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<document ID-DOI="10.11646/zootaxa.5270.3.4" ID-GBIF-Dataset="ec485338-4b66-4557-be6c-b53ac23a4678" ID-ISSN="1175-5326" ID-Zenodo-Dep="7860353" ID-ZooBank="D3EE46BD-0755-4730-AF3C-ACD92085311A" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1682061938543" checkinUser="plazi" docAuthor="Valdés, Ángel, Feliciano, Kendall &amp; Malaquias, Manuel A. E." docDate="2023" docId="03C38798FF9746748ED72A2E4393FE3F" docLanguage="en" docName="zootaxa.5270.3.4.pdf" docOrigin="Zootaxa 5270 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5270.3.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pupa pascuana Raines 2003" docType="treatment" docVersion="3" lastPageNumber="498" masterDocId="FFFAFFE0FF8D466F8E402C1C413AFFB6" masterDocTitle="The genus Pupa Röding, 1798 (Mollusca, Gastropoda, Acteonidae) in New Caledonia with notes on Recent species" masterLastPageNumber="506" masterPageNumber="471" pageNumber="497" updateTime="1682367318127" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>The genus Pupa Röding, 1798 (Mollusca, Gastropoda, Acteonidae) in New Caledonia with notes on Recent species</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Valdés, Ángel</mods:namePart>
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<mods:affiliation>Department of Biological Sciences, California State Polytechnic University, 3801 West Temple Avenue, Pomona, California 91768, USA. &amp; aavaldes @ cpp. edu; https: // orcid. org / 0000 - 0002 - 2347 - 4896</mods:affiliation>
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<mods:namePart>Feliciano, Kendall</mods:namePart>
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<mods:affiliation>Department of Biological Sciences, California State Polytechnic University, 3801 West Temple Avenue, Pomona, California 91768, USA. &amp; https: // orcid. org / 0000 - 0003 - 0536 - 2628</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Malaquias, Manuel A. E.</mods:namePart>
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<mods:affiliation>Department of Natural History, University Museum of Bergen, University of Bergen, PB 7800, N- 5020 Bergen, Norway. https: // orcid. org / 0000 - 0002 - 9668 - 945 X</mods:affiliation>
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<paragraph blockId="26.[151,504,1585,1647]" box="[151,504,1585,1612]" pageId="26" pageNumber="497">
<heading bold="true" box="[151,504,1585,1612]" fontSize="11" level="1" pageId="26" pageNumber="497" reason="1">
<taxonomicName ID-CoL="4QJHX" authority="Raines, 2003" authorityName="Raines" authorityYear="2003" box="[151,504,1585,1612]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="26" pageNumber="497" phylum="Mollusca" rank="species" species="pascuana">
<emphasis bold="true" box="[151,504,1585,1612]" pageId="26" pageNumber="497">
<emphasis bold="true" box="[151,338,1586,1612]" italics="true" pageId="26" pageNumber="497">Pupa pascuana</emphasis>
<bibRefCitation author="Raines, B. K." box="[345,504,1585,1611]" pageId="26" pageNumber="497" pagination="51 - 53" refId="ref25687" refString="Raines, B. K. (2003) Pupa pascuana: A new species of Acteonidae from Easter Island. La Conchiglia, 34, 51 - 53." type="journal article" year="2003">Raines, 2003</bibRefCitation>
</emphasis>
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<paragraph blockId="26.[151,504,1585,1647]" box="[151,267,1621,1647]" pageId="26" pageNumber="497">
(
<figureCitation box="[159,259,1621,1647]" captionStart="FIGURE 16" captionStartId="28.[152,255,768,792]" captionTargetBox="[151,1436,181,745]" captionTargetId="figure-24@28.[151,1436,181,745]" captionTargetPageId="28" captionText="FIGURE 16. Photographs of shell specimens of Pupa niecaensis (Barnard, 1963) (A), Pupa pascuana Raines, 2003 (B) and Pupa charlesi sp. nov. (CE). A. Holotype of Pupa niecaensis (Barnard, 1963) (SAM A6553), ©SAM, photo by R. Adams. B. Holotype of Pupa pascuana Raines, 2003 (LACM 2954). CE. Specimens of Pupa charlesi sp. nov. from Koumac, New Caledonia, (C) MHNH IM-2013-86154, (D) MHNH IM-2013-86159, (E) MHNH IM-2013-86158." figureDoi="http://doi.org/10.5281/zenodo.7860408" httpUri="https://zenodo.org/record/7860408/files/figure.png" pageId="26" pageNumber="497">Fig. 16B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection box="[151,1189,1690,1714]" pageId="26" pageNumber="497" type="reference_group">
<paragraph blockId="26.[151,1189,1690,1714]" box="[151,1189,1690,1714]" pageId="26" pageNumber="497">
<treatmentCitationGroup box="[151,1189,1690,1714]" pageId="26" pageNumber="497">
<treatmentCitation author="Raines, B. K." box="[151,488,1690,1714]" page="52" pageId="26" pageNumber="497" year="2003">
<taxonomicName ID-CoL="4QJHX" authority="Raines, 2003: 52" authorityName="Raines" authorityPageNumber="52" authorityYear="2003" box="[151,488,1690,1714]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="26" pageNumber="497" phylum="Mollusca" rank="species" species="pascuana">
<emphasis box="[151,308,1691,1714]" italics="true" pageId="26" pageNumber="497">Pupa pascuana</emphasis>
<materialsCitation box="[314,488,1690,1714]" location="Tahai" municipality="Easter Island" pageId="26" pageNumber="497" specimenCount="1">
<bibRefCitation author="Raines, B. K." box="[314,488,1690,1714]" pageId="26" pageNumber="497" pagination="51 - 53" refId="ref25687" refString="Raines, B. K. (2003) Pupa pascuana: A new species of Acteonidae from Easter Island. La Conchiglia, 34, 51 - 53." type="journal article" year="2003">Raines, 2003: 52</bibRefCitation>
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, figs 12.
<typeStatus box="[593,643,1690,1714]" pageId="26" pageNumber="497">Type</typeStatus>
locality: off the western coast of
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,
<collectingMunicipality box="[1050,1185,1690,1714]" pageId="26" pageNumber="497">Easter Island</collectingMunicipality>
.
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</paragraph>
</subSubSection>
<subSubSection lastPageId="27" lastPageNumber="498" pageId="26" pageNumber="497" type="materials_examined">
<paragraph blockId="26.[151,1436,1760,2039]" box="[151,873,1760,1787]" pageId="26" pageNumber="497">
<emphasis bold="true" box="[151,318,1760,1786]" pageId="26" pageNumber="497">Type material</emphasis>
.
<materialsCitation box="[332,873,1760,1787]" collectionCode="LACM" pageId="26" pageNumber="497" specimenCode="LACM 2954" specimenCount="1" typeStatus="holotype">
<typeStatus box="[332,439,1760,1786]" pageId="26" pageNumber="497">Holotype</typeStatus>
, dry shell,
<quantity box="[561,639,1760,1787]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="26" pageNumber="497" unit="mm" value="11.0">11 mm</quantity>
long (
<specimenCode box="[710,861,1760,1787]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="26" pageNumber="497" type="Museum">LACM 2954</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph blockId="26.[151,1436,1760,2039]" lastBlockId="27.[151,1436,151,393]" lastPageId="27" lastPageNumber="498" pageId="26" pageNumber="497">
<emphasis bold="true" box="[199,459,1796,1822]" pageId="26" pageNumber="497">External morphology</emphasis>
. Live animal unknown. Shell solid, narrow, very elongate, widest at midlengh, with slightly convex, nearly parallel sides, rounded to elongate anterior end (
<figureCitation box="[982,1088,1832,1858]" captionStart="FIGURE 16" captionStartId="28.[152,255,768,792]" captionTargetBox="[151,1436,181,745]" captionTargetId="figure-24@28.[151,1436,181,745]" captionTargetPageId="28" captionText="FIGURE 16. Photographs of shell specimens of Pupa niecaensis (Barnard, 1963) (A), Pupa pascuana Raines, 2003 (B) and Pupa charlesi sp. nov. (CE). A. Holotype of Pupa niecaensis (Barnard, 1963) (SAM A6553), ©SAM, photo by R. Adams. B. Holotype of Pupa pascuana Raines, 2003 (LACM 2954). CE. Specimens of Pupa charlesi sp. nov. from Koumac, New Caledonia, (C) MHNH IM-2013-86154, (D) MHNH IM-2013-86159, (E) MHNH IM-2013-86158." figureDoi="http://doi.org/10.5281/zenodo.7860408" httpUri="https://zenodo.org/record/7860408/files/figure.png" pageId="26" pageNumber="497">Fig. 16B</figureCitation>
). Body whorl large, about 3/4 of total length. Spire conical, with 5 whorls. Suture channeled. Aperture elongate, wider anteriorly, narrowing gradually towards posterior end, ending at 4/5 of the posterior end of first whorl. Columellar margin thickened, slightly oblique, with large, channeled anterior fold starting at anterior end of aperture; minute, simple posterior fold located at aperture mid-length, separated from anterior fold by wide gap. Protoconch globose,
<quantity box="[1211,1297,1976,2003]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="26" pageNumber="497" unit="mm" value="0.1">0.1 mm</quantity>
in diameter, with ~1.5 whorls. Sculpture composed of numerous punctuated spiral grooves. Punctuations conspicuous, irregular, oval, situated next to each other, often fused together, within each groove. Grooves separated by gaps slightly wider than grooves. Shell color whitish to pale cream with irregular orange spots all over the surface.
</paragraph>
<paragraph blockId="27.[151,1436,151,393]" box="[199,1151,222,249]" pageId="27" pageNumber="498">
<emphasis bold="true" box="[199,414,222,249]" pageId="27" pageNumber="498">Geographic range</emphasis>
. Possibly endemic to Easter Island (
<bibRefCitation author="Raines, B. K." box="[821,972,223,249]" pageId="27" pageNumber="498" pagination="51 - 53" refId="ref25687" refString="Raines, B. K. (2003) Pupa pascuana: A new species of Acteonidae from Easter Island. La Conchiglia, 34, 51 - 53." type="journal article" year="2003">Raines, 2003</bibRefCitation>
; present paper).
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="498" type="discussion">
<paragraph blockId="27.[151,1436,151,393]" pageId="27" pageNumber="498">
<emphasis bold="true" box="[199,308,259,285]" pageId="27" pageNumber="498">Remarks</emphasis>
.
<bibRefCitation author="Raines, B. K." box="[323,482,259,285]" pageId="27" pageNumber="498" pagination="51 - 53" refId="ref25687" refString="Raines, B. K. (2003) Pupa pascuana: A new species of Acteonidae from Easter Island. La Conchiglia, 34, 51 - 53." type="journal article" year="2003">Raines (2003)</bibRefCitation>
introduced the name
<taxonomicName authority="Raines, 2003" authorityName="Raines" authorityYear="2003" box="[729,1060,259,285]" class="Gastropoda" family="Acteonidae" genus="Pupa" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="27" pageNumber="498" phylum="Mollusca" rank="species" species="pascuana">
<emphasis box="[729,904,259,284]" italics="true" pageId="27" pageNumber="498">Pupa pascuana</emphasis>
<bibRefCitation author="Raines, B. K." box="[912,1060,259,285]" pageId="27" pageNumber="498" pagination="51 - 53" refId="ref25687" refString="Raines, B. K. (2003) Pupa pascuana: A new species of Acteonidae from Easter Island. La Conchiglia, 34, 51 - 53." type="journal article" year="2003">Raines, 2003</bibRefCitation>
</taxonomicName>
based on several shells collected on Easter Island. Based on the examination of the
<typeStatus box="[721,819,295,321]" pageId="27" pageNumber="498">holotype</typeStatus>
(LACM 2954;
<figureCitation box="[996,1095,295,321]" captionStart="FIGURE 16" captionStartId="28.[152,255,768,792]" captionTargetBox="[151,1436,181,745]" captionTargetId="figure-24@28.[151,1436,181,745]" captionTargetPageId="28" captionText="FIGURE 16. Photographs of shell specimens of Pupa niecaensis (Barnard, 1963) (A), Pupa pascuana Raines, 2003 (B) and Pupa charlesi sp. nov. (CE). A. Holotype of Pupa niecaensis (Barnard, 1963) (SAM A6553), ©SAM, photo by R. Adams. B. Holotype of Pupa pascuana Raines, 2003 (LACM 2954). CE. Specimens of Pupa charlesi sp. nov. from Koumac, New Caledonia, (C) MHNH IM-2013-86154, (D) MHNH IM-2013-86159, (E) MHNH IM-2013-86158." figureDoi="http://doi.org/10.5281/zenodo.7860408" httpUri="https://zenodo.org/record/7860408/files/figure.png" pageId="27" pageNumber="498">Fig. 16B</figureCitation>
) it appears to be distinct from other species examined herein and probably constitutes an endemic species to Easter Island, but this needs to be confirmed with molecular and anatomical data.
</paragraph>
</subSubSection>
</treatment>
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