon heterogenum</emphasis>
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BENICIO, PIRES, DA ROSA, SPIEKERMANN, UHL et JASPER
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</taxonomicName>
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<taxonomicNameLabel id="A22D5799FE32FFF82028FF37AF0DFF53" box="[549,635,184,207]" pageId="3" pageNumber="254" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD536F0FE32FFF8229DFF68AC15FEA1" blockId="3.[106,745,231,317]" pageId="3" pageNumber="254">
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H o l o t y p e. UFSM 12032 (Repository: Coleção Paleontológica da Universidade Federal de Santa Maria,
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<collectingRegion id="49AEF812FE32FFF820CFFE89AC70FEA1" country="Brazil" name="Rio Grande do Sul" pageId="3" pageNumber="254">Rio Grande do Sul</collectingRegion>
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,
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<collectingCountry id="F37D7660FE32FFF8231CFEA9AC21FEA1" box="[273,343,294,317]" name="Brazil" pageId="3" pageNumber="254">Brazil</collectingCountry>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C370657BFE32FFF8229DFEDAAED4FDD3" pageId="3" pageNumber="254" type="description">
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<paragraph id="8BD536F0FE32FFF8229DFEDAAC8CFE10" blockId="3.[106,745,341,396]" pageId="3" pageNumber="254">
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E t y m o l o g y. The specific name,
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<emphasis id="B91EEAE2FE32FFF82019FEDAAFD3FEF0" box="[532,677,341,364]" italics="true" pageId="3" pageNumber="254">
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<taxonomicName id="4C6A4D73FE32FFF82019FEDAAFD7FEF0" authorityName="BENICIO, PIRES, DA ROSA, SPIEKERMANN, UHL et JASPER" authorityYear="2016" box="[532,673,341,364]" class="Magnoliopsida" family="Fabaceae" genus="Itaquixylon" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="254" phylum="Tracheophyta" rank="species" species="heterogenum">heterogenum</taxonomicName>
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,
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</emphasis>
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is due to the presence of heterocellular rays.
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</paragraph>
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<paragraph id="8BD536F0FE32FFF8229DFE2BAF19FE27" blockId="3.[144,623,420,443]" box="[144,623,420,443]" pageId="3" pageNumber="254">T y p e h o r i z o n. Touro Passo Formation.</paragraph>
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<paragraph id="8BD536F0FE32FFF8229DFE5CAF12FE76" blockId="3.[144,612,467,490]" box="[144,612,467,490]" pageId="3" pageNumber="254">A g e. Late Pleistocene, 14,600 years (TL).</paragraph>
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<paragraph id="8BD536F0FE32FFF8229DFD8EAC81FDA4" blockId="3.[106,745,513,568]" pageId="3" pageNumber="254">
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<materialsCitation id="3B023CADFE32FFF8229DFD8EAC85FDA4" ID-GBIF-Occurrence="3340942301" collectionCode="T" country="Brazil" latitude="-29.39886" location="Itaqui" longLatPrecision="1" longitude="-56.70164" pageId="3" pageNumber="254" specimenCount="1" stateProvince="Rio Grande do Sul">
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<collectionCode id="ED7BAE35FE32FFF8229DFD8EADD6FD84" box="[144,160,513,536]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="254">T</collectionCode>
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y p e l o c a l i t y. (
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<geoCoordinate id="EE5E5037FE32FFF82379FD8EAC8DFD84" box="[372,507,513,536]" degrees="29" direction="south" minutes="23" orientation="latitude" pageId="3" pageNumber="254" precision="1" seconds="55.9" value="-29.39886">29º23'55.9"S</geoCoordinate>
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/
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<geoCoordinate id="EE5E5037FE32FFF8200EFD8EAFEEFD84" box="[515,664,513,536]" degrees="56" direction="west" minutes="42" orientation="longitude" pageId="3" pageNumber="254" precision="1" seconds="05.9" value="-56.70164">56º42'05.9"W</geoCoordinate>
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),
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<location id="8EB5602BFE32FFF820A1FD8EAF9FFD84" LSID="urn:lsid:plazi:treatment:03C387E6FE32FFF82286FF18AED4FDD3:8EB5602BFE32FFF820A1FD8EAF9FFD84" box="[684,745,513,536]" country="Brazil" latitude="-29.39886" longLatPrecision="1" longitude="-56.70164" name="Itaqui" pageId="3" pageNumber="254" stateProvince="Rio Grande do Sul">Itaqui</location>
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city,
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<collectingRegion id="49AEF812FE32FFF82290FDAEACD5FDA4" box="[157,419,545,568]" country="Brazil" name="Rio Grande do Sul" pageId="3" pageNumber="254">Rio Grande do Sul State</collectingRegion>
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,
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<collectingCountry id="F37D7660FE32FFF823A3FDAEAC85FDA4" box="[430,499,545,568]" name="Brazil" pageId="3" pageNumber="254">Brazil</collectingCountry>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BD536F0FE32FFF8229DFDDFADC3FBD8" blockId="3.[106,745,592,1092]" pageId="3" pageNumber="254">D i a g n o s i s. Angiosperm wood, diffuse porosity, numerous vessels in tangential bands; vessels generally solitary, sometimes in radial multiples of 3 to 5, circular to oval in outline, with narrow diameter and short length, helical thickenings absent, tyloses present; simple perforation plates; alternate intervessel pits, vestured, circular, small with lenticular irregular aperture slightly oblique; vessel-ray pits and vessel-parenchyma pits bordered; libriform fibres, non-septate, extremely short and narrow, with simple minute pits; paratracheal axial parenchyma, vasicentric and scanty and apotracheal axial parenchyma diffuse; rays are nume. rous, exclusively heterocellular, composed of procumbent and square marginal cells, short and extremely narrow, simple pits of minute size; mostly uniseriate, less frequently biseriate and triseriate, rarely tetraseriate; aggregate rays are absent.</paragraph>
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<paragraph id="8BD536F0FE32FFF8229DFBD3AED4FDD3" blockId="3.[106,745,1116,2099]" lastBlockId="3.[820,1459,152,591]" pageId="3" pageNumber="254">D e s c r i p t i o n. Distinctive growth rings, alternately separated by light and dark bands (Pl. 1, Fig. 1), marked by radially flattened thick-walled latewood fibres versus thin-walled earlywood fibers. Absence of stratification in secondary xylem elements. Wood diffuse-porous (Pl. 1, Figs 1–2). Vessels in tangential bands (Pl. 1, Figs 1–2), numerous (16–36/mm²), occupying 25.6% of the cross section (Pl. 1, Figs 1–2). Vessels generally solitary, sometimes in radial multiples of 3 to 5, circular to oval in outline (Pl. 1, Figs 2–3). Mean tangential diameter 65.5 (32–96) µm, mean length 186.5 (48–360) µm. Helical thickenings in vessel elements are absent. Simple perforation plates (Pl. 2, Fig. 4), intervessel pits alternate, small 4.1 (2–8) µm, vestured, with lenticular irregular slightly oblique apertures (Pl. 1, Figs 4–5). Vessel-ray pits with distinct borders, rounded and bordered (Pl. 1, Figs 4–5). Vessel-parenchyma pits are similar to vessel-ray pits. Tyloses (Pl. 1, Figs 2–6) and prismatic crystals are present in the marginal square ray cells (Pl. 2, Figs 4–6). Fibres libriform, non-septate, with simple and minutely bordered pits 2.3 (1–4) µm (Pl. 2, Figs 1–4), occupying 45% of the wood. The fibres are extremely short, mean length 162 (30–360) µm, narrow 3.2 (1–8) µm and thin-walled 1.9 (1–4) µm, with a narrow diameter of 3.9 (2–8) µm. The paratracheal axial parenchyma is vasicentric and scanty, occupying only 5.45% of total wood volume (Pl. 1, Figs 2–3). The apotracheal axial parenchyma is diffuse, with dispersed cells between the fibers (Pl. 1, Fig. 2–3). Axial parenchyma cells are rectangular (Pl. 2, Fig. 2), mean length 36.9 (16–80) µm and mean width 18.5 (8–32) µm, constituting a series with mean length 364.8 (176–896) µm, composed of 13 (3–6) cells. Rays are numerous, occupying 24% of the volume of the wood (Pl. 1, Figs 1–2). They are mostly uniseriate (44%), less frequent biseriate (27%), triseriate (21%) and tetraseriate (8%) (Pl. 2, Figs 1–3). Aggregate rays are absent. Rays are heterocellular, composed of procumbent cells and 1–4 rows of square marginal cells, most commonly 2 rows (Pl. 2, Figs 4–5). Uniseriate rays extremely short, mean length 204.9 (112–424) µm, and extremely narrow, mean width 12.4 (8–24) µm, 7 to 27 cells high (Pl. 2, Figs 1–3). The multiseriate rays are narrow, mean width 26.8 (16–40) µm, 1 to 4 cells wide, and very short, mean length 241.5 (120–556) µm, 8 to 27 cells high (Pl. 2, Figs 1–3). In tangential section, the ray cells are circular to oval (Pl. 2, Figs 1–3) with minute simple pits, ranging from 1 to 4 µm.</paragraph>
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</subSubSection>
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</treatment>
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