treatments-xml/data/03/C3/87/03C387C4FFAEFFACB4C1369F8596A878.xml

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<mods:title id="C0C35924DCFA13CBC428E527A95B36CF">New species of the Corumbataia cuestae group (Siluriformes: Loricariidae) from the Rio Tocantins basin, with comments on its phylogenetic relationships</mods:title>
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<mods:namePart id="8F1DB30B933DE5636B5AEE2CDD63C77C">Thimotheo, Mateus Giovanni Rocha</mods:namePart>
<mods:affiliation id="9959FE17374E160A50776EC7EDAEB821">Laboratório de Ictiologia, Setor de Zoologia, Campus de Botucatu, Universidade Estadual Paulista, IBB / UNESP, 18618 - 970 Botucatu, SP, Brazil.</mods:affiliation>
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<mods:namePart id="7C4820E2207317D9BEB4DFA081B5FD67">Benine, Ricardo Cardoso</mods:namePart>
<mods:affiliation id="E21CAA7854A5F752CE0B8D2DD3805D69">Laboratório de Ictiologia, Setor de Zoologia, Campus de Botucatu, Universidade Estadual Paulista, IBB / UNESP, 18618 - 970 Botucatu, SP, Brazil.</mods:affiliation>
<mods:nameIdentifier id="833972B3440231FFD979F15AE6A79279" type="email">ricardo.benine@unesp.br</mods:nameIdentifier>
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<mods:namePart id="2D65447D622E495EC933E98B560FA2A1">Oliveira, Claudio</mods:namePart>
<mods:affiliation id="DAEB46D10F13DE249F7D0CBD5FAB921F">Laboratório de Biologiae Genética de Peixes, Departamento de Biologia Estrutural e Funcional, Instituto de Biociências, Universidade Estadual Paulista, IBB / UNESP, R. Prof. Dr. Antonio C. W. Zanin, 250, 18618 - 689 Botucatu, SP, Brazil.</mods:affiliation>
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<mods:namePart id="A66CE495D1F7DA2E46E725D598D8FFE2">Silva, Gabriel de Souza da Costa e</mods:namePart>
<mods:affiliation id="D288B2847000CAB0558724A172A4C598">Laboratório de Ictiologia, Setor de Zoologia, Campus de Botucatu, Universidade Estadual Paulista, IBB / UNESP, 18618 - 970 Botucatu, SP, Brazil.</mods:affiliation>
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<mods:title id="28E5F1616FF375F8D0182C32F4D8C10B">Neotrop. Ichthyol.</mods:title>
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<taxonomicName id="4C6A4D51FFAEFFA1B4C1369F825EAA88" authority="Thimotheo &amp; Benine &amp; Oliveira &amp; Silva, 2020" authorityName="Thimotheo &amp; Benine &amp; Oliveira &amp; Silva" authorityYear="2020" box="[744,1058,190,214]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="acanthodela" status="sp. nov.">
<emphasis id="B91EEAC0FFAEFFA1B4C1369F825EAA88" bold="true" box="[744,1058,190,214]" pageId="6" pageNumber="7">Corumbataia acanthodela</emphasis>
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, 
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urn:lsid:zoobank.org:act: 
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(
<figureCitation id="13512A57FFAEFFA1B5B9377285A8AB35" box="[912,980,339,363]" captionStart="FIGURE 2" captionStartId="6.[488,561,1836,1856]" captionTargetBox="[667,1295,1095,1777]" captionTargetId="figure-28@6.[618,1329,1068,1814]" captionTargetPageId="6" captionText="FIGURE 2 | Corumbataia acanthodela, holotype, MZUSP 125794, male 27.9 mm SL, from Rio Maranhão, Rio Tocantins basin, Niquelândia, Goiás, Brazil." figureDoi="http://doi.org/10.5281/zenodo.10960537" httpUri="https://zenodo.org/record/10960537/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
, 
<tableCitation id="C6E80369FFAEFFA1B5F637728254AB35" box="[991,1064,339,363]" captionStart="TABLE 3" captionStartId="8.[136,198,1014,1034]" captionTargetId="graphics-150@8.[136,1462,1127,1678]" captionTargetPageId="8" captionText="TABLE 3 | Morphometrics of the holotype and 27 paratypes of Corumbataia acanthodela. SD = standard deviation." httpUri="http://table.plazi.org/id/DF15665AFFA0FFAFB6A135D782DBAE55" pageId="6" pageNumber="7" tableUuid="DF15665AFFA0FFAFB6A135D782DBAE55">Tab. 3</tableCitation>
)
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<taxonomicName id="4C6A4D51FFAEFFA1B42437BF8591ABE8" authority="Bristki, 1997: 237" authorityName="Bristki" authorityPageNumber="237" authorityYear="1997" box="[525,1005,414,438]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="tocantinensis">
<emphasis id="B91EEAC0FFAEFFA1B42437BF855EABE8" box="[525,802,414,438]" italics="true" pageId="6" pageNumber="7">Corumbataia tocantinensis</emphasis>
Bristki, 1997:237
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(original description; in part, 
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, 
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, 
<specimenCode id="DBCC9EA9FFAEFFA1B56737E285E9AB85" box="[846,917,451,475]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="6" pageNumber="7">50158</specimenCode>
; UFRJ 3000, 3001, 2999; 
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, 
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). — 
<bibRefCitation id="EFFB4B23FFAEFFA1B4FE37C985D1A85E" author="Carvalho" box="[727,941,488,512]" firstAuthor="Carvalho" page="554" pageId="6" pageNumber="7" pagination="552 - 57" refId="ref7175" refString="• Carvalho TP. A new species of Corumbataia (Siluriformes: Loricariidae: Hypoptopomatinae) from upper Rio Tocantins Basin, Central Brazil. Copeia. 2008; 3: 552 - 57. https: // doi. org / 10.1643 / CI- 07 - 064" type="journal article" year="2008">
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, 2008:554
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(diagnosis, fig. 2A, MCN 13462).
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.
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<specimenCode id="DBCC9EA9FFAEFFA1B4A434128535A815" box="[653,841,563,587]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="6" pageNumber="7">MZUSP 125794</specimenCode>
, 
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SL, 
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, 
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, 
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, 
<location id="8EB56009FFAEFFA1B7C1347984F0A82E" LSID="urn:lsid:plazi:treatment:03C387C4FFAEFFACB4C1369F8596A878:8EB56009FFAEFFA1B7C1347984F0A82E" box="[488,652,600,624]" county="Niquelandia" latitude="-14.510917" longLatPrecision="1" longitude="-48.68611" municipality="Rio do Peixe" name="Rio Maranhao" pageId="6" pageNumber="7" stateProvince="Goias">Rio Maranhão</location>
, 
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basin, 
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, 
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<collectingDate id="EF90E9FAFFAEFFA1B2FE34788313A82F" box="[1239,1391,601,625]" pageId="6" pageNumber="7" value="2012-11-23">23 Nov 2012</collectingDate>
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, 
<collectorName id="269F5304FFAEFFA1B3553478845AA8C8" pageId="6" pageNumber="7">B. F. Melo</collectorName>
, 
<collectorName id="269F5304FFAEFFA1B418345F84A8A8C8" box="[561,724,638,662]" pageId="6" pageNumber="7">J. H. Martinez</collectorName>
, 
<collectorName id="269F5304FFAEFFA1B4C9345F850AA8C8" box="[736,886,638,662]" pageId="6" pageNumber="7">G. S. C. Silva</collectorName>
&amp; 
<collectorName id="269F5304FFAEFFA1B5BE345F826CA8C8" box="[919,1040,638,662]" pageId="6" pageNumber="7">R. Devidé.</collectorName>
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<materialsCitation id="3B023C8FFFAEFFA1B42434828521A8BE" collectingDate="2014-08-05" collectingDateMax="2017-11-25" collectingDateMin="2014-08-05" collectionCode="R" collectorName="B. F. Melo &amp; C. Oliveira &amp; G. S. C. Silva &amp; M. Taylor. &amp; Devide, B. F &amp; Melo, C &amp; Araya, G. S. C. &amp; Niquelandia &amp; Rio &amp; M. Taylor" country="Brazil" county="All" latitude="-14.638027" location="Goias" longLatPrecision="1" longitude="-48.76642" municipality="Niquelandia" pageId="6" pageNumber="7" specimenCode="LBP 17153" specimenCount="10" stateProvince="Goias" typeStatus="paratype">
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<typeStatus id="54D18870FFAEFFA1B424348284FEA8E5" box="[525,642,675,699]" pageId="6" pageNumber="7" type="paratype">Paratypes</typeStatus>
.
</emphasis>
All from 
<collectingCountry id="F37D7642FFAEFFA1B4C53482854CA8E5" box="[748,816,675,699]" name="Brazil" pageId="6" pageNumber="7">Brazil</collectingCountry>
, 
<collectingRegion id="49AEF830FFAEFFA1B51034828500A8E5" box="[825,892,675,699]" country="Brazil" name="Goias" pageId="6" pageNumber="7">Goiás</collectingRegion>
: 
<specimenCode id="DBCC9EA9FFAEFFA1B5AD3485827DA8E5" box="[900,1025,675,699]" pageId="6" pageNumber="7">LBP 17153</specimenCode>
, 
<specimenCount id="9D6CFD5BFFAEFFA1B2233482825BA8E5" box="[1034,1063,675,699]" pageId="6" pageNumber="7">10</specimenCount>
, 28.9– 
<quantity id="4C929B37FFAEFFA1B258348282ADA8E5" box="[1137,1233,675,699]" metricMagnitude="-2" metricUnit="m" metricValue="1.68" pageId="6" pageNumber="7" unit="mm" value="16.8">16.8 mm</quantity>
SL, 2 c&amp;s, 
<quantity id="4C929B37FFAEFFA1B36C34828469A8BE" metricMagnitude="-2" metricUnit="m" metricValue="2.625" metricValueMax="2.6399999999999997" metricValueMin="2.61" pageId="6" pageNumber="7" unit="mm" value="26.25" valueMax="26.4" valueMin="26.1">26.1–26.4 mm</quantity>
SL, collected with holotype.
</materialsCitation>
<materialsCitation id="3B023C8FFFAEFFA1B54D34E88447A90E" collectingDate="2014-08-05" collectingDateMax="2017-11-25" collectingDateMin="2014-08-05" collectionCode="R" collectorName="B. F. Melo &amp; C. Oliveira &amp; G. S. C. Silva &amp; M. Taylor." country="Brazil" county="All" latitude="-14.638027" location="Rio Maranhao" longLatPrecision="1" longitude="-48.76642" municipality="Niquelandia" pageId="6" pageNumber="7" specimenCode="LBP 19095" specimenCount="10" stateProvince="Goias" typeStatus="paratype">
<specimenCode id="DBCC9EA9FFAEFFA1B54D34E8859AA8BE" box="[868,998,712,736]" pageId="6" pageNumber="7">LBP 19095</specimenCode>
, 
<specimenCount id="9D6CFD5BFFAEFFA1B5DA34E8826CA8BF" box="[1011,1040,713,737]" pageId="6" pageNumber="7">10</specimenCount>
, 27.3– 
<quantity id="4C929B37FFAEFFA1B27734E982BDA8BE" box="[1118,1217,712,736]" metricMagnitude="-2" metricUnit="m" metricValue="1.61" pageId="6" pageNumber="7" unit="mm" value="16.1">16.1 mm</quantity>
SL, 2 c&amp;s, 29.4– 
<quantity id="4C929B37FFAEFFA1B3AA34E98469A95B" metricMagnitude="-2" metricUnit="m" metricValue="2.65" pageId="6" pageNumber="7" unit="mm" value="26.5">26.5 mm</quantity>
SL (tissues 77006–10), 
<collectingRegion id="49AEF830FFAEFFA1B50F34CF8514A958" box="[806,872,750,774]" country="Brazil" name="Goias" pageId="6" pageNumber="7">Goiás</collectingRegion>
, 
<collectingMunicipality id="6BB1ACA8FFAEFFA1B55F34CF827FA958" box="[886,1027,750,774]" pageId="6" pageNumber="7">Niquelândia</collectingMunicipality>
, 
<location id="8EB56009FFAEFFA1B23834CF82CBA958" LSID="urn:lsid:plazi:treatment:03C387C4FFAEFFACB4C1369F8596A878:8EB56009FFAEFFA1B23834CF82CBA958" box="[1041,1207,750,774]" country="Brazil" county="All" latitude="-14.638027" longLatPrecision="1" longitude="-48.76642" municipality="Niquelandia" name="Rio Maranhao" pageId="6" pageNumber="7" stateProvince="Goias">Rio Maranhão</location>
, 
<location id="8EB56009FFAEFFA1B2EC34CF8315A958" LSID="urn:lsid:plazi:treatment:03C387C4FFAEFFACB4C1369F8596A878:8EB56009FFAEFFA1B2EC34CF8315A958" box="[1221,1385,750,774]" country="Brazil" county="All" latitude="-14.638027" longLatPrecision="1" longitude="-48.76642" municipality="Niquelandia" name="Rio Tocantins" pageId="6" pageNumber="7" stateProvince="Goias">Rio Tocantins</location>
basin, 
<geoCoordinate id="EE5E5015FFAEFFA1B7C135328406A975" box="[488,634,787,811]" degrees="14" direction="south" minutes="38" orientation="latitude" pageId="6" pageNumber="7" precision="1" seconds="16.9" value="-14.638027">14°38’16.9’’S</geoCoordinate>
<geoCoordinate id="EE5E5015FFAEFFA1B4AB35328550A975" box="[642,812,787,811]" degrees="48" direction="west" minutes="45" orientation="longitude" pageId="6" pageNumber="7" precision="1" seconds="59.1" value="-48.76642">48°45’59.1” W</geoCoordinate>
, 
<date id="FFD41012FFAEFFA1B510353285C2A975" box="[825,958,787,811]" pageId="6" pageNumber="7" value="2014-08-05">
<collectingDate id="EF90E9FAFFAEFFA1B510353285C2A975" box="[825,958,787,811]" pageId="6" pageNumber="7" value="2014-08-05">5 Aug 2014</collectingDate>
</date>
, 
<collectorName id="269F5304FFAEFFA1B5E335358238A975" box="[970,1092,787,811]" pageId="6" pageNumber="7">B. F. Melo</collectorName>
, 
<collectorName id="269F5304FFAEFFA1B279353282ACA975" box="[1104,1232,787,811]" pageId="6" pageNumber="7">C. Oliveira</collectorName>
, 
<collectorName id="269F5304FFAEFFA1B2F53532830DA975" box="[1244,1393,787,811]" pageId="6" pageNumber="7">G. S. C. Silva</collectorName>
&amp; 
<collectorName id="269F5304FFAEFFA1B3BA35358447A90E" pageId="6" pageNumber="7">M. Taylor.</collectorName>
</materialsCitation>
<materialsCitation id="3B023C8FFFAEFFA1B4683518857FA9C5" collectingDate="2017-11-25" collectingDateMax="2017-11-25" collectingDateMin="2014-08-05" collectionCode="R" collectorName="R. Devide &amp; B. F. Melo &amp; C. Araya &amp; G. S. C. Silva" country="Brazil" county="All" latitude="-14.638027" location="Rio Vaivem" longLatPrecision="1" longitude="-48.76642" municipality="Mara Rosa" pageId="6" pageNumber="7" specimenCode="LBP 25634" specimenCount="1" stateProvince="Goias" typeStatus="paratype">
<specimenCode id="DBCC9EA9FFAEFFA1B468351884BCA90E" box="[577,704,824,848]" pageId="6" pageNumber="7">LBP 25634</specimenCode>
, 
<specimenCount id="9D6CFD5BFFAEFFA1B4E2351884A5A90F" box="[715,729,825,849]" pageId="6" pageNumber="7">1</specimenCount>
, 
<quantity id="4C929B37FFAEFFA1B4CF35188535A90E" box="[742,841,825,849]" metricMagnitude="-2" metricUnit="m" metricValue="2.84" pageId="6" pageNumber="7" unit="mm" value="28.4">28.4 mm</quantity>
SL, 
<collectingRegion id="49AEF830FFAEFFA1B553351985C0A90E" box="[890,956,824,848]" country="Brazil" name="Goias" pageId="6" pageNumber="7">Goiás</collectingRegion>
, 
<collectingMunicipality id="6BB1ACA8FFAEFFA1B5EE35188243A90E" box="[967,1087,825,848]" pageId="6" pageNumber="7">Mara Rosa</collectingMunicipality>
, 
<location id="8EB56009FFAEFFA1B263351982AFA90E" LSID="urn:lsid:plazi:treatment:03C387C4FFAEFFACB4C1369F8596A878:8EB56009FFAEFFA1B263351982AFA90E" box="[1098,1235,824,848]" country="Brazil" county="All" latitude="-14.638027" longLatPrecision="1" longitude="-48.76642" municipality="Mara Rosa" name="Rio Vaivem" pageId="6" pageNumber="7" stateProvince="Goias">Rio Vaivém</location>
, 
<location id="8EB56009FFAEFFA1B2F4351983FCA90E" LSID="urn:lsid:plazi:treatment:03C387C4FFAEFFACB4C1369F8596A878:8EB56009FFAEFFA1B2F4351983FCA90E" box="[1245,1408,824,848]" country="Brazil" county="All" latitude="-14.638027" longLatPrecision="1" longitude="-48.76642" municipality="Mara Rosa" name="Rio Maranhao" pageId="6" pageNumber="7" stateProvince="Goias">Rio Maranhão</location>
, 
<location id="8EB56009FFAEFFA1B3A335198425A928" LSID="urn:lsid:plazi:treatment:03C387C4FFAEFFACB4C1369F8596A878:8EB56009FFAEFFA1B3A335198425A928" country="Brazil" county="All" latitude="-14.638027" longLatPrecision="1" longitude="-48.76642" municipality="Mara Rosa" name="Rio Tocantins" pageId="6" pageNumber="7" stateProvince="Goias">Rio Tocantins</location>
basin, 
<geoCoordinate id="EE5E5015FFAEFFA1B483357F8539A928" box="[682,837,862,886]" degrees="14" direction="south" minutes="03" orientation="latitude" pageId="6" pageNumber="7" precision="1" seconds="45.10" value="-14.062528">14º03’45.10”S</geoCoordinate>
<geoCoordinate id="EE5E5015FFAEFFA1B567357F8580A928" box="[846,1020,862,886]" degrees="49" direction="west" minutes="05" orientation="longitude" pageId="6" pageNumber="7" precision="1" seconds="34.25" value="-49.092846">49º05’34.25”W</geoCoordinate>
, 
<date id="FFD41012FFAEFFA1B223357F82DEA928" box="[1034,1186,862,886]" pageId="6" pageNumber="7" value="2017-11-25">
<collectingDate id="EF90E9FAFFAEFFA1B223357F82DEA928" box="[1034,1186,862,886]" pageId="6" pageNumber="7" value="2017-11-25">25 Nov 2017</collectingDate>
</date>
, 
<collectorName id="269F5304FFAEFFA1B286357F8359A928" box="[1199,1317,862,886]" pageId="6" pageNumber="7">R. Devidé</collectorName>
, 
<collectorName id="269F5304FFAEFFA1B318357F83D3A928" box="[1329,1455,862,886]" pageId="6" pageNumber="7">B. F. Melo</collectorName>
, 
<collectorName id="269F5304FFAEFFA1B7C135A2842DA9C5" box="[488,593,899,923]" pageId="6" pageNumber="7">C. Araya</collectorName>
, 
<collectorName id="269F5304FFAEFFA1B47635A28482A9C5" box="[607,766,899,923]" pageId="6" pageNumber="7">G. S. C. Silva</collectorName>
.
</materialsCitation>
<materialsCitation id="3B023C8FFFAEFFA1B52235A585DDA9B8" collectingDate="2014-08-05" collectingDateMax="2017-11-25" collectingDateMin="2014-08-05" collectionCode="R" collectorName="B. F. Melo &amp; C. Oliveira &amp; G. S. C. Silva &amp; M. Taylor" country="Brazil" county="All" latitude="-14.638027" location="Rio Maranhao" longLatPrecision="1" longitude="-48.76642" municipality="Niquelandia" pageId="6" pageNumber="7" specimenCode="NUP 22694" specimenCount="3" stateProvince="Goias" typeStatus="paratype">
<specimenCode id="DBCC9EA9FFAEFFA1B52235A585E4A9C5" box="[779,920,899,923]" pageId="6" pageNumber="7">NUP 22694</specimenCode>
, 
<specimenCount id="9D6CFD5BFFAEFFA1B58F35A285C9A9C5" box="[934,949,899,923]" pageId="6" pageNumber="7">3</specimenCount>
, 
<quantity id="4C929B37FFAEFFA1B5ED35A28215A9C5" box="[964,1129,899,923]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.4899999999999998" metricValueMin="1.71" pageId="6" pageNumber="7" unit="mm" value="21.0" valueMax="24.9" valueMin="17.1">17.1–24.9 mm</quantity>
SL, 
<collectingRegion id="49AEF830FFAEFFA1B28935A2829EA9C5" box="[1184,1250,899,923]" country="Brazil" name="Goias" pageId="6" pageNumber="7">Goiás</collectingRegion>
, 
<collectingMunicipality id="6BB1ACA8FFAEFFA1B2D935A28301A9C5" box="[1264,1405,899,923]" pageId="6" pageNumber="7">Niquelândia</collectingMunicipality>
, 
<location id="8EB56009FFAEFFA1B3A235A28420A99E" LSID="urn:lsid:plazi:treatment:03C387C4FFAEFFACB4C1369F8596A878:8EB56009FFAEFFA1B3A235A28420A99E" country="Brazil" county="All" latitude="-14.638027" longLatPrecision="1" longitude="-48.76642" municipality="Niquelandia" name="Rio Maranhao" pageId="6" pageNumber="7" stateProvince="Goias">Rio Maranhão</location>
, 
<location id="8EB56009FFAEFFA1B44E35898574A99E" LSID="urn:lsid:plazi:treatment:03C387C4FFAEFFACB4C1369F8596A878:8EB56009FFAEFFA1B44E35898574A99E" box="[615,776,936,960]" country="Brazil" county="All" latitude="-14.638027" longLatPrecision="1" longitude="-48.76642" municipality="Niquelandia" name="Rio Tocantins" pageId="6" pageNumber="7" stateProvince="Goias">Rio Tocantins</location>
basin, 
<geoCoordinate id="EE5E5015FFAEFFA1B57E35898596A99E" box="[855,1002,936,960]" degrees="14" direction="south" minutes="38" orientation="latitude" pageId="6" pageNumber="7" precision="1" seconds="16.9" value="-14.638027">14°38’16.9’’S</geoCoordinate>
<geoCoordinate id="EE5E5015FFAEFFA1B5DB358982EAA99E" box="[1010,1174,936,960]" degrees="48" direction="west" minutes="45" orientation="longitude" pageId="6" pageNumber="7" precision="1" seconds="59.1" value="-48.76642">48°45’59.1”W</geoCoordinate>
, 
<date id="FFD41012FFAEFFA1B28B35898354A99F" box="[1186,1320,936,961]" pageId="6" pageNumber="7" value="2014-08-05">
<collectingDate id="EF90E9FAFFAEFFA1B28B35898354A99F" box="[1186,1320,936,961]" pageId="6" pageNumber="7" value="2014-08-05">5 Aug 2014</collectingDate>
</date>
, 
<collectorName id="269F5304FFAEFFA1B31D358883D3A99E" box="[1332,1455,936,960]" pageId="6" pageNumber="7">B. F. Melo</collectorName>
, 
<collectorName id="269F5304FFAEFFA1B7C135EF8414A9B8" box="[488,616,974,998]" pageId="6" pageNumber="7">C. Oliveira</collectorName>
, 
<collectorName id="269F5304FFAEFFA1B45D35EF8575A9B8" box="[628,777,974,998]" pageId="6" pageNumber="7">G. S. C. Silva</collectorName>
&amp; 
<collectorName id="269F5304FFAEFFA1B50235EF85DDA9B8" box="[811,929,974,998]" pageId="6" pageNumber="7">M. Taylor</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF15665AFFAEFFA1B7C1310D85D5AD3B" ID-DOI="http://doi.org/10.5281/zenodo.10960537" ID-Zenodo-Dep="10960537" httpUri="https://zenodo.org/record/10960537/files/figure.png" pageId="6" pageNumber="7" startId="6.[488,561,1836,1856]" targetBox="[667,1295,1095,1777]" targetPageId="6" targetType="figure">
<paragraph id="8BD536D2FFAEFFA1B7C1310D85D5AD3B" blockId="6.[488,1436,1836,1893]" pageId="6" pageNumber="7">
<emphasis id="B91EEAC0FFAEFFA1B7C1310D8547AD1E" bold="true" box="[488,827,1836,1856]" pageId="6" pageNumber="7">
FIGURE 2 | 
<taxonomicName id="4C6A4D51FFAEFFA1B470310D8547AD1E" authorityName="Thimotheo &amp; Benine &amp; Oliveira &amp; Silva" authorityYear="2020" box="[601,827,1836,1856]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="acanthodela">
<emphasis id="B91EEAC0FFAEFFA1B470310D8547AD1E" bold="true" box="[601,827,1836,1856]" italics="true" pageId="6" pageNumber="7">Corumbataia acanthodela</emphasis>
</taxonomicName>
</emphasis>
, holotype, MZUSP 125794, male 27.9 mm SL, from Rio Maranhão, Rio Tocantins basin, Niquelândia, Goiás, Brazil.
</paragraph>
</caption>
<subSubSection id="C3706559FFAFFFA0B424369F8326A99E" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="8BD536D2FFAFFFA0B424369F8326A99E" blockId="7.[486,1459,190,961]" pageId="7" pageNumber="8">
<emphasis id="B91EEAC0FFAFFFA0B424369F84F2AA88" bold="true" box="[525,654,190,214]" pageId="7" pageNumber="8">Diagnosis.</emphasis>
The new species differs from all congeners, except for the members of 
<taxonomicName id="4C6A4D51FFAFFFA0B7C036C284C2AAA5" authorityName=", LBP" authorityYear="1309" box="[489,702,227,251]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cuestae">
<emphasis id="B91EEAC0FFAFFFA0B7C036C284C2AAA5" box="[489,702,227,251]" italics="true" pageId="7" pageNumber="8">Corumbataia cuestae</emphasis>
</taxonomicName>
group by having a conspicuous pair of tufts with enlarged odontodes on the tip of the supraoccipital (
<emphasis id="B91EEAC0FFAFFFA0B56137288521AB7E" box="[840,861,265,288]" italics="true" pageId="7" pageNumber="8">vs</emphasis>
. absence of crest of enlarged odontodes in 
<taxonomicName id="4C6A4D51FFAFFFA0B31F372883D0AB7E" baseAuthorityName="Carvalho, Lehmann &amp; Reis" baseAuthorityYear="2008" box="[1334,1452,265,288]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="anosteos">
<emphasis id="B91EEAC0FFAFFFA0B31F372883D0AB7E" box="[1334,1452,265,288]" italics="true" pageId="7" pageNumber="8">C. anosteos</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C6A4D51FFAFFFA0B7C1370F8427AB18" baseAuthorityName="Roxo, Silva, Ochoa &amp; Zawadzki" baseAuthorityYear="2017" box="[488,603,302,326]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="canoeiro">
<emphasis id="B91EEAC0FFAFFFA0B7C1370F8427AB18" box="[488,603,302,326]" italics="true" pageId="7" pageNumber="8">C. canoeiro</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C6A4D51FFAFFFA0B4B1370F8569AB18" authorityName="Carvalho" authorityYear="2008" box="[664,789,302,326]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="veadeiros">
<emphasis id="B91EEAC0FFAFFFA0B4B1370F8569AB18" box="[664,789,302,326]" italics="true" pageId="7" pageNumber="8">C. veadeiros</emphasis>
</taxonomicName>
); by the absence of an adipose fin or single series of platelets at adipose fin position (
<emphasis id="B91EEAC0FFAFFFA0B4DA37758574AB35" box="[755,776,340,363]" italics="true" pageId="7" pageNumber="8">vs</emphasis>
. presence of an adipose fin in 
<taxonomicName id="4C6A4D51FFAFFFA0B24E377582A1AB35" baseAuthorityName="Roxo, Silva, Ochoa &amp; Zawadzki" baseAuthorityYear="2017" box="[1127,1245,340,363]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="canoeiro">
<emphasis id="B91EEAC0FFAFFFA0B24E377582A1AB35" box="[1127,1245,340,363]" italics="true" pageId="7" pageNumber="8">C. canoeiro</emphasis>
</taxonomicName>
, or single series of platelets in 
<taxonomicName id="4C6A4D51FFAFFFA0B44737588495ABCE" baseAuthorityName="Carvalho, Lehmann &amp; Reis" baseAuthorityYear="2008" box="[622,745,377,400]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="anosteos">
<emphasis id="B91EEAC0FFAFFFA0B44737588495ABCE" box="[622,745,377,400]" italics="true" pageId="7" pageNumber="8">C. anosteos</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6A4D51FFAFFFA0B501375885D5ABCE" authorityName="Carvalho" authorityYear="2008" box="[808,937,376,400]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="veadeiros">
<emphasis id="B91EEAC0FFAFFFA0B501375885D5ABCE" box="[808,937,376,400]" italics="true" pageId="7" pageNumber="8">C. veadeiros</emphasis>
</taxonomicName>
); by anastomosis of the infraorbital and otic sensory canals over the pterotic-supracleithrum (
<figureCitation id="13512A57FFAFFFA0B22A37BF822BABE8" box="[1027,1111,414,438]" captionStart="FIGURE 3" captionStartId="7.[136,209,1836,1856]" captionTargetBox="[161,1436,1261,1757]" captionTargetId="figure-409@7.[135,1459,1216,1781]" captionTargetPageId="7" captionText="FIGURE 3 | Infraorbital series of Corumbataia acanthodela, LBP 19095, paratype (A), and C. anosteos, LBP 17125 (B). io1-io5 (Infraorbitals); sp (sphenotic); cpt (pterotic-supracleithrum). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10960539" httpUri="https://zenodo.org/record/10960539/files/figure.png" pageId="7" pageNumber="8">Fig. 3A</figureCitation>
) (
<emphasis id="B91EEAC0FFAFFFA0B24737BE82FFABE8" box="[1134,1155,415,438]" italics="true" pageId="7" pageNumber="8">vs</emphasis>
. anastomosis of infraorbital and otic sensory canals over the sphenotic in 
<taxonomicName id="4C6A4D51FFAFFFA0B5C637E5821BAB85" baseAuthorityName="Carvalho, Lehmann &amp; Reis" baseAuthorityYear="2008" box="[1007,1127,452,475]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="anosteos">
<emphasis id="B91EEAC0FFAFFFA0B5C637E5821BAB85" box="[1007,1127,452,475]" italics="true" pageId="7" pageNumber="8">C. anosteos</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C6A4D51FFAFFFA0B25C37E58296AB85" baseAuthorityName="Roxo, Silva, Ochoa &amp; Zawadzki" baseAuthorityYear="2017" box="[1141,1258,452,475]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="canoeiro">
<emphasis id="B91EEAC0FFAFFFA0B25C37E58296AB85" box="[1141,1258,452,475]" italics="true" pageId="7" pageNumber="8">C. canoeiro</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C6A4D51FFAFFFA0B30437E583D0AB85" authorityName="Carvalho" authorityYear="2008" box="[1325,1452,451,475]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="veadeiros">
<emphasis id="B91EEAC0FFAFFFA0B30437E583D0AB85" box="[1325,1452,451,475]" italics="true" pageId="7" pageNumber="8">C. veadeiros</emphasis>
</taxonomicName>
) (
<figureCitation id="13512A57FFAFFFA0B7D937C9843BA85F" box="[496,583,488,513]" captionStart="FIGURE 3" captionStartId="7.[136,209,1836,1856]" captionTargetBox="[161,1436,1261,1757]" captionTargetId="figure-409@7.[135,1459,1216,1781]" captionTargetPageId="7" captionText="FIGURE 3 | Infraorbital series of Corumbataia acanthodela, LBP 19095, paratype (A), and C. anosteos, LBP 17125 (B). io1-io5 (Infraorbitals); sp (sphenotic); cpt (pterotic-supracleithrum). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10960539" httpUri="https://zenodo.org/record/10960539/files/figure.png" pageId="7" pageNumber="8">Fig. 3B</figureCitation>
). Additionally, the new species differs from all species of 
<taxonomicName id="4C6A4D51FFAFFFA0B2F337C983CFA85E" authorityName=", LBP" authorityYear="1309" box="[1242,1459,488,512]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cuestae">
<emphasis id="B91EEAC0FFAFFFA0B2F337C983CFA85E" box="[1242,1459,488,512]" italics="true" pageId="7" pageNumber="8">Corumbataia cuestae</emphasis>
</taxonomicName>
group by having a small naked area on snout tip (
<emphasis id="B91EEAC0FFAFFFA0B21B342E823BA878" box="[1074,1095,527,550]" italics="true" pageId="7" pageNumber="8">vs</emphasis>
. large naked area on snout tip, 
<figureCitation id="13512A57FFAFFFA0B7C134128451A815" box="[488,557,563,587]" captionStart="FIGURE 4" captionStartId="8.[487,560,865,885]" captionTargetBox="[510,1453,449,800]" captionTargetId="figure-129@8.[488,1459,432,819]" captionTargetPageId="8" captionText="FIGURE 4 | Frontal view of snout tip showing a naked area without odontodes. A. Corumbataia acanthodela, LBP 19095, paratype; B. Corumbataia cuestae, LBP 1309. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10960541" httpUri="https://zenodo.org/record/10960541/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
). Moreover, 
<taxonomicName id="4C6A4D51FFAFFFA0B4EB341285B1A815" authorityName="Thimotheo &amp; Benine &amp; Oliveira &amp; Silva" authorityYear="2020" box="[706,973,563,587]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="acanthodela">
<emphasis id="B91EEAC0FFAFFFA0B4EB341285B1A815" box="[706,973,563,587]" italics="true" pageId="7" pageNumber="8">Corumbataia acanthodela</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C6A4D51FFAFFFA0B249341582B6A815" authorityName=", LBP" authorityYear="1309" box="[1120,1226,564,587]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cuestae">
<emphasis id="B91EEAC0FFAFFFA0B249341582B6A815" box="[1120,1226,564,587]" italics="true" pageId="7" pageNumber="8">C. cuestae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C6A4D51FFAFFFA0B2F034158357A815" authorityName="Silva, Roxo, Souza &amp; Oliveira" authorityYear="2018" box="[1241,1323,563,587]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="liliai">
<emphasis id="B91EEAC0FFAFFFA0B2F034158357A815" box="[1241,1323,563,587]" italics="true" pageId="7" pageNumber="8">C. liliai</emphasis>
</taxonomicName>
, 
<emphasis id="B91EEAC0FFAFFFA0B312341583CFA815" box="[1339,1459,563,587]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="4C6A4D51FFAFFFA0B312341583D3A815" authorityName="Silva, Roxo, Souza &amp; Oliveira" authorityYear="2018" box="[1339,1455,563,587]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="lucianoi">C. lucianoi</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C6A4D51FFAFFFA0B431347884FCA82E" box="[536,640,600,624]" family="Loricariidae" genus="Rhinolekos" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="britskii">
<emphasis id="B91EEAC0FFAFFFA0B431347884FCA82E" box="[536,640,600,624]" italics="true" pageId="7" pageNumber="8">C. britskii</emphasis>
</taxonomicName>
by having abdominal platelets reaching the mid-ventral lateral plates (
<emphasis id="B91EEAC0FFAFFFA0B3BE347883D0A82E" box="[1431,1452,601,624]" italics="true" pageId="7" pageNumber="8">vs</emphasis>
. abdominal platelets far from reaching mid-ventral lateral plates); from 
<taxonomicName id="4C6A4D51FFAFFFA0B2C0345F832CA8C8" authorityName=", LBP" authorityYear="1309" box="[1257,1360,638,662]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cuestae">
<emphasis id="B91EEAC0FFAFFFA0B2C0345F832CA8C8" box="[1257,1360,638,662]" italics="true" pageId="7" pageNumber="8">C. cuestae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C6A4D51FFAFFFA0B374345F83D0A8C8" authorityName="Silva, Roxo, Souza &amp; Oliveira" authorityYear="2018" box="[1373,1452,638,662]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="liliai">
<emphasis id="B91EEAC0FFAFFFA0B374345F83D0A8C8" box="[1373,1452,638,662]" italics="true" pageId="7" pageNumber="8">C. liliai</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C6A4D51FFAFFFA0B43F348584CBA8E5" authorityName="Bristki" authorityYear="1997" box="[534,695,676,699]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="tocantinensis">
<emphasis id="B91EEAC0FFAFFFA0B43F348584CBA8E5" box="[534,695,676,699]" italics="true" pageId="7" pageNumber="8">C. tocantinensis</emphasis>
</taxonomicName>
by having two rounded and more anteriorly positioned hyaline areas on the caudal-fin (
<figureCitation id="13512A57FFAFFFA0B49D34E9848AA8BF" box="[692,758,712,737]" captionStart="FIGURE 2" captionStartId="6.[488,561,1836,1856]" captionTargetBox="[667,1295,1095,1777]" captionTargetId="figure-28@6.[618,1329,1068,1814]" captionTargetPageId="6" captionText="FIGURE 2 | Corumbataia acanthodela, holotype, MZUSP 125794, male 27.9 mm SL, from Rio Maranhão, Rio Tocantins basin, Niquelândia, Goiás, Brazil." figureDoi="http://doi.org/10.5281/zenodo.10960537" httpUri="https://zenodo.org/record/10960537/files/figure.png" pageId="7" pageNumber="8">Fig. 2</figureCitation>
) (
<emphasis id="B91EEAC0FFAFFFA0B52734E8855FA8BE" box="[782,803,713,736]" italics="true" pageId="7" pageNumber="8">vs</emphasis>
. absence of two smaller rounded hyaline areas, see fig. 
<quantity id="4C929B37FFAFFFA0B3A134E883CFA8BE" box="[1416,1459,712,737]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="7" pageNumber="8" unit="in" value="3.0">3 in</quantity>
<bibRefCitation id="EFFB4B23FFAFFFA0B7C134CF84E5A958" author="Silva" box="[488,665,750,774]" etAl="et al." firstAuthor="Silva" pageId="7" pageNumber="8" pagination="317 - 30" refId="ref8357" refString="• Silva GSC, Roxo FF, Souza CS, Oliveira C. Two new species of Corumbataia (Hypoptopomatinae: Loricariidae) from Rio Corrente, upper Rio Parana basin, Brazil. Zootaxa. 2018; 4483 (2): 317 - 30. https: // doi. org / 10.11646 / zootaxa. 4483.2.5" type="journal article" year="2018">
Silva 
<emphasis id="B91EEAC0FFAFFFA0B40B34CE842FA958" box="[546,595,750,774]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
, 2018
</bibRefCitation>
). It differs from 
<taxonomicName id="4C6A4D51FFAFFFA0B56134CF8595A958" authorityName="Bristki" authorityYear="1997" box="[840,1001,750,774]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="tocantinensis">
<emphasis id="B91EEAC0FFAFFFA0B56134CF8595A958" box="[840,1001,750,774]" italics="true" pageId="7" pageNumber="8">C. tocantinensis</emphasis>
</taxonomicName>
by having the anterior profile of the head rounded in dorsal view (
<emphasis id="B91EEAC0FFAFFFA0B4D43535856EA975" box="[765,786,788,811]" italics="true" pageId="7" pageNumber="8">vs</emphasis>
. elliptical) and from 
<taxonomicName id="4C6A4D51FFAFFFA0B5D63535820EA975" authorityName="Silva, Roxo, Souza &amp; Oliveira" authorityYear="2018" box="[1023,1138,787,811]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="lucianoi">
<emphasis id="B91EEAC0FFAFFFA0B5D63535820EA975" box="[1023,1138,787,811]" italics="true" pageId="7" pageNumber="8">C. lucianoi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6A4D51FFAFFFA0B28235358287A975" authorityName="Silva, Roxo, Souza &amp; Oliveira" authorityYear="2018" box="[1195,1275,787,811]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="liliai">
<emphasis id="B91EEAC0FFAFFFA0B28235358287A975" box="[1195,1275,787,811]" italics="true" pageId="7" pageNumber="8">C. liliai</emphasis>
</taxonomicName>
by the presence of plates on dorsal portion of snout (
<emphasis id="B91EEAC0FFAFFFA0B553351885F3A90E" box="[890,911,825,848]" italics="true" pageId="7" pageNumber="8">vs</emphasis>
. presence of a broad naked area without plates or odontodes on dorsal portion of snout). It differs from 
<taxonomicName id="4C6A4D51FFAFFFA0B216357F82DBA928" box="[1087,1191,862,886]" family="Loricariidae" genus="Rhinolekos" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="britskii">
<emphasis id="B91EEAC0FFAFFFA0B216357F82DBA928" box="[1087,1191,862,886]" italics="true" pageId="7" pageNumber="8">C. britskii</emphasis>
</taxonomicName>
, 
<emphasis id="B91EEAC0FFAFFFA0B29C357F8350A928" box="[1205,1324,862,886]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="4C6A4D51FFAFFFA0B29C357F8354A928" authorityName="Silva, Roxo, Souza &amp; Oliveira" authorityYear="2018" box="[1205,1320,862,886]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="lucianoi">C. lucianoi</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C6A4D51FFAFFFA0B34A357F83CFA928" authorityName="Silva, Roxo, Souza &amp; Oliveira" authorityYear="2018" box="[1379,1459,862,886]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="liliai">
<emphasis id="B91EEAC0FFAFFFA0B34A357F83CFA928" box="[1379,1459,862,886]" italics="true" pageId="7" pageNumber="8">C. liliai</emphasis>
</taxonomicName>
by the presence of 28–29 vertebrae (
<emphasis id="B91EEAC0FFAFFFA0B5AB35A585EBA9C5" box="[898,919,900,923]" italics="true" pageId="7" pageNumber="8">vs</emphasis>
. 27) and from 
<taxonomicName id="4C6A4D51FFAFFFA0B26935A582D5A9C5" box="[1088,1193,899,923]" family="Loricariidae" genus="Rhinolekos" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="britskii">
<emphasis id="B91EEAC0FFAFFFA0B26935A582D5A9C5" box="[1088,1193,899,923]" italics="true" pageId="7" pageNumber="8">C. britskii</emphasis>
</taxonomicName>
by having lower lip far from reaching pectoral girdle line (
<emphasis id="B91EEAC0FFAFFFA0B558358885FAA99E" box="[881,902,937,960]" italics="true" pageId="7" pageNumber="8">vs</emphasis>
. reaching pectoral girdle transversal line).
</paragraph>
</subSubSection>
<subSubSection id="C3706559FFAFFFADB42435D285DFAAA5" lastPageId="10" lastPageNumber="11" pageId="7" pageNumber="8" type="description">
<paragraph id="8BD536D2FFAFFFAFB42435D283D6AB35" blockId="7.[486,1459,1011,1147]" lastBlockId="8.[485,1461,190,364]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<emphasis id="B91EEAC0FFAFFFA0B42435D284DEAE55" bold="true" box="[525,674,1011,1035]" pageId="7" pageNumber="8">Description.</emphasis>
Morphometric and meristic data are summarized in 
<tableCitation id="C6E80369FFAFFFA0B2C835D28356AE55" box="[1249,1322,1011,1035]" captionStart="TABLE 3" captionStartId="8.[136,198,1014,1034]" captionTargetId="graphics-150@8.[136,1462,1127,1678]" captionTargetPageId="8" captionText="TABLE 3 | Morphometrics of the holotype and 27 paratypes of Corumbataia acanthodela. SD = standard deviation." httpUri="http://table.plazi.org/id/DF15665AFFA0FFAFB6A135D782DBAE55" pageId="7" pageNumber="8" tableUuid="DF15665AFFA0FFAFB6A135D782DBAE55">Tab. 3</tableCitation>
. Small-sized loricariid (maximum 
<quantity id="4C929B37FFAFFFA0B4FB32388549AE6E" box="[722,821,1049,1073]" metricMagnitude="-2" metricUnit="m" metricValue="2.73" pageId="7" pageNumber="8" unit="mm" value="27.3">27.3 mm</quantity>
SL). Snout rounded in dorsal view. Dorsal profile of head ascending convexly approximately 45° to parieto-supraoccipital. Eyes relatively small (15.3–21.8% of HL), dorsolaterally positioned, just posterior to midpoint of head. Iris operculum present and poorly developed. Mouth moderate in size; oral disk ellipsoid with papillae randomly distributed. Lower lip larger than upper lip, not reaching cleithrum; its border fringed; lower lip inner surface covered with small papillae, similar in size. Maxillary barbel adnate to lower lip. Teeth slender with two cusps; central cusp larger than lateral cusp. Premaxillary teeth 29–42 (mode 37). Dentary teeth 22–37 (mode 32).
</paragraph>
<caption id="DF15665AFFAFFFA0B6A1310D84BFAD3B" ID-DOI="http://doi.org/10.5281/zenodo.10960539" ID-Zenodo-Dep="10960539" httpUri="https://zenodo.org/record/10960539/files/figure.png" pageId="7" pageNumber="8" startId="7.[136,209,1836,1856]" targetBox="[161,1436,1261,1757]" targetPageId="7" targetType="figure">
<paragraph id="8BD536D2FFAFFFA0B6A1310D84BFAD3B" blockId="7.[136,1425,1836,1893]" pageId="7" pageNumber="8">
<emphasis id="B91EEAC0FFAFFFA0B6A1310D868FAD1E" bold="true" box="[136,243,1836,1856]" pageId="7" pageNumber="8">FIGURE 3 |</emphasis>
Infraorbital series of 
<taxonomicName id="4C6A4D51FFAFFFA0B7EB310D84AFAD1E" authority=", LBP" authorityName="LBP" box="[450,723,1836,1856]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="acanthodela">
<emphasis id="B91EEAC0FFAFFFA0B7EB310D84DFAD1E" bold="true" box="[450,675,1836,1856]" italics="true" pageId="7" pageNumber="8">Corumbataia acanthodela</emphasis>
, LBP
</taxonomicName>
19095, paratype (
<emphasis id="B91EEAC0FFAFFFA0B55D310D85F8AD61" bold="true" box="[884,900,1836,1855]" pageId="7" pageNumber="8">A</emphasis>
), and 
<taxonomicName id="4C6A4D51FFAFFFA0B597310D8233AD1E" authority=", LBP" authorityName="LBP" box="[958,1103,1836,1856]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="anosteos">
<emphasis id="B91EEAC0FFAFFFA0B597310D8263AD1E" bold="true" box="[958,1055,1836,1856]" italics="true" pageId="7" pageNumber="8">C. anosteos</emphasis>
, LBP
</taxonomicName>
17125 (
<emphasis id="B91EEAC0FFAFFFA0B2A4310D82E7AD61" bold="true" box="[1165,1179,1836,1855]" pageId="7" pageNumber="8">B</emphasis>
). io1-io5 (Infraorbitals); sp (sphenotic); cpt (pterotic-supracleithrum). Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="DF15665AFFA0FFAFB7CE3540837FA9C5" ID-DOI="http://doi.org/10.5281/zenodo.10960541" ID-Zenodo-Dep="10960541" httpUri="https://zenodo.org/record/10960541/files/figure.png" pageId="8" pageNumber="9" startId="8.[487,560,865,885]" targetBox="[510,1453,449,800]" targetPageId="8" targetType="figure">
<paragraph id="8BD536D2FFA0FFAFB7CE3540837FA9C5" blockId="8.[487,1378,865,923]" pageId="8" pageNumber="9">
<emphasis id="B91EEAC0FFA0FFAFB7CE3540842FA92B" bold="true" box="[487,595,865,885]" pageId="8" pageNumber="9">FIGURE 4 |</emphasis>
Frontal view of snout tip showing a naked area without odontodes. 
<emphasis id="B91EEAC0FFA0FFAFB2FD35408298A92A" bold="true" box="[1236,1252,865,884]" pageId="8" pageNumber="9">A</emphasis>
. 
<taxonomicName id="4C6A4D51FFA0FFAFB2C6354084FCA9C5" authority=", LBP" authorityName="LBP" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="acanthodela">
<emphasis id="B91EEAC0FFA0FFAFB2C63540842CA9C5" bold="true" italics="true" pageId="8" pageNumber="9">Corumbataia acanthodela</emphasis>
, LBP
</taxonomicName>
19095, paratype; 
<emphasis id="B91EEAC0FFA0FFAFB50835A68553A9C4" bold="true" box="[801,815,903,922]" pageId="8" pageNumber="9">B</emphasis>
. 
<taxonomicName id="4C6A4D51FFA0FFAFB51335A6822CA9C5" authority=", LBP 1309" authorityName=", LBP" authorityYear="1309" box="[826,1104,903,923]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="cuestae">
<emphasis id="B91EEAC0FFA0FFAFB51335A6858CA9C5" bold="true" box="[826,1008,903,923]" italics="true" pageId="8" pageNumber="9">Corumbataia cuestae</emphasis>
, LBP 1309
</taxonomicName>
. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="DF15665AFFA0FFAFB6A135D782DBAE55" ID-Table-UUID="DF15665AFFA0FFAFB6A135D782DBAE55" box="[136,1191,1014,1035]" httpUri="http://table.plazi.org/id/DF15665AFFA0FFAFB6A135D782DBAE55" pageId="8" pageNumber="9" startId="8.[136,198,1014,1034]" targetBox="[364,1386,1076,1884]" targetIsTable="true" targetPageId="8" targetType="table">
<paragraph id="8BD536D2FFA0FFAFB6A135D782DBAE55" blockId="8.[136,1191,1014,1035]" box="[136,1191,1014,1035]" pageId="8" pageNumber="9">
<emphasis id="B91EEAC0FFA0FFAFB6A135D78694AE54" bold="true" box="[136,232,1014,1034]" pageId="8" pageNumber="9">TABLE 3 |</emphasis>
Morphometrics of the holotype and 27 paratypes of 
<taxonomicName id="4C6A4D51FFA0FFAFB4FD35D685C9AE55" box="[724,949,1015,1035]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="acanthodela">
<emphasis id="B91EEAC0FFA0FFAFB4FD35D685C9AE55" bold="true" box="[724,949,1015,1035]" italics="true" pageId="8" pageNumber="9">Corumbataia acanthodela</emphasis>
</taxonomicName>
. SD = standard deviation.
</paragraph>
</caption>
<paragraph id="8BD536D2FFA0FFAFB45132158316AD02" pageId="8" pageNumber="9">
<table id="F96AC472FFA00058B74532158316AD02" box="[364,1386,1076,1884]" gridcols="5" gridrows="27" pageId="8" pageNumber="9">
<tr id="355A3490FFA00058B74532158316AE18" box="[364,1386,1076,1094]" gridrow="0" pageId="8" pageNumber="9" rowspan-0="1">
<th id="768B5DECFFA00058B45132158518AE18" box="[632,868,1076,1094]" gridcol="1" gridrow="0" pageId="8" pageNumber="9">
<emphasis id="B91EEAC0FFA0FFAFB45132158518AE18" bold="true" box="[632,868,1076,1094]" pageId="8" pageNumber="9">Holotype Min</emphasis>
</th>
<th id="768B5DECFFA00058B5C232158268AE18" box="[1003,1044,1076,1094]" gridcol="2" gridrow="0" pageId="8" pageNumber="9">
<emphasis id="B91EEAC0FFA0FFAFB5C232158268AE1A" bold="true" box="[1003,1044,1076,1092]" pageId="8" pageNumber="9">Max</emphasis>
</th>
<th id="768B5DECFFA00058B2BC321582B5AE18" box="[1173,1225,1076,1094]" gridcol="3" gridrow="0" pageId="8" pageNumber="9">
<emphasis id="B91EEAC0FFA0FFAFB2BC321582B5AE1A" bold="true" box="[1173,1225,1076,1092]" pageId="8" pageNumber="9">Mean</emphasis>
</th>
<th id="768B5DECFFA00058B37932158316AE18" box="[1360,1386,1076,1094]" gridcol="4" gridrow="0" pageId="8" pageNumber="9">
<emphasis id="B91EEAC0FFA0FFAFB37832158316AE18" bold="true" box="[1361,1386,1076,1094]" pageId="8" pageNumber="9">SD</emphasis>
</th>
</tr>
<tr id="355A3490FFA00058B74532738316AE3A" box="[364,1386,1106,1124]" gridrow="1" pageId="8" pageNumber="9">
<th id="768B5DECFFA00058B74532738438AE3A" box="[364,580,1106,1124]" gridcol="0" gridrow="1" pageId="8" pageNumber="9">Standard length</th>
<td id="768B5DECFFA00058B45132738518AE3A" box="[632,868,1106,1124]" gridcol="1" gridrow="1" pageId="8" pageNumber="9">27.9 16.6</td>
<td id="768B5DECFFA00058B5C232738268AE3A" box="[1003,1044,1106,1124]" gridcol="2" gridrow="1" pageId="8" pageNumber="9">30.8</td>
<td id="768B5DECFFA00058B2BC327382B5AE3A" box="[1173,1225,1106,1124]" gridcol="3" gridrow="1" pageId="8" pageNumber="9">23.0</td>
<td id="768B5DECFFA00058B37932738316AE3A" box="[1360,1386,1106,1124]" gridcol="4" gridrow="1" pageId="8" pageNumber="9">4.3</td>
</tr>
<tr id="355A3490FFA00058B745324E8316AEDF" box="[364,1386,1135,1153]" gridrow="2" pageId="8" pageNumber="9">
<th id="768B5DECFFA00058B745324E8316AEDF" box="[364,1386,1135,1153]" colspan="5" colspanRight="4" gridcol="0" gridrow="2" pageId="8" pageNumber="9">
<emphasis id="B91EEAC0FFA0FFAFB4A5325185CDAEDE" bold="true" box="[652,945,1135,1153]" pageId="8" pageNumber="9">Percentages of standard length</emphasis>
</th>
</tr>
<tr id="355A3490FFA00058B74532AC8316AEC1" box="[364,1386,1165,1183]" gridrow="3" pageId="8" pageNumber="9">
<th id="768B5DECFFA00058B74532AC8438AEC1" box="[364,580,1165,1183]" gridcol="0" gridrow="3" pageId="8" pageNumber="9">Predorsal length</th>
<td id="768B5DECFFA00058B45132AC8518AEC1" box="[632,868,1165,1183]" gridcol="1" gridrow="3" pageId="8" pageNumber="9">44.2 40.4</td>
<td id="768B5DECFFA00058B5C232AC8268AEC1" box="[1003,1044,1165,1183]" gridcol="2" gridrow="3" pageId="8" pageNumber="9">50.3</td>
<td id="768B5DECFFA00058B2BC32AC82B5AEC1" box="[1173,1225,1165,1183]" gridcol="3" gridrow="3" pageId="8" pageNumber="9">44.2</td>
<td id="768B5DECFFA00058B37932AC8316AEC1" box="[1360,1386,1165,1183]" gridcol="4" gridrow="3" pageId="8" pageNumber="9">2.0</td>
</tr>
<tr id="355A3490FFA00058B745328A8316AEE3" box="[364,1386,1195,1213]" gridrow="4" pageId="8" pageNumber="9">
<th id="768B5DECFFA00058B745328A8438AEE3" box="[364,580,1195,1213]" gridcol="0" gridrow="4" pageId="8" pageNumber="9">Preanal length</th>
<td id="768B5DECFFA00058B451328A8518AEE3" box="[632,868,1195,1213]" gridcol="1" gridrow="4" pageId="8" pageNumber="9">61.5 55.9</td>
<td id="768B5DECFFA00058B5C2328A8268AEE3" box="[1003,1044,1195,1213]" gridcol="2" gridrow="4" pageId="8" pageNumber="9">63.2</td>
<td id="768B5DECFFA00058B2BC328A82B5AEE3" box="[1173,1225,1195,1213]" gridcol="3" gridrow="4" pageId="8" pageNumber="9">59.7</td>
<td id="768B5DECFFA00058B379328A8316AEE3" box="[1360,1386,1195,1213]" gridcol="4" gridrow="4" pageId="8" pageNumber="9">1.9</td>
</tr>
<tr id="355A3490FFA00058B74532E88316AE85" box="[364,1386,1225,1243]" gridrow="5" pageId="8" pageNumber="9">
<th id="768B5DECFFA00058B74532E88438AE85" box="[364,580,1225,1243]" gridcol="0" gridrow="5" pageId="8" pageNumber="9">Head length</th>
<td id="768B5DECFFA00058B45132E88518AE85" box="[632,868,1225,1243]" gridcol="1" gridrow="5" pageId="8" pageNumber="9">31.7 28.8</td>
<td id="768B5DECFFA00058B5C232E88268AE85" box="[1003,1044,1225,1243]" gridcol="2" gridrow="5" pageId="8" pageNumber="9">35.1</td>
<td id="768B5DECFFA00058B2BC32E882B5AE85" box="[1173,1225,1225,1243]" gridcol="3" gridrow="5" pageId="8" pageNumber="9">31.7</td>
<td id="768B5DECFFA00058B37932E88316AE85" box="[1360,1386,1225,1243]" gridcol="4" gridrow="5" pageId="8" pageNumber="9">1.6</td>
</tr>
<tr id="355A3490FFA00058B74532C68316AEA7" box="[364,1386,1255,1273]" gridrow="6" pageId="8" pageNumber="9">
<th id="768B5DECFFA00058B74532C68438AEA7" box="[364,580,1255,1273]" gridcol="0" gridrow="6" pageId="8" pageNumber="9">Cleithral width</th>
<td id="768B5DECFFA00058B45132C68518AEA7" box="[632,868,1255,1273]" gridcol="1" gridrow="6" pageId="8" pageNumber="9">27.5 22.7</td>
<td id="768B5DECFFA00058B5C232C68268AEA7" box="[1003,1044,1255,1273]" gridcol="2" gridrow="6" pageId="8" pageNumber="9">30.0</td>
<td id="768B5DECFFA00058B2BC32C682B5AEA7" box="[1173,1225,1255,1273]" gridcol="3" gridrow="6" pageId="8" pageNumber="9">27.6</td>
<td id="768B5DECFFA00058B37932C68316AEA7" box="[1360,1386,1255,1273]" gridcol="4" gridrow="6" pageId="8" pageNumber="9">1.6</td>
</tr>
<tr id="355A3490FFA00058B74533248316AF49" box="[364,1386,1285,1303]" gridrow="7" pageId="8" pageNumber="9">
<th id="768B5DECFFA00058B74533248438AF49" box="[364,580,1285,1303]" gridcol="0" gridrow="7" pageId="8" pageNumber="9">Dorsal-fin spine length</th>
<td id="768B5DECFFA00058B45133248518AF49" box="[632,868,1285,1303]" gridcol="1" gridrow="7" pageId="8" pageNumber="9">24.9 18.6</td>
<td id="768B5DECFFA00058B5C233248268AF49" box="[1003,1044,1285,1303]" gridcol="2" gridrow="7" pageId="8" pageNumber="9">25.9</td>
<td id="768B5DECFFA00058B2BC332482B5AF49" box="[1173,1225,1285,1303]" gridcol="3" gridrow="7" pageId="8" pageNumber="9">22.6</td>
<td id="768B5DECFFA00058B37933248316AF49" box="[1360,1386,1285,1303]" gridcol="4" gridrow="7" pageId="8" pageNumber="9">1.9</td>
</tr>
<tr id="355A3490FFA00058B74533088316AF65" box="[364,1386,1321,1339]" gridrow="8" pageId="8" pageNumber="9">
<th id="768B5DECFFA00058B74533088438AF65" box="[364,580,1321,1339]" gridcol="0" gridrow="8" pageId="8" pageNumber="9">Base of dorsal-fin length</th>
<td id="768B5DECFFA00058B45133088518AF65" box="[632,868,1321,1339]" gridcol="1" gridrow="8" pageId="8" pageNumber="9">14.2 11.9</td>
<td id="768B5DECFFA00058B5C233088268AF65" box="[1003,1044,1321,1339]" gridcol="2" gridrow="8" pageId="8" pageNumber="9">15.4</td>
<td id="768B5DECFFA00058B2BC330882B5AF65" box="[1173,1225,1321,1339]" gridcol="3" gridrow="8" pageId="8" pageNumber="9">13.7</td>
<td id="768B5DECFFA00058B37933088316AF65" box="[1360,1386,1321,1339]" gridcol="4" gridrow="8" pageId="8" pageNumber="9">0.8</td>
</tr>
<tr id="355A3490FFA00058B745336C8316AF01" box="[364,1386,1357,1375]" gridrow="9" pageId="8" pageNumber="9">
<th id="768B5DECFFA00058B745336C8438AF01" box="[364,580,1357,1375]" gridcol="0" gridrow="9" pageId="8" pageNumber="9">Thorax length</th>
<td id="768B5DECFFA00058B451336C8518AF01" box="[632,868,1357,1375]" gridcol="1" gridrow="9" pageId="8" pageNumber="9">19.5 12.5</td>
<td id="768B5DECFFA00058B5C2336C8268AF01" box="[1003,1044,1357,1375]" gridcol="2" gridrow="9" pageId="8" pageNumber="9">18.4</td>
<td id="768B5DECFFA00058B2BC336C82B5AF01" box="[1173,1225,1357,1375]" gridcol="3" gridrow="9" pageId="8" pageNumber="9">16.5</td>
<td id="768B5DECFFA00058B379336C8316AF01" box="[1360,1386,1357,1375]" gridcol="4" gridrow="9" pageId="8" pageNumber="9">1.5</td>
</tr>
<tr id="355A3490FFA00058B745334A8316AF23" box="[364,1386,1387,1405]" gridrow="10" pageId="8" pageNumber="9">
<th id="768B5DECFFA00058B745334A8438AF23" box="[364,580,1387,1405]" gridcol="0" gridrow="10" pageId="8" pageNumber="9">Pectoral-fin length</th>
<td id="768B5DECFFA00058B451334A8518AF23" box="[632,868,1387,1405]" gridcol="1" gridrow="10" pageId="8" pageNumber="9">25.7 18.1</td>
<td id="768B5DECFFA00058B5C2334A8268AF23" box="[1003,1044,1387,1405]" gridcol="2" gridrow="10" pageId="8" pageNumber="9">25.7</td>
<td id="768B5DECFFA00058B2BC334A82B5AF23" box="[1173,1225,1387,1405]" gridcol="3" gridrow="10" pageId="8" pageNumber="9">20.5</td>
<td id="768B5DECFFA00058B379334A8316AF23" box="[1360,1386,1387,1405]" gridcol="4" gridrow="10" pageId="8" pageNumber="9">1.6</td>
</tr>
<tr id="355A3490FFA00058B74533A88316AFC5" box="[364,1386,1417,1435]" gridrow="11" pageId="8" pageNumber="9">
<th id="768B5DECFFA00058B74533A88438AFC5" box="[364,580,1417,1435]" gridcol="0" gridrow="11" pageId="8" pageNumber="9">Abdomen length</th>
<td id="768B5DECFFA00058B45133A88518AFC5" box="[632,868,1417,1435]" gridcol="1" gridrow="11" pageId="8" pageNumber="9">26.1 20.4</td>
<td id="768B5DECFFA00058B5C233A88268AFC5" box="[1003,1044,1417,1435]" gridcol="2" gridrow="11" pageId="8" pageNumber="9">26.8</td>
<td id="768B5DECFFA00058B2BC33A882B5AFC5" box="[1173,1225,1417,1435]" gridcol="3" gridrow="11" pageId="8" pageNumber="9">23.5</td>
<td id="768B5DECFFA00058B37933A88316AFC5" box="[1360,1386,1417,1435]" gridcol="4" gridrow="11" pageId="8" pageNumber="9">1.5</td>
</tr>
<tr id="355A3490FFA00058B74533868316AFE7" box="[364,1386,1447,1465]" gridrow="12" pageId="8" pageNumber="9">
<th id="768B5DECFFA00058B74533868438AFE7" box="[364,580,1447,1465]" gridcol="0" gridrow="12" pageId="8" pageNumber="9">Pelvic-fin spine length</th>
<td id="768B5DECFFA00058B45133868518AFE7" box="[632,868,1447,1465]" gridcol="1" gridrow="12" pageId="8" pageNumber="9">19.9 17.3</td>
<td id="768B5DECFFA00058B5C233868268AFE7" box="[1003,1044,1447,1465]" gridcol="2" gridrow="12" pageId="8" pageNumber="9">23.4</td>
<td id="768B5DECFFA00058B2BC338682B5AFE7" box="[1173,1225,1447,1465]" gridcol="3" gridrow="12" pageId="8" pageNumber="9">21.0</td>
<td id="768B5DECFFA00058B37933868316AFE7" box="[1360,1386,1447,1465]" gridcol="4" gridrow="12" pageId="8" pageNumber="9">1.6</td>
</tr>
<tr id="355A3490FFA00058B74533E58316AF88" box="[364,1386,1476,1494]" gridrow="13" pageId="8" pageNumber="9">
<th id="768B5DECFFA00058B74533E58438AF88" box="[364,580,1476,1494]" gridcol="0" gridrow="13" pageId="8" pageNumber="9">Anal-fin spine length</th>
<td id="768B5DECFFA00058B45133E58518AF88" box="[632,868,1476,1494]" gridcol="1" gridrow="13" pageId="8" pageNumber="9">19.2 16.1</td>
<td id="768B5DECFFA00058B5C233E58268AF88" box="[1003,1044,1476,1494]" gridcol="2" gridrow="13" pageId="8" pageNumber="9">21.1</td>
<td id="768B5DECFFA00058B2BC33E582B5AF88" box="[1173,1225,1476,1494]" gridcol="3" gridrow="13" pageId="8" pageNumber="9">18.8</td>
<td id="768B5DECFFA00058B37933E58316AF88" box="[1360,1386,1476,1494]" gridcol="4" gridrow="13" pageId="8" pageNumber="9">1.3</td>
</tr>
<tr id="355A3490FFA00058B74533C38316AFAA" box="[364,1386,1506,1524]" gridrow="14" pageId="8" pageNumber="9">
<th id="768B5DECFFA00058B74533C38438AFAA" box="[364,580,1506,1524]" gridcol="0" gridrow="14" pageId="8" pageNumber="9">Lower cd spine</th>
<td id="768B5DECFFA00058B45133C38518AFAA" box="[632,868,1506,1524]" gridcol="1" gridrow="14" pageId="8" pageNumber="9">24.7 22.8</td>
<td id="768B5DECFFA00058B5C233C38268AFAA" box="[1003,1044,1506,1524]" gridcol="2" gridrow="14" pageId="8" pageNumber="9">34.8</td>
<td id="768B5DECFFA00058B2BC33C382B5AFAA" box="[1173,1225,1506,1524]" gridcol="3" gridrow="14" pageId="8" pageNumber="9">28.7</td>
<td id="768B5DECFFA00058B37933C38316AFAA" box="[1360,1386,1506,1524]" gridcol="4" gridrow="14" pageId="8" pageNumber="9">3.7</td>
</tr>
<tr id="355A3490FFA00058B74530218316AC4C" box="[364,1386,1536,1554]" gridrow="15" pageId="8" pageNumber="9">
<th id="768B5DECFFA00058B74530218438AC4C" box="[364,580,1536,1554]" gridcol="0" gridrow="15" pageId="8" pageNumber="9">Caudal peduncle depth</th>
<td id="768B5DECFFA00058B45130218518AC4C" box="[632,868,1536,1554]" gridcol="1" gridrow="15" pageId="8" pageNumber="9">11.1 10.2</td>
<td id="768B5DECFFA00058B5C230218268AC4C" box="[1003,1044,1536,1554]" gridcol="2" gridrow="15" pageId="8" pageNumber="9">12.8</td>
<td id="768B5DECFFA00058B2BC302182B5AC4C" box="[1173,1225,1536,1554]" gridcol="3" gridrow="15" pageId="8" pageNumber="9">11.3</td>
<td id="768B5DECFFA00058B37930218316AC4C" box="[1360,1386,1536,1554]" gridcol="4" gridrow="15" pageId="8" pageNumber="9">0.6</td>
</tr>
<tr id="355A3490FFA00058B745303F8316AC6E" box="[364,1386,1566,1584]" gridrow="16" pageId="8" pageNumber="9">
<th id="768B5DECFFA00058B745303F8438AC6E" box="[364,580,1566,1584]" gridcol="0" gridrow="16" pageId="8" pageNumber="9">Caudal peduncle length</th>
<td id="768B5DECFFA00058B451303F8518AC6E" box="[632,868,1566,1584]" gridcol="1" gridrow="16" pageId="8" pageNumber="9">32.2 29.3</td>
<td id="768B5DECFFA00058B5C2303F8268AC6E" box="[1003,1044,1566,1584]" gridcol="2" gridrow="16" pageId="8" pageNumber="9">36.6</td>
<td id="768B5DECFFA00058B2BC303F82B5AC6E" box="[1173,1225,1566,1584]" gridcol="3" gridrow="16" pageId="8" pageNumber="9">33.7</td>
<td id="768B5DECFFA00058B379303F8316AC6E" box="[1360,1386,1566,1584]" gridcol="4" gridrow="16" pageId="8" pageNumber="9">2.1</td>
</tr>
<tr id="355A3490FFA00058B745301D8316AC10" box="[364,1386,1596,1614]" gridrow="17" pageId="8" pageNumber="9">
<th id="768B5DECFFA00058B745301D8438AC10" box="[364,580,1596,1614]" gridcol="0" gridrow="17" pageId="8" pageNumber="9">Anal width</th>
<td id="768B5DECFFA00058B451301D8518AC10" box="[632,868,1596,1614]" gridcol="1" gridrow="17" pageId="8" pageNumber="9">14.2 10.6</td>
<td id="768B5DECFFA00058B5C2301D8268AC10" box="[1003,1044,1596,1614]" gridcol="2" gridrow="17" pageId="8" pageNumber="9">14.9</td>
<td id="768B5DECFFA00058B2BC301D82B5AC10" box="[1173,1225,1596,1614]" gridcol="3" gridrow="17" pageId="8" pageNumber="9">13.4</td>
<td id="768B5DECFFA00058B379301D8316AC10" box="[1360,1386,1596,1614]" gridcol="4" gridrow="17" pageId="8" pageNumber="9">1.1</td>
</tr>
<tr id="355A3490FFA00058B745307B8316AC32" box="[364,1386,1626,1644]" gridrow="18" pageId="8" pageNumber="9">
<th id="768B5DECFFA00058B745307B8438AC32" box="[364,580,1626,1644]" gridcol="0" gridrow="18" pageId="8" pageNumber="9">Snout-opercle length</th>
<td id="768B5DECFFA00058B451307B8518AC32" box="[632,868,1626,1644]" gridcol="1" gridrow="18" pageId="8" pageNumber="9">24.3 20.4</td>
<td id="768B5DECFFA00058B5C2307B8268AC32" box="[1003,1044,1626,1644]" gridcol="2" gridrow="18" pageId="8" pageNumber="9">25.8</td>
<td id="768B5DECFFA00058B2BC307B82B5AC32" box="[1173,1225,1626,1644]" gridcol="3" gridrow="18" pageId="8" pageNumber="9">22.6</td>
<td id="768B5DECFFA00058B379307B8316AC32" box="[1360,1386,1626,1644]" gridcol="4" gridrow="18" pageId="8" pageNumber="9">1.2</td>
</tr>
<tr id="355A3490FFA00058B74530598316ACD4" box="[364,1386,1656,1674]" gridrow="19" pageId="8" pageNumber="9">
<th id="768B5DECFFA00058B74530598316ACD4" box="[364,1386,1656,1674]" colspan="5" colspanRight="4" gridcol="0" gridrow="19" pageId="8" pageNumber="9">
<emphasis id="B91EEAC0FFA0FFAFB4B6305885E1ACD7" bold="true" box="[671,925,1656,1674]" pageId="8" pageNumber="9">Percentages of head length</emphasis>
</th>
</tr>
<tr id="355A3490FFA00058B74530B78316ACF6" box="[364,1386,1686,1704]" gridrow="20" pageId="8" pageNumber="9">
<th id="768B5DECFFA00058B74530B78438ACF6" box="[364,580,1686,1704]" gridcol="0" gridrow="20" pageId="8" pageNumber="9">Head width</th>
<td id="768B5DECFFA00058B45130B78518ACF6" box="[632,868,1686,1704]" gridcol="1" gridrow="20" pageId="8" pageNumber="9">86.7 76.1</td>
<td id="768B5DECFFA00058B5C230B78268ACF6" box="[1003,1044,1686,1704]" gridcol="2" gridrow="20" pageId="8" pageNumber="9">90.3</td>
<td id="768B5DECFFA00058B2BC30B782B5ACF6" box="[1173,1225,1686,1704]" gridcol="3" gridrow="20" pageId="8" pageNumber="9">83.3</td>
<td id="768B5DECFFA00058B37930B78316ACF6" box="[1360,1386,1686,1704]" gridcol="4" gridrow="20" pageId="8" pageNumber="9">3.8</td>
</tr>
<tr id="355A3490FFA00058B74530958316AC98" box="[364,1386,1716,1734]" gridrow="21" pageId="8" pageNumber="9">
<th id="768B5DECFFA00058B74530958438AC98" box="[364,580,1716,1734]" gridcol="0" gridrow="21" pageId="8" pageNumber="9">Head depth</th>
<td id="768B5DECFFA00058B45130958518AC98" box="[632,868,1716,1734]" gridcol="1" gridrow="21" pageId="8" pageNumber="9">71.2 66.5</td>
<td id="768B5DECFFA00058B5C230958268AC98" box="[1003,1044,1716,1734]" gridcol="2" gridrow="21" pageId="8" pageNumber="9">76.3</td>
<td id="768B5DECFFA00058B2BC309582B5AC98" box="[1173,1225,1716,1734]" gridcol="3" gridrow="21" pageId="8" pageNumber="9">70.3</td>
<td id="768B5DECFFA00058B37930958316AC98" box="[1360,1386,1716,1734]" gridcol="4" gridrow="21" pageId="8" pageNumber="9">2.8</td>
</tr>
<tr id="355A3490FFA00058B74530F38316ACBA" box="[364,1386,1746,1764]" gridrow="22" pageId="8" pageNumber="9">
<th id="768B5DECFFA00058B74530F38438ACBA" box="[364,580,1746,1764]" gridcol="0" gridrow="22" pageId="8" pageNumber="9">Snout length</th>
<td id="768B5DECFFA00058B45130F38518ACBA" box="[632,868,1746,1764]" gridcol="1" gridrow="22" pageId="8" pageNumber="9">56.9 51.2</td>
<td id="768B5DECFFA00058B5C230F38268ACBA" box="[1003,1044,1746,1764]" gridcol="2" gridrow="22" pageId="8" pageNumber="9">59.8</td>
<td id="768B5DECFFA00058B2BC30F382B5ACBA" box="[1173,1225,1746,1764]" gridcol="3" gridrow="22" pageId="8" pageNumber="9">55.5</td>
<td id="768B5DECFFA00058B37930F38316ACBA" box="[1360,1386,1746,1764]" gridcol="4" gridrow="22" pageId="8" pageNumber="9">1.9</td>
</tr>
<tr id="355A3490FFA00058B74530D18316AD5C" box="[364,1386,1776,1794]" gridrow="23" pageId="8" pageNumber="9">
<th id="768B5DECFFA00058B74530D18438AD5C" box="[364,580,1776,1794]" gridcol="0" gridrow="23" pageId="8" pageNumber="9">Interorbital width</th>
<td id="768B5DECFFA00058B45130D18518AD5C" box="[632,868,1776,1794]" gridcol="1" gridrow="23" pageId="8" pageNumber="9">46.1 41.8</td>
<td id="768B5DECFFA00058B5C230D18268AD5C" box="[1003,1044,1776,1794]" gridcol="2" gridrow="23" pageId="8" pageNumber="9">48.5</td>
<td id="768B5DECFFA00058B2BC30D182B5AD5C" box="[1173,1225,1776,1794]" gridcol="3" gridrow="23" pageId="8" pageNumber="9">44.3</td>
<td id="768B5DECFFA00058B37930D18316AD5C" box="[1360,1386,1776,1794]" gridcol="4" gridrow="23" pageId="8" pageNumber="9">1.5</td>
</tr>
<tr id="355A3490FFA00058B745312F8316AD7E" box="[364,1386,1806,1824]" gridrow="24" pageId="8" pageNumber="9">
<th id="768B5DECFFA00058B745312F8438AD7E" box="[364,580,1806,1824]" gridcol="0" gridrow="24" pageId="8" pageNumber="9">Orbital diameter</th>
<td id="768B5DECFFA00058B451312F8518AD7E" box="[632,868,1806,1824]" gridcol="1" gridrow="24" pageId="8" pageNumber="9">19.5 15.3</td>
<td id="768B5DECFFA00058B5C2312F8268AD7E" box="[1003,1044,1806,1824]" gridcol="2" gridrow="24" pageId="8" pageNumber="9">21.8</td>
<td id="768B5DECFFA00058B2BC312F82B5AD7E" box="[1173,1225,1806,1824]" gridcol="3" gridrow="24" pageId="8" pageNumber="9">18.5</td>
<td id="768B5DECFFA00058B379312F8316AD7E" box="[1360,1386,1806,1824]" gridcol="4" gridrow="24" pageId="8" pageNumber="9">1.6</td>
</tr>
<tr id="355A3490FFA00058B745310D8316AD60" box="[364,1386,1836,1854]" gridrow="25" pageId="8" pageNumber="9">
<th id="768B5DECFFA00058B745310D8438AD60" box="[364,580,1836,1854]" gridcol="0" gridrow="25" pageId="8" pageNumber="9">Suborbital depth</th>
<td id="768B5DECFFA00058B451310D8518AD60" box="[632,868,1836,1854]" gridcol="1" gridrow="25" pageId="8" pageNumber="9">29.5 22.9</td>
<td id="768B5DECFFA00058B5C2310D8268AD60" box="[1003,1044,1836,1854]" gridcol="2" gridrow="25" pageId="8" pageNumber="9">34.1</td>
<td id="768B5DECFFA00058B2BC310D82B5AD60" box="[1173,1225,1836,1854]" gridcol="3" gridrow="25" pageId="8" pageNumber="9">27.2</td>
<td id="768B5DECFFA00058B379310D8316AD60" box="[1360,1386,1836,1854]" gridcol="4" gridrow="25" pageId="8" pageNumber="9">2.3</td>
</tr>
<tr id="355A3490FFA00058B745316B8316AD02" box="[364,1386,1866,1884]" gridrow="26" pageId="8" pageNumber="9">
<th id="768B5DECFFA00058B745316B8438AD02" box="[364,580,1866,1884]" gridcol="0" gridrow="26" pageId="8" pageNumber="9">Mandibular ramus</th>
<td id="768B5DECFFA00058B451316B8518AD02" box="[632,868,1866,1884]" gridcol="1" gridrow="26" pageId="8" pageNumber="9">11.3 10.6</td>
<td id="768B5DECFFA00058B5C2316B8268AD02" box="[1003,1044,1866,1884]" gridcol="2" gridrow="26" pageId="8" pageNumber="9">15.4</td>
<td id="768B5DECFFA00058B2BC316B82B5AD02" box="[1173,1225,1866,1884]" gridcol="3" gridrow="26" pageId="8" pageNumber="9">13.1</td>
<td id="768B5DECFFA00058B379316B8316AD02" box="[1360,1386,1866,1884]" gridcol="4" gridrow="26" pageId="8" pageNumber="9">1.5</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD536D2FFA1FFAEB424369F833EA82E" blockId="9.[488,1461,190,1371]" pageId="9" pageNumber="10">
Lower surface of head naked. Head lacking ridges. Parieto-supraoccipital process elevated and with conspicuous pair of tufts of hypertrophied odontodes in specimens of all examined sizes. Predorsal region without ridges. Body elongate and compressed at caudal peduncle. Greatest body width at cleithral region, progressively narrowing anteriorly towards snout tip and posteriorly towards caudal-fin. Head and trunk covered by dermal plates, except for naked area around dorsal-fin insertion. Body dorsoventrally compressed. Dorsal profile convex from snout tip to posterior margin of parieto-supraoccipital and slightly concave from that point to dorsal-fin origin. Dorsal profile slightly concave and descending from dorsal-fin origin to first upper procurrent caudal-fin ray, rising posteriorly to insertion of caudal fin. Greatest body depth at unbranched dorsal-fin ray insertion. Cleithrum and coracoid exposed in ventral view, covered by odontodes. 
<emphasis id="B91EEAC0FFA1FFAEB4433478857DA82F" box="[618,769,601,625]" italics="true" pageId="9" pageNumber="10">Arrector fossae</emphasis>
completely enclosed by ventral lamina of coracoid.
</paragraph>
<paragraph id="8BD536D2FFA1FFAEB424345F8419A99E" blockId="9.[488,1461,190,1371]" pageId="9" pageNumber="10">Ventral profile straight and descending from snout tip to opercular region, slightly convex from opercular region to anal-fin origin and slightly concave from that point to lower procurrent caudal-fin ray origin. Lateral surface of body entirely covered by plates; dorsal series with 23–24 plates. Mid-dorsal plate series truncated with 18– 19 plates not reaching end of caudal peduncle. Lateral plate series with 22–23 plates. Lateral line incomplete, with gaps in line of pores along mid-length of body. Mid-ventral series of lateral plates well-developed, reaching middle of caudal peduncle (17– 19 plates). Ventral plates series with 20–21 plates. Body plates covered with minute odontodes.</paragraph>
<paragraph id="8BD536D2FFA1FFAEB42435EF85E5AF05" blockId="9.[488,1461,190,1371]" pageId="9" pageNumber="10">Dorsal fin ii,7; its origin slightly posterior to vertical with pelvic-fin origin. Unbranched rays of dorsal fin slightly convex. Tip of adpressed dorsal-fin rays surpassing anal-fin origin. Dorsal-fin spinelet small and rounded (4 c&amp;s). Pectoral-fin rays i,6; tip of adpressed pectoral fin surpassing pelvic-fin origin. Pectoral-fin axillary slit small. Pelvic-fin rays i,5. Distal margin of pelvic fin straight to slightly convex; tip of adpressed pelvic-fin ray reaching anal-fin origin in mature males, but not in females. Adipose fin absent. Anal-fin rays i,5; distal margin slightly convex. Caudal-fin rays i,7–i,7. Slightly emarginated; unbranched rays of same size. Rays of all fins covered with pointed odontodes. Lowest body depth at caudal peduncle. Caudal peduncle ellipsoid in cross section, rounded dorsally and ventrally. Hypurals elements fused in a unique hypural plate. Total vertebrae 28 or 29 (4 c&amp;s).</paragraph>
<paragraph id="8BD536D2FFA1FFAEB42433AF822AACA8" blockId="9.[486,1460,1422,1782]" pageId="9" pageNumber="10">
<emphasis id="B91EEAC0FFA1FFAEB42433AF84A4AFF8" bold="true" box="[525,728,1422,1446]" pageId="9" pageNumber="10">Color in alcohol.</emphasis>
Background color of dorsal region of head and trunk brown. Four dark brown saddles along dorsal portion of body: first at dorsal-fin origin, second at end of dorsal-fin base, third at middle of caudal peduncle, and fourth reaching anteriormost caudal procurrent ray. Unpigmented portion of snout appears as two parallel hyaline stripes from rostral plate to nares. Mid-lateral dark brown stripe extending from tip of snout to caudal peduncle. Ventral portion of body almost entirely yellowish, except for dark, randomly-distributed chromatophores and for a concentration of chromatophores at anal-fin origin. Dorsal, pectoral, and pelvic fins with dark, irregularly distributed chromatophores. Caudal-fin brown, with two smaller rounded hyaline areas anteriorly, and two large rounded hyaline areas posteriorly (
<figureCitation id="13512A57FFA1FFAEB22430FF8235ACA8" box="[1037,1097,1758,1782]" captionStart="FIGURE 1" captionStartId="5.[204,277,1276,1296]" captionTargetBox="[487,1459,880,1889]" captionTargetId="figure-430@5.[594,1352,880,1888]" captionTargetPageId="5" captionText="FIGURE 1 | A. Maximum likelihood – ML (LogL = -3132.01) tree of Corumbataia species using TN93+G+I nucleotide substitution model and based on the analysis of partial sequence of cytochrome oxidase C subunit I (COI). Numbers at nodes are bootstrap values based on 1000 pseudoreplicates. Values below 50% are not shown. B. Majority rule consensus tree obtained in Bayesian analysis using GTR+I. Numbers below branches are posterior probabilities obtained from 10000 trees." figureDoi="http://doi.org/10.5281/zenodo.10998354" httpUri="https://zenodo.org/record/10998354/files/figure.png" pageId="9" pageNumber="10">Fig.1</figureCitation>
).
</paragraph>
<paragraph id="8BD536D2FFA1FFADB424310985DFAAA5" blockId="9.[486,1457,1832,1894]" lastBlockId="10.[488,1459,190,251]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<emphasis id="B91EEAC0FFA1FFAEB42431098481AD1E" bold="true" box="[525,765,1832,1856]" pageId="9" pageNumber="10">Sexual dimorphism.</emphasis>
Adult males possess a papilla located posterior to the urogenital opening; an unbranched pelvic-fin ray that supports dermal flap along its dorsal surface; and a wider head (84.5–88.6% 
<emphasis id="B91EEAC0FFA2FFADB572369E850CAA88" box="[859,880,191,214]" italics="true" pageId="10" pageNumber="11">vs</emphasis>
. 80.6–85.3 of HL in female) with hypertrophied odontodes on its lateral margin (
<figureCitation id="13512A57FFA2FFADB57A36C285E9AAA5" box="[851,917,227,251]" captionStart="FIGURE 5" captionStartId="10.[488,561,902,922]" captionTargetBox="[487,1459,506,869]" captionTargetId="figure-200@10.[488,1459,507,871]" captionTargetPageId="10" captionText="FIGURE 5 | Hypertrophied odontodes on the lateral margins of head in Corumbataia acanthodela, paratypes, male (left), NUP 22694 and female (right), LBP 19095. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10960543" httpUri="https://zenodo.org/record/10960543/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3706559FFA2FFADB424370F8282ABE8" pageId="10" pageNumber="11" type="distribution">
<paragraph id="8BD536D2FFA2FFADB424370F8282ABE8" blockId="10.[487,1460,302,438]" pageId="10" pageNumber="11">
<emphasis id="B91EEAC0FFA2FFADB424370F8533AB18" bold="true" box="[525,847,302,326]" pageId="10" pageNumber="11">Geographical distribution.</emphasis>
<taxonomicName id="4C6A4D51FFA2FFADB57F370F8223AB18" box="[854,1119,302,326]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="acanthodela">
<emphasis id="B91EEAC0FFA2FFADB57F370F8223AB18" box="[854,1119,302,326]" italics="true" pageId="10" pageNumber="11">Corumbataia acanthodela</emphasis>
</taxonomicName>
is known from 
<collectorName id="269F5304FFA2FFADB33C370F83D3AB1B" box="[1301,1455,302,326]" pageId="10" pageNumber="11">Rio dos Patos</collectorName>
, 
<location id="8EB56009FFA2FFADB7C137728405AB35" LSID="urn:lsid:plazi:treatment:03C387C4FFAEFFACB4C1369F8596A878:8EB56009FFA2FFADB7C137728405AB35" box="[488,633,339,363]" country="Brazil" municipality="Rio Vaivem" name="Rio do Peixe" pageId="10" pageNumber="11" stateProvince="Maranhao">Rio do Peixe</location>
, and 
<collectingMunicipality id="6BB1ACA8FFA2FFADB49D37728540AB35" box="[692,828,339,363]" pageId="10" pageNumber="11">Rio Vaivém</collectingMunicipality>
, all tributaries of the Rio 
<collectingRegion id="49AEF830FFA2FFADB27D377282B4AB35" box="[1108,1224,339,363]" country="Brazil" name="Maranhao" pageId="10" pageNumber="11">Maranhão</collectingRegion>
, in the headwaters of the Rio 
<collectingRegion id="49AEF830FFA2FFADB46A375984C8ABCE" box="[579,692,376,400]" country="Brazil" name="Tocantins" pageId="10" pageNumber="11">Tocantins</collectingRegion>
basin, 
<collectingRegion id="49AEF830FFA2FFADB52D3759853AABCE" box="[772,838,376,400]" country="Brazil" name="Goias" pageId="10" pageNumber="11">Goiás</collectingRegion>
, Central 
<collectingCountry id="F37D7642FFA2FFADB5863759858DABCE" box="[943,1009,376,400]" name="Brazil" pageId="10" pageNumber="11">Brazil</collectingCountry>
(
<figureCitation id="13512A57FFA2FFADB2283759823AABCE" box="[1025,1094,376,400]" captionStart="FIGURE 6" captionStartId="10.[488,561,1761,1781]" captionTargetBox="[490,1458,1044,1719]" captionTargetId="figure-202@10.[476,1467,1029,1739]" captionTargetPageId="10" captionText="FIGURE 6 | Distribution of Corumbataia species. Yellow circles: C. acanthodela, Rio Maranhão; Green circle: C. anosteos, Rio Piçarras; Blue circle: C. britskii, Rio Sucuriu; White circle: C. canoeiro, Rio Paranã; Orange circles: C. cuestae, Rio Tietê; Purple circle: C. liliai, Rio Correntes; Brown circles: C. lucianoi, Rio Correntes; Red circle: C. tocantinensis, Rio Vermelho; Pink circle: C. veadeiros, Rio Paranã." figureDoi="http://doi.org/10.5281/zenodo.10960545" httpUri="https://zenodo.org/record/10960545/files/figure.png" pageId="10" pageNumber="11">Fig. 6</figureCitation>
). The new species was found in clear water of fast-flowing stream with rocks and marginal vegetation.
</paragraph>
</subSubSection>
<caption id="DF15665AFFA2FFADB7C135A78291A99E" ID-DOI="http://doi.org/10.5281/zenodo.10960543" ID-Zenodo-Dep="10960543" httpUri="https://zenodo.org/record/10960543/files/figure.png" pageId="10" pageNumber="11" startId="10.[488,561,902,922]" targetBox="[487,1459,506,869]" targetPageId="10" targetType="figure">
<paragraph id="8BD536D2FFA2FFADB7C135A78291A99E" blockId="10.[488,1394,902,960]" pageId="10" pageNumber="11">
<emphasis id="B91EEAC0FFA2FFADB7C135A7842FA9C4" bold="true" box="[488,595,902,922]" pageId="10" pageNumber="11">FIGURE 5 |</emphasis>
Hypertrophied odontodes on the lateral margins of head in 
<taxonomicName id="4C6A4D51FFA2FFADB2A535A68311A9C5" box="[1164,1389,903,923]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="acanthodela">
<emphasis id="B91EEAC0FFA2FFADB2A535A68311A9C5" bold="true" box="[1164,1389,903,923]" italics="true" pageId="10" pageNumber="11">Corumbataia acanthodela</emphasis>
</taxonomicName>
, paratypes, male (left), NUP 22694 and female (right), LBP 19095. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="DF15665AFFA2FFADB7C130C08359AD3B" ID-DOI="http://doi.org/10.5281/zenodo.10960545" ID-Zenodo-Dep="10960545" httpUri="https://zenodo.org/record/10960545/files/figure.png" pageId="10" pageNumber="11" startId="10.[488,561,1761,1781]" targetBox="[490,1458,1044,1719]" targetPageId="10" targetType="figure">
<paragraph id="8BD536D2FFA2FFADB7C130C08359AD3B" blockId="10.[488,1451,1761,1893]" pageId="10" pageNumber="11">
<emphasis id="B91EEAC0FFA2FFADB7C130C0842FACAB" bold="true" box="[488,595,1761,1781]" pageId="10" pageNumber="11">FIGURE 6 |</emphasis>
Distribution of 
<taxonomicName id="4C6A4D51FFA2FFADB4C230C085D1ACAB" box="[747,941,1761,1781]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B91EEAC0FFA2FFADB4C230C08522ACAB" bold="true" box="[747,862,1761,1781]" italics="true" pageId="10" pageNumber="11">Corumbataia</emphasis>
species.
</taxonomicName>
Yellow circles: 
<taxonomicName id="4C6A4D51FFA2FFADB21730C0829BACAB" authority=", Rio" authorityName="Rio" box="[1086,1255,1761,1781]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="acanthodela">
<emphasis id="B91EEAC0FFA2FFADB21730C082C2ACAB" bold="true" box="[1086,1214,1761,1781]" italics="true" pageId="10" pageNumber="11">C. acanthodela</emphasis>
, Rio
</taxonomicName>
Maranhão; Green circle: 
<taxonomicName id="4C6A4D51FFA2FFADB40E3126857BAD45" authority=", Rio Picarras" authorityName="Rio Picarras" box="[551,775,1799,1819]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="anosteos">
<emphasis id="B91EEAC0FFA2FFADB40E312684F4AD45" bold="true" box="[551,648,1799,1819]" italics="true" pageId="10" pageNumber="11">C. anosteos</emphasis>
, Rio Piçarras
</taxonomicName>
; Blue circle: 
<taxonomicName id="4C6A4D51FFA2FFADB55731268231AD45" authority=", Rio Sucuriu" authorityName="Rio Sucuriu" box="[894,1101,1799,1819]" family="Loricariidae" genus="Rhinolekos" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="britskii">
<emphasis id="B91EEAC0FFA2FFADB557312685AEAD45" bold="true" box="[894,978,1799,1819]" italics="true" pageId="10" pageNumber="11">C. britskii</emphasis>
, Rio Sucuriu
</taxonomicName>
; White circle: 
<taxonomicName id="4C6A4D51FFA2FFADB2FA31268321AD45" authority=", Rio" authorityName="Rio" box="[1235,1373,1799,1819]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="canoeiro">
<emphasis id="B91EEAC0FFA2FFADB2FA31268348AD45" bold="true" box="[1235,1332,1799,1819]" italics="true" pageId="10" pageNumber="11">C. canoeiro</emphasis>
, Rio
</taxonomicName>
Paranã; Orange circles: 
<taxonomicName id="4C6A4D51FFA2FFADB452310D854CAD1E" authority=", Rio Tiete" authorityName="Rio Tiete" box="[635,816,1836,1856]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="cuestae">
<emphasis id="B91EEAC0FFA2FFADB452310D84ACAD1E" bold="true" box="[635,720,1836,1856]" italics="true" pageId="10" pageNumber="11">C. cuestae</emphasis>
, Rio Tietê
</taxonomicName>
; Purple circle: 
<taxonomicName id="4C6A4D51FFA2FFADB592310D82F9AD1E" authority=", Rio Correntes" authorityName="Rio Correntes" box="[955,1157,1836,1856]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="liliai">
<emphasis id="B91EEAC0FFA2FFADB592310D858BAD1E" bold="true" box="[955,1015,1836,1856]" italics="true" pageId="10" pageNumber="11">C. liliai</emphasis>
, Rio Correntes
</taxonomicName>
; Brown circles: 
<taxonomicName id="4C6A4D51FFA2FFADB330310D843BAD3B" authority=", Rio Correntes" authorityName="Rio Correntes" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="lucianoi">
<emphasis id="B91EEAC0FFA2FFADB330310D8309AD1E" bold="true" box="[1305,1397,1836,1856]" italics="true" pageId="10" pageNumber="11">C. lucianoi</emphasis>
, Rio Correntes
</taxonomicName>
; Red circle: 
<taxonomicName id="4C6A4D51FFA2FFADB49E317085B0AD3B" authority=", Rio Vermelho" authorityName="Rio Vermelho" box="[695,972,1873,1893]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="tocantinensis">
<emphasis id="B91EEAC0FFA2FFADB49E31708543AD3B" bold="true" box="[695,831,1873,1893]" italics="true" pageId="10" pageNumber="11">C. tocantinensis</emphasis>
, Rio Vermelho
</taxonomicName>
; Pink circle: 
<taxonomicName id="4C6A4D51FFA2FFADB26C317082ABAD3B" authority=", Rio" authorityName="Rio" box="[1093,1239,1873,1893]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="veadeiros">
<emphasis id="B91EEAC0FFA2FFADB26C317082D1AD3B" bold="true" box="[1093,1197,1873,1893]" italics="true" pageId="10" pageNumber="11">C. veadeiros</emphasis>
, Rio
</taxonomicName>
Paranã.
</paragraph>
</caption>
<subSubSection id="C3706559FFA3FFACB424369F850BAB7E" pageId="11" pageNumber="12" type="etymology">
<paragraph id="8BD536D2FFA3FFACB424369F850BAB7E" blockId="11.[488,1459,190,289]" pageId="11" pageNumber="12">
<emphasis id="B91EEAC0FFA3FFACB424369F84E4AA88" bold="true" box="[525,664,190,214]" pageId="11" pageNumber="12">Etymology.</emphasis>
The specific epithet 
<taxonomicName id="4C6A4D51FFA3FFACB5AC369F8582AA88" box="[901,1022,190,214]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="acanthodela">
<emphasis id="B91EEAC0FFA3FFACB5AC369F8582AA88" box="[901,1022,190,214]" italics="true" pageId="11" pageNumber="12">acanthodela</emphasis>
</taxonomicName>
is from Greek 
<emphasis id="B91EEAC0FFA3FFACB283369F8280AA88" box="[1194,1276,190,214]" italics="true" pageId="11" pageNumber="12">akantha</emphasis>
meaning thorn, and 
<emphasis id="B91EEAC0FFA3FFACB43236C28433AAA5" box="[539,591,227,251]" italics="true" pageId="11" pageNumber="12">delos</emphasis>
meaning conspicuous, in reference to the hypertrophied odontodes in the head of mature males. An adjective.
</paragraph>
</subSubSection>
<subSubSection id="C3706559FFA3FFACB42437728596A878" pageId="11" pageNumber="12" type="conservation">
<paragraph id="8BD536D2FFA3FFACB42437728596A878" blockId="11.[488,1461,339,550]" pageId="11" pageNumber="12">
<emphasis id="B91EEAC0FFA3FFACB4243772857DAB35" bold="true" box="[525,769,339,363]" pageId="11" pageNumber="12">Conservation status.</emphasis>
<taxonomicName id="4C6A4D51FFA3FFACB5203772826EAB35" box="[777,1042,339,363]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="acanthodela">
<emphasis id="B91EEAC0FFA3FFACB5203772826EAB35" box="[777,1042,339,363]" italics="true" pageId="11" pageNumber="12">Corumbataia acanthodela</emphasis>
</taxonomicName>
is known from three localities in the Rio 
<collectingRegion id="49AEF830FFA3FFACB433375984F0ABCE" box="[538,652,376,400]" country="Brazil" name="Maranhao" pageId="11" pageNumber="12">Maranhão</collectingRegion>
basin. The areas where the specimens were collected are mountainous and relatively well preserved, which do not qualify it for threatened status. Therefore, 
<taxonomicName id="4C6A4D51FFA3FFACB7C137E584F8AB85" box="[488,644,451,475]" family="Loricariidae" genus="Corumbataia" kingdom="Animalia" order="Siluriformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="acanthodela">
<emphasis id="B91EEAC0FFA3FFACB7C137E584F8AB85" box="[488,644,451,475]" italics="true" pageId="11" pageNumber="12">C. acanthodela</emphasis>
</taxonomicName>
is here recommended to be categorized as Least Concern (
<collectionCode id="ED7BAE17FFA3FFACB31037E28323AB85" box="[1337,1375,451,475]" pageId="11" pageNumber="12">LC</collectionCode>
) under the categories and criteria of the International Union for Conservation Nature (
<collectionCode id="ED7BAE17FFA3FFACB34137C883CFA85F" box="[1384,1459,489,513]" pageId="11" pageNumber="12">IUCN</collectionCode>
Standards and Petitions Subcommittee, 2019).
</paragraph>
</subSubSection>
</treatment>
</document>