340 lines
75 KiB
XML
340 lines
75 KiB
XML
<document id="799E42F5D9DD098DF3ADF82F2A64F9C5" ID-DOI="10.5281/zenodo.5341505" ID-ISSN="0374-1036" ID-Zenodo-Dep="5341505" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630190714458" checkinUser="felipe" docAuthor="Cassis, Gerasimos & Symonds, Celia" docDate="2008" docId="03C387BADC31FF92BC5BFE866AF6FE3A" docLanguage="en" docName="ActaEntMusNatPra.2008.48.2.433-484.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 48 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Inoma Hacker 1927" docType="treatment" docVersion="4" lastPageNumber="440" masterDocId="FFFAFFC2DC35FF95BE56FF92685EFF8E" masterDocTitle="Systematics, biogeography and host associations of the lace bug genus Inoma (Hemiptera: Heteroptera: Tingidae)" masterLastPageNumber="484" masterPageNumber="433" pageNumber="437" updateTime="1699181464015" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A97411307865E55BDB07129A6A055B58">Systematics, biogeography and host associations of the lace bug genus Inoma (Hemiptera: Heteroptera: Tingidae)</mods:title>
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<mods:namePart id="3CAF11D9CC03BAF2F998FC9E71E24991">Cassis, Gerasimos</mods:namePart>
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<mods:namePart id="84A257B2C5BD3F537C3150AC4E73C6D5">Symonds, Celia</mods:namePart>
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<mods:title id="6AFBCD3FB7AC020AD9425E9E12C3CEFE">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:part id="016E48732B9247CE7394D78067992D6B">
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<mods:date id="1E6FF1CD8E7519ED92A54AE4219216A1">2008</mods:date>
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<mods:number id="451530A8767E173C2F73BB6CE19F6B72">2008-12-08</mods:number>
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<mods:number id="26C4A202F52F4D3DCF62302E20369908">48</mods:number>
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<mods:identifier id="C4A0103EAFBD9F103FC3B50F01C9C3DC" type="DOI">10.5281/zenodo.5341505</mods:identifier>
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<paragraph id="8BD536ACDC31FF91BC5BFE866AAEFEA5" blockId="4.[525,752,275,300]" box="[525,752,275,300]" pageId="4" pageNumber="437">
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<heading id="D09D81C0DC31FF91BC5BFE866AAEFEA5" bold="true" box="[525,752,275,300]" centered="true" fontSize="8" level="2" pageId="4" pageNumber="437" reason="3">
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<taxonomicName id="4C6A4D2FDC31FF91BC5BFE866AAEFEA5" ID-CoL="54HV" authority="Hacker, 1927" authorityName="Hacker" authorityYear="1927" box="[525,752,275,300]" class="Insecta" family="Tingidae" genus="Inoma" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="437" phylum="Arthropoda" rank="genus">
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<emphasis id="B91EEABEDC31FF91BC5BFE866AAEFEA5" bold="true" box="[525,752,275,300]" pageId="4" pageNumber="437">
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<emphasis id="B91EEABEDC31FF91BC5BFE866A08FEA5" bold="true" box="[525,598,276,299]" italics="true" pageId="4" pageNumber="437">Inoma</emphasis>
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<bibRefCitation id="EFFB4B5DDC31FF91BC0AFE866AAEFEA5" author="HACKER H." box="[604,752,275,300]" pageId="4" pageNumber="437" pagination="19 - 32" refId="ref28582" refString="HACKER H. 1927: New Tingitoidea (Hemiptera) in the Queensland Museum. Memoirs of Queensland Museum 9: 19 - 32 + pls. vi-x." type="journal article" year="1927">Hacker, 1927</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3706527DC31FF91BEC1FEA86942FE0C" pageId="4" pageNumber="437" type="reference_group">
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<paragraph id="8BD536ACDC31FF91BEC1FEA86993FEC3" blockId="4.[151,1126,313,386]" box="[151,461,313,333]" pageId="4" pageNumber="437">
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<treatmentCitationGroup id="AB7A1182DC31FF91BEC1FEA86993FEC3" box="[151,461,313,333]" pageId="4" pageNumber="437">
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<treatmentCitation id="0ACB10BDDC31FF91BEC1FEA8693BFEC2" author="HACKER H." box="[151,357,313,333]" page="25" pageId="4" pageNumber="437" year="1927">
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<taxonomicName id="4C6A4D2FDC31FF91BEC1FEA8693BFEC2" ID-CoL="54HV" authority="Hacker, 1927: 25" authorityName="Hacker" authorityPageNumber="25" authorityYear="1927" box="[151,357,313,333]" class="Insecta" family="Tingidae" genus="Inoma" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="437" phylum="Arthropoda" rank="genus">
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<emphasis id="B91EEABEDC31FF91BEC1FEA86890FEC2" box="[151,206,314,332]" italics="true" pageId="4" pageNumber="437">Inoma</emphasis>
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<bibRefCitation id="EFFB4B5DDC31FF91BE85FEAB693BFEC2" author="HACKER H." box="[211,357,313,333]" pageId="4" pageNumber="437" pagination="19 - 32" refId="ref28582" refString="HACKER H. 1927: New Tingitoidea (Hemiptera) in the Queensland Museum. Memoirs of Queensland Museum 9: 19 - 32 + pls. vi-x." type="journal article" year="1927">Hacker, 1927: 25</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(gen. nov.).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BD536ACDC31FF91BEC1FEC76942FE0C" blockId="4.[151,1126,313,386]" pageId="4" pageNumber="437">
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<treatmentCitationGroup id="AB7A1182DC31FF91BEC1FEC76942FE0C" pageId="4" pageNumber="437">
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<treatmentCitation id="0ACB10BDDC31FF91BEC1FEC76992FEE9" author="DRAKE C. J. & RUHOFF F. A." box="[151,460,340,360]" page="62" pageId="4" pageNumber="437" year="1960">
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<taxonomicName id="4C6A4D2FDC31FF91BEC1FEC769F0FEE9" ID-CoL="54HV" authority=": DRAKE & RUHOFF (1960)" authorityName=": DRAKE & RUHOFF" authorityYear="1960" box="[151,430,340,360]" class="Insecta" family="Tingidae" genus="Inoma" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="437" phylum="Arthropoda" rank="genus">
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<emphasis id="B91EEABEDC31FF91BEC1FEC76890FEE9" box="[151,206,341,359]" italics="true" pageId="4" pageNumber="437">Inoma</emphasis>
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:
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<bibRefCitation id="EFFB4B5DDC31FF91BE8FFEC769F0FEE9" author="DRAKE C. J. & RUHOFF F. A." box="[217,430,340,360]" pageId="4" pageNumber="437" pagination="1 - 105" refId="ref28368" refString="DRAKE C. J. & RUHOFF F. A. 1960: Lace-bug genera of the world (Hemiptera: Tingidae). Proceedings of the United States National Museum 112: 1 - 105 + pls. i-ix." type="journal article" year="1960">DRAKE & RUHOFF (1960)</bibRefCitation>
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</taxonomicName>
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: 62
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||
</treatmentCitation>
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(list);
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<treatmentCitation id="0ACB10BDDC31FF91BC50FEC76B5DFEE9" author="DRAKE C. J. & RUHOFF F. A." box="[518,771,340,360]" page="249" pageId="4" pageNumber="437" year="1965">
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<bibRefCitation id="EFFB4B5DDC31FF91BC50FEC76A85FEE9" author="DRAKE C. J. & RUHOFF F. A." box="[518,731,340,360]" pageId="4" pageNumber="437" pagination="1 - 634" refId="ref28409" refString="DRAKE C. J. & RUHOFF F. A. 1965: Lacebugs of the world: a catalog (Hemiptera: Tingidae). Bulletin of the United States National Museum 243: i-viii + 1 - 634 pp + 56 pls." type="book chapter" year="1965">DRAKE & RUHOFF (1965)</bibRefCitation>
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: 249
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||
</treatmentCitation>
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(catalogue);
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<treatmentCitation id="0ACB10BDDC31FF91BD23FEC66C38FEE9" author="CASSIS G. & GROSS G. F." box="[885,1126,340,359]" page="417" pageId="4" pageNumber="437" year="1995">
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<bibRefCitation id="EFFB4B5DDC31FF91BD23FEC66C60FEE9" author="CASSIS G. & GROSS G. F." box="[885,1086,340,359]" pageId="4" pageNumber="437" refId="ref27525" refString="CASSIS G. & GROSS G. F. 1995: Hemiptera: Heteroptera (Coleorrhyncha to Cimicomorpha). In: HOUSTON W. W. K. & MAYNARD G. V. (eds.): Zoological Catalogue of Australia. Volume 27.3 (A). ABRS, Melbourne, xv + 506 pp." type="book" year="1995">CASSIS & GROSS (1995)</bibRefCitation>
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: 417
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||
</treatmentCitation>
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(catalogue).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3706527DC31FF91BEC1FE086B65FE3D" box="[151,827,410,435]" pageId="4" pageNumber="437" type="type_taxon">
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<paragraph id="8BD536ACDC31FF91BEC1FE086B65FE3D" blockId="4.[151,827,410,435]" box="[151,827,410,435]" pageId="4" pageNumber="437">
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<emphasis id="B91EEABEDC31FF91BEC1FE086975FE3C" bold="true" box="[151,299,410,434]" pageId="4" pageNumber="437">
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<typeStatus id="54D1880EDC31FF91BEC1FE086891FE3C" box="[151,207,410,434]" pageId="4" pageNumber="437">Type</typeStatus>
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species.
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</emphasis>
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||
<taxonomicName id="4C6A4D2FDC31FF91BF64FE096AC3FE3C" ID-CoL="3PRXN" authority="Hacker, 1927" authorityName="Hacker" authorityYear="1927" box="[306,669,410,434]" class="Insecta" family="Tingidae" genus="Inoma" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="437" phylum="Arthropoda" rank="species" species="multispinosa">
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<emphasis id="B91EEABEDC31FF91BF64FE096A5AFE3C" box="[306,516,411,434]" italics="true" pageId="4" pageNumber="437">Inoma multispinosa</emphasis>
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<bibRefCitation id="EFFB4B5DDC31FF91BC5DFE086AC3FE3C" author="HACKER H." box="[523,669,410,434]" pageId="4" pageNumber="437" pagination="19 - 32" refId="ref28582" refString="HACKER H. 1927: New Tingitoidea (Hemiptera) in the Queensland Museum. Memoirs of Queensland Museum 9: 19 - 32 + pls. vi-x." type="journal article" year="1927">Hacker, 1927</bibRefCitation>
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</taxonomicName>
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, by monotypy.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3706527DC31FF91BEC1FE5B6B3DFCAF" pageId="4" pageNumber="437" type="diagnosis">
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<paragraph id="8BD536ACDC31FF91BEC1FE5B6B3DFCAF" blockId="4.[151,1127,457,801]" pageId="4" pageNumber="437">
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<emphasis id="B91EEABEDC31FF91BEC1FE5B6952FE6F" bold="true" box="[151,268,457,481]" pageId="4" pageNumber="437">Diagnosis.</emphasis>
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<taxonomicName id="4C6A4D2FDC31FF91BF46FE59690AFE6C" authorityName=": DRAKE & RUHOFF" authorityYear="1960" box="[272,340,459,482]" class="Insecta" family="Tingidae" genus="Inoma" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="437" phylum="Arthropoda" rank="genus">
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<emphasis id="B91EEABEDC31FF91BF46FE59690AFE6C" box="[272,340,459,482]" italics="true" pageId="4" pageNumber="437">Inoma</emphasis>
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</taxonomicName>
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is recognised by the following combination of characters: small to medium size, males (BL 1.81-2.96), females (BL 1.9-2.94); brachypterous and macropterous morphs; ovoid body; mottled or variegated colouration; with or without woolly or scale-like, pale setae, often densely distributed; all species with setiferous tubercles on anterior margin and crest of collum, pronotal carinae, margins of paranota and hemelytra, and carinate margins of discoidal area; antennae slender, elongate, AI subequal to AII; head with five elongate, porrect spines, paired frontal and occipital spines, medial spine sometimes forked; pronotum tricarinate, carinae uniseriate, subequal in height and not highly elevated; pronotal collum small to moderately enlarged, subtriangular, dorsally keeled; paranota and costal areas well developed, uniseriate or biseriate; legs slender, elongate; and, male aedeagus with a well developed, heavily sclerotised, distal U-shaped endosomal sclerite.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3706527DC31FF93BEC1FCAA6C10FDBA" lastPageId="6" lastPageNumber="439" pageId="4" pageNumber="437" type="description">
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<paragraph id="8BD536ACDC31FF91BEC1FCAA6A8FFC5E" blockId="4.[151,1127,824,1553]" pageId="4" pageNumber="437">
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<emphasis id="B91EEABEDC31FF91BEC1FCAA6963FCDE" bold="true" box="[151,317,824,848]" pageId="4" pageNumber="437">Redescription.</emphasis>
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COLOURATION. Varying from cream, yellow brown, to orange and red brown, to dark brown (
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<figureCitation id="13512A29DC31FF91BFDFFCCA698CFCFE" box="[393,466,856,880]" captionStart="Fig" captionStartId="10.[151,182,1545,1564]" captionTargetBox="[178,1096,215,1509]" captionTargetId="figure-10@10.[177,1102,215,1512]" captionTargetPageId="10" captionText="Fig. 1. Habitus photographs of Inoma species: I. arrernte sp. nov. (macropters and brachypters), I. breviseta sp. nov., I. fuscata sp. nov. (also lateral view), I. innamincka sp. nov. and I. kalbarri sp. nov. (macropters and female brachypter). Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341507" httpUri="https://zenodo.org/record/5341507/files/figure.png" pageId="4" pageNumber="437">Figs. 1</figureCitation>
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,
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<figureCitation id="13512A29DC31FF91BF89FCCA69B1FCFE" box="[479,495,856,880]" captionStart="Fig" captionStartId="11.[113,144,1518,1537]" captionTargetBox="[118,1089,218,1490]" captionTargetId="figure-13@11.[116,1090,215,1490]" captionTargetPageId="11" captionText="Fig. 2. Habitus photographs of Inoma species:I.multispinosa Hacker, 1927 (macropters and brachypters), I. silveirae sp. nov. (macropters and brachypters, also in lateral view), I. solusa sp. nov. and I. stysi sp. nov. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341509" httpUri="https://zenodo.org/record/5341509/files/figure.png" pageId="4" pageNumber="437">2</figureCitation>
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). Often with mottled or patterned colour pattern on dorsum, more so on hemelytra. Head and callar region of pronotum predominantly darkened, with posterior lobe of pronotum lighter. AIV and tarsi dark brown, almost black. Venter generally darker and more uniform in colouration than dorsum.
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</paragraph>
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<paragraph id="8BD536ACDC31FF90BEE3FC4A6B33FE7A" blockId="4.[151,1127,824,1553]" lastBlockId="5.[113,1090,219,1588]" lastPageId="5" lastPageNumber="438" pageId="4" pageNumber="437">
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VESTITURE. Most species with covering of silvery to golden, woolly or scale-like setae, intermixed either with elongate, curly, woolly setae, or short, flattened, lanceolate, scale-like setae; sometimes with setiferous tubercles only. All species with setiferous tubercles in single or multiple (alternating or opposing) rows, mutiple rows most often along lateral paranotal and costal margins; either major (e.g.,
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<figureCitation id="13512A29DC31FF91BC5EFBCA6A19FBFE" box="[520,583,1112,1136]" captionStart="Fig" captionStartId="21.[113,144,1546,1565]" captionTargetBox="[141,1061,217,1503]" captionTargetId="figure-13@21.[141,1061,217,1504]" captionTargetPageId="21" captionText="Fig. 5. Inoma fuscata sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (♀, 30 μm); c – head lateral (♀, 30 μm); d – pronotum dorsal (♀, 100 μm); e – hemelytra (♀, 100 μm); f – tarsal claw (♀, 3 μm); g – pygophore dorsal (J, 30 μm); h – abdomen ventral (♀, 100 μm). Scales in brackets, measured in μm." figureDoi="http://doi.org/10.5281/zenodo.5341515" httpUri="https://zenodo.org/record/5341515/files/figure.png" pageId="4" pageNumber="437">Fig. 5</figureCitation>
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a-e), or minor setiferous tubercles (e.g.,
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<figureCitation id="13512A29DC31FF91BDA1FBCA6C69FBFE" box="[1015,1079,1112,1136]" captionStart="Fig" captionStartId="20.[151,182,1488,1507]" captionTargetBox="[198,1079,219,1449]" captionTargetId="figure-10@20.[198,1079,219,1449]" captionTargetPageId="20" captionText="Fig. 4. Inoma arrernte sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (♀, 30 μm); c – head lateral (♀, 30 μm); d – pronotum dorsal (♀, 100 μm); e – hemelytra (J, 100 μm); f – hemelytra (♀, 100 μm); g – pygophore dorsal (J, 30 μm); h – abdomen ventral (♀, 30 μm). Scales in brackets, measured in μm. FS, frontal spine; MS, medial spine; OS, occipital spine; Bu, bucculae; CaR, callar region; Co, collum; Pa, paranota; MC, median carina; LC, lateral carina; PL, posterior lobe; R+M, radius and media vein; Cu, cubitus vein; CoA, costal area; ScA, subcostal area; SuA, sutural area; DiA, discoidal area; RP, right paramere; LP, left paramere; Py, Pygophore; SgP, subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.5341513" httpUri="https://zenodo.org/record/5341513/files/figure.png" pageId="4" pageNumber="437">Fig. 4</figureCitation>
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a-e). Head: setae when present, woolly or scale-like, dense and adpressed (e.g.,
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<figureCitation id="13512A29DC31FF91BDE7FBEA6BAEFB1E" box="[945,1008,1144,1168]" captionStart="Fig" captionStartId="20.[151,182,1488,1507]" captionTargetBox="[198,1079,219,1449]" captionTargetId="figure-10@20.[198,1079,219,1449]" captionTargetPageId="20" captionText="Fig. 4. Inoma arrernte sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (♀, 30 μm); c – head lateral (♀, 30 μm); d – pronotum dorsal (♀, 100 μm); e – hemelytra (J, 100 μm); f – hemelytra (♀, 100 μm); g – pygophore dorsal (J, 30 μm); h – abdomen ventral (♀, 30 μm). Scales in brackets, measured in μm. FS, frontal spine; MS, medial spine; OS, occipital spine; Bu, bucculae; CaR, callar region; Co, collum; Pa, paranota; MC, median carina; LC, lateral carina; PL, posterior lobe; R+M, radius and media vein; Cu, cubitus vein; CoA, costal area; ScA, subcostal area; SuA, sutural area; DiA, discoidal area; RP, right paramere; LP, left paramere; Py, Pygophore; SgP, subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.5341513" httpUri="https://zenodo.org/record/5341513/files/figure.png" pageId="4" pageNumber="437">Fig. 4</figureCitation>
|
||
a-c); minor setiferous tubercles present around base or on cephalic spines (e.g.,
|
||
<figureCitation id="13512A29DC31FF91BD3CFB0A6B9EFB3E" box="[874,960,1176,1200]" captionStart="Fig" captionStartId="20.[151,182,1488,1507]" captionTargetBox="[198,1079,219,1449]" captionTargetId="figure-10@20.[198,1079,219,1449]" captionTargetPageId="20" captionText="Fig. 4. Inoma arrernte sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (♀, 30 μm); c – head lateral (♀, 30 μm); d – pronotum dorsal (♀, 100 μm); e – hemelytra (J, 100 μm); f – hemelytra (♀, 100 μm); g – pygophore dorsal (J, 30 μm); h – abdomen ventral (♀, 30 μm). Scales in brackets, measured in μm. FS, frontal spine; MS, medial spine; OS, occipital spine; Bu, bucculae; CaR, callar region; Co, collum; Pa, paranota; MC, median carina; LC, lateral carina; PL, posterior lobe; R+M, radius and media vein; Cu, cubitus vein; CoA, costal area; ScA, subcostal area; SuA, sutural area; DiA, discoidal area; RP, right paramere; LP, left paramere; Py, Pygophore; SgP, subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.5341513" httpUri="https://zenodo.org/record/5341513/files/figure.png" pageId="4" pageNumber="437">Figs. 4a</figureCitation>
|
||
,
|
||
<figureCitation id="13512A29DC31FF91BD9DFB0A6BB9FB3E" box="[971,999,1176,1200]" captionStart="Fig" captionStartId="21.[113,144,1546,1565]" captionTargetBox="[141,1061,217,1503]" captionTargetId="figure-13@21.[141,1061,217,1504]" captionTargetPageId="21" captionText="Fig. 5. Inoma fuscata sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (♀, 30 μm); c – head lateral (♀, 30 μm); d – pronotum dorsal (♀, 100 μm); e – hemelytra (♀, 100 μm); f – tarsal claw (♀, 3 μm); g – pygophore dorsal (J, 30 μm); h – abdomen ventral (♀, 100 μm). Scales in brackets, measured in μm." figureDoi="http://doi.org/10.5281/zenodo.5341515" httpUri="https://zenodo.org/record/5341515/files/figure.png" pageId="4" pageNumber="437">5a</figureCitation>
|
||
). Antennae: AI and AII glabrous or with setae as on head and in a single ring around segment (e.g.,
|
||
<figureCitation id="13512A29DC31FF91BA64FB2A68EDFB7E" captionStart="Fig" captionStartId="20.[151,182,1488,1507]" captionTargetBox="[198,1079,219,1449]" captionTargetId="figure-10@20.[198,1079,219,1449]" captionTargetPageId="20" captionText="Fig. 4. Inoma arrernte sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (♀, 30 μm); c – head lateral (♀, 30 μm); d – pronotum dorsal (♀, 100 μm); e – hemelytra (J, 100 μm); f – hemelytra (♀, 100 μm); g – pygophore dorsal (J, 30 μm); h – abdomen ventral (♀, 30 μm). Scales in brackets, measured in μm. FS, frontal spine; MS, medial spine; OS, occipital spine; Bu, bucculae; CaR, callar region; Co, collum; Pa, paranota; MC, median carina; LC, lateral carina; PL, posterior lobe; R+M, radius and media vein; Cu, cubitus vein; CoA, costal area; ScA, subcostal area; SuA, sutural area; DiA, discoidal area; RP, right paramere; LP, left paramere; Py, Pygophore; SgP, subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.5341513" httpUri="https://zenodo.org/record/5341513/files/figure.png" pageId="4" pageNumber="437">Figs. 4a</figureCitation>
|
||
,
|
||
<figureCitation id="13512A29DC31FF91BEEAFB4A6886FB7E" box="[188,216,1240,1264]" captionStart="Fig" captionStartId="21.[113,144,1546,1565]" captionTargetBox="[141,1061,217,1503]" captionTargetId="figure-13@21.[141,1061,217,1504]" captionTargetPageId="21" captionText="Fig. 5. Inoma fuscata sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (♀, 30 μm); c – head lateral (♀, 30 μm); d – pronotum dorsal (♀, 100 μm); e – hemelytra (♀, 100 μm); f – tarsal claw (♀, 3 μm); g – pygophore dorsal (J, 30 μm); h – abdomen ventral (♀, 100 μm). Scales in brackets, measured in μm." figureDoi="http://doi.org/10.5281/zenodo.5341515" httpUri="https://zenodo.org/record/5341515/files/figure.png" pageId="4" pageNumber="437">5a</figureCitation>
|
||
); AIII glabrous or with minor setiferous tubercles (with an elongate, erect terminal seta – straight or recurved), and short, bristle-like setae (e.g.,
|
||
<figureCitation id="13512A29DC31FF91BCB1FB6A6B6BFA9E" box="[743,821,1272,1296]" captionStart="Fig" captionStartId="23.[113,144,1277,1296]" captionTargetBox="[113,1087,219,1238]" captionTargetId="figure-13@23.[113,1087,219,1239]" captionTargetPageId="23" captionText="Fig. 7. Inoma kalbarri sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (J, 30 μm); c – head lateral (♀, 30 μm); d – pronotum dorsal (♀, 100 μm); e – hemelytra (♀, 100 μm); f – genital segment of abdomen ventral (♀, 100 μm). Scales in brackets, measured in μm. Hc, hypocostal vein." figureDoi="http://doi.org/10.5281/zenodo.5341519" httpUri="https://zenodo.org/record/5341519/files/figure.png" pageId="4" pageNumber="437">Fig. 7b</figureCitation>
|
||
); AIV always with both short and elongate bristle-like simple setae. Pronotum: setae when present, woolly or scale-like, most dense on pronotal callar region, less dense on collum, paranota, pronotal longitudinal carinae and posterior lobe (e.g.,
|
||
<figureCitation id="13512A29DC31FF91BFBBFACA6A65FAFE" box="[493,571,1368,1392]" captionStart="Fig" captionStartId="20.[151,182,1488,1507]" captionTargetBox="[198,1079,219,1449]" captionTargetId="figure-10@20.[198,1079,219,1449]" captionTargetPageId="20" captionText="Fig. 4. Inoma arrernte sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (♀, 30 μm); c – head lateral (♀, 30 μm); d – pronotum dorsal (♀, 100 μm); e – hemelytra (J, 100 μm); f – hemelytra (♀, 100 μm); g – pygophore dorsal (J, 30 μm); h – abdomen ventral (♀, 30 μm). Scales in brackets, measured in μm. FS, frontal spine; MS, medial spine; OS, occipital spine; Bu, bucculae; CaR, callar region; Co, collum; Pa, paranota; MC, median carina; LC, lateral carina; PL, posterior lobe; R+M, radius and media vein; Cu, cubitus vein; CoA, costal area; ScA, subcostal area; SuA, sutural area; DiA, discoidal area; RP, right paramere; LP, left paramere; Py, Pygophore; SgP, subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.5341513" httpUri="https://zenodo.org/record/5341513/files/figure.png" pageId="4" pageNumber="437">Fig. 4d</figureCitation>
|
||
); minor or major setiferous tubercles always present along lateral margins of paranota, margins of carinae, anterior margin and dorsal keel of collum (e.g.,
|
||
<figureCitation id="13512A29DC31FF91BF73FA0A6932FA3E" box="[293,364,1432,1456]" captionStart="Fig" captionStartId="21.[113,144,1546,1565]" captionTargetBox="[141,1061,217,1503]" captionTargetId="figure-13@21.[141,1061,217,1504]" captionTargetPageId="21" captionText="Fig. 5. Inoma fuscata sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (♀, 30 μm); c – head lateral (♀, 30 μm); d – pronotum dorsal (♀, 100 μm); e – hemelytra (♀, 100 μm); f – tarsal claw (♀, 3 μm); g – pygophore dorsal (J, 30 μm); h – abdomen ventral (♀, 100 μm). Scales in brackets, measured in μm." figureDoi="http://doi.org/10.5281/zenodo.5341515" httpUri="https://zenodo.org/record/5341515/files/figure.png" pageId="4" pageNumber="437">Fig 5d</figureCitation>
|
||
), also present along ventral margin of paranota (e.g.,
|
||
<figureCitation id="13512A29DC31FF91BDFEFA0A6BABFA3E" box="[936,1013,1432,1456]" captionStart="Fig" captionStartId="21.[113,144,1546,1565]" captionTargetBox="[141,1061,217,1503]" captionTargetId="figure-13@21.[141,1061,217,1504]" captionTargetPageId="21" captionText="Fig. 5. Inoma fuscata sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (♀, 30 μm); c – head lateral (♀, 30 μm); d – pronotum dorsal (♀, 100 μm); e – hemelytra (♀, 100 μm); f – tarsal claw (♀, 3 μm); g – pygophore dorsal (J, 30 μm); h – abdomen ventral (♀, 100 μm). Scales in brackets, measured in μm." figureDoi="http://doi.org/10.5281/zenodo.5341515" httpUri="https://zenodo.org/record/5341515/files/figure.png" pageId="4" pageNumber="437">Fig. 5c</figureCitation>
|
||
). Thoracic pleura and sterna: setae when present, woolly or scale-like, generally denser on pleura, sparser on supracoxal lobes and sterna; posteroventral margin of proepimeron with a few minor or major setiferous tubercles (e.g.,
|
||
<figureCitation id="13512A29DC31FF91BFB8FA6A6A65F99E" box="[494,571,1528,1552]" captionStart="Fig" captionStartId="22.[151,182,1274,1293]" captionTargetBox="[152,1126,218,1237]" captionTargetId="figure-10@22.[151,1126,218,1237]" captionTargetPageId="22" captionText="Fig. 6. Inoma innamincka sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (♀, 30 μm); c – head lateral (♀, 30 μm); d – pronotum dorsal (♀, 100 μm); e – hemelytra (♀, 100 μm); f – genital segment of abdomen ventral (♀, 30 μm). Scales in brackets, measured in μm." figureDoi="http://doi.org/10.5281/zenodo.5341517" httpUri="https://zenodo.org/record/5341517/files/figure.png" pageId="4" pageNumber="437">Fig. 6c</figureCitation>
|
||
); sternal carinae with one or two rows of setae,
|
||
<typeStatus id="54D1880EDC31FF91BA6FFA6B6C39F99F" box="[1081,1127,1529,1553]" pageId="4" pageNumber="437">type</typeStatus>
|
||
variable, rarely with minor setiferous tubercles. Legs: femora and tibiae with minor setiferous tubercles, setae when present, elongate and bristle-like with very small tuberculate base (as in antennae) (e.g.,
|
||
<figureCitation id="13512A29DC30FF90BF4FFE89692EFEBD" box="[281,368,283,307]" captionStart="Fig" captionStartId="23.[113,144,1277,1296]" captionTargetBox="[113,1087,219,1238]" captionTargetId="figure-13@23.[113,1087,219,1239]" captionTargetPageId="23" captionText="Fig. 7. Inoma kalbarri sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (J, 30 μm); c – head lateral (♀, 30 μm); d – pronotum dorsal (♀, 100 μm); e – hemelytra (♀, 100 μm); f – genital segment of abdomen ventral (♀, 100 μm). Scales in brackets, measured in μm. Hc, hypocostal vein." figureDoi="http://doi.org/10.5281/zenodo.5341519" httpUri="https://zenodo.org/record/5341519/files/figure.png" pageId="5" pageNumber="438">Figs. 7b</figureCitation>
|
||
,
|
||
<figureCitation id="13512A29DC30FF90BF2CFE8969C9FEBD" box="[378,407,283,307]" captionStart="Fig" captionStartId="24.[151,182,1497,1516]" captionTargetBox="[195,1083,219,1457]" captionTargetId="figure-10@24.[194,1084,219,1458]" captionTargetPageId="24" captionText="Fig. 8. Inoma silveirae sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (♀, 30 μm); c – head lateral (♀, 20 μm); d – pronotum dorsal (♀, 100 μm); e – macropterous hemelytra (J, 100 μm); f – brachypterous hemelytra (♀, 100 μm); g – pygophore dorsal (J, 30 μm); h – genital segment of abdomen ventral (♀, 100 μm). Scales in brackets, measured in μm." figureDoi="http://doi.org/10.5281/zenodo.5341521" httpUri="https://zenodo.org/record/5341521/files/figure.png" pageId="5" pageNumber="438">8b</figureCitation>
|
||
,
|
||
<figureCitation id="13512A29DC30FF90BFF6FE8969E0FEBD" box="[416,446,283,307]" captionStart="Fig" captionStartId="25.[113,144,1512,1531]" captionTargetBox="[155,1049,219,1466]" captionTargetId="figure-13@25.[154,1049,219,1467]" captionTargetPageId="25" captionText="Fig. 9. Inoma stysi sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (J, 30 μm); c – head lateral (♀, 30 μm); d – pronotum dorsal (♀, 100 μm); e – hemelytra (♀, 200 μm); f – tarsal claw (♀, 3 μm); g – pygophore dorsal (J, 30 μm); h – abdomen ventral (♀, 100 μm). Scales in brackets, measured in μm." figureDoi="http://doi.org/10.5281/zenodo.5341523" httpUri="https://zenodo.org/record/5341523/files/figure.png" pageId="5" pageNumber="438">9h</figureCitation>
|
||
). Hemelytra: (e.g.,
|
||
<figureCitation id="13512A29DC30FF90BCDEFE896A90FEBD" box="[648,718,283,307]" captionStart="Fig" captionStartId="20.[151,182,1488,1507]" captionTargetBox="[198,1079,219,1449]" captionTargetId="figure-10@20.[198,1079,219,1449]" captionTargetPageId="20" captionText="Fig. 4. Inoma arrernte sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (♀, 30 μm); c – head lateral (♀, 30 μm); d – pronotum dorsal (♀, 100 μm); e – hemelytra (J, 100 μm); f – hemelytra (♀, 100 μm); g – pygophore dorsal (J, 30 μm); h – abdomen ventral (♀, 30 μm). Scales in brackets, measured in μm. FS, frontal spine; MS, medial spine; OS, occipital spine; Bu, bucculae; CaR, callar region; Co, collum; Pa, paranota; MC, median carina; LC, lateral carina; PL, posterior lobe; R+M, radius and media vein; Cu, cubitus vein; CoA, costal area; ScA, subcostal area; SuA, sutural area; DiA, discoidal area; RP, right paramere; LP, left paramere; Py, Pygophore; SgP, subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.5341513" httpUri="https://zenodo.org/record/5341513/files/figure.png" pageId="5" pageNumber="438">Figs. 4</figureCitation>
|
||
e-f, 5e) setae when present, woolly, less dense than on pronotal callar region, moderately distributed over discoidal and subcostal areas, very sparse on costal area; minor or major setiferous tubercles present along margin of costal area, on carinate R+M and cubitus veins bounding discoidal and sutural areas, on veins tubercles often recurved, terminal setae generally more elongate than on costal margins and paranota. Abdomen: setae when present, with either scale-like setae (e.g.
|
||
<figureCitation id="13512A29DC30FF90BDF6FE296BA6FE5D" box="[928,1016,443,467]" captionStart="Fig" captionStartId="20.[151,182,1488,1507]" captionTargetBox="[198,1079,219,1449]" captionTargetId="figure-10@20.[198,1079,219,1449]" captionTargetPageId="20" captionText="Fig. 4. Inoma arrernte sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (♀, 30 μm); c – head lateral (♀, 30 μm); d – pronotum dorsal (♀, 100 μm); e – hemelytra (J, 100 μm); f – hemelytra (♀, 100 μm); g – pygophore dorsal (J, 30 μm); h – abdomen ventral (♀, 30 μm). Scales in brackets, measured in μm. FS, frontal spine; MS, medial spine; OS, occipital spine; Bu, bucculae; CaR, callar region; Co, collum; Pa, paranota; MC, median carina; LC, lateral carina; PL, posterior lobe; R+M, radius and media vein; Cu, cubitus vein; CoA, costal area; ScA, subcostal area; SuA, sutural area; DiA, discoidal area; RP, right paramere; LP, left paramere; Py, Pygophore; SgP, subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.5341513" httpUri="https://zenodo.org/record/5341513/files/figure.png" pageId="5" pageNumber="438">Figs. 4h</figureCitation>
|
||
,
|
||
<figureCitation id="13512A29DC30FF90BA57FE296C43FE5D" box="[1025,1053,443,467]" captionStart="Fig" captionStartId="21.[113,144,1546,1565]" captionTargetBox="[141,1061,217,1503]" captionTargetId="figure-13@21.[141,1061,217,1504]" captionTargetPageId="21" captionText="Fig. 5. Inoma fuscata sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (♀, 30 μm); c – head lateral (♀, 30 μm); d – pronotum dorsal (♀, 100 μm); e – hemelytra (♀, 100 μm); f – tarsal claw (♀, 3 μm); g – pygophore dorsal (J, 30 μm); h – abdomen ventral (♀, 100 μm). Scales in brackets, measured in μm." figureDoi="http://doi.org/10.5281/zenodo.5341515" httpUri="https://zenodo.org/record/5341515/files/figure.png" pageId="5" pageNumber="438">5h</figureCitation>
|
||
) or hair-like setae (e.g.,
|
||
<figureCitation id="13512A29DC30FF90BF1DFE4E69C0FE7A" box="[331,414,476,500]" captionStart="Fig" captionStartId="23.[113,144,1277,1296]" captionTargetBox="[113,1087,219,1238]" captionTargetId="figure-13@23.[113,1087,219,1239]" captionTargetPageId="23" captionText="Fig. 7. Inoma kalbarri sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (J, 30 μm); c – head lateral (♀, 30 μm); d – pronotum dorsal (♀, 100 μm); e – hemelytra (♀, 100 μm); f – genital segment of abdomen ventral (♀, 100 μm). Scales in brackets, measured in μm. Hc, hypocostal vein." figureDoi="http://doi.org/10.5281/zenodo.5341519" httpUri="https://zenodo.org/record/5341519/files/figure.png" pageId="5" pageNumber="438">Figs. 7f</figureCitation>
|
||
,
|
||
<figureCitation id="13512A29DC30FF90BFFFFE4E6998FE7A" box="[425,454,476,500]" captionStart="Fig" captionStartId="25.[113,144,1512,1531]" captionTargetBox="[155,1049,219,1466]" captionTargetId="figure-13@25.[154,1049,219,1467]" captionTargetPageId="25" captionText="Fig. 9. Inoma stysi sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (J, 30 μm); c – head lateral (♀, 30 μm); d – pronotum dorsal (♀, 100 μm); e – hemelytra (♀, 200 μm); f – tarsal claw (♀, 3 μm); g – pygophore dorsal (J, 30 μm); h – abdomen ventral (♀, 100 μm). Scales in brackets, measured in μm." figureDoi="http://doi.org/10.5281/zenodo.5341523" httpUri="https://zenodo.org/record/5341523/files/figure.png" pageId="5" pageNumber="438">9h</figureCitation>
|
||
), short or elongate, densely distributed.
|
||
</paragraph>
|
||
<paragraph id="8BD536ACDC30FF93BEC6FE6E6A65FE1D" blockId="5.[113,1090,219,1588]" lastBlockId="6.[151,1127,219,564]" lastPageId="6" lastPageNumber="439" pageId="5" pageNumber="438">
|
||
STRUCTURE. Macropterous and brachypterous forms. Head: five spines present, spines mostly as long and often longer than AI (e.g.,
|
||
<figureCitation id="13512A29DC30FF90BC0BFD8E6AF4FDBD" box="[605,682,539,564]" captionStart="Fig" captionStartId="21.[113,144,1546,1565]" captionTargetBox="[141,1061,217,1503]" captionTargetId="figure-13@21.[141,1061,217,1504]" captionTargetPageId="21" captionText="Fig. 5. Inoma fuscata sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (♀, 30 μm); c – head lateral (♀, 30 μm); d – pronotum dorsal (♀, 100 μm); e – hemelytra (♀, 100 μm); f – tarsal claw (♀, 3 μm); g – pygophore dorsal (J, 30 μm); h – abdomen ventral (♀, 100 μm). Scales in brackets, measured in μm." figureDoi="http://doi.org/10.5281/zenodo.5341515" httpUri="https://zenodo.org/record/5341515/files/figure.png" pageId="5" pageNumber="438">Fig. 5a</figureCitation>
|
||
), rarely shorter than AI (e.g.,
|
||
<figureCitation id="13512A29DC30FF90BDB1FD8E6C6AFDBA" box="[999,1076,540,564]" captionStart="Fig" captionStartId="20.[151,182,1488,1507]" captionTargetBox="[198,1079,219,1449]" captionTargetId="figure-10@20.[198,1079,219,1449]" captionTargetPageId="20" captionText="Fig. 4. Inoma arrernte sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (♀, 30 μm); c – head lateral (♀, 30 μm); d – pronotum dorsal (♀, 100 μm); e – hemelytra (J, 100 μm); f – hemelytra (♀, 100 μm); g – pygophore dorsal (J, 30 μm); h – abdomen ventral (♀, 30 μm). Scales in brackets, measured in μm. FS, frontal spine; MS, medial spine; OS, occipital spine; Bu, bucculae; CaR, callar region; Co, collum; Pa, paranota; MC, median carina; LC, lateral carina; PL, posterior lobe; R+M, radius and media vein; Cu, cubitus vein; CoA, costal area; ScA, subcostal area; SuA, sutural area; DiA, discoidal area; RP, right paramere; LP, left paramere; Py, Pygophore; SgP, subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.5341513" httpUri="https://zenodo.org/record/5341513/files/figure.png" pageId="5" pageNumber="438">Fig. 4a</figureCitation>
|
||
), semi-erect to erect; frontal spines inserted behind antennal bases, either slightly divergent, parallel or slightly convergent, not contiguous or crossing; medial spine straight or apex forked; occipital spines strongly arcuate laterally, or sometimes straight and slightly divergent apically; bucculae prominent, extending slightly beyond head, closed in front, broadly rounded anteriorly laterally (
|
||
<figureCitation id="13512A29DC30FF90BFF3FD2E69ADFD5A" box="[421,499,700,724]" captionStart="Fig" captionStartId="20.[151,182,1488,1507]" captionTargetBox="[198,1079,219,1449]" captionTargetId="figure-10@20.[198,1079,219,1449]" captionTargetPageId="20" captionText="Fig. 4. Inoma arrernte sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (♀, 30 μm); c – head lateral (♀, 30 μm); d – pronotum dorsal (♀, 100 μm); e – hemelytra (J, 100 μm); f – hemelytra (♀, 100 μm); g – pygophore dorsal (J, 30 μm); h – abdomen ventral (♀, 30 μm). Scales in brackets, measured in μm. FS, frontal spine; MS, medial spine; OS, occipital spine; Bu, bucculae; CaR, callar region; Co, collum; Pa, paranota; MC, median carina; LC, lateral carina; PL, posterior lobe; R+M, radius and media vein; Cu, cubitus vein; CoA, costal area; ScA, subcostal area; SuA, sutural area; DiA, discoidal area; RP, right paramere; LP, left paramere; Py, Pygophore; SgP, subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.5341513" httpUri="https://zenodo.org/record/5341513/files/figure.png" pageId="5" pageNumber="438">Fig. 4b</figureCitation>
|
||
). Labium: usually extending to mesocoxae; sometimes extending onto abdomen. Antennae: elongate; AI cylindrical, short, of greater diameter than other segments; AII equal to or at least 2/3 length of AI; AIII elongate; AIV short but longer than AI+AII, clavate, base sometimes elongate. Pronotum: (e.g.,
|
||
<figureCitation id="13512A29DC30FF90BD70FC8E6B21FCBA" box="[806,895,796,820]" captionStart="Fig" captionStartId="20.[151,182,1488,1507]" captionTargetBox="[198,1079,219,1449]" captionTargetId="figure-10@20.[198,1079,219,1449]" captionTargetPageId="20" captionText="Fig. 4. Inoma arrernte sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (♀, 30 μm); c – head lateral (♀, 30 μm); d – pronotum dorsal (♀, 100 μm); e – hemelytra (J, 100 μm); f – hemelytra (♀, 100 μm); g – pygophore dorsal (J, 30 μm); h – abdomen ventral (♀, 30 μm). Scales in brackets, measured in μm. FS, frontal spine; MS, medial spine; OS, occipital spine; Bu, bucculae; CaR, callar region; Co, collum; Pa, paranota; MC, median carina; LC, lateral carina; PL, posterior lobe; R+M, radius and media vein; Cu, cubitus vein; CoA, costal area; ScA, subcostal area; SuA, sutural area; DiA, discoidal area; RP, right paramere; LP, left paramere; Py, Pygophore; SgP, subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.5341513" httpUri="https://zenodo.org/record/5341513/files/figure.png" pageId="5" pageNumber="438">Figs. 4d</figureCitation>
|
||
,
|
||
<figureCitation id="13512A29DC30FF90BDDCFC896BFBFCBD" box="[906,933,795,819]" captionStart="Fig" captionStartId="21.[113,144,1546,1565]" captionTargetBox="[141,1061,217,1503]" captionTargetId="figure-13@21.[141,1061,217,1504]" captionTargetPageId="21" captionText="Fig. 5. Inoma fuscata sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (♀, 30 μm); c – head lateral (♀, 30 μm); d – pronotum dorsal (♀, 100 μm); e – hemelytra (♀, 100 μm); f – tarsal claw (♀, 3 μm); g – pygophore dorsal (J, 30 μm); h – abdomen ventral (♀, 100 μm). Scales in brackets, measured in μm." figureDoi="http://doi.org/10.5281/zenodo.5341515" httpUri="https://zenodo.org/record/5341515/files/figure.png" pageId="5" pageNumber="438">5d</figureCitation>
|
||
) broad, lateral margins rounded, callar region flattened or convex (lateral views in
|
||
<figureCitation id="13512A29DC30FF90BD60FCAE6B23FCDA" box="[822,893,828,852]" captionStart="Fig" captionStartId="10.[151,182,1545,1564]" captionTargetBox="[178,1096,215,1509]" captionTargetId="figure-10@10.[177,1102,215,1512]" captionTargetPageId="10" captionText="Fig. 1. Habitus photographs of Inoma species: I. arrernte sp. nov. (macropters and brachypters), I. breviseta sp. nov., I. fuscata sp. nov. (also lateral view), I. innamincka sp. nov. and I. kalbarri sp. nov. (macropters and female brachypter). Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341507" httpUri="https://zenodo.org/record/5341507/files/figure.png" pageId="5" pageNumber="438">Figs. 1</figureCitation>
|
||
&
|
||
<figureCitation id="13512A29DC30FF90BDCBFCAE6BF2FCDA" box="[925,940,828,852]" captionStart="Fig" captionStartId="11.[113,144,1518,1537]" captionTargetBox="[118,1089,218,1490]" captionTargetId="figure-13@11.[116,1090,215,1490]" captionTargetPageId="11" captionText="Fig. 2. Habitus photographs of Inoma species:I.multispinosa Hacker, 1927 (macropters and brachypters), I. silveirae sp. nov. (macropters and brachypters, also in lateral view), I. solusa sp. nov. and I. stysi sp. nov. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341509" httpUri="https://zenodo.org/record/5341509/files/figure.png" pageId="5" pageNumber="438">2</figureCitation>
|
||
), closely punctate, posterior lobe subequal to anterior portion, pointed and reticulated; collum weakly to moderately elevated and sub-triangular in shape, strongly keeled medially, truncate anteriorly, covering half of head; pronotum tricarinate, carinae elevated, but not highly so, uniseriate, areolae subquadrate and small or large, straight over most of length but lateral carinae slightly convergent at posterior end and terminating anteriorly at calli, median carina percurrent to collum, lateral carinae of equal height to median carina; paranota narrow and linear, obliquely extended or upright, uniseriate or biseriate, areolae subquadrate and small or large. Thoracic sterna: prosternal carinae barely visible, meso and metasternal carinae strongly elevated, parallel (or very slightly rounded) and of equal width. Metathoracic gland: orifice obsolete. Legs: slender, elongate, tibiae longer than femora. Hemelytra: macropters (e.g.,
|
||
<figureCitation id="13512A29DC30FF90BD92FBEE6C51FB1D" box="[964,1039,1147,1172]" captionStart="Fig" captionStartId="21.[113,144,1546,1565]" captionTargetBox="[141,1061,217,1503]" captionTargetId="figure-13@21.[141,1061,217,1504]" captionTargetPageId="21" captionText="Fig. 5. Inoma fuscata sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (♀, 30 μm); c – head lateral (♀, 30 μm); d – pronotum dorsal (♀, 100 μm); e – hemelytra (♀, 100 μm); f – tarsal claw (♀, 3 μm); g – pygophore dorsal (J, 30 μm); h – abdomen ventral (♀, 100 μm). Scales in brackets, measured in μm." figureDoi="http://doi.org/10.5281/zenodo.5341515" httpUri="https://zenodo.org/record/5341515/files/figure.png" pageId="5" pageNumber="438">Fig. 5e</figureCitation>
|
||
,
|
||
<figureCitation id="13512A29DC30FF90BA4DFBEE6C6AFB1A" box="[1051,1076,1148,1172]" captionStart="Fig" captionStartId="24.[151,182,1497,1516]" captionTargetBox="[195,1083,219,1457]" captionTargetId="figure-10@24.[194,1084,219,1458]" captionTargetPageId="24" captionText="Fig. 8. Inoma silveirae sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (♀, 30 μm); c – head lateral (♀, 20 μm); d – pronotum dorsal (♀, 100 μm); e – macropterous hemelytra (J, 100 μm); f – brachypterous hemelytra (♀, 100 μm); g – pygophore dorsal (J, 30 μm); h – genital segment of abdomen ventral (♀, 100 μm). Scales in brackets, measured in μm." figureDoi="http://doi.org/10.5281/zenodo.5341521" httpUri="https://zenodo.org/record/5341521/files/figure.png" pageId="5" pageNumber="438">8f</figureCitation>
|
||
), brachypters (e.g.,
|
||
<figureCitation id="13512A29DC30FF90BF63FB0E692BFB3A" box="[309,373,1180,1204]" captionStart="Fig" captionStartId="20.[151,182,1488,1507]" captionTargetBox="[198,1079,219,1449]" captionTargetId="figure-10@20.[198,1079,219,1449]" captionTargetPageId="20" captionText="Fig. 4. Inoma arrernte sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (♀, 30 μm); c – head lateral (♀, 30 μm); d – pronotum dorsal (♀, 100 μm); e – hemelytra (J, 100 μm); f – hemelytra (♀, 100 μm); g – pygophore dorsal (J, 30 μm); h – abdomen ventral (♀, 30 μm). Scales in brackets, measured in μm. FS, frontal spine; MS, medial spine; OS, occipital spine; Bu, bucculae; CaR, callar region; Co, collum; Pa, paranota; MC, median carina; LC, lateral carina; PL, posterior lobe; R+M, radius and media vein; Cu, cubitus vein; CoA, costal area; ScA, subcostal area; SuA, sutural area; DiA, discoidal area; RP, right paramere; LP, left paramere; Py, Pygophore; SgP, subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.5341513" httpUri="https://zenodo.org/record/5341513/files/figure.png" pageId="5" pageNumber="438">Fig. 4</figureCitation>
|
||
e-f, 8e); cubitus and R+M veins carinate; costal area uniseriate or biseriate, areolae large and subquadrate; subcostal area wide, at least half width of discoidal area, extending to just before forewing apex; discoidal area large, lanceolate in shape, junction of cubitus and R+M veins medially located on hemelytra; sutural area small, areole size generally large, areolae small in brachypters; hypocosta as wide as costal area or narrow. Male genitalia: pygophore subquadrate, ventral margin of opening expanded posteriorly with a concave or sinuous margin (e.g.,
|
||
<figureCitation id="13512A29DC30FF90BFB5FACE6A6CFAFA" box="[483,562,1372,1396]" captionStart="Fig" captionStartId="20.[151,182,1488,1507]" captionTargetBox="[198,1079,219,1449]" captionTargetId="figure-10@20.[198,1079,219,1449]" captionTargetPageId="20" captionText="Fig. 4. Inoma arrernte sp. nov. a – head dorsal (♀, 30 μm); b – head ventral (♀, 30 μm); c – head lateral (♀, 30 μm); d – pronotum dorsal (♀, 100 μm); e – hemelytra (J, 100 μm); f – hemelytra (♀, 100 μm); g – pygophore dorsal (J, 30 μm); h – abdomen ventral (♀, 30 μm). Scales in brackets, measured in μm. FS, frontal spine; MS, medial spine; OS, occipital spine; Bu, bucculae; CaR, callar region; Co, collum; Pa, paranota; MC, median carina; LC, lateral carina; PL, posterior lobe; R+M, radius and media vein; Cu, cubitus vein; CoA, costal area; ScA, subcostal area; SuA, sutural area; DiA, discoidal area; RP, right paramere; LP, left paramere; Py, Pygophore; SgP, subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.5341513" httpUri="https://zenodo.org/record/5341513/files/figure.png" pageId="5" pageNumber="438">
|
||
Fig.
|
||
<quantity id="4C929B49DC30FF90BC42FACE6A6CFAFA" box="[532,562,1372,1396]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="5" pageNumber="438" unit="g" value="4.0">4g</quantity>
|
||
</figureCitation>
|
||
); parameres simple, sickle-shaped (e.g.,
|
||
<figureCitation id="13512A29DC30FF90BDB7FACE6C75FAFA" box="[993,1067,1372,1396]" captionStart="Fig" captionStartId="28.[151,181,1484,1503]" captionTargetBox="[195,1082,220,1456]" captionTargetId="figure-10@28.[193,1084,219,1456]" captionTargetPageId="28" captionText="Fig.10. Male genitalia of Inoma species. a-c, m – I.multispinosa Hacker,1927; d-f,k – I. fuscata sp. nov.; g – I. kalbarri sp. nov.; h-i – I. silveirae sp. nov.; j – I. arrernte sp. nov.; l – I. innamincka sp. nov. a, d, h – right paramere, dorsal view; b, e, i – left paramere, dorsal view; c, f – aedeagus, dorsal view; g – aedeagus, ventral view; j-m – paired basal endosomal sclerites. Scale = 0.1 mm. Ap, apophysis; SL, sensory lobe; PBES, paired basal endosomal sclerites; DP, dorsal plate; DUES, dorsal U-shaped endosomal sclerite; dm, distal margin; bb, basal branch; cl, cleft; MPP, medial portion of phallotheca." figureDoi="http://doi.org/10.5281/zenodo.5341525" httpUri="https://zenodo.org/record/5341525/files/figure.png" pageId="5" pageNumber="438">Fig. 10</figureCitation>
|
||
ab, d-e, h-i), with short bristle-like setae on entire inner margin and sometimes also on outer margin of apophysis, mostly with a few more elongate setae on sensory lobe, sometimes also with minute setae on dorsal surface (
|
||
<figureCitation id="13512A29DC30FF90BC54FA2E6A0EFA5A" box="[514,592,1468,1492]" captionStart="Fig" captionStartId="28.[151,181,1484,1503]" captionTargetBox="[195,1082,220,1456]" captionTargetId="figure-10@28.[193,1084,219,1456]" captionTargetPageId="28" captionText="Fig.10. Male genitalia of Inoma species. a-c, m – I.multispinosa Hacker,1927; d-f,k – I. fuscata sp. nov.; g – I. kalbarri sp. nov.; h-i – I. silveirae sp. nov.; j – I. arrernte sp. nov.; l – I. innamincka sp. nov. a, d, h – right paramere, dorsal view; b, e, i – left paramere, dorsal view; c, f – aedeagus, dorsal view; g – aedeagus, ventral view; j-m – paired basal endosomal sclerites. Scale = 0.1 mm. Ap, apophysis; SL, sensory lobe; PBES, paired basal endosomal sclerites; DP, dorsal plate; DUES, dorsal U-shaped endosomal sclerite; dm, distal margin; bb, basal branch; cl, cleft; MPP, medial portion of phallotheca." figureDoi="http://doi.org/10.5281/zenodo.5341525" httpUri="https://zenodo.org/record/5341525/files/figure.png" pageId="5" pageNumber="438">Fig. 10</figureCitation>
|
||
h-i), sensory lobe rounded (
|
||
<figureCitation id="13512A29DC30FF90BD2BFA2E6B94FA5A" box="[893,970,1468,1492]" captionStart="Fig" captionStartId="28.[151,181,1484,1503]" captionTargetBox="[195,1082,220,1456]" captionTargetId="figure-10@28.[193,1084,219,1456]" captionTargetPageId="28" captionText="Fig.10. Male genitalia of Inoma species. a-c, m – I.multispinosa Hacker,1927; d-f,k – I. fuscata sp. nov.; g – I. kalbarri sp. nov.; h-i – I. silveirae sp. nov.; j – I. arrernte sp. nov.; l – I. innamincka sp. nov. a, d, h – right paramere, dorsal view; b, e, i – left paramere, dorsal view; c, f – aedeagus, dorsal view; g – aedeagus, ventral view; j-m – paired basal endosomal sclerites. Scale = 0.1 mm. Ap, apophysis; SL, sensory lobe; PBES, paired basal endosomal sclerites; DP, dorsal plate; DUES, dorsal U-shaped endosomal sclerite; dm, distal margin; bb, basal branch; cl, cleft; MPP, medial portion of phallotheca." figureDoi="http://doi.org/10.5281/zenodo.5341525" httpUri="https://zenodo.org/record/5341525/files/figure.png" pageId="5" pageNumber="438">Fig. 10</figureCitation>
|
||
d-e, h-i) or sometimes slightly angular (
|
||
<figureCitation id="13512A29DC30FF90BFF4FA4E69B1FA7A" box="[418,495,1500,1524]" captionStart="Fig" captionStartId="28.[151,181,1484,1503]" captionTargetBox="[195,1082,220,1456]" captionTargetId="figure-10@28.[193,1084,219,1456]" captionTargetPageId="28" captionText="Fig.10. Male genitalia of Inoma species. a-c, m – I.multispinosa Hacker,1927; d-f,k – I. fuscata sp. nov.; g – I. kalbarri sp. nov.; h-i – I. silveirae sp. nov.; j – I. arrernte sp. nov.; l – I. innamincka sp. nov. a, d, h – right paramere, dorsal view; b, e, i – left paramere, dorsal view; c, f – aedeagus, dorsal view; g – aedeagus, ventral view; j-m – paired basal endosomal sclerites. Scale = 0.1 mm. Ap, apophysis; SL, sensory lobe; PBES, paired basal endosomal sclerites; DP, dorsal plate; DUES, dorsal U-shaped endosomal sclerite; dm, distal margin; bb, basal branch; cl, cleft; MPP, medial portion of phallotheca." figureDoi="http://doi.org/10.5281/zenodo.5341525" httpUri="https://zenodo.org/record/5341525/files/figure.png" pageId="5" pageNumber="438">Fig. 10</figureCitation>
|
||
a-b), apophysis elongate and acuminate with a rounded apex, tubular in cross section with rounded margins; aedeagus with a large, curved, inverted U-shaped sclerite in apical tubular portion of endosoma (
|
||
<figureCitation id="13512A29DC30FF90BC8AF98E6B02F9BA" box="[732,860,1564,1588]" captionStart="Fig" captionStartId="28.[151,181,1484,1503]" captionTargetBox="[195,1082,220,1456]" captionTargetId="figure-10@28.[193,1084,219,1456]" captionTargetPageId="28" captionText="Fig.10. Male genitalia of Inoma species. a-c, m – I.multispinosa Hacker,1927; d-f,k – I. fuscata sp. nov.; g – I. kalbarri sp. nov.; h-i – I. silveirae sp. nov.; j – I. arrernte sp. nov.; l – I. innamincka sp. nov. a, d, h – right paramere, dorsal view; b, e, i – left paramere, dorsal view; c, f – aedeagus, dorsal view; g – aedeagus, ventral view; j-m – paired basal endosomal sclerites. Scale = 0.1 mm. Ap, apophysis; SL, sensory lobe; PBES, paired basal endosomal sclerites; DP, dorsal plate; DUES, dorsal U-shaped endosomal sclerite; dm, distal margin; bb, basal branch; cl, cleft; MPP, medial portion of phallotheca." figureDoi="http://doi.org/10.5281/zenodo.5341525" httpUri="https://zenodo.org/record/5341525/files/figure.png" pageId="5" pageNumber="438">Fig. 10c,f,g</figureCitation>
|
||
), heavily sclerotised, some subtle variation in size and shape of cleft and length of the basal branches of sclerite; sometimes also with small, paired endosomal sclerites positioned basally near dorsal plate (e.g.,
|
||
<figureCitation id="13512A29DC33FF93BE83FE896961FEBD" box="[213,319,283,307]" captionStart="Fig" captionStartId="28.[151,181,1484,1503]" captionTargetBox="[195,1082,220,1456]" captionTargetId="figure-10@28.[193,1084,219,1456]" captionTargetPageId="28" captionText="Fig.10. Male genitalia of Inoma species. a-c, m – I.multispinosa Hacker,1927; d-f,k – I. fuscata sp. nov.; g – I. kalbarri sp. nov.; h-i – I. silveirae sp. nov.; j – I. arrernte sp. nov.; l – I. innamincka sp. nov. a, d, h – right paramere, dorsal view; b, e, i – left paramere, dorsal view; c, f – aedeagus, dorsal view; g – aedeagus, ventral view; j-m – paired basal endosomal sclerites. Scale = 0.1 mm. Ap, apophysis; SL, sensory lobe; PBES, paired basal endosomal sclerites; DP, dorsal plate; DUES, dorsal U-shaped endosomal sclerite; dm, distal margin; bb, basal branch; cl, cleft; MPP, medial portion of phallotheca." figureDoi="http://doi.org/10.5281/zenodo.5341525" httpUri="https://zenodo.org/record/5341525/files/figure.png" pageId="6" pageNumber="439">Fig. 10c,f</figureCitation>
|
||
), small with shape varying from elongate-ovate (
|
||
<figureCitation id="13512A29DC33FF93BD06FE896BE3FEBD" box="[848,957,283,307]" captionStart="Fig" captionStartId="28.[151,181,1484,1503]" captionTargetBox="[195,1082,220,1456]" captionTargetId="figure-10@28.[193,1084,219,1456]" captionTargetPageId="28" captionText="Fig.10. Male genitalia of Inoma species. a-c, m – I.multispinosa Hacker,1927; d-f,k – I. fuscata sp. nov.; g – I. kalbarri sp. nov.; h-i – I. silveirae sp. nov.; j – I. arrernte sp. nov.; l – I. innamincka sp. nov. a, d, h – right paramere, dorsal view; b, e, i – left paramere, dorsal view; c, f – aedeagus, dorsal view; g – aedeagus, ventral view; j-m – paired basal endosomal sclerites. Scale = 0.1 mm. Ap, apophysis; SL, sensory lobe; PBES, paired basal endosomal sclerites; DP, dorsal plate; DUES, dorsal U-shaped endosomal sclerite; dm, distal margin; bb, basal branch; cl, cleft; MPP, medial portion of phallotheca." figureDoi="http://doi.org/10.5281/zenodo.5341525" httpUri="https://zenodo.org/record/5341525/files/figure.png" pageId="6" pageNumber="439">Fig. 10f,k</figureCitation>
|
||
), sub-triangular (
|
||
<figureCitation id="13512A29DC33FF93BEC9FEA96946FEDA" box="[159,280,315,340]" captionStart="Fig" captionStartId="28.[151,181,1484,1503]" captionTargetBox="[195,1082,220,1456]" captionTargetId="figure-10@28.[193,1084,219,1456]" captionTargetPageId="28" captionText="Fig.10. Male genitalia of Inoma species. a-c, m – I.multispinosa Hacker,1927; d-f,k – I. fuscata sp. nov.; g – I. kalbarri sp. nov.; h-i – I. silveirae sp. nov.; j – I. arrernte sp. nov.; l – I. innamincka sp. nov. a, d, h – right paramere, dorsal view; b, e, i – left paramere, dorsal view; c, f – aedeagus, dorsal view; g – aedeagus, ventral view; j-m – paired basal endosomal sclerites. Scale = 0.1 mm. Ap, apophysis; SL, sensory lobe; PBES, paired basal endosomal sclerites; DP, dorsal plate; DUES, dorsal U-shaped endosomal sclerite; dm, distal margin; bb, basal branch; cl, cleft; MPP, medial portion of phallotheca." figureDoi="http://doi.org/10.5281/zenodo.5341525" httpUri="https://zenodo.org/record/5341525/files/figure.png" pageId="6" pageNumber="439">Fig. 10c,m</figureCitation>
|
||
) and semi-circular (
|
||
<figureCitation id="13512A29DC33FF93BFADFEA96A3FFEDD" box="[507,609,315,339]" captionStart="Fig" captionStartId="28.[151,181,1484,1503]" captionTargetBox="[195,1082,220,1456]" captionTargetId="figure-10@28.[193,1084,219,1456]" captionTargetPageId="28" captionText="Fig.10. Male genitalia of Inoma species. a-c, m – I.multispinosa Hacker,1927; d-f,k – I. fuscata sp. nov.; g – I. kalbarri sp. nov.; h-i – I. silveirae sp. nov.; j – I. arrernte sp. nov.; l – I. innamincka sp. nov. a, d, h – right paramere, dorsal view; b, e, i – left paramere, dorsal view; c, f – aedeagus, dorsal view; g – aedeagus, ventral view; j-m – paired basal endosomal sclerites. Scale = 0.1 mm. Ap, apophysis; SL, sensory lobe; PBES, paired basal endosomal sclerites; DP, dorsal plate; DUES, dorsal U-shaped endosomal sclerite; dm, distal margin; bb, basal branch; cl, cleft; MPP, medial portion of phallotheca." figureDoi="http://doi.org/10.5281/zenodo.5341525" httpUri="https://zenodo.org/record/5341525/files/figure.png" pageId="6" pageNumber="439">Fig. 10j,l</figureCitation>
|
||
); medial portion of phallotheca mostly entire dorsally, rarely divided (
|
||
<figureCitation id="13512A29DC33FF93BFC8FEC969A3FEFA" box="[414,509,347,372]" captionStart="Fig" captionStartId="28.[151,181,1484,1503]" captionTargetBox="[195,1082,220,1456]" captionTargetId="figure-10@28.[193,1084,219,1456]" captionTargetPageId="28" captionText="Fig.10. Male genitalia of Inoma species. a-c, m – I.multispinosa Hacker,1927; d-f,k – I. fuscata sp. nov.; g – I. kalbarri sp. nov.; h-i – I. silveirae sp. nov.; j – I. arrernte sp. nov.; l – I. innamincka sp. nov. a, d, h – right paramere, dorsal view; b, e, i – left paramere, dorsal view; c, f – aedeagus, dorsal view; g – aedeagus, ventral view; j-m – paired basal endosomal sclerites. Scale = 0.1 mm. Ap, apophysis; SL, sensory lobe; PBES, paired basal endosomal sclerites; DP, dorsal plate; DUES, dorsal U-shaped endosomal sclerite; dm, distal margin; bb, basal branch; cl, cleft; MPP, medial portion of phallotheca." figureDoi="http://doi.org/10.5281/zenodo.5341525" httpUri="https://zenodo.org/record/5341525/files/figure.png" pageId="6" pageNumber="439">
|
||
Fig.
|
||
<quantity id="4C929B49DC33FF93BF87FECE69A3FEFA" box="[465,509,348,372]" metricMagnitude="-2" metricUnit="kg" metricValue="1.0" pageId="6" pageNumber="439" unit="g" value="10.0">10g</quantity>
|
||
</figureCitation>
|
||
); dorsal plate of aedeagus simple, broadly U-shaped with rounded distal margin (e.g.,
|
||
<figureCitation id="13512A29DC33FF93BF92FEE96A71FE1D" box="[452,559,379,403]" captionStart="Fig" captionStartId="28.[151,181,1484,1503]" captionTargetBox="[195,1082,220,1456]" captionTargetId="figure-10@28.[193,1084,219,1456]" captionTargetPageId="28" captionText="Fig.10. Male genitalia of Inoma species. a-c, m – I.multispinosa Hacker,1927; d-f,k – I. fuscata sp. nov.; g – I. kalbarri sp. nov.; h-i – I. silveirae sp. nov.; j – I. arrernte sp. nov.; l – I. innamincka sp. nov. a, d, h – right paramere, dorsal view; b, e, i – left paramere, dorsal view; c, f – aedeagus, dorsal view; g – aedeagus, ventral view; j-m – paired basal endosomal sclerites. Scale = 0.1 mm. Ap, apophysis; SL, sensory lobe; PBES, paired basal endosomal sclerites; DP, dorsal plate; DUES, dorsal U-shaped endosomal sclerite; dm, distal margin; bb, basal branch; cl, cleft; MPP, medial portion of phallotheca." figureDoi="http://doi.org/10.5281/zenodo.5341525" httpUri="https://zenodo.org/record/5341525/files/figure.png" pageId="6" pageNumber="439">Fig. 10c,f</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD536ACDC33FF93BEE3FE096C10FDBA" blockId="6.[151,1127,219,564]" pageId="6" pageNumber="439">Females: minor sexual dimorphism; differing from males as follows: eyes smaller; AIII less pilose with only elongate minor setiferous tubercles, lacking short simple setae; hemelytra slightly shorter and broader (both brachypters and macropters); sometimes slight colour variation. Female genitalia: membraneous, with paired pseudospermathecae, basally derived at junction of lateral oviducts, each with short duct and distal capitate seminal sac (Fig. 11).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3706527DC33FF93BEC1FDD86C45FD0D" pageId="6" pageNumber="439" type="biology_ecology">
|
||
<paragraph id="8BD536ACDC33FF93BEC1FDD86C45FD0D" blockId="6.[151,1126,586,643]" pageId="6" pageNumber="439">
|
||
<emphasis id="B91EEABEDC33FF93BEC1FDD86A6BFDEC" bold="true" box="[151,565,586,611]" pageId="6" pageNumber="439">Biology and host plant relationships.</emphasis>
|
||
<taxonomicName id="4C6A4D2FDC33FF93BC6CFDDE6A20FDED" authorityName=": DRAKE & RUHOFF" authorityYear="1960" box="[570,638,588,611]" class="Insecta" family="Tingidae" genus="Inoma" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91EEABEDC33FF93BC6CFDDE6A20FDED" box="[570,638,588,611]" italics="true" pageId="6" pageNumber="439">Inoma</emphasis>
|
||
</taxonomicName>
|
||
is found mostly on plant species of the angiosperm order
|
||
<taxonomicName id="4C6A4D2FDC33FF93BF4BFDF969DDFD0D" box="[285,387,619,643]" class="Magnoliopsida" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="439" phylum="Tracheophyta" rank="order">Asterales</taxonomicName>
|
||
, often belonging to the families
|
||
<taxonomicName id="4C6A4D2FDC33FF93BC8DFDF96B32FD0D" authorityName="R.Brown" authorityYear="1810" box="[731,876,619,643]" class="Magnoliopsida" family="Myoporaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="439" phylum="Tracheophyta" rank="family">Myoporaceae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6A4D2FDC33FF93BDF6FDF96C49FD0D" box="[928,1047,619,643]" class="Magnoliopsida" family="Lamiaceae" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="439" phylum="Tracheophyta" rank="family">Lamiaceae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3706527DC33FF93BEC1FD0B6C1DFD5F" pageId="6" pageNumber="439" type="distribution">
|
||
<paragraph id="8BD536ACDC33FF93BEC1FD0B6C1DFD5F" blockId="6.[151,1127,665,722]" pageId="6" pageNumber="439">
|
||
<emphasis id="B91EEABEDC33FF93BEC1FD0B6979FD3F" bold="true" box="[151,295,665,689]" pageId="6" pageNumber="439">Distribution.</emphasis>
|
||
<taxonomicName id="4C6A4D2FDC33FF93BF7AFD096931FD3C" authorityName=": DRAKE & RUHOFF" authorityYear="1960" box="[300,367,667,690]" class="Insecta" family="Tingidae" genus="Inoma" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91EEABEDC33FF93BF7AFD096931FD3C" box="[300,367,667,690]" italics="true" pageId="6" pageNumber="439">Inoma</emphasis>
|
||
</taxonomicName>
|
||
is comprised of nine species, and is largely found in arid
|
||
<collectingCountry id="F37D763CDC33FF93BDE0FD086C47FD3C" box="[950,1049,666,690]" name="Australia" pageId="6" pageNumber="439">Australia</collectingCountry>
|
||
, except for
|
||
<taxonomicName id="4C6A4D2FDC33FF93BEEBFD296905FD5C" authorityName="Hacker" authorityYear="1927" box="[189,347,699,722]" class="Insecta" family="Tingidae" genus="Inoma" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="439" phylum="Arthropoda" rank="species" species="multispinosa">
|
||
<emphasis id="B91EEABEDC33FF93BEEBFD296905FD5C" box="[189,347,699,722]" italics="true" pageId="6" pageNumber="439">I. multispinosa</emphasis>
|
||
</taxonomicName>
|
||
which is known from temperate
|
||
<collectingCountry id="F37D763CDC33FF93BCEDFD286B7EFD5C" box="[699,800,698,722]" name="Australia" pageId="6" pageNumber="439">Australia</collectingCountry>
|
||
, in the southeastern corner.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3706527DC33FF92BEC1FD7B6AF6FE3A" lastPageId="7" lastPageNumber="440" pageId="6" pageNumber="439" type="discussion">
|
||
<paragraph id="8BD536ACDC33FF93BEC1FD7B6A29FBAF" blockId="6.[151,1127,745,1601]" pageId="6" pageNumber="439">
|
||
<emphasis id="B91EEABEDC33FF93BEC1FD7B6958FC8F" bold="true" box="[151,262,745,769]" pageId="6" pageNumber="439">Remarks.</emphasis>
|
||
<taxonomicName id="4C6A4D2FDC33FF93BF5BFD78690FFC8F" authorityName=": DRAKE & RUHOFF" authorityYear="1960" box="[269,337,746,769]" class="Insecta" family="Tingidae" genus="Inoma" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91EEABEDC33FF93BF5BFD78690FFC8F" box="[269,337,746,769]" italics="true" pageId="6" pageNumber="439">Inoma</emphasis>
|
||
</taxonomicName>
|
||
has been redefined, with a detailed definition of characters, for inclusion of new species described herein. Characters which are now more broadly defined and show infrageneric variation include: 1) condition of setiferous tubercles (formerly referred to as spines by
|
||
<bibRefCitation id="EFFB4B5DDC33FF93BF57FCDB69F2FCEF" author="HACKER H." box="[257,428,841,865]" pageId="6" pageNumber="439" pagination="19 - 32" refId="ref28582" refString="HACKER H. 1927: New Tingitoidea (Hemiptera) in the Queensland Museum. Memoirs of Queensland Museum 9: 19 - 32 + pls. vi-x." type="journal article" year="1927">HACKER (1927))</bibRefCitation>
|
||
; 2) setal characters of pilose species; 3) paranota and costal areas, now either biseriate or uniseriate; 4) body colouration, including variation in patterning; 5) orientation of frontal cephalic spines, which may be weakly divergent, parallel or slightly converging; 6) elevation of collum, which although consistently keeled is not always sharply elevated, as indicated in the original description, with some species having only a weakly elevated collum; and, 7) elevation of pronotal callar region, which is not always convex, but sometimes almost flat or very slightly raised.
|
||
</paragraph>
|
||
<paragraph id="8BD536ACDC33FF93BEE3FBB86A5FFA8E" blockId="6.[151,1127,745,1601]" pageId="6" pageNumber="439">
|
||
<taxonomicName id="4C6A4D2FDC33FF93BEE3FBB868A7FBCF" authorityName=": DRAKE & RUHOFF" authorityYear="1960" box="[181,249,1066,1089]" class="Insecta" family="Tingidae" genus="Inoma" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91EEABEDC33FF93BEE3FBB868A7FBCF" box="[181,249,1066,1089]" italics="true" pageId="6" pageNumber="439">Inoma</emphasis>
|
||
</taxonomicName>
|
||
is aligned with the speciose genus
|
||
<taxonomicName id="4C6A4D2FDC33FF93BCD3FBB86A98FBCF" authorityName="Fabricius" authorityYear="1803" box="[645,710,1066,1089]" class="Insecta" family="Tingidae" genus="Tingis" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91EEABEDC33FF93BCD3FBB86A98FBCF" box="[645,710,1066,1089]" italics="true" pageId="6" pageNumber="439">Tingis</emphasis>
|
||
</taxonomicName>
|
||
, and putatively closely related to and possibly synonymous to
|
||
<taxonomicName id="4C6A4D2FDC33FF93BFC1FBD86A47FBEF" authorityName="Stal" authorityYear="1873" box="[407,537,1098,1121]" class="Insecta" family="Tingidae" genus="Lasiacantha" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91EEABEDC33FF93BFC1FBD86A47FBEF" box="[407,537,1098,1121]" italics="true" pageId="6" pageNumber="439">Lasiacantha</emphasis>
|
||
</taxonomicName>
|
||
, the latter a diverse taxon, with species in Europe, Africa,
|
||
<collectingCountry id="F37D763CDC33FF93BEC1FBFB6891FB0F" box="[151,207,1129,1153]" name="India" pageId="6" pageNumber="439">India</collectingCountry>
|
||
, southeast Asia and
|
||
<collectingCountry id="F37D763CDC33FF93BFCEFBFB69A5FB0F" box="[408,507,1129,1153]" name="Australia" pageId="6" pageNumber="439">Australia</collectingCountry>
|
||
. Morphological similarities between
|
||
<taxonomicName id="4C6A4D2FDC33FF93BD22FBF86BE9FB0F" authorityName=": DRAKE & RUHOFF" authorityYear="1960" box="[884,951,1130,1153]" class="Insecta" family="Tingidae" genus="Inoma" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91EEABEDC33FF93BD22FBF86BE9FB0F" box="[884,951,1130,1153]" italics="true" pageId="6" pageNumber="439">Inoma</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6A4D2FDC33FF93BDB0FBF86C39FB0F" authorityName="Stal" authorityYear="1873" box="[998,1127,1130,1153]" class="Insecta" family="Tingidae" genus="Lasiacantha" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91EEABEDC33FF93BDB0FBF86C39FB0F" box="[998,1127,1130,1153]" italics="true" pageId="6" pageNumber="439">Lasiacantha</emphasis>
|
||
</taxonomicName>
|
||
include: 1) shape and form of hemelytra; 2) elongate cephalic spines with laterally arcuate occipital spines; 3) sometimes bifurcate medial spine; 4) outline of head and bucculae; 6) mostly elevated pronotal collum; 7) marginal setiferous tubercles; 8) often dense vestiture; and, 9) slender legs and antennae.
|
||
</paragraph>
|
||
<paragraph id="8BD536ACDC33FF93BEE3FA9B6B8DFA6E" blockId="6.[151,1127,745,1601]" pageId="6" pageNumber="439">
|
||
Nonetheless, we have found a number of characters of
|
||
<taxonomicName id="4C6A4D2FDC33FF93BD5CFA986B10FAAF" authorityName=": DRAKE & RUHOFF" authorityYear="1960" box="[778,846,1290,1313]" class="Insecta" family="Tingidae" genus="Inoma" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91EEABEDC33FF93BD5CFA986B10FAAF" box="[778,846,1290,1313]" italics="true" pageId="6" pageNumber="439">Inoma</emphasis>
|
||
</taxonomicName>
|
||
which distinguish it from
|
||
<taxonomicName id="4C6A4D2FDC33FF93BEC1FAB86945FACF" authorityName="Stal" authorityYear="1873" box="[151,283,1322,1345]" class="Insecta" family="Tingidae" genus="Lasiacantha" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91EEABEDC33FF93BEC1FAB86945FACF" box="[151,283,1322,1345]" italics="true" pageId="6" pageNumber="439">Lasiacantha</emphasis>
|
||
</taxonomicName>
|
||
, with the most critical being: 1) strongly keeled pronotal collum, subtriangular in shape, more moderately elevated; 2) setiferous tubercles along medial keel and anterior margin of the collum; 3) pronotal carinae distinctly elevated, but not greatly enlarged, 4) medial pronotal carina always same height as lateral carinae; 5) linear, narrow paranota, equal width throughout and at most two areolae wide; and, 6) costal area at most two areolae rows wide; and, 7) lack of very fine, hook-like setae on dorsum, as found in
|
||
<taxonomicName id="4C6A4D2FDC33FF93BD1EFA586B92FA6F" authorityName="Stal" authorityYear="1873" box="[840,972,1482,1505]" class="Insecta" family="Tingidae" genus="Lasiacantha" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91EEABEDC33FF93BD1EFA586B92FA6F" box="[840,972,1482,1505]" italics="true" pageId="6" pageNumber="439">Lasiacantha</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD536ACDC33FF93BEE3FA7B6A1AF9CF" blockId="6.[151,1127,745,1601]" pageId="6" pageNumber="439">
|
||
<bibRefCitation id="EFFB4B5DDC33FF93BEE3FA7B6906F98F" author="HACKER H." box="[181,344,1513,1537]" pageId="6" pageNumber="439" pagination="19 - 32" refId="ref28582" refString="HACKER H. 1927: New Tingitoidea (Hemiptera) in the Queensland Museum. Memoirs of Queensland Museum 9: 19 - 32 + pls. vi-x." type="journal article" year="1927">HACKER (1927)</bibRefCitation>
|
||
noted an affinity of
|
||
<taxonomicName id="4C6A4D2FDC33FF93BC6BFA786ADFF98F" authorityName=": DRAKE & RUHOFF" authorityYear="1960" box="[573,641,1514,1537]" class="Insecta" family="Tingidae" genus="Inoma" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91EEABEDC33FF93BC6BFA786ADFF98F" box="[573,641,1514,1537]" italics="true" pageId="6" pageNumber="439">Inoma</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C6A4D2FDC33FF93BC94FA786BE2F98F" authority="Distant, 1903" authorityName="Distant" authorityYear="1903" box="[706,956,1513,1537]" class="Insecta" family="Tingidae" genus="Urentius" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91EEABEDC33FF93BC94FA786B40F98F" box="[706,798,1514,1537]" italics="true" pageId="6" pageNumber="439">Urentius</emphasis>
|
||
Distant, 1903
|
||
</taxonomicName>
|
||
, but apart from superficial similarities, with the marginal setiferous tubercles and general body shape, this genus is possibly more distantly related.
|
||
</paragraph>
|
||
<paragraph id="8BD536ACDC32FF92BEC6FF4F6AF6FE3A" blockId="7.[113,1089,220,436]" pageId="7" pageNumber="440">
|
||
<taxonomicName id="4C6A4D2FDC32FF92BEC6FF4F69E5FF7A" authority="Drake, 1942" authorityName="Drake" authorityYear="1942" box="[144,443,220,244]" class="Insecta" family="Tingidae" genus="Inoma" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="440" phylum="Arthropoda" rank="species" species="angusta">
|
||
<emphasis id="B91EEABEDC32FF92BEC6FF4F6970FF7A" box="[144,302,221,244]" italics="true" pageId="7" pageNumber="440">Inoma angusta</emphasis>
|
||
Drake, 1942
|
||
</taxonomicName>
|
||
, is considered here as species
|
||
<emphasis id="B91EEABEDC32FF92BCACFF4F6BD3FF7A" box="[762,909,221,244]" italics="true" pageId="7" pageNumber="440">incertae sedis</emphasis>
|
||
, because it lacks the aforementioned diagnostic characters, including the dorsally rounded collum, dorsal crest and setiferous tubercles on anterior margin and dorsal surface. In addition,
|
||
<taxonomicName id="4C6A4D2FDC32FF92BDFEFE8F6C4AFEBA" authorityName="Drake" authorityYear="1942" box="[936,1044,285,308]" class="Insecta" family="Tingidae" genus="Inoma" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="440" phylum="Arthropoda" rank="species" species="angusta">
|
||
<emphasis id="B91EEABEDC32FF92BDFEFE8F6C4AFEBA" box="[936,1044,285,308]" italics="true" pageId="7" pageNumber="440">I. angusta</emphasis>
|
||
</taxonomicName>
|
||
has incomplete and very narrow paranota, and has a rather elongate linear, narrow form rather than being either elongate-ovoid to ovoid. We have refrained from treating this species any further in this work, pending a more thorough examination of genus-group boundaries in Australian lace bugs, but here remove it from
|
||
<emphasis id="B91EEABEDC32FF92BC0AFE0F6AF6FE3A" box="[604,680,413,436]" italics="true" pageId="7" pageNumber="440">
|
||
<taxonomicName id="4C6A4D2FDC32FF92BC0AFE0F6AFAFE3A" authorityName=": DRAKE & RUHOFF" authorityYear="1960" box="[604,676,413,436]" class="Insecta" family="Tingidae" genus="Inoma" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="440" phylum="Arthropoda" rank="genus">Inoma</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |