184 lines
22 KiB
XML
184 lines
22 KiB
XML
<document id="AA3D95B5B5AA805D0F2ADB772B4E3236" ID-DOI="10.11646/palaeoentomology.6.3.13" ID-ISSN="2624-2834" ID-Zenodo-Dep="8209295" ID-ZooBank="12D99FD9-32AF-4F30-AF15-35CD80E7A586" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1691005736286" checkinUser="plazi" docAuthor="PANKOWSKI, MAXIMILIAN G. & Fanti, FABRIZIO FANTI" docDate="2023" docId="03C387DBD02CFFAD87B2FDADE20FFE41" docLanguage="en" docName="palaeoentomology.6.3.13.pdf" docOrigin="Palaeoentomology 6 (3)" docSource="http://dx.doi.org/10.11646/palaeoentomology.6.3.13" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Podistra madelineae PANKOWSKI & Fanti 2023, sp. nov." docType="treatment" docVersion="1" lastPageNumber="304" masterDocId="FFFAFFA3D02FFFA9873AFFE3E744FFF4" masterDocTitle="Six new species of fossil soldier beetles (Coleoptera: Cantharidae) from Eocene Baltic amber" masterLastPageNumber="312" masterPageNumber="300" pageNumber="303" updateTime="1691482849966" updateUser="tatiana">
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<mods:title id="CF7BB8E4A3BFCE3712B961991F6A7550">Six new species of fossil soldier beetles (Coleoptera: Cantharidae) from Eocene Baltic amber</mods:title>
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<mods:namePart id="28DFF6C01BD1B5BBA55AFB6D8A716CBE">PANKOWSKI, MAXIMILIAN G.</mods:namePart>
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<mods:namePart id="699CB51C10E738A11C7C7F3704E617B5">Fanti, FABRIZIO FANTI</mods:namePart>
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<mods:typeOfResource id="17D9F259901024A909EEB3546A511E21">text</mods:typeOfResource>
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<mods:title id="73F65647F5EE606771900D462E1D8A34">Palaeoentomology</mods:title>
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<mods:part id="DA7079CAAE553CBA12672729A173B261">
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<mods:date id="E07F0AA1F714D01E5F4F96A818873B02">2023</mods:date>
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<mods:number id="66DACBBB38A7FCD047C9452A70EB1FC3">2023-06-29</mods:number>
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<mods:url id="C2681BC458BC504282EAB53792AA5046">http://dx.doi.org/10.11646/palaeoentomology.6.3.13</mods:url>
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<mods:classification id="30CAA00090404480ECBBF99EE3288A25">journal article</mods:classification>
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<mods:identifier id="C1C47BD5697E2584645D76B045F3D070" type="DOI">10.11646/palaeoentomology.6.3.13</mods:identifier>
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<subSubSection id="C3706546D02CFFAA87B2FDADE690FD9C" box="[136,468,590,617]" pageId="3" pageNumber="303" type="nomenclature">
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<paragraph id="8BD536CDD02CFFAA87B2FDADE690FD9C" blockId="3.[136,468,590,652]" box="[136,468,590,617]" pageId="3" pageNumber="303">
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<heading id="D09D81A1D02CFFAA87B2FDADE690FD9C" bold="true" box="[136,468,590,617]" fontSize="10" level="2" pageId="3" pageNumber="303" reason="3">
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<emphasis id="B91EEADFD02CFFAA87B2FDADE690FD9C" bold="true" box="[136,468,590,617]" pageId="3" pageNumber="303">
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<taxonomicName id="4C6A4D4ED02CFFAA87B2FDADE630FD9D" authority="PANKOWSKI & Fanti, 2023" authorityName="PANKOWSKI & Fanti" authorityYear="2023" box="[136,372,590,617]" class="Insecta" family="Cantharidae" genus="Podistra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="303" phylum="Arthropoda" rank="species" species="madelineae" status="sp. nov.">
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<emphasis id="B91EEADFD02CFFAA87B2FDADE630FD9D" bold="true" box="[136,372,590,617]" italics="true" pageId="3" pageNumber="303">Podistra madelineae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22D57A4D02CFFAA8641FDADE690FD9C" box="[379,468,590,616]" pageId="3" pageNumber="303" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3706546D02CFFAA87B2FD91E799FD78" box="[136,221,626,652]" pageId="3" pageNumber="303" type="description">
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<paragraph id="8BD536CDD02CFFAA87B2FD91E799FD78" blockId="3.[136,468,590,652]" box="[136,221,626,652]" pageId="3" pageNumber="303">
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(
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<figureCitation id="13512A48D02CFFAA87AAFD91E790FD78" box="[144,212,626,652]" captionStart="FIGURE 2" captionStartId="3.[136,235,1998,2022]" captionTargetBox="[157,1436,1203,1974]" captionTargetId="figure-394@3.[151,1436,1203,1974]" captionTargetPageId="3" captionText="FIGURE 2. Podistra madelineae sp. nov. in Baltic amber. Holotype (USNM PAL 787824). A, B, Lateral views, scale bars = 0.5 mm. C, Dorsal view, scale bar = 0.5 mm. D, Detail of last abdominal segments, scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8209299" httpUri="https://zenodo.org/record/8209299/files/figure.png" pageId="3" pageNumber="303">Fig. 2</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3706546D02CFFAA87B2FD59E644FC90" pageId="3" pageNumber="303" type="materials_examined">
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<paragraph id="8BD536CDD02CFFAA87B2FD59E630FCE8" blockId="3.[136,778,698,1157]" pageId="3" pageNumber="303">
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<materialsCitation id="3B023C90D02CFFAA87B2FD59E630FCE8" collectionCode="USNM" county="Smithsonian National Museum of Natural History" location="Female" municipality="Baltic" pageId="3" pageNumber="303" specimenCode="PAL 787824" specimenCount="1" typeStatus="holotype">
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<emphasis id="B91EEADFD02CFFAA87B2FD59E7B8FD20" bold="true" box="[136,252,698,724]" pageId="3" pageNumber="303">
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<typeStatus id="54D1886FD02CFFAA87B2FD59E7B3FD20" box="[136,247,698,724]" pageId="3" pageNumber="303">Holotype</typeStatus>
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.
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</emphasis>
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<location id="8EB56016D02CFFAA863CFD59E61FFD20" LSID="urn:lsid:plazi:treatment:03C387DBD02CFFAD87B2FDADE20FFE41:8EB56016D02CFFAA863CFD59E61FFD20" box="[262,347,698,724]" county="Smithsonian National Museum of Natural History" municipality="Baltic" name="Female" pageId="3" pageNumber="303">Female</location>
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, inclusion in
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<collectingMunicipality id="6BB1ACB7D02CFFAA86C0FD59E579FD20" box="[506,573,698,724]" pageId="3" pageNumber="303">Baltic</collectingMunicipality>
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amber, housed at the
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<collectingCounty id="62B44E41D02CFFAA878DFD3DE442FD0C" box="[183,774,734,761]" pageId="3" pageNumber="303">Smithsonian National Museum of Natural History</collectingCounty>
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,
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<collectionCode id="ED7BAE08D02CFFAA87B2FCE1E79DFCE8" box="[136,217,770,796]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="303" type="Museum">USNM</collectionCode>
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<specimenCode id="DBCC9EB6D02CFFAA87DAFCE1E634FCE8" box="[224,368,770,796]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="303" type="Museum">PAL 787824</specimenCode>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BD536CDD02CFFAA8787FCC5E644FC90" blockId="3.[136,778,698,1157]" pageId="3" pageNumber="303">Syninclusions: Air bubbles, detritus and botanical fragments.</paragraph>
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</subSubSection>
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<subSubSection id="C3706546D02CFFAA8787FC8DE558FC24" pageId="3" pageNumber="303" type="etymology">
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<paragraph id="8BD536CDD02CFFAA8787FC8DE558FC24" blockId="3.[136,778,698,1157]" pageId="3" pageNumber="303">
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<emphasis id="B91EEADFD02CFFAA8787FC8DE600FC7C" bold="true" box="[189,324,878,904]" pageId="3" pageNumber="303">Etymology.</emphasis>
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Species named in honor of Madeline Pankowski, the first author’s younger sister and valued partner in palaeontological projects.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3706546D02CFFAA8787FC39E388FDA7" pageId="3" pageNumber="303" type="diagnosis">
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<paragraph id="8BD536CDD02CFFAA8787FC39E229FEF7" blockId="3.[136,778,698,1157]" lastBlockId="3.[810,1452,158,1156]" pageId="3" pageNumber="303">
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<emphasis id="B91EEADFD02CFFAA8787FC39E67CFC00" bold="true" box="[189,312,986,1012]" pageId="3" pageNumber="303">Diagnosis.</emphasis>
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The last maxillary palpomere securiform, the long elytra and the rectangular pronotum place this taxon in the genus
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<taxonomicName id="4C6A4D4ED02CFFAA8665FBC1E6FBFBC8" authorityName="Motschulsky" authorityYear="1839" box="[351,447,1058,1084]" class="Insecta" family="Cantharidae" genus="Podistra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="303" phylum="Arthropoda" rank="genus">
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<emphasis id="B91EEADFD02CFFAA8665FBC1E6FBFBC8" box="[351,447,1058,1084]" italics="true" pageId="3" pageNumber="303">Podistra</emphasis>
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</taxonomicName>
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. Based on the absence of the basal tooth of claws but with an obtuse enlargement, the species could be placed in the subgenus
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<taxonomicName id="4C6A4D4ED02CFFAA8564FB89E447FB70" box="[606,771,1130,1156]" class="Mucoromycetes" family="Cunninghamellaceae" genus="Pseudoabsidia" kingdom="Fungi" order="Mucorales" pageId="3" pageNumber="303" phylum="Zygomycota" rank="genus">
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<emphasis id="B91EEADFD02CFFAA8564FB89E447FB70" box="[606,771,1130,1156]" italics="true" pageId="3" pageNumber="303">Pseudoabsidia</emphasis>
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</taxonomicName>
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. However, this subgenus has recently been synonymized with the subgenus
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<taxonomicName id="4C6A4D4ED02CFFAA8332FF20E202FF2A" authority="(Kazantsev, 2023)" baseAuthorityName="Kazantsev" baseAuthorityYear="2023" box="[1032,1350,195,222]" class="Insecta" family="Cantharidae" genus="Podistra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="303" phylum="Arthropoda" rank="genus">
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<emphasis id="B91EEADFD02CFFAA8332FF20E32CFF29" box="[1032,1128,195,221]" italics="true" pageId="3" pageNumber="303">Podistra</emphasis>
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(
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<bibRefCitation id="EFFB4B3CD02CFFAA8341FF27E27AFF2A" author="Kazantsev, S. V." box="[1147,1342,196,222]" pageId="3" pageNumber="303" pagination="389 - 400" refId="ref7798" refString="Kazantsev, S. V. (2023) Do the Palaearctic subgenera of Podistra Motschulsky, 1839 really occur in North America: description of P. (Pidostria), subgen. nov. (Coleoptera: Cantharidae). Zootaxa, 5263 (3), 389 - 400. https: // doi. org / 10.11646 / zootaxa. 5263.3.4" type="journal article" year="2023">Kazantsev, 2023</bibRefCitation>
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)
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</taxonomicName>
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, and the taxonomy of the subgenera remains largely unclear.
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</paragraph>
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<paragraph id="8BD536CDD02CFFAA8465FEEDE436FE4A" blockId="3.[810,1452,158,1156]" pageId="3" pageNumber="303">
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The new species is most similar to
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<taxonomicName id="4C6A4D4ED02CFFAA83D1FEEDE4EAFEBA" authority="Fanti, 2021" authorityName="Fanti" authorityYear="2021" class="Insecta" family="Cantharidae" genus="Podistra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="303" phylum="Arthropoda" rank="species" species="guthriei">
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<emphasis id="B91EEADFD02CFFAA83D1FEEDE2E8FEDC" box="[1259,1452,270,296]" italics="true" pageId="3" pageNumber="303">Podistra guthriei</emphasis>
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<bibRefCitation id="EFFB4B3CD02CFFAA8410FED7E4EAFEBA" author="Fanti, F." box="[810,942,308,334]" pageId="3" pageNumber="303" pagination="1 - 9" refId="ref7176" refString="Fanti, F. (2021) Two new Malthodes and a new Podistra from Eocene Baltic amber (Coleoptera, Cantharidae). Baltic Journal of Coleopterology, 21 (1), 1 - 9." type="journal article" year="2021">Fanti, 2021</bibRefCitation>
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</taxonomicName>
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but is significantly larger than that species (
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<quantity id="4C929B28D02CFFAA8408FEBAE4C9FE80" box="[818,909,345,372]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="303" unit="mm" value="7.5">7.5 mm</quantity>
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<emphasis id="B91EEADFD02CFFAA84A2FEB9E4EBFE87" box="[920,943,346,371]" italics="true" pageId="3" pageNumber="303">vs</emphasis>
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.
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<quantity id="4C929B28D02CFFAA84F8FEBAE364FE87" box="[962,1056,345,372]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="3" pageNumber="303" unit="mm" value="5.7">5.7 mm</quantity>
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). Antennomeres III–V of the new species also are not subequal, unlike in
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<taxonomicName id="4C6A4D4ED02CFFAA83D9FE9CE42AFE4A" authority="(Fanti, 2021)" baseAuthorityName="Fanti" baseAuthorityYear="2021" class="Insecta" family="Cantharidae" genus="Podistra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="303" phylum="Arthropoda" rank="species" species="guthriei">
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||
<emphasis id="B91EEADFD02CFFAA83D9FE9CE3B0FE6C" box="[1251,1268,383,408]" italics="true" pageId="3" pageNumber="303">P</emphasis>
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.
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<emphasis id="B91EEADFD02CFFAA8238FE9DE21FFE6C" box="[1282,1371,382,408]" italics="true" pageId="3" pageNumber="303">guthriei</emphasis>
|
||
(
|
||
<bibRefCitation id="EFFB4B3CD02CFFAA8253FE9DE422FE4A" author="Fanti, F." pageId="3" pageNumber="303" pagination="1 - 9" refId="ref7176" refString="Fanti, F. (2021) Two new Malthodes and a new Podistra from Eocene Baltic amber (Coleoptera, Cantharidae). Baltic Journal of Coleopterology, 21 (1), 1 - 9." type="journal article" year="2021">Fanti, 2021</bibRefCitation>
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)
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</taxonomicName>
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||
.
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</paragraph>
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<paragraph id="8BD536CDD02CFFAA8465FE2AE388FDA7" blockId="3.[810,1452,158,1156]" pageId="3" pageNumber="303">
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<emphasis id="B91EEADFD02CFFAA8465FE2AE325FE17" bold="true" box="[863,1121,457,483]" pageId="3" pageNumber="303">Locality and horizon.</emphasis>
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Amber mine in the Yantarny settlement, Sambian Peninsula,
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<collectingRegion id="49AEF82FD02CFFAA83FAFE0DE2E3FDFC" box="[1216,1447,494,520]" country="Russia" name="Kaliningrad" pageId="3" pageNumber="303">Kaliningrad region</collectingRegion>
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,
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<collectingCountry id="F37D765DD02CFFAA8410FDF7E43CFDDA" box="[810,888,532,558]" name="Russia" pageId="3" pageNumber="303">Russia</collectingCountry>
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. Middle Eocene (Lutetian) (47.8–41.2 Ma) to late Eocene (Priabonian) (37.8–33.9 Ma).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3706546D02CFFAD8465FDBDE3CBFED0" lastPageId="4" lastPageNumber="304" pageId="3" pageNumber="303" type="description">
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||
<paragraph id="8BD536CDD02CFFAA8465FDBDE34AFD1D" blockId="3.[810,1452,158,1156]" pageId="3" pageNumber="303">
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<emphasis id="B91EEADFD02CFFAA8465FDBDE4B5FD8C" bold="true" box="[863,1009,606,632]" pageId="3" pageNumber="303">Description.</emphasis>
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Female, defined on the basis of the short antennae and the last sternite small and rounded. Body length: about
|
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<quantity id="4C929B28D02CFFAA833DFD4AE31BFD30" box="[1031,1119,681,708]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="303" unit="mm" value="7.5">7.5 mm</quantity>
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. Head black, other body parts entirely dark brown.
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</paragraph>
|
||
<paragraph id="8BD536CDD02CFFAD8465FD17E3CBFED0" blockId="3.[810,1452,158,1156]" lastBlockId="4.[810,1452,158,437]" lastPageId="4" lastPageNumber="304" pageId="3" pageNumber="303">Head not completely exposed, wide, rounded, equipped with scattered short setae and without punctation. Eyes relatively small, rounded, located in upper lateral part of the head. Mandibles falciform. Maxillary palpi 4-segmented, with the last palpomere securiform. Labial palpi 3-segmented with the last palpomere securiform. Antennae 11-segmented, filiform, short, surpassing the humeral region of the elytra, inserted close to the upper part of the eyes, all antennomeres pubescent with short and long setae; antennomere I elongate, club-shaped, enlarged apically; antennomere II short, slightly elongate, rather robust, about 2.4–2.5 times shorter than antennomere I; antennomere III about 1.4 times longer than antennomere II; antennomere IV robust, longer than previous one; antennomeres V–X subequal and shorter than previous ones; antennomere XI filiform, elongate, rounded apically. Pronotum longer than wide, rectangular, about as wide as head, surface slightly granulous and equipped with several setae and slightly bulged in the posterior part, anterior margin rounded and not bordered, posterior margin and sides straight and slightly bordered, propleura small and rounded. Scutellum very wide, triangular shaped, with apex slightly rounded, equipped with scattered setae. Elytra wider than pronotum, elongate, covering and slightly surpassing the last abdominal segments, surface slightly wrinkled and equipped with scattered and long setae, parallel-sided, rounded apically. Hind wings semitransparent, slightly longer than elytra. Legs short, relatively robust, densely pubescent; coxae robust, elongate, rounded apically; trochanters elongate with rounded apex; femora slightly enlarged, straight, cylindrical; tibiae thin, cylindrical, with apical spur that is short and robust, pro- and mesotibiae shorter than pro- and mesofemora, metatibiae slightly longer than metafemora. First tarsomere elongate and robust; second tarsomere shorter than first tarsomere; third tarsomere triangularshaped, with apical margin straight; tarsomere IV conspicuously widened, very shortly bilobed; tarsomere V thin, elongate, curved; claws simple without teeth but with an obtuse enlargement at base. Metasternum elongate, equipped with scattered long setae, rounded apically. Sternites transverse, wide, pubescent. Last sternite short, small, rounded. Male unknown.</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF156645D02CFFAA87B2F82DE339F7FE" ID-DOI="http://doi.org/10.5281/zenodo.8209299" ID-Zenodo-Dep="8209299" httpUri="https://zenodo.org/record/8209299/files/figure.png" pageId="3" pageNumber="303" startId="3.[136,235,1998,2022]" targetBox="[157,1436,1203,1974]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BD536CDD02CFFAA87B2F82DE339F7FE" blockId="3.[136,1452,1998,2059]" pageId="3" pageNumber="303">
|
||
<emphasis id="B91EEADFD02CFFAA87B2F82DE640F812" bold="true" box="[136,260,1998,2022]" pageId="3" pageNumber="303">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C6A4D4ED02CFFAA8631F82DE69AF812" authority="PANKOWSKI & Fanti, 2023" authorityName="PANKOWSKI & Fanti" authorityYear="2023" box="[267,478,1998,2022]" class="Insecta" family="Cantharidae" genus="Podistra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="303" phylum="Arthropoda" rank="species" species="madelineae" status="sp. nov.">
|
||
<emphasis id="B91EEADFD02CFFAA8631F82DE69AF812" box="[267,478,1998,2022]" italics="true" pageId="3" pageNumber="303">Podistra madelineae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91EEADFD02CFFAA86DFF82DE572F812" bold="true" box="[485,566,1998,2022]" pageId="3" pageNumber="303">
|
||
<taxonomicNameLabel id="A22D57A4D02CFFAA86DFF82DE572F812" box="[485,566,1998,2022]" pageId="3" pageNumber="303" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in Baltic amber. Holotype (USNM PAL 787824).
|
||
<emphasis id="B91EEADFD02CFFAA8302F82DE30EF812" bold="true" box="[1080,1098,1998,2022]" pageId="3" pageNumber="303">A</emphasis>
|
||
,
|
||
<emphasis id="B91EEADFD02CFFAA836DF82DE32CF812" bold="true" box="[1111,1128,1998,2022]" pageId="3" pageNumber="303">B</emphasis>
|
||
, Lateral views, scale bars = 0.5 mm.
|
||
<emphasis id="B91EEADFD02CFFAA8786F811E78AF7FE" bold="true" box="[188,206,2034,2058]" pageId="3" pageNumber="303">C</emphasis>
|
||
, Dorsal view, scale bar = 0.5 mm.
|
||
<emphasis id="B91EEADFD02CFFAA850AF811E506F7FE" bold="true" box="[560,578,2034,2058]" pageId="3" pageNumber="303">D</emphasis>
|
||
, Detail of last abdominal segments, scale bar = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3706546D02BFFAD8465FECCE20FFE41" pageId="4" pageNumber="304" type="discussion">
|
||
<paragraph id="8BD536CDD02BFFAD8465FECCE20FFE41" blockId="4.[810,1452,158,437]" pageId="4" pageNumber="304">
|
||
<emphasis id="B91EEADFD02BFFAD8465FECCE497FEBD" bold="true" box="[863,979,303,329]" pageId="4" pageNumber="304">Remarks.</emphasis>
|
||
The yellow, nearly square amber piece measures approximately 12 × 11 ×
|
||
<quantity id="4C929B28D02BFFAD8387FEB0E246FE99" box="[1213,1282,339,365]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="304" unit="mm" value="8.0">8 mm</quantity>
|
||
. The specimen is bent. One posterior leg is not preserved and the last sternite is partially covered by a white emulsion.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |