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<document id="DF485125101485B468CCC3556FAE55AF" ID-CLB-Dataset="46025" ID-DOI="10.11646/zootaxa.3786.1.2" ID-GBIF-Dataset="ef8a0c66-3103-4a40-8302-0a1a4055811a" ID-ISSN="1175-5326" ID-Zenodo-Dep="231537" ID-ZooBank="09DC7D2C-B15B-4DC8-9D89-FDC0E97D2776" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460407886034" checkinUser="plazi" docAuthor="Kluge, Nikita J." docDate="2014" docId="03C2A127FFAFF34F6380FF658243EFF9" docLanguage="en" docName="zt03786p056.pdf" docOrigin="Zootaxa 3786 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Choroterpes (Dilatognathus) bogori Kluge, 2014, sp.n." docType="treatment" docVersion="9" masterDocId="FFFBD95FFFABF3476317FFF2816EEE71" masterDocTitle="Indonesian species of Dilatognathus Kluge 2012 (Ephemeroptera, Leptophlebiidae, Choroterpes s. l.) and species-specific sexual dimorphism in development of maxilla" updateTime="1698300041941" updateUser="ExternalLinkService">
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<mods:title id="8ECCD0AB14E8361EB02BD697F8120996">Indonesian species of Dilatognathus Kluge 2012 (Ephemeroptera, Leptophlebiidae, Choroterpes s. l.) and species-specific sexual dimorphism in development of maxilla</mods:title>
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<mods:namePart id="6721C55FAF1D7C00604F70E17E1397DD">Kluge, Nikita J.</mods:namePart>
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<mods:date id="45D9B6C1747D5FFCF67406B59FA12EEA">2014</mods:date>
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<treatment id="03C2A127FFAFF34F6380FF658243EFF9" ID-DOI="http://doi.org/10.5281/zenodo.6137922" ID-GBIF-Taxon="119422426" ID-Zenodo-Dep="6137922" LSID="urn:lsid:plazi:treatment:03C2A127FFAFF34F6380FF658243EFF9" httpUri="http://treatment.plazi.org/id/03C2A127FFAFF34F6380FF658243EFF9" lastPageId="8" pageId="4">
<subSubSection id="C37143BAFFAFF3436380FF658348EF46" pageId="4" type="nomenclature">
<paragraph id="8BD41031FFAFF3436380FF6583E4EEC3" blockId="4.[151,650,151,211]" box="[151,650,151,178]" pageId="4">
<heading id="D09CA75DFFAFF3436380FF6583E4EEC3" bold="true" box="[151,650,151,178]" fontSize="11" level="1" pageId="4" reason="1">
<emphasis id="B91FCC23FFAFF3436380FF6583E4EEC3" bold="true" box="[151,650,151,178]" pageId="4">
<taxonomicName id="4C6B6BB2FFAFF3436380FF658326EEC0" box="[151,584,151,178]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="4" phylum="Arthropoda" rank="species" species="bogori" status="sp. nov." subGenus="Dilatognathus">
<emphasis id="B91FCC23FFAFF3436380FF658045EEC0" bold="true" box="[151,299,151,177]" italics="true" pageId="4">Choroterpes</emphasis>
(
<emphasis id="B91FCC23FFAFF343622BFF658087EEC0" bold="true" box="[316,489,151,177]" italics="true" pageId="4">Dilatognathus</emphasis>
)
<emphasis id="B91FCC23FFAFF34362EDFF658326EEC0" bold="true" box="[506,584,151,177]" italics="true" pageId="4">bogori</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C7158FFAFF3436158FF6A83E4EEC3" box="[591,650,152,178]" pageId="4" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD41031FFAFF3436380FF488323EEA2" blockId="4.[151,650,151,211]" box="[151,589,186,211]" pageId="4">
(
<figureCitation id="13500CB4FFAFF3436388FF48804EEEA2" box="[159,288,186,211]" captionStart-0="FIGURES 16 18" captionStart-1="FIGURES 19 28" captionStartId-0="4.[151,264,1940,1963]" captionStartId-1="5.[151,264,1784,1807]" captionTargetBox-0="[193,1408,353,1907]" captionTargetBox-1="[155,1436,207,1738]" captionTargetId-0="figure@4.[193,1437,353,1908]" captionTargetId-1="figure@5.[151,1436,193,1763]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 16 18. Choroterpes (Dilatognathus) bogori, maxilla of penultimolarva at various stages of transformation preceding molt to ultimolarva (slides in Canadian balsam): 16, female, early stage; 17, female, middle stage; 18, male, late stage." captionText-1="FIGURES 19 28. Choroterpes (Dilatognathus) spp. 19 24, Ch. (D.) bogori: 19 and 20, apex of maxilla in pen-penpenultimate larval instar (ventral view; tusk, ventro-apical flange and vestiges of pectinate ventral setae are shown, other setae not shown): 19, male; 20, female; 21 and 21, genitals of male imago, ventral view (holotype); 23, tergalius III (second existent); 24, middle part of anterior margin of labrum, ventral view. 25 26, genitals and enlarged penis of Choroterpides exiguus (Eaton 1884 [Thraulus]) (from Eaton 1884: Pl. 13, Fig. 20 * 2). 27, Choroterpes (Dilatognathus) major, tergalius III (second existent). 28, Choroterpes (Dilatognathus) minor, the same." httpUri-0="https://zenodo.org/record/231539/files/figure.png" httpUri-1="https://zenodo.org/record/231540/files/figure.png" pageId="4">Figs 1624</figureCitation>
,
<figureCitation id="13500CB4FFAFF343623CFF49801DEEA2" box="[299,371,187,211]" captionStart="FIGURES 29 33" captionStartId="6.[151,264,1213,1236]" captionTargetBox="[151,1415,193,1192]" captionTargetId="figure@6.[151,1436,193,1192]" captionTargetPageId="6" captionText="FIGURES 29 33. Choroterpes (Dilatognathus) spp. 29, Ch. (D.) bogori, abdomen of male imago, spread on slide (holotype); 30, Ch. (D.) cataractae, the same (holotype); 31, Ch. (D.) bogori, hind leg of male imago (holotype); 32, Ch. (D.) cataractae, the same (holotype); 33, Ch. (D.) major, hind leg of mature larva with developing subimaginal leg." httpUri="https://zenodo.org/record/231541/files/figure.png" pageId="4">29, 31</figureCitation>
,
<figureCitation id="13500CB4FFAFF3436268FF49832BEEA2" box="[383,581,186,211]" captionStart-0="FIGURES 34 39" captionStart-1="FIGURES 40 43" captionStartId-0="8.[151,264,1934,1957]" captionStartId-1="9.[151,264,1169,1192]" captionTargetBox-0="[187,1399,447,1886]" captionTargetBox-1="[151,1436,193,1148]" captionTargetId-0="figure@8.[187,1400,447,1913]" captionTargetId-1="figure@9.[151,1436,193,1149]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURES 34 39. Choroterpes (Dilatognathus) spp. 34 37, median part of postrior margin of abdominal tergum of ultimolarva: 34 35, Ch. (D.) bogori; 36 37, Ch. (D.) cataractae; 34 and 36, tergum II; 35 and 37, tergum VII. 38 39, Ch. (D.) bogori: 38, exuviae of right half of subimaginal mesonotum; 39, egg." captionText-1="FIGURES 40 43. Choroterpes (Dilatognathus) bogori sp. n., eggs. 40, 41, total view; 42, 43, two poles of egg in Fig. 41." httpUri-0="https://zenodo.org/record/231542/files/figure.png" httpUri-1="https://zenodo.org/record/231543/files/figure.png" pageId="4">34, 35, 38, 3943</figureCitation>
)
</paragraph>
<paragraph id="8BD41031FFAFF3436380FEF18348EF46" blockId="4.[151,983,258,311]" pageId="4">
= &quot;
<taxonomicName id="4C6B6BB2FFAFF34363A1FEF080F4EF69" authority="Ulmer" authorityName="Ulmer" authorityYear="1939" box="[182,410,258,280]" class="Insecta" family="Leptophlebiidae" genus="Choroterpides" kingdom="Animalia" order="Ephemeroptera" pageId="4" phylum="Arthropoda" rank="species" species="exiguus">
<emphasis id="B91FCC23FFAFF34363A1FEF080F4EF69" box="[182,410,258,280]" italics="true" pageId="4">Choroterpides exiguus</emphasis>
</taxonomicName>
&quot; sensu
<bibRefCitation id="EFFA6DC0FFAFF34362FDFEF0830AEF69" author="Ulmer" box="[490,612,258,280]" pageId="4" refString="Ulmer, G. (1939) Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln. Archiv fur Hydrobiologie, Supplement 16, 443 - 692." type="book chapter" year="1939">Ulmer 1939</bibRefCitation>
(non
<taxonomicName id="4C6B6BB2FFAFF3436189FEF082A5EF69" authority="Eaton 1884" authorityName="Eaton" authorityYear="1884" box="[670,971,258,280]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="4" phylum="Arthropoda" rank="species" species="exiguus">
<emphasis id="B91FCC23FFAFF3436189FEF08223EF69" box="[670,845,258,280]" italics="true" pageId="4">Thraulus exiguus</emphasis>
Eaton 1884
</taxonomicName>
); = Dilatognathus/g(1) sp.2:
<bibRefCitation id="EFFA6DC0FFAFF34362BFFED3834CEF46" author="Kluge" box="[424,546,289,311]" pageId="4" refString="Kluge, N. J. (2012) Contribution to the knowledge of Choroterpes (Ephemeroptera, Leptophlebiidae). Russian Entomological Journal, 21 (3), 273 - 306." type="journal article" year="2012">Kluge 2012</bibRefCitation>
.
</paragraph>
</subSubSection>
<caption id="DF1440B9FFAFF3436380F86681BCE996" httpUri="https://zenodo.org/record/231539/files/figure.png" pageId="4" targetBox="[193,1408,353,1907]" targetPageId="4">
<paragraph id="8BD41031FFAFF3436380F86681BCE996" blockId="4.[151,1436,1940,2023]" pageId="4">
<emphasis id="B91FCC23FFAFF3436380F8668035E9D8" bold="true" box="[151,347,1940,1963]" pageId="4">FIGURES 1618.</emphasis>
<taxonomicName id="4C6B6BB2FFAFF343627EF8668389E9DB" box="[361,743,1940,1962]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="4" phylum="Arthropoda" rank="species" species="bogori" subGenus="Dilatognathus">
<emphasis id="B91FCC23FFAFF343627EF8668089E9DB" box="[361,487,1940,1962]" italics="true" pageId="4">Choroterpes</emphasis>
(
<emphasis id="B91FCC23FFAFF34362EBF86683E3E9DB" box="[508,653,1940,1962]" italics="true" pageId="4">Dilatognathus</emphasis>
)
<emphasis id="B91FCC23FFAFF34361B3F8668389E9DB" box="[676,743,1940,1962]" italics="true" pageId="4">bogori</emphasis>
</taxonomicName>
, maxilla of penultimolarva at various stages of transformation preceding molt to ultimolarva (slides in Canadian balsam): 16, female, early stage; 17, female, middle stage; 18, male, late stage.
</paragraph>
</caption>
<caption id="DF1440B9FFAEF3426380F90A83FFE9D9" httpUri="https://zenodo.org/record/231540/files/figure.png" pageId="5" targetBox="[155,1436,207,1738]" targetPageId="5">
<paragraph id="8BD41031FFAEF3426380F90A83FFE9D9" blockId="5.[151,1437,1784,1960]" pageId="5">
<emphasis id="B91FCC23FFAEF3426380F90A8034E97C" bold="true" box="[151,346,1784,1807]" pageId="5">FIGURES 1928.</emphasis>
<taxonomicName id="4C6B6BB2FFAEF3426271F90A83FCE97F" authority="Dilatognathus" authorityName="Dilatognathus" box="[358,658,1784,1806]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="5" phylum="Arthropoda" rank="genus">
<emphasis id="B91FCC23FFAEF3426271F90A808AE97F" box="[358,484,1784,1806]" italics="true" pageId="5">Choroterpes</emphasis>
(
<emphasis id="B91FCC23FFAEF34262EEF90A83E4E97F" box="[505,650,1784,1806]" italics="true" pageId="5">Dilatognathus</emphasis>
)
</taxonomicName>
spp. 1924,
<taxonomicName id="4C6B6BB2FFAEF342603FF90A82BCE97F" box="[808,978,1784,1807]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="5" phylum="Arthropoda" rank="species" species="bogori" subGenus="Dilatognathus">
<emphasis id="B91FCC23FFAEF342603FF90A8223E97F" box="[808,845,1784,1806]" italics="true" pageId="5">Ch.</emphasis>
(
<emphasis id="B91FCC23FFAEF3426076F9088214E97E" box="[865,890,1786,1807]" italics="true" pageId="5">D.</emphasis>
)
<emphasis id="B91FCC23FFAEF3426098F90A82BCE97F" box="[911,978,1784,1806]" italics="true" pageId="5">bogori</emphasis>
</taxonomicName>
: 19 and 20, apex of maxilla in pen-penpenultimate larval instar (ventral view; tusk, ventro-apical flange and vestiges of pectinate ventral setae are shown, other setae not shown): 19, male; 20, female; 21 and 21, genitals of male imago, ventral view (holotype); 23, tergalius III (second existent); 24, middle part of anterior margin of labrum, ventral view. 2526, genitals and enlarged penis of
<taxonomicName id="4C6B6BB2FFAEF3426779F8A6843CE91B" box="[1134,1362,1876,1898]" class="Insecta" family="Leptophlebiidae" genus="Choroterpides" kingdom="Animalia" order="Ephemeroptera" pageId="5" phylum="Arthropoda" rank="species" species="exiguus">
<emphasis id="B91FCC23FFAEF3426779F8A6843CE91B" box="[1134,1362,1876,1898]" italics="true" pageId="5">Choroterpides exiguus</emphasis>
</taxonomicName>
(Eaton 1884 [
<taxonomicName id="4C6B6BB2FFAEF34263CEF881805AE9F8" box="[217,308,1907,1929]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="5" phylum="Arthropoda" rank="genus">
<emphasis id="B91FCC23FFAEF34263CEF881805AE9F8" box="[217,308,1907,1929]" italics="true" pageId="5">Thraulus</emphasis>
</taxonomicName>
]) (from Eaton 1884: Pl. 13, Fig. 20*2). 27,
<taxonomicName id="4C6B6BB2FFAEF34261E2F8818530E9F8" box="[757,1118,1907,1929]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="5" phylum="Arthropoda" rank="species" species="major" subGenus="Dilatognathus">
<emphasis id="B91FCC23FFAEF34261E2F881821CE9F8" box="[757,882,1907,1929]" italics="true" pageId="5">Choroterpes</emphasis>
(
<emphasis id="B91FCC23FFAEF3426095F881857DE9F8" box="[898,1043,1907,1929]" italics="true" pageId="5">Dilatognathus</emphasis>
)
<emphasis id="B91FCC23FFAEF3426735F8868530E9F8" box="[1058,1118,1908,1929]" italics="true" pageId="5">major</emphasis>
</taxonomicName>
, tergalius III (second existent). 28,
<taxonomicName id="4C6B6BB2FFAEF34263AAF860834BE9D9" box="[189,549,1938,1960]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="5" phylum="Arthropoda" rank="species" species="minor" subGenus="Dilatognathus">
<emphasis id="B91FCC23FFAEF34263AAF8608055E9D9" box="[189,315,1938,1960]" italics="true" pageId="5">Choroterpes</emphasis>
(
<emphasis id="B91FCC23FFAEF342625EF86080B4E9D9" box="[329,474,1938,1960]" italics="true" pageId="5">Dilatognathus</emphasis>
)
<emphasis id="B91FCC23FFAEF34262FEF861834BE9D9" box="[489,549,1939,1960]" italics="true" pageId="5">minor</emphasis>
</taxonomicName>
, the same.
</paragraph>
</caption>
<caption id="DF1440B9FFADF3416380FB4F85EFEB60" httpUri="https://zenodo.org/record/231541/files/figure.png" pageId="6" targetBox="[151,1415,193,1192]" targetPageId="6">
<paragraph id="8BD41031FFADF3416380FB4F85EFEB60" blockId="6.[151,1436,1213,1297]" pageId="6">
<emphasis id="B91FCC23FFADF3416380FB4F803AEAA2" bold="true" box="[151,340,1213,1236]" pageId="6">FIGURES 2933.</emphasis>
<taxonomicName id="4C6B6BB2FFADF341624DFB4F83EEEAA2" authority="Dilatognathus" authorityName="Dilatognathus" box="[346,640,1213,1235]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="6" phylum="Arthropoda" rank="genus">
<emphasis id="B91FCC23FFADF341624DFB4F80B6EAA2" box="[346,472,1213,1235]" italics="true" pageId="6">Choroterpes</emphasis>
(
<emphasis id="B91FCC23FFADF34162F0FB4F8316EAA2" box="[487,632,1213,1235]" italics="true" pageId="6">Dilatognathus</emphasis>
)
</taxonomicName>
spp. 29,
<taxonomicName id="4C6B6BB2FFADF34161CAFB4F8215EAA2" box="[733,891,1213,1236]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="6" phylum="Arthropoda" rank="species" species="bogori" subGenus="Dilatognathus">
<emphasis id="B91FCC23FFADF34161CAFB4F826CEAA2" box="[733,770,1213,1235]" italics="true" pageId="6">Ch.</emphasis>
(
<emphasis id="B91FCC23FFADF3416007FB4D8244EAA5" box="[784,810,1215,1236]" italics="true" pageId="6">D.</emphasis>
)
<emphasis id="B91FCC23FFADF341602FFB4F8215EAA2" box="[824,891,1213,1235]" italics="true" pageId="6">bogori</emphasis>
</taxonomicName>
, abdomen of male imago, spread on slide (holotype); 30,
<taxonomicName id="4C6B6BB2FFADF34163AAFB2E80EAEA83" box="[189,388,1244,1266]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="6" phylum="Arthropoda" rank="species" species="cataractae" subGenus="Dilatognathus">
<emphasis id="B91FCC23FFADF34163AAFB2E818FEA83" box="[189,225,1244,1266]" italics="true" pageId="6">Ch.</emphasis>
(
<emphasis id="B91FCC23FFADF34163E7FB2F8064EA83" box="[240,266,1245,1266]" italics="true" pageId="6">D.</emphasis>
)
<emphasis id="B91FCC23FFADF341620FFB2F80EAEA83" box="[280,388,1245,1266]" italics="true" pageId="6">cataractae</emphasis>
</taxonomicName>
, the same (holotype); 31,
<taxonomicName id="4C6B6BB2FFADF3416187FB2E8241EA83" box="[656,815,1244,1266]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="6" phylum="Arthropoda" rank="species" species="bogori" subGenus="Dilatognathus">
<emphasis id="B91FCC23FFADF3416187FB2E83DAEA83" box="[656,692,1244,1266]" italics="true" pageId="6">Ch.</emphasis>
(
<emphasis id="B91FCC23FFADF34161D3FB2F83B3EA83" box="[708,733,1245,1266]" italics="true" pageId="6">D.</emphasis>
)
<emphasis id="B91FCC23FFADF34161FBFB2E8241EA83" box="[748,815,1244,1266]" italics="true" pageId="6">bogori</emphasis>
</taxonomicName>
, hind leg of male imago (holotype); 32,
<taxonomicName id="4C6B6BB2FFADF34167D9FB2E84FBEA83" box="[1230,1429,1244,1266]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="6" phylum="Arthropoda" rank="species" species="cataractae" subGenus="Dilatognathus">
<emphasis id="B91FCC23FFADF34167D9FB2E859DEA83" box="[1230,1267,1244,1266]" italics="true" pageId="6">Ch.</emphasis>
(
<emphasis id="B91FCC23FFADF3416616FB2F8475EA83" box="[1281,1307,1245,1266]" italics="true" pageId="6">D.</emphasis>
)
<emphasis id="B91FCC23FFADF341663EFB2F84FBEA83" box="[1321,1429,1245,1266]" italics="true" pageId="6">cataractae</emphasis>
</taxonomicName>
, the same (holotype); 33,
<taxonomicName id="4C6B6BB2FFADF3416284FB098347EB60" box="[403,553,1275,1297]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="6" phylum="Arthropoda" rank="species" species="major" subGenus="Dilatognathus">
<emphasis id="B91FCC23FFADF3416284FB0980D6EB60" box="[403,440,1275,1297]" italics="true" pageId="6">Ch.</emphasis>
(
<emphasis id="B91FCC23FFADF34162D1FB0E80B1EB60" box="[454,479,1276,1297]" italics="true" pageId="6">D.</emphasis>
)
<emphasis id="B91FCC23FFADF34162FAFB0E8347EB60" box="[493,553,1276,1297]" italics="true" pageId="6">major</emphasis>
</taxonomicName>
, hind leg of mature larva with developing subimaginal leg.
</paragraph>
</caption>
<subSubSection id="C37143BAFFADF3406380FACA8285E85E" lastPageId="7" pageId="6" type="materials_examined">
<paragraph id="8BD41031FFADF3416380FACA81ADEB98" blockId="6.[151,1436,1336,2024]" pageId="6">
<emphasis id="B91FCC23FFADF3416380FACA8068EB20" bold="true" box="[151,262,1336,1361]" pageId="6">Material.</emphasis>
<typeStatus id="54D0AE93FFADF341621AFACA8014EB20" box="[269,378,1336,1361]" pageId="6" type="holotype">Holotype</typeStatus>
: L-S-I♂ {No. [XXIII] (5)},
<collectingCountry id="F37C50A1FFADF34161AEFACB823AEB20" box="[697,852,1337,1361]" name="Indonesia" pageId="6">INDONESIA</collectingCountry>
,
<collectionCode id="ED7A88F4FFADF3416077FACB82C4EB20" box="[864,938,1337,1361]" pageId="6">JAVA</collectionCode>
, Bogor, river Ci Sadane,
<date id="FFD536F1FFADF34167DFFACB8455EB20" box="[1224,1339,1337,1361]" pageId="6" value="2012-09-03">3.IX.2012</date>
, coll. N. Kluge.
<typeStatus id="54D0AE93FFADF34163FDFAAF8035EB04" box="[234,347,1373,1397]" pageId="6" type="paratype">Paratypes</typeStatus>
: the same locality and date: 4 L-S-I♂, 1 L-S♂, 4 L-S-I♀,
<specimenCount id="9D6DDBB8FFADF34160FDFAAF8538EB04" box="[1002,1110,1372,1397]" pageId="6" type="larva">98 larvae</specimenCount>
: 20 L n♂, 38 L n♀, 9 L n-
<specimenCount id="9D6DDBB8FFADF3416640FA99841DEB05" box="[1367,1395,1373,1401]" pageId="6" type="generic">1♂</specimenCount>
, 20 L n-1♀, 4 L n-
<specimenCount id="9D6DDBB8FFADF341620EFA608059EBEA" box="[281,311,1412,1440]" pageId="6" type="generic">2♂</specimenCount>
, 5 L n-2♀, 1 L n-
<specimenCount id="9D6DDBB8FFADF34162F6FA608091EBEA" box="[481,511,1412,1440]" pageId="6" type="generic">3♂</specimenCount>
, 1 L n-3♀; river Ci Liwung in botanic garden of Bogor,
<date id="FFD536F1FFADF3416795FA718599EBEA" box="[1154,1271,1411,1435]" pageId="6" value="2012-09-02">2.IX.2012</date>
: 1 L-S-I♂,
<specimenCount id="9D6DDBB8FFADF3416697FA71818EEBB2" pageId="6" type="larva">19 larvae</specimenCount>
: 5 L n♂, 9 L n♀, 1 L n-
<specimenCount id="9D6DDBB8FFADF34162D5FA4A80B0EBB3" box="[450,478,1451,1478]" pageId="6" type="generic">1♂</specimenCount>
, 2 L n-1♀, 2 L n-2♀; the same locality,
<date id="FFD536F1FFADF3416066FA588287EBB3" box="[881,1001,1450,1474]" pageId="6" value="2008-02-24">24.II.2008</date>
, at light, coll. V. Ivanov:
<date id="FFD536F1FFADF3416612FA58844AEBB3" box="[1285,1316,1450,1474]" pageId="6">2 I</date>
♂, 4 S♂, 1 S♀.
</paragraph>
<paragraph id="8BD41031FFADF34163D0FA0684FEE840" blockId="6.[151,1436,1336,2024]" pageId="6">
<emphasis id="B91FCC23FFADF34163D0FA06800DE87C" bold="true" box="[199,355,1524,1549]" pageId="6">Descriptions.</emphasis>
<emphasis id="B91FCC23FFADF3416266FA0480D2E87C" box="[369,444,1526,1549]" italics="true" pageId="6">Larva.</emphasis>
CUTICULAR COLORATION: Cuticle of head, pronotum and mesonotum brown, with diffusive markings; cuticle of legs and abdomen light brownish, nearly unicolorous, without contrasting markings.
</paragraph>
<paragraph id="8BD41031FFADF34163D0F9CF841DE894" blockId="6.[151,1436,1336,2024]" pageId="6">
HYPODERMAL COLORATION: General color light brown. Head and thorax with dark maculae. Legs light ocher; each femur with large brown macula or band at middle, sometimes with smaller brown macula at apex, which is separated from middle macula (as in imago,
<figureCitation id="13500CB4FFADF3416187F976838DE8EC" box="[656,739,1668,1693]" captionStart="FIGURES 29 33" captionStartId="6.[151,264,1213,1236]" captionTargetBox="[151,1415,193,1192]" captionTargetId="figure@6.[151,1436,193,1192]" captionTargetPageId="6" captionText="FIGURES 29 33. Choroterpes (Dilatognathus) spp. 29, Ch. (D.) bogori, abdomen of male imago, spread on slide (holotype); 30, Ch. (D.) cataractae, the same (holotype); 31, Ch. (D.) bogori, hind leg of male imago (holotype); 32, Ch. (D.) cataractae, the same (holotype); 33, Ch. (D.) major, hind leg of mature larva with developing subimaginal leg." httpUri="https://zenodo.org/record/231541/files/figure.png" pageId="6">Fig. 31</figureCitation>
). Abdominal terga with brown pigmentation, median-anterior part of each tergum ocher (as in imago,
<figureCitation id="13500CB4FFADF3416175F95A83D8E8B0" box="[610,694,1704,1729]" captionStart="FIGURES 29 33" captionStartId="6.[151,264,1213,1236]" captionTargetBox="[151,1415,193,1192]" captionTargetId="figure@6.[151,1436,193,1192]" captionTargetPageId="6" captionText="FIGURES 29 33. Choroterpes (Dilatognathus) spp. 29, Ch. (D.) bogori, abdomen of male imago, spread on slide (holotype); 30, Ch. (D.) cataractae, the same (holotype); 31, Ch. (D.) bogori, hind leg of male imago (holotype); 32, Ch. (D.) cataractae, the same (holotype); 33, Ch. (D.) major, hind leg of mature larva with developing subimaginal leg." httpUri="https://zenodo.org/record/231541/files/figure.png" pageId="6">Fig. 29</figureCitation>
). Dorsal lamella of each tergalius light brown, with main trachea bordered by dark brown and side tracheae poorly visible; ventral lamella lighter brownish, with lighter trachea.
</paragraph>
<paragraph id="8BD41031FFADF34063D0F90383A6ED66" blockId="6.[151,1436,1336,2024]" lastBlockId="7.[151,1436,151,2016]" lastPageId="7" pageId="6">
<collectionCode id="ED7A88F4FFADF34163D0F903807FE978" box="[199,273,1777,1801]" pageId="6">SHAPE</collectionCode>
AND SETATION: Labrum as in
<taxonomicName id="4C6B6BB2FFADF341619EF902821EE978" box="[649,880,1776,1801]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="6" phylum="Arthropoda" rank="species" species="cataractae" subGenus="Dilatognathus">
<emphasis id="B91FCC23FFADF341619EF90283C4E978" box="[649,682,1776,1801]" italics="true" pageId="6">Ch</emphasis>
. (
<emphasis id="B91FCC23FFADF34161D0F90083B5E978" box="[711,731,1778,1801]" italics="true" pageId="6">D</emphasis>
.)
<emphasis id="B91FCC23FFADF34161EFF900821EE978" box="[760,880,1778,1801]" italics="true" pageId="6">cataractae</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFFA6DC0FFADF3416092F90285E5E978" author="Kluge" box="[901,1163,1776,1801]" pageId="6" refString="Kluge, N. J. (2012) Contribution to the knowledge of Choroterpes (Ephemeroptera, Leptophlebiidae). Russian Entomological Journal, 21 (3), 273 - 306." type="journal article" year="2012">Kluge 2012: Fig. 133</bibRefCitation>
): median incision very shallow, with shallow convexity in middle and 5 shallow denticles ventrad of it (
<figureCitation id="13500CB4FFADF3416735F8E68519E95C" box="[1058,1143,1812,1837]" captionStart="FIGURES 19 28" captionStartId="5.[151,264,1784,1807]" captionTargetBox="[155,1436,207,1738]" captionTargetId="figure@5.[151,1436,193,1763]" captionTargetPageId="5" captionText="FIGURES 19 28. Choroterpes (Dilatognathus) spp. 19 24, Ch. (D.) bogori: 19 and 20, apex of maxilla in pen-penpenultimate larval instar (ventral view; tusk, ventro-apical flange and vestiges of pectinate ventral setae are shown, other setae not shown): 19, male; 20, female; 21 and 21, genitals of male imago, ventral view (holotype); 23, tergalius III (second existent); 24, middle part of anterior margin of labrum, ventral view. 25 26, genitals and enlarged penis of Choroterpides exiguus (Eaton 1884 [Thraulus]) (from Eaton 1884: Pl. 13, Fig. 20 * 2). 27, Choroterpes (Dilatognathus) major, tergalius III (second existent). 28, Choroterpes (Dilatognathus) minor, the same." httpUri="https://zenodo.org/record/231540/files/figure.png" pageId="6">Fig. 24</figureCitation>
); proximal transverse row of setae (shifted proximally, as in all
<emphasis id="B91FCC23FFADF341612DF8CA83B4E920" box="[570,730,1848,1873]" italics="true" pageId="6">Dilatognathus</emphasis>
) interrupted medially; instead of distal transverse row, there is a wide stripe of irregularly situated setae, smaller than setae of the proximal transverse row; most part of dorsal surface with irregularly situated long and thin setae, somewhat smaller than setae of the distal stripe. Maxilla without dentiseta at least beginning from instar L n-3; with maxillary tusk at least beginning from instar L n-3; without ventro-apical flange beginning from instar L n-2. In both sexes maxilla of pen-pen-penultimolarva (L n-3) with wellexpressed maxillary tusk; ventro-apical flange either small (
<figureCitation id="13500CB4FFACF3406058FF6582CAEEC1" box="[847,932,151,176]" captionStart="FIGURES 19 28" captionStartId="5.[151,264,1784,1807]" captionTargetBox="[155,1436,207,1738]" captionTargetId="figure@5.[151,1436,193,1763]" captionTargetPageId="5" captionText="FIGURES 19 28. Choroterpes (Dilatognathus) spp. 19 24, Ch. (D.) bogori: 19 and 20, apex of maxilla in pen-penpenultimate larval instar (ventral view; tusk, ventro-apical flange and vestiges of pectinate ventral setae are shown, other setae not shown): 19, male; 20, female; 21 and 21, genitals of male imago, ventral view (holotype); 23, tergalius III (second existent); 24, middle part of anterior margin of labrum, ventral view. 25 26, genitals and enlarged penis of Choroterpides exiguus (Eaton 1884 [Thraulus]) (from Eaton 1884: Pl. 13, Fig. 20 * 2). 27, Choroterpes (Dilatognathus) major, tergalius III (second existent). 28, Choroterpes (Dilatognathus) minor, the same." httpUri="https://zenodo.org/record/231540/files/figure.png" pageId="7">Fig. 20</figureCitation>
) or smoothed out (
<figureCitation id="13500CB4FFACF3406790FF6585B0EEC1" box="[1159,1246,151,176]" captionStart="FIGURES 19 28" captionStartId="5.[151,264,1784,1807]" captionTargetBox="[155,1436,207,1738]" captionTargetId="figure@5.[151,1436,193,1763]" captionTargetPageId="5" captionText="FIGURES 19 28. Choroterpes (Dilatognathus) spp. 19 24, Ch. (D.) bogori: 19 and 20, apex of maxilla in pen-penpenultimate larval instar (ventral view; tusk, ventro-apical flange and vestiges of pectinate ventral setae are shown, other setae not shown): 19, male; 20, female; 21 and 21, genitals of male imago, ventral view (holotype); 23, tergalius III (second existent); 24, middle part of anterior margin of labrum, ventral view. 25 26, genitals and enlarged penis of Choroterpides exiguus (Eaton 1884 [Thraulus]) (from Eaton 1884: Pl. 13, Fig. 20 * 2). 27, Choroterpes (Dilatognathus) major, tergalius III (second existent). 28, Choroterpes (Dilatognathus) minor, the same." httpUri="https://zenodo.org/record/231540/files/figure.png" pageId="7">Fig. 19</figureCitation>
). In next instars (from L n-2 to L n) ventro-apical flange not expressed, and maxillary tusk becomes progressively longer at each instar (
<figureCitation id="13500CB4FFACF3406388FF10819DEE8A" box="[159,243,226,251]" captionStart="FIGURES 16 18" captionStartId="4.[151,264,1940,1963]" captionTargetBox="[193,1408,353,1907]" captionTargetId="figure@4.[193,1437,353,1908]" captionTargetPageId="4" captionText="FIGURES 16 18. Choroterpes (Dilatognathus) bogori, maxilla of penultimolarva at various stages of transformation preceding molt to ultimolarva (slides in Canadian balsam): 16, female, early stage; 17, female, middle stage; 18, male, late stage." httpUri="https://zenodo.org/record/231539/files/figure.png" pageId="7">Fig. 18</figureCitation>
). Three vestiges of pectinate setae present in all instars. Legs as in
<taxonomicName id="4C6B6BB2FFACF34060FDFF1085A9EE8A" box="[1002,1223,226,251]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="7" phylum="Arthropoda" rank="species" species="cataractae" subGenus="Dilatognathus">
<emphasis id="B91FCC23FFACF34060FDFF108565EE8A" box="[1002,1035,226,251]" italics="true" pageId="7">Ch</emphasis>
. (
<emphasis id="B91FCC23FFACF3406734FF168559EE8A" box="[1059,1079,228,251]" italics="true" pageId="7">D</emphasis>
.)
<emphasis id="B91FCC23FFACF3406758FF1685A9EE8A" box="[1103,1223,228,251]" italics="true" pageId="7">cataractae</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFFA6DC0FFACF34067C1FF108193EF6E" author="Kluge" pageId="7" refString="Kluge, N. J. (2012) Contribution to the knowledge of Choroterpes (Ephemeroptera, Leptophlebiidae). Russian Entomological Journal, 21 (3), 273 - 306." type="journal article" year="2012">Kluge 2012: Figs 147151</bibRefCitation>
): Femur of fore leg widest proximally, femora of middle and hind legs widest at middle. Femora with irregularly situated stout setae of various lengths; dorsal side of fore femur with few setae, dorsal side of middle and hind femora with many setae. Ventral side of hind femur with irregular row of curved bipectinate setae. Stout pointed bipectinate setae on inner side of fore tibia situated densely and irregularly. Inner side of middle and hind tibia with sparse row of stout setae; dorsal side of middle tibia with row of longer setae; outer side of hind tibia with stout setae of variable length, as on outer side of femur. Outer sides of all tibiae with irregularly situated thin hairs, whose length exceed tibia width. Abdominal segments VIII and IX with posterolateral spines short and obtuse. In all instars including ultimolarva, hind margins of abdominal terga III without regular row of denticles (
<figureCitation id="13500CB4FFACF340662EFDF084E1EC6A" box="[1337,1423,514,539]" captionStart="FIGURES 34 39" captionStartId="8.[151,264,1934,1957]" captionTargetBox="[187,1399,447,1886]" captionTargetId="figure@8.[187,1400,447,1913]" captionTargetPageId="8" captionText="FIGURES 34 39. Choroterpes (Dilatognathus) spp. 34 37, median part of postrior margin of abdominal tergum of ultimolarva: 34 35, Ch. (D.) bogori; 36 37, Ch. (D.) cataractae; 34 and 36, tergum II; 35 and 37, tergum VII. 38 39, Ch. (D.) bogori: 38, exuviae of right half of subimaginal mesonotum; 39, egg." httpUri="https://zenodo.org/record/231542/files/figure.png" pageId="7">Fig. 34</figureCitation>
); terga IIIVII with row of very small, thin, spine-like denticles only in middle part (
<figureCitation id="13500CB4FFACF340672FFDD485E5EC4E" box="[1080,1163,550,575]" captionStart="FIGURES 34 39" captionStartId="8.[151,264,1934,1957]" captionTargetBox="[187,1399,447,1886]" captionTargetId="figure@8.[187,1400,447,1913]" captionTargetPageId="8" captionText="FIGURES 34 39. Choroterpes (Dilatognathus) spp. 34 37, median part of postrior margin of abdominal tergum of ultimolarva: 34 35, Ch. (D.) bogori; 36 37, Ch. (D.) cataractae; 34 and 36, tergum II; 35 and 37, tergum VII. 38 39, Ch. (D.) bogori: 38, exuviae of right half of subimaginal mesonotum; 39, egg." httpUri="https://zenodo.org/record/231542/files/figure.png" pageId="7">Fig. 35</figureCitation>
); terga VIIIX with row of somewhat longer denticles; sterna lack denticles. Tergalii IIVII with apical processes gradually narrowed toward apex (
<figureCitation id="13500CB4FFACF3406225FD9C80E9ECF6" box="[306,391,622,647]" captionStart="FIGURES 19 28" captionStartId="5.[151,264,1784,1807]" captionTargetBox="[155,1436,207,1738]" captionTargetId="figure@5.[151,1436,193,1763]" captionTargetPageId="5" captionText="FIGURES 19 28. Choroterpes (Dilatognathus) spp. 19 24, Ch. (D.) bogori: 19 and 20, apex of maxilla in pen-penpenultimate larval instar (ventral view; tusk, ventro-apical flange and vestiges of pectinate ventral setae are shown, other setae not shown): 19, male; 20, female; 21 and 21, genitals of male imago, ventral view (holotype); 23, tergalius III (second existent); 24, middle part of anterior margin of labrum, ventral view. 25 26, genitals and enlarged penis of Choroterpides exiguus (Eaton 1884 [Thraulus]) (from Eaton 1884: Pl. 13, Fig. 20 * 2). 27, Choroterpes (Dilatognathus) major, tergalius III (second existent). 28, Choroterpes (Dilatognathus) minor, the same." httpUri="https://zenodo.org/record/231540/files/figure.png" pageId="7">Fig. 23</figureCitation>
); tergalii IIV of equal size and shape, tergalius VI smaller, tergalius VII smallest and lacks costal process of dorsal lamella; anal-proximal projection of ventral lamella poorly expressed on tergalius II, better developed on tergalii IIIVII. Protogonostyli of male larva short and narrowly separated; protopenes convergent, completely hidden under sternum XI; female larva has hind margin of abdominal sternum IX slightly incised [as in
<taxonomicName id="4C6B6BB2FFACF3406380FD0C801CED66" box="[151,370,766,791]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="7" phylum="Arthropoda" rank="species" species="cataractae" subGenus="Dilatognathus">
<emphasis id="B91FCC23FFACF3406380FD0C81D6ED66" box="[151,184,766,791]" italics="true" pageId="7">Ch</emphasis>
. (
<emphasis id="B91FCC23FFACF34063D8FCF2818DED66" box="[207,227,768,791]" italics="true" pageId="7">D</emphasis>
.)
<emphasis id="B91FCC23FFACF34063EDFCF2801CED66" box="[250,370,768,791]" italics="true" pageId="7">cataractae</emphasis>
</taxonomicName>
] (
<bibRefCitation id="EFFA6DC0FFACF340629DFD0C83D5ED66" author="Kluge" box="[394,699,766,791]" pageId="7" refString="Kluge, N. J. (2012) Contribution to the knowledge of Choroterpes (Ephemeroptera, Leptophlebiidae). Russian Entomological Journal, 21 (3), 273 - 306." type="journal article" year="2012">Kluge 2012: Figs 143, 144</bibRefCitation>
).
</paragraph>
<paragraph id="8BD41031FFACF34063D0FCD08224EDD6" blockId="7.[151,1436,151,2016]" pageId="7">
<emphasis id="B91FCC23FFACF34063D0FCD08050ED4A" box="[199,318,802,827]" italics="true" pageId="7">Subimago.</emphasis>
CUTICULAR COLORATION: Cuticle of thorax at most light with brown sclerites; mesonotum at most brown; most part of medioscutum and submedioscutum brown, posterior scutal protuberance brown (
<figureCitation id="13500CB4FFACF3406620FCB484E1ED2E" box="[1335,1423,838,863]" captionStart="FIGURES 34 39" captionStartId="8.[151,264,1934,1957]" captionTargetBox="[187,1399,447,1886]" captionTargetId="figure@8.[187,1400,447,1913]" captionTargetPageId="8" captionText="FIGURES 34 39. Choroterpes (Dilatognathus) spp. 34 37, median part of postrior margin of abdominal tergum of ultimolarva: 34 35, Ch. (D.) bogori; 36 37, Ch. (D.) cataractae; 34 and 36, tergum II; 35 and 37, tergum VII. 38 39, Ch. (D.) bogori: 38, exuviae of right half of subimaginal mesonotum; 39, egg." httpUri="https://zenodo.org/record/231542/files/figure.png" pageId="7">Fig. 38</figureCitation>
). Cuticle of legs light, with diffusive brownish longitudinal stripes. Wings light brownish. Cuticle of all abdominal terga and sterna colorless; cuticle of gonostyli light brownish.
</paragraph>
<paragraph id="8BD41031FFACF34063D0FC4185FFEDBA" blockId="7.[151,1436,151,2016]" box="[199,1169,946,971]" pageId="7">HYPODERMAL COLORATION: Abdomen and femora with brown coloration as in imago.</paragraph>
<paragraph id="8BD41031FFACF34063D0FC2A853BEA62" blockId="7.[151,1436,151,2016]" pageId="7">TEXTURE: On each leg 1st tarsomere (shortened and fused with tibia) covered by microtrichiae of the same shape as microtrichiae on tibia; 2nd5th tarsomeres covered by pointed microlepides.</paragraph>
<paragraph id="8BD41031FFACF34063D0FBD28243EBEE" blockId="7.[151,1436,151,2016]" pageId="7">
<emphasis id="B91FCC23FFACF34063D0FBD28033EA46" box="[199,349,1054,1079]" italics="true" pageId="7">Imago, male.</emphasis>
Head light brown with ocher; dorsal eyes not elevated, brownish-orange. Thorax light brown with ocher. Fore and hind wings with membrane nearly colorless, evenly tinged with light ocher; veins from light brown to colorless; pterostigma with 915 simple cross veins.
<collectingCountry id="F37C50A1FFACF3406048FB9482F9EA0E" box="[863,919,1126,1151]" name="India" pageId="7">Hind</collectingCountry>
wing with blunt costal projection. Femora of all legs light ocher-brown with contrasting dark brown band in middle (
<figureCitation id="13500CB4FFACF34060F0FB788550EAD2" box="[999,1086,1162,1187]" captionStart="FIGURES 29 33" captionStartId="6.[151,264,1213,1236]" captionTargetBox="[151,1415,193,1192]" captionTargetId="figure@6.[151,1436,193,1192]" captionTargetPageId="6" captionText="FIGURES 29 33. Choroterpes (Dilatognathus) spp. 29, Ch. (D.) bogori, abdomen of male imago, spread on slide (holotype); 30, Ch. (D.) cataractae, the same (holotype); 31, Ch. (D.) bogori, hind leg of male imago (holotype); 32, Ch. (D.) cataractae, the same (holotype); 33, Ch. (D.) major, hind leg of mature larva with developing subimaginal leg." httpUri="https://zenodo.org/record/231541/files/figure.png" pageId="7">Fig. 31</figureCitation>
). On fore leg tibia and tarsus entirely light brown. On middle and hind leg tibia lighter, with brown base (
<figureCitation id="13500CB4FFACF3406714FB5C8536EAB6" box="[1027,1112,1198,1223]" captionStart="FIGURES 29 33" captionStartId="6.[151,264,1213,1236]" captionTargetBox="[151,1415,193,1192]" captionTargetId="figure@6.[151,1436,193,1192]" captionTargetPageId="6" captionText="FIGURES 29 33. Choroterpes (Dilatognathus) spp. 29, Ch. (D.) bogori, abdomen of male imago, spread on slide (holotype); 30, Ch. (D.) cataractae, the same (holotype); 31, Ch. (D.) bogori, hind leg of male imago (holotype); 32, Ch. (D.) cataractae, the same (holotype); 33, Ch. (D.) major, hind leg of mature larva with developing subimaginal leg." httpUri="https://zenodo.org/record/231541/files/figure.png" pageId="7">Fig. 31</figureCitation>
). Abdominal terga IIVII in anterior-median part light ocher, in posterior and lateral parts brown with ocher blanks (
<figureCitation id="13500CB4FFACF340677AFB2085AFEA9A" box="[1133,1217,1234,1259]" captionStart="FIGURES 29 33" captionStartId="6.[151,264,1213,1236]" captionTargetBox="[151,1415,193,1192]" captionTargetId="figure@6.[151,1436,193,1192]" captionTargetPageId="6" captionText="FIGURES 29 33. Choroterpes (Dilatognathus) spp. 29, Ch. (D.) bogori, abdomen of male imago, spread on slide (holotype); 30, Ch. (D.) cataractae, the same (holotype); 31, Ch. (D.) bogori, hind leg of male imago (holotype); 32, Ch. (D.) cataractae, the same (holotype); 33, Ch. (D.) major, hind leg of mature larva with developing subimaginal leg." httpUri="https://zenodo.org/record/231541/files/figure.png" pageId="7">Fig. 29</figureCitation>
). Abdominal sterna IVIII light. Abdominal sternum IX and styliger ocher with brown, gonostyli light brown. Styliger with nearly straight posterior margin. 1st+2nd segments of gonostylus with inner margin angulated. Penis lobes apically pressed together; each lobe with apical projection moderately long, conic, slightly bent medially-dorsally, rounded apically; denticles absent, instead of them few round concavities on dorsal side of penis close to apex (
<figureCitation id="13500CB4FFACF340662EFA9084E1EB0A" box="[1337,1423,1378,1403]" captionStart="FIGURES 19 28" captionStartId="5.[151,264,1784,1807]" captionTargetBox="[155,1436,207,1738]" captionTargetId="figure@5.[151,1436,193,1763]" captionTargetPageId="5" captionText="FIGURES 19 28. Choroterpes (Dilatognathus) spp. 19 24, Ch. (D.) bogori: 19 and 20, apex of maxilla in pen-penpenultimate larval instar (ventral view; tusk, ventro-apical flange and vestiges of pectinate ventral setae are shown, other setae not shown): 19, male; 20, female; 21 and 21, genitals of male imago, ventral view (holotype); 23, tergalius III (second existent); 24, middle part of anterior margin of labrum, ventral view. 25 26, genitals and enlarged penis of Choroterpides exiguus (Eaton 1884 [Thraulus]) (from Eaton 1884: Pl. 13, Fig. 20 * 2). 27, Choroterpes (Dilatognathus) major, tergalius III (second existent). 28, Choroterpes (Dilatognathus) minor, the same." httpUri="https://zenodo.org/record/231540/files/figure.png" pageId="7">Fig. 21</figureCitation>
). Caudalii light brown, with dark brown ring at each joining.
</paragraph>
<paragraph id="8BD41031FFACF34063D0FA5E843EEBB2" blockId="7.[151,1436,151,2016]" box="[199,1360,1450,1475]" pageId="7">
<emphasis id="B91FCC23FFACF34063D0FA5E8001EBB2" box="[199,367,1450,1475]" italics="true" pageId="7">Imago, female.</emphasis>
Coloration as in male. Posterior plate of abdominal sternum IX apically slightly incised.
</paragraph>
<paragraph id="8BD41031FFACF34063D0FA228267E87A" blockId="7.[151,1436,151,2016]" pageId="7">
<emphasis id="B91FCC23FFACF34063D0FA228195EB96" box="[199,251,1488,1511]" italics="true" pageId="7">Egg.</emphasis>
Oval, 0.15 mm length. Chorion with evenly dispersed papillae and ridges going from one papilla to another; papillae small, on one pole larger (
<figureCitation id="13500CB4FFACF340616AFA008392E87A" box="[637,764,1522,1547]" captionStart-0="FIGURES 34 39" captionStart-1="FIGURES 40 43" captionStartId-0="8.[151,264,1934,1957]" captionStartId-1="9.[151,264,1169,1192]" captionTargetBox-0="[187,1399,447,1886]" captionTargetBox-1="[151,1436,193,1148]" captionTargetId-0="figure@8.[187,1400,447,1913]" captionTargetId-1="figure@9.[151,1436,193,1149]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURES 34 39. Choroterpes (Dilatognathus) spp. 34 37, median part of postrior margin of abdominal tergum of ultimolarva: 34 35, Ch. (D.) bogori; 36 37, Ch. (D.) cataractae; 34 and 36, tergum II; 35 and 37, tergum VII. 38 39, Ch. (D.) bogori: 38, exuviae of right half of subimaginal mesonotum; 39, egg." captionText-1="FIGURES 40 43. Choroterpes (Dilatognathus) bogori sp. n., eggs. 40, 41, total view; 42, 43, two poles of egg in Fig. 41." httpUri-0="https://zenodo.org/record/231542/files/figure.png" httpUri-1="https://zenodo.org/record/231543/files/figure.png" pageId="7">Fig. 3943</figureCitation>
).
</paragraph>
<paragraph id="8BD41031FFACF34063D0F9EA8285E85E" blockId="7.[151,1436,151,2016]" box="[199,1003,1558,1583]" pageId="7">
<emphasis id="B91FCC23FFACF34063D0F9EA8026E85E" box="[199,328,1560,1583]" italics="true" pageId="7">Dimension.</emphasis>
Size of both sexes greatly varies: fore wing length
<quantity id="4C93BDD4FFACF3406091F9E48288E85E" box="[902,998,1558,1583]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="7" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">58 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C37143BAFFACF34063D0F9C883E0E976" pageId="7" type="distribution">
<paragraph id="8BD41031FFACF34063D0F9C880CEE823" blockId="7.[151,1436,151,2016]" box="[199,416,1594,1619]" pageId="7">
<emphasis id="B91FCC23FFACF34063D0F9C88037E822" bold="true" box="[199,345,1594,1619]" pageId="7">Distribution</emphasis>
. Java.
</paragraph>
<paragraph id="8BD41031FFACF34063D0F9AC83E0E976" blockId="7.[151,1436,151,2016]" pageId="7">
<emphasis id="B91FCC23FFACF34063D0F9AC8044E806" bold="true" box="[199,298,1630,1655]" pageId="7">Habitat.</emphasis>
Larvae were collected in rivers Ci Sadane and Ci Liwung at places with stony bottom and fast current. This species is most abundant upstream of the city, where water pollution is moderate; but is also present in center of Bogor (Ci Liwung in botanic garden), where the river is extremely polluted. In this respect,
<taxonomicName id="4C6B6BB2FFACF340662BF954818CE892" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="7" phylum="Arthropoda" rank="species" species="bogori" subGenus="Dilatognathus">
<emphasis id="B91FCC23FFACF340662BF9548433E8CE" box="[1340,1373,1702,1727]" italics="true" pageId="7">Ch</emphasis>
. (
<emphasis id="B91FCC23FFACF3406660F95A84E5E8CE" box="[1399,1419,1704,1727]" italics="true" pageId="7">D</emphasis>
.)
<emphasis id="B91FCC23FFACF3406380F938818CE892" box="[151,226,1738,1763]" italics="true" pageId="7">bogori</emphasis>
</taxonomicName>
markedly differs from
<taxonomicName id="4C6B6BB2FFACF34062F9F93883A3E892" box="[494,717,1738,1763]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="7" phylum="Arthropoda" rank="species" species="cataractae" subGenus="Dilatognathus">
<emphasis id="B91FCC23FFACF34062F9F9388361E892" box="[494,527,1738,1763]" italics="true" pageId="7">Ch</emphasis>
. (
<emphasis id="B91FCC23FFACF340613FF93E8352E892" box="[552,572,1740,1763]" italics="true" pageId="7">D</emphasis>
.)
<emphasis id="B91FCC23FFACF3406142F93E83A3E892" box="[597,717,1740,1763]" italics="true" pageId="7">cataractae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6B6BB2FFACF3406010F93882DFE892" box="[775,945,1738,1763]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="7" phylum="Arthropoda" rank="species" species="major" subGenus="Dilatognathus">
<emphasis id="B91FCC23FFACF3406010F9388246E892" box="[775,808,1738,1763]" italics="true" pageId="7">Ch</emphasis>
. (
<emphasis id="B91FCC23FFACF3406055F93E8238E892" box="[834,854,1740,1763]" italics="true" pageId="7">D</emphasis>
.)
<emphasis id="B91FCC23FFACF3406078F93E82DFE892" box="[879,945,1740,1763]" italics="true" pageId="7">major</emphasis>
</taxonomicName>
, whose larvae were found only in selected areas of clean rivers under certain waterfalls.
</paragraph>
</subSubSection>
<subSubSection id="C37143BAFFACF34F63D0F8E08243EFF9" lastPageId="8" pageId="7" type="diagnosis">
<paragraph id="8BD41031FFACF34F63D0F8E08243EFF9" blockId="7.[151,1436,151,2016]" lastBlockId="8.[151,1436,151,392]" lastPageId="8" pageId="7">
<emphasis id="B91FCC23FFACF34063D0F8E0802FE95A" bold="true" box="[199,321,1810,1835]" pageId="7">Diagnosis.</emphasis>
Male and female imagoes differ from
<taxonomicName id="4C6B6BB2FFACF3406003F8E08294E95A" box="[788,1018,1810,1835]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="7" phylum="Arthropoda" rank="species" species="cataractae" subGenus="Dilatognathus">
<emphasis id="B91FCC23FFACF3406003F8E0825BE95A" box="[788,821,1810,1835]" italics="true" pageId="7">Ch</emphasis>
. (
<emphasis id="B91FCC23FFACF3406046F8E6820BE95A" box="[849,869,1812,1835]" italics="true" pageId="7">D</emphasis>
.)
<emphasis id="B91FCC23FFACF3406095F8E68294E95A" box="[898,1018,1812,1835]" italics="true" pageId="7">cataractae</emphasis>
</taxonomicName>
by absence of brown coloration in proximal part of subcostal field of fore wing. Larva of penultimate and previous instars differs from
<taxonomicName id="4C6B6BB2FFACF340662CF8C48061E902" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="7" phylum="Arthropoda" rank="species" species="cataractae" subGenus="Dilatognathus">
<emphasis id="B91FCC23FFACF340662CF8C48432E93E" box="[1339,1372,1846,1871]" italics="true" pageId="7">Ch</emphasis>
. (
<emphasis id="B91FCC23FFACF3406660F8CA84E5E93E" box="[1399,1419,1848,1871]" italics="true" pageId="7">D</emphasis>
.)
<emphasis id="B91FCC23FFACF3406380F8AE8061E902" box="[151,271,1884,1907]" italics="true" pageId="7">cataractae</emphasis>
</taxonomicName>
by structure of maxilla (compare
<figureCitation id="13500CB4FFACF3406183F8A8827AE902" box="[660,788,1882,1907]" captionStart="FIGURES 19 28" captionStartId="5.[151,264,1784,1807]" captionTargetBox="[155,1436,207,1738]" captionTargetId="figure@5.[151,1436,193,1763]" captionTargetPageId="5" captionText="FIGURES 19 28. Choroterpes (Dilatognathus) spp. 19 24, Ch. (D.) bogori: 19 and 20, apex of maxilla in pen-penpenultimate larval instar (ventral view; tusk, ventro-apical flange and vestiges of pectinate ventral setae are shown, other setae not shown): 19, male; 20, female; 21 and 21, genitals of male imago, ventral view (holotype); 23, tergalius III (second existent); 24, middle part of anterior margin of labrum, ventral view. 25 26, genitals and enlarged penis of Choroterpides exiguus (Eaton 1884 [Thraulus]) (from Eaton 1884: Pl. 13, Fig. 20 * 2). 27, Choroterpes (Dilatognathus) major, tergalius III (second existent). 28, Choroterpes (Dilatognathus) minor, the same." httpUri="https://zenodo.org/record/231540/files/figure.png" pageId="7">Figs 19, 20</figureCitation>
and 613): pen-pen-penultimolarva of both sexes differs by presence of maxillary tusk and absence of dentiseta; pen-penultimolarva of both sexes differs by absence of dentiseta and ventro-apical flange; male penultimolarva differs by absence of dentiseta and ventro-apical flange. Imagoes, subimagoes and larvae differ from
<taxonomicName id="4C6B6BB2FFACF34061B4F834823FE9AE" box="[675,849,1990,2015]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="7" phylum="Arthropoda" rank="species" species="major" subGenus="Dilatognathus">
<emphasis id="B91FCC23FFACF34061B4F83483AAE9AE" box="[675,708,1990,2015]" italics="true" pageId="7">Ch</emphasis>
. (
<emphasis id="B91FCC23FFACF34061C9F83A839CE9AE" box="[734,754,1992,2015]" italics="true" pageId="7">D</emphasis>
.)
<emphasis id="B91FCC23FFACF3406019F83A823FE9AE" box="[782,849,1992,2015]" italics="true" pageId="7">major</emphasis>
</taxonomicName>
by light femora with well-expressed dark band at middle (compare
<figureCitation id="13500CB4FFA3F34F6271FF658370EEC1" box="[358,542,151,176]" captionStart="FIGURES 29 33" captionStartId="6.[151,264,1213,1236]" captionTargetBox="[151,1415,193,1192]" captionTargetId="figure@6.[151,1436,193,1192]" captionTargetPageId="6" captionText="FIGURES 29 33. Choroterpes (Dilatognathus) spp. 29, Ch. (D.) bogori, abdomen of male imago, spread on slide (holotype); 30, Ch. (D.) cataractae, the same (holotype); 31, Ch. (D.) bogori, hind leg of male imago (holotype); 32, Ch. (D.) cataractae, the same (holotype); 33, Ch. (D.) major, hind leg of mature larva with developing subimaginal leg." httpUri="https://zenodo.org/record/231541/files/figure.png" pageId="8">Figs 31 and 33</figureCitation>
). Imagoes, subimagoes and larvae differ from
<taxonomicName id="4C6B6BB2FFA3F34F675DFF658594EEC1" box="[1098,1274,151,176]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="8" phylum="Arthropoda" rank="species" species="major" subGenus="Dilatognathus">
<emphasis id="B91FCC23FFA3F34F675DFF658505EEC1" box="[1098,1131,151,176]" italics="true" pageId="8">Ch</emphasis>
. (
<emphasis id="B91FCC23FFA3F34F6790FF6B85F5EEC1" box="[1159,1179,153,176]" italics="true" pageId="8">D</emphasis>
.)
<emphasis id="B91FCC23FFA3F34F67A0FF6B8594EEC1" box="[1207,1274,153,176]" italics="true" pageId="8">major</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6B6BB2FFA3F34F662DFF658061EEA5" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="8" phylum="Arthropoda" rank="species" species="cataractae" subGenus="Dilatognathus">
<emphasis id="B91FCC23FFA3F34F662DFF658435EEC1" box="[1338,1371,151,176]" italics="true" pageId="8">Ch</emphasis>
. (
<emphasis id="B91FCC23FFA3F34F6660FF6B84E5EEC1" box="[1399,1419,153,176]" italics="true" pageId="8">D</emphasis>
.)
<emphasis id="B91FCC23FFA3F34F6380FF4F8061EEA5" box="[151,271,189,212]" italics="true" pageId="8">cataractae</emphasis>
</taxonomicName>
by light median part of abdominal terga (compare
<figureCitation id="13500CB4FFA3F34F605AFF4E8296EEA5" box="[845,1016,188,213]" captionStart="FIGURES 29 33" captionStartId="6.[151,264,1213,1236]" captionTargetBox="[151,1415,193,1192]" captionTargetId="figure@6.[151,1436,193,1192]" captionTargetPageId="6" captionText="FIGURES 29 33. Choroterpes (Dilatognathus) spp. 29, Ch. (D.) bogori, abdomen of male imago, spread on slide (holotype); 30, Ch. (D.) cataractae, the same (holotype); 31, Ch. (D.) bogori, hind leg of male imago (holotype); 32, Ch. (D.) cataractae, the same (holotype); 33, Ch. (D.) major, hind leg of mature larva with developing subimaginal leg." httpUri="https://zenodo.org/record/231541/files/figure.png" pageId="8">Figs 29 and 30</figureCitation>
). Larva of last instar differs from
<taxonomicName id="4C6B6BB2FFA3F34F6663FF4E802AEE89" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="8" phylum="Arthropoda" rank="species" species="cataractae" subGenus="Dilatognathus">
<emphasis id="B91FCC23FFA3F34F6663FF4E84FBEEA4" box="[1396,1429,188,213]" italics="true" pageId="8">Ch</emphasis>
. (
<emphasis id="B91FCC23FFA3F34F63B7FF1381DAEE89" box="[160,180,225,248]" italics="true" pageId="8">D</emphasis>
.)
<emphasis id="B91FCC23FFA3F34F63DBFF13802AEE89" box="[204,324,225,248]" italics="true" pageId="8">cataractae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6B6BB2FFA3F34F626BFF2D834AEE89" box="[380,548,223,248]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="8" phylum="Arthropoda" rank="species" species="major" subGenus="Dilatognathus">
<emphasis id="B91FCC23FFA3F34F626BFF2D80F3EE89" box="[380,413,223,248]" italics="true" pageId="8">Ch</emphasis>
. (
<emphasis id="B91FCC23FFA3F34F62A2FF1380A7EE89" box="[437,457,225,248]" italics="true" pageId="8">D</emphasis>
.)
<emphasis id="B91FCC23FFA3F34F62F6FF13834AEE89" box="[481,548,225,248]" italics="true" pageId="8">major</emphasis>
</taxonomicName>
by absence of regular row of spines on posterior margins of abdominal terga I and II and by smaller spines on abdominal terga IIIIX (compare
<figureCitation id="13500CB4FFA3F34F6092FEF685E9EF6C" box="[901,1159,260,285]" captionStart="FIGURES 34 39" captionStartId="8.[151,264,1934,1957]" captionTargetBox="[187,1399,447,1886]" captionTargetId="figure@8.[187,1400,447,1913]" captionTargetPageId="8" captionText="FIGURES 34 39. Choroterpes (Dilatognathus) spp. 34 37, median part of postrior margin of abdominal tergum of ultimolarva: 34 35, Ch. (D.) bogori; 36 37, Ch. (D.) cataractae; 34 and 36, tergum II; 35 and 37, tergum VII. 38 39, Ch. (D.) bogori: 38, exuviae of right half of subimaginal mesonotum; 39, egg." httpUri="https://zenodo.org/record/231542/files/figure.png" pageId="8">Figs 3435 and 3637</figureCitation>
). Larva differs from
<taxonomicName id="4C6B6BB2FFA3F34F6663FEF68074EF31" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="8" phylum="Arthropoda" rank="species" species="nigella" subGenus="Dilatognathus">
<emphasis id="B91FCC23FFA3F34F6663FEF684FBEF6C" box="[1396,1429,260,285]" italics="true" pageId="8">Ch</emphasis>
. (
<emphasis id="B91FCC23FFA3F34F63B7FEDB81DAEF31" box="[160,180,297,320]" italics="true" pageId="8">D</emphasis>
.)
<emphasis id="B91FCC23FFA3F34F63DBFED58074EF31" box="[204,282,295,320]" italics="true" pageId="8">nigella</emphasis>
</taxonomicName>
in shape of labrum, and from
<taxonomicName id="4C6B6BB2FFA3F34F617AFED5827AEF31" box="[621,788,295,320]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="8" phylum="Arthropoda" rank="species" species="minor" subGenus="Dilatognathus">
<emphasis id="B91FCC23FFA3F34F617AFED583E0EF31" box="[621,654,295,320]" italics="true" pageId="8">Ch</emphasis>
. (
<emphasis id="B91FCC23FFA3F34F61B1FEDB83D4EF31" box="[678,698,297,320]" italics="true" pageId="8">D</emphasis>
.)
<emphasis id="B91FCC23FFA3F34F61C6FEDB827AEF31" box="[721,788,297,320]" italics="true" pageId="8">minor</emphasis>
</taxonomicName>
in shape of tergalii. Structure of egg (
<figureCitation id="13500CB4FFA3F34F67A9FED5847EEF31" box="[1214,1296,295,320]" captionStart="FIGURES 34 39" captionStartId="8.[151,264,1934,1957]" captionTargetBox="[187,1399,447,1886]" captionTargetId="figure@8.[187,1400,447,1913]" captionTargetPageId="8" captionText="FIGURES 34 39. Choroterpes (Dilatognathus) spp. 34 37, median part of postrior margin of abdominal tergum of ultimolarva: 34 35, Ch. (D.) bogori; 36 37, Ch. (D.) cataractae; 34 and 36, tergum II; 35 and 37, tergum VII. 38 39, Ch. (D.) bogori: 38, exuviae of right half of subimaginal mesonotum; 39, egg." httpUri="https://zenodo.org/record/231542/files/figure.png" pageId="8">Fig. 39</figureCitation>
) is the same as in
<taxonomicName id="4C6B6BB2FFA3F34F63C6FEBE80C2EF15" box="[209,428,332,357]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="8" phylum="Arthropoda" rank="species" species="cataractae" subGenus="Dilatognathus">
<emphasis id="B91FCC23FFA3F34F63C6FEBE819CEF14" box="[209,242,332,357]" italics="true" pageId="8">Ch</emphasis>
. (
<emphasis id="B91FCC23FFA3F34F621EFEBF8073EF15" box="[265,285,333,356]" italics="true" pageId="8">D</emphasis>
.)
<emphasis id="B91FCC23FFA3F34F6223FEBF80C2EF15" box="[308,428,333,356]" italics="true" pageId="8">cataractae</emphasis>
</taxonomicName>
and differs from
<taxonomicName id="4C6B6BB2FFA3F34F617AFEBE8270EF14" box="[621,798,332,357]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="8" phylum="Arthropoda" rank="species" species="nigella" subGenus="Dilatognathus">
<emphasis id="B91FCC23FFA3F34F617AFEBE83E0EF14" box="[621,654,332,357]" italics="true" pageId="8">Ch</emphasis>
. (
<emphasis id="B91FCC23FFA3F34F61B2FEBF83D7EF15" box="[677,697,333,356]" italics="true" pageId="8">D</emphasis>
.)
<emphasis id="B91FCC23FFA3F34F61C7FEBE8270EF14" box="[720,798,332,357]" italics="true" pageId="8">nigella</emphasis>
</taxonomicName>
, whose papillae have equal size on all egg surface (
<bibRefCitation id="EFFA6DC0FFA3F34F664AFEBE80E3EFF9" author="Kang" pageId="8" refString="Kang, S. C. &amp; Yang, C. T. (1994) Leptophlebiidae of Taiwan (Ephemeroptera). Journal of Taiwan Museum, 47 (1), 57 - 81." type="journal article" year="1994">Kang &amp; Yang 1994: Fig. 12</bibRefCitation>
); eggs of other species are unknown.
</paragraph>
</subSubSection>
</treatment>
</document>