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<mods:title id="08EBA8FE251D057DDC55176BC9DC9315">Taxonomic revision of the genus Deltepilissus Pereira, 1949 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini)</mods:title>
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<mods:affiliation id="28139480FBC952E6EB7ED2F0F1D38103">Universidade Federal Rural de Pernambuco, Unidade Acadêmica de Serra Talhada. Avenida Gregório Ferraz Nogueira, s / n. Serra Talhada, PE, 56909 - 535, Brazil. &amp; Universidade Federal do Pará, Instituto de Ciências Biológicas, setor de Zoologia, Campus Belém. Rua Augusto Corrêa, 01, Guamá. Belém, PA, 66075 - 110, Brazil.</mods:affiliation>
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<taxonomicName id="4C33E0E3FFD8FFAF09C9E001FDF8FEB9" ID-CoL="34K62" authority="(Harold, 1880)" baseAuthorityName="Harold" baseAuthorityYear="1880" box="[151,625,314,341]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="diabolicus">
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<emphasis id="B9474772FFD8FFAF09C9E001FE3DFEB8" bold="true" box="[151,436,314,341]" italics="true" pageId="4" pageNumber="87">Deltepilissus diabolicus</emphasis>
(
<bibRefCitation id="EFA2E691FFD8FFAF089DE001FDE1FEB9" author="Harold, E." box="[451,616,314,340]" pageId="4" pageNumber="87" pagination="148 - 171" refId="ref9058" refString="Harold, E. (1880) Einige Neue Coleopteren. Mittheilungen des Munchener Entomologischen Vereins, 4, 148 - 171." type="journal article" year="1880">Harold, 1880</bibRefCitation>
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(
<figureCitation id="130887E5FFD8FFAF09C1E065FE9DFE94" box="[159,276,350,377]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="4" pageNumber="87">Figs 1A–I</figureCitation>
, 
<figureCitation id="130887E5FFD8FFAF0841E064FD96FE94" box="[287,543,351,377]" captionStart="FIGURE 3" captionStartId="9.[151,250,1773,1797]" captionTargetBox="[231,1367,208,1740]" captionTargetId="figure-19@9.[214,1374,181,1750]" captionTargetPageId="9" captionText="FIGURE 3. Details of the male genitalia in Deltepilissus. A–C, Deltepilissus diabolicus: A, parameres, dorsal view; B, parameres, ventral view; C, aedeagus, lateral view. D–F. Deltepilissus infernalis: D, parameres, dorsal view; E, parameres, ventral view; F, aedeagus, lateral view. G, Deltepilissus diabolicus superior right peripheral (SRP) endophallite. H, Deltepilissus infernalis superior right peripheral (SRP) endophallite. I, Deltepilissus diabolicus frontolateral peripheral (FLP) endophallite. J, Deltepilissus infernalis frontolateral peripheral (FLP) endophallite. K, Deltepilissus diabolicus additional endophallites (AE). L, Deltepilissus infernalis additional endophallites (AE). M, Deltepilissus diabolicus complex of axial and subaxial (A+SA) endophallites. N, Deltepilissus infernalis complex of axial and subaxial (A+SA) endophallites." figureDoi="http://doi.org/10.5281/zenodo.6388915" httpUri="https://zenodo.org/record/6388915/files/figure.png" pageId="4" pageNumber="87">3A–C, 3G, 3I, 3K, 3M</figureCitation>
, 
<figureCitation id="130887E5FFD8FFAF0B75E064FDF9FE94" box="[555,624,351,377]" captionStart="FIGURE 4" captionStartId="10.[151,250,1910,1934]" captionTargetBox="[278,1321,181,1886]" captionTargetId="figure-30@10.[267,1321,181,1886]" captionTargetPageId="10" captionText="FIGURE 4. Type specimens in Deltepilissus and respective label data. A–B, Lectotype of Canthon diabolicus Harold, 1880: A, body, dorsal view; B, labels. C–D, Lectotype of Canthon infernalis Harold, 1880: C, body, dorsal view; D, labels. E–H, holotype of Deltepilissus travassosi Pereira, 1949: E, body, dorsal view; F, head; G, left protibia, dorsal view; H, protibial and tarsal claws. Photographs A–D provided by Fernando Z. Vaz-de-Mello." figureDoi="http://doi.org/10.5281/zenodo.6388917" httpUri="https://zenodo.org/record/6388917/files/figure.png" pageId="4" pageNumber="87">4A–B</figureCitation>
, 
<figureCitation id="130887E5FFD8FFAF0B22E065FD49FE94" box="[636,704,350,377]" captionStart="FIGURE 5" captionStartId="11.[151,250,1103,1127]" captionTargetBox="[151,1436,181,1066]" captionTargetId="figure-19@11.[151,1436,181,1080]" captionTargetPageId="11" captionText="FIGURE 5. Known distribution of species of Deltepilissus. A, Records plotted on terrestrial biomes layer; B, records plotted on elevational range layer." figureDoi="http://doi.org/10.5281/zenodo.6388921" httpUri="https://zenodo.org/record/6388921/files/figure.png" pageId="4" pageNumber="87">5A–B</figureCitation>
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<treatmentCitation id="0A92BD71FFD8FFAF09C9E098FDA0FE56" author="Harold, E." box="[151,553,419,443]" page="150" pageId="4" pageNumber="87" year="1880">
<taxonomicName id="4C33E0E3FFD8FFAF09C9E098FDA0FE56" ID-CoL="QPC8" authority="Harold 1880: 150" authorityName="Harold" authorityPageNumber="150" authorityYear="1880" box="[151,553,419,443]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="diabolicus">
<emphasis id="B9474772FFD8FFAF09C9E098FEEAFE56" box="[151,355,419,443]" italics="true" pageId="4" pageNumber="87">Canthon diabolicus</emphasis>
<bibRefCitation id="EFA2E691FFD8FFAF0833E098FDA0FE56" author="Harold, E." box="[365,553,419,443]" pageId="4" pageNumber="87" pagination="148 - 171" refId="ref9058" refString="Harold, E. (1880) Einige Neue Coleopteren. Mittheilungen des Munchener Entomologischen Vereins, 4, 148 - 171." type="journal article" year="1880">Harold 1880: 150</bibRefCitation>
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(original description); 
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<bibRefCitation id="EFA2E691FFD8FFAF0A7EE098FC48FE56" author="Gillet, J. J. E." box="[800,961,419,443]" pageId="4" pageNumber="87" refId="ref8874" refString="Gillet, J. J. E. (1911) Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin, Germany, 100 pp." type="book" year="1911">Gillet 1911: 29</bibRefCitation>
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(catalog); 
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<bibRefCitation id="EFA2E691FFD8FFAF0D69E098FB7FFE56" author="Schmidt, A." box="[1079,1270,419,443]" pageId="4" pageNumber="87" pagination="61 - 103" refId="ref9438" refString="Schmidt, A. (1922) 1. Bestimmungstabelle der mir bekannten Canthon - Arten. 2. Verbreitungsgebiete der Canthon - Arten. 3. Neubeschreibungen von Canthon, Saprositis, Mendidius, Euparia und Ataenius. Archiv fur Naturgeschichte Abteilung A, 88 (3), 61 - 103." type="journal article" year="1922">Schmidt 1922: 67</bibRefCitation>
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, 74 (taxonomic remarks, distribution); 
<treatmentCitation id="0A92BD71FFD8FFAF08EBE0F9FD0EFE37" author="Balthasar, V." box="[437,647,450,474]" page="201" pageId="4" pageNumber="87" year="1939">
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(identification key); 
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<bibRefCitation id="EFA2E691FFD8FFAF0A3BE0F9FBDEFE37" author="Blackwelder, R." box="[869,1111,450,474]" pageId="4" pageNumber="87" pagination="189 - 341" refId="ref8489" refString="Blackwelder, R. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America. Part 2. Bulletin of the United States National Museum, 185, 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 199</bibRefCitation>
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(checklist); 
<treatmentCitation id="0A92BD71FFD8FFAF0D84E0F9FEA5FE15" author="Vulcano, M. A. &amp; Pereira, F." page="611" pageId="4" pageNumber="87" year="1964">
<bibRefCitation id="EFA2E691FFD8FFAF0D84E0F9FEA5FE15" author="Vulcano, M. A. &amp; Pereira, F." pageId="4" pageNumber="87" pagination="570 - 685" refId="ref10315" refString="Vulcano, M. A. &amp; Pereira, F. (1964) Catalogue of the Canthonini (Col. Scarab.) inhabiting the Western Hemisphere. Entomologische Arbeiten aus dem Museum G. Frey, 15, 570 - 685." type="journal article" year="1964">Vulcano &amp; Pereira 1964: 611</bibRefCitation>
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(catalog).
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<treatmentCitation id="0A92BD71FFD8FFAF09C9E0C4FD49FDFA" author="Halffter, G. &amp; Martinez, A." box="[151,704,511,535]" page="66" pageId="4" pageNumber="87" year="1977">
<taxonomicName id="4C33E0E3FFD8FFAF09C9E0C4FD49FDFA" ID-CoL="34K62" authority=": Halffter &amp; Martinez 1977: 66" authorityName=": Halffter &amp; Martinez" authorityPageNumber="66" authorityYear="1977" box="[151,704,511,535]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="diabolicus">
<emphasis id="B9474772FFD8FFAF09C9E0C4FE03FDFA" box="[151,394,511,535]" italics="true" pageId="4" pageNumber="87">Deltepilissus diabolicus</emphasis>
: 
<bibRefCitation id="EFA2E691FFD8FFAF08C9E0C4FD49FDFA" author="Halffter, G. &amp; Martinez, A." box="[407,704,511,535]" pageId="4" pageNumber="87" pagination="29 - 107" refId="ref8982" refString="Halffter, G. &amp; Martinez, A. (1977) Revision monografica de los Canthonina Americanos, IV parte. Clave para generos y subgeneros. Folia Entomologica Mexicana, 38, 29 - 107." type="journal article" year="1977">Halffter &amp; Martínez 1977: 66</bibRefCitation>
</taxonomicName>
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(transferred 
<taxonomicName id="4C33E0E3FFD8FFAF0A1DE0C4FC43FDFA" ID-CoL="34K62" authorityName=": Halffter &amp; Martinez" authorityYear="1977" box="[835,970,511,535]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="diabolicus">
<emphasis id="B9474772FFD8FFAF0A1DE0C4FC43FDFA" box="[835,970,511,535]" italics="true" pageId="4" pageNumber="87">C. diabolicus</emphasis>
</taxonomicName>
to the genus 
<taxonomicName id="4C33E0E3FFD8FFAF0D11E0C4FB5BFDFA" ID-CoL="42SD" authorityName="Pereira" authorityYear="1949" box="[1103,1234,511,535]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis id="B9474772FFD8FFAF0D11E0C4FB5BFDFA" box="[1103,1234,511,535]" italics="true" pageId="4" pageNumber="87">Deltepilissus</emphasis>
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, checklist); Vaz-de-Mello 2000: 192 (checklist); 
<treatmentCitation id="0A92BD71FFD8FFAF08A8E325FD2CFDDB" author="Krajcik, M." box="[502,677,542,566]" page="47" pageId="4" pageNumber="87" year="2006">
<bibRefCitation id="EFA2E691FFD8FFAF08A8E325FD2CFDDB" author="Krajcik, M." box="[502,677,542,566]" pageId="4" pageNumber="87" pagination="1 - 189" refId="ref9158" refString="Krajcik, M. (2006) Checklist of Scarabeoidea of the World. 1. Scarabaeinae (Coleoptera: Scarabaeidae: Scarabaeinae). Animma. X, Supplement 3, 1 - 189." type="journal article" year="2006">Krajcik 2006: 47</bibRefCitation>
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(checklist); 
<treatmentCitation id="0A92BD71FFD8FFAF0A76E325FC5EFDDB" author="Krajcik, M." box="[808,983,542,566]" page="88" pageId="4" pageNumber="87" year="2012">
<bibRefCitation id="EFA2E691FFD8FFAF0A76E325FC5EFDDB" author="Krajcik, M." box="[808,983,542,566]" pageId="4" pageNumber="87" pagination="1 - 278" refId="ref9194" refString="Krajcik, M. (2012) Checklist of the World Scarabaeoidea. Animma. X. Supplement 5, 1 - 278." type="journal article" year="2012">Krajcik 2012: 88</bibRefCitation>
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(checklist); 
<bibRefCitation id="EFA2E691FFD8FFAF0D07E325FABAFDDB" author="Schoolmeesters, P." box="[1113,1331,542,566]" pageId="4" pageNumber="87" pagination="2405 - 8858" refId="ref9492" refString="Schoolmeesters, P. (2020) Scarabs: world Scarabaeidae database (version 2019 - 11 - 02). In: Roskov Y. et al., (Eds.), Species 2000 &amp; ITIS Catalogue of Life, 2020 - 02 - 24. ISSN 2405 - 8858. Species 2000: Naturalis, Leiden, the Netherlands. Available from: http: // www. catalogueoflife. org / col (accessed 12 January 2022)" type="book chapter" year="2020">Schoolmeesters 2020</bibRefCitation>
(catalog); 
<bibRefCitation id="EFA2E691FFD8FFAF0999E307FE05FDB9" author="Vaz-de-Mello, F." box="[199,396,572,596]" pageId="4" pageNumber="87" refId="ref9975" refString="Vaz-de-Mello, F. (2021) Scarabaeinae. In: Catalogo Taxonomico da Fauna do Brasil. Available from: http: // fauna. jbrj. gov. br / fauna / faunadobrasil / 127498 (accessed 5 March 2021)" type="url" year="2021">Vaz-de-Mello 2021</bibRefCitation>
(checklist).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C329C8EBFFD8FFAF09C9E3B9FBE4FCC0" pageId="4" pageNumber="87" type="diagnosis">
<paragraph id="8B8C9B60FFD8FFAF09C9E3B9FBE4FCC0" blockId="4.[151,1437,642,2037]" pageId="4" pageNumber="87">
<emphasis id="B9474772FFD8FFAF09C9E3B9FE83FD71" bold="true" box="[151,266,642,668]" pageId="4" pageNumber="87">Diagnosis</emphasis>
. 
<taxonomicName id="4C33E0E3FFD8FFAF0844E3B9FDA0FD71" authorityName=": Halffter &amp; Martinez" authorityYear="1977" box="[282,553,642,668]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="diabolicus">
<emphasis id="B9474772FFD8FFAF0844E3B9FDA0FD71" box="[282,553,642,668]" italics="true" pageId="4" pageNumber="87">Deltepilissus diabolicus</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName id="4C33E0E3FFD8FFAF0A3CE3B8FC7BFD71" baseAuthorityName="Harold" baseAuthorityYear="1880" box="[866,1010,642,668]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="infernalis">
<emphasis id="B9474772FFD8FFAF0A3CE3B8FC7BFD71" box="[866,1010,642,668]" italics="true" pageId="4" pageNumber="87">D. infernalis</emphasis>
</taxonomicName>
by the head approximately 1.2 times wider than long (
<figureCitation id="130887E5FFD8FFAF0806E39DFE26FD2C" box="[344,431,678,705]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="4" pageNumber="87">Fig. 1C</figureCitation>
); punctures of central portion of pronotum closely set, spaced by less than their diameter (
<figureCitation id="130887E5FFD8FFAF09FEE3F1FF73FD08" box="[160,250,714,741]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="4" pageNumber="87">Fig. 1A</figureCitation>
); body length about 
<quantity id="4CCB3685FFD8FFAF08B7E3F1FDB5FD08" box="[489,572,714,741]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="87" unit="mm" value="15.0">15 mm</quantity>
. Furthermore, males have wide and rounded protibial spur, lacking denticle apically (
<figureCitation id="130887E5FFD8FFAF085EE3D5FED0FCE4" box="[256,345,750,777]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="4" pageNumber="87">Fig. 1G</figureCitation>
); and rounded metatibial spur apically, with almost inconspicuous inner denticle (
<figureCitation id="130887E5FFD8FFAF0DACE3D5FAC5FCE4" box="[1266,1356,750,777]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="4" pageNumber="87">Fig. 1H</figureCitation>
). Apex of parameres are also distinctly curved inward in dorsal and ventral views (
<figureCitation id="130887E5FFD8FFAF0ABFE229FBE9FCC0" box="[993,1120,786,813]" captionStart="FIGURE 3" captionStartId="9.[151,250,1773,1797]" captionTargetBox="[231,1367,208,1740]" captionTargetId="figure-19@9.[214,1374,181,1750]" captionTargetPageId="9" captionText="FIGURE 3. Details of the male genitalia in Deltepilissus. A–C, Deltepilissus diabolicus: A, parameres, dorsal view; B, parameres, ventral view; C, aedeagus, lateral view. D–F. Deltepilissus infernalis: D, parameres, dorsal view; E, parameres, ventral view; F, aedeagus, lateral view. G, Deltepilissus diabolicus superior right peripheral (SRP) endophallite. H, Deltepilissus infernalis superior right peripheral (SRP) endophallite. I, Deltepilissus diabolicus frontolateral peripheral (FLP) endophallite. J, Deltepilissus infernalis frontolateral peripheral (FLP) endophallite. K, Deltepilissus diabolicus additional endophallites (AE). L, Deltepilissus infernalis additional endophallites (AE). M, Deltepilissus diabolicus complex of axial and subaxial (A+SA) endophallites. N, Deltepilissus infernalis complex of axial and subaxial (A+SA) endophallites." figureDoi="http://doi.org/10.5281/zenodo.6388915" httpUri="https://zenodo.org/record/6388915/files/figure.png" pageId="4" pageNumber="87">Figs 3A–B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C329C8EBFFD8FFAF0999E20DFBEAF8F0" pageId="4" pageNumber="87" type="description">
<paragraph id="8B8C9B60FFD8FFAF0999E20DFA15F984" blockId="4.[151,1437,642,2037]" pageId="4" pageNumber="87">
<emphasis id="B9474772FFD8FFAF0999E20DFD71FCBC" bold="true" box="[199,760,822,849]" pageId="4" pageNumber="87">Redescription. Colour and tegument sculpture</emphasis>
(
<figureCitation id="130887E5FFD8FFAF0A54E20DFC02FCBC" box="[778,907,822,849]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="4" pageNumber="87">Figs 1A–B</figureCitation>
). Body, in general, black. Head and pronotum with light copper or green sheen. Body surface densely punctate. 
<emphasis id="B9474772FFD8FFAF0A3DE260FC30FC98" bold="true" box="[867,953,859,885]" pageId="4" pageNumber="87">Length</emphasis>
. 
<quantity id="4CCB3685FFD8FFAF0A9BE261FB9FFC98" box="[965,1046,858,885]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="87" unit="mm" value="15.0">15 mm</quantity>
. 
<emphasis id="B9474772FFD8FFAF0D41E260FBD6FC98" bold="true" box="[1055,1119,859,885]" pageId="4" pageNumber="87">Head</emphasis>
(
<figureCitation id="130887E5FFD8FFAF0D32E261FB4BFC98" box="[1132,1218,858,885]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="4" pageNumber="87">Fig. 1C</figureCitation>
). Densely punctate. Foveiform punctures approximately equally spaced to their diameter; punctures around clypeal teeth shallower, more spaced and smaller. Clypeus with two widely spaced, small teeth. Surface slightly depressed behind clypeal teeth. Clypeo-genal suture visible. 
<emphasis id="B9474772FFD8FFAF0887E2FCFDBBFC0C" bold="true" box="[473,562,967,993]" pageId="4" pageNumber="87">Thorax</emphasis>
. Disc of pronotum with dense closely set foveiform punctures, approximately spaced by less than their diameter; punctures denser on posterocentral portion (
<figureCitation id="130887E5FFD8FFAF0D47E2D1FBFAFBE8" box="[1049,1139,1002,1029]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="4" pageNumber="87">Fig. 1A</figureCitation>
). Hypomera not excavate, with sparse setose punctures on anterior and posterolateral portions (
<figureCitation id="130887E5FFD8FFAF0AFBE535FB89FBC4" box="[933,1024,1038,1065]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="4" pageNumber="87">Fig. 1D</figureCitation>
). Hypomera with oblique carina not reaching lateral margin; carina about half hypomeral width. Mesoventrite almost smooth, with sparse punctures. Metaventrite punctures variable in size, smaller and denser medially (
<figureCitation id="130887E5FFD8FFAF0A9AE56DFB97FB9C" box="[964,1054,1110,1137]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="4" pageNumber="87">Fig. 1B</figureCitation>
). 
<emphasis id="B9474772FFD8FFAF0D69E56CFB0AFB9C" bold="true" box="[1079,1155,1111,1137]" pageId="4" pageNumber="87">Elytra</emphasis>
(
<figureCitation id="130887E5FFD8FFAF0DC9E56DFB7AFB9C" box="[1175,1267,1110,1137]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="4" pageNumber="87">Fig. 1A</figureCitation>
). Punctures of elytral striae spaced by once or twice their diameter on 1st, 2nd and 3rd striae; punctures more spaced on lateral striae. Elytral striae wider, carinate margins clearly separated. Seventh stria effaced basally. Eighth and ninth striae almost fused, only visible on posterior half. Elytral interstriae with dense, close and foveiform punctures, generally spaced by less than their diameter. Pseudepipleural carina complete, visible on entire length of epipleuron. 
<emphasis id="B9474772FFD8FFAF09C9E431FE85FAC9" bold="true" box="[151,268,1290,1316]" pageId="4" pageNumber="87">Abdomen</emphasis>
(
<figureCitation id="130887E5FFD8FFAF0845E431FEF9FAC8" box="[283,368,1290,1317]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="4" pageNumber="87">Fig. 1E</figureCitation>
). Punctures denser on lateral surface of ventrites. Pygidium rounded apically, densely punctate, separated from propygidium by carina (
<figureCitation id="130887E5FFD8FFAF0B07E415FD27FAA4" box="[601,686,1326,1353]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="4" pageNumber="87">Fig. 1F</figureCitation>
). Pygidium with sulcus on basal and apical edges (
<figureCitation id="130887E5FFD8FFAF0DACE415FACEFAA4" box="[1266,1351,1326,1353]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="4" pageNumber="87">Fig. 1F</figureCitation>
). 
<emphasis id="B9474772FFD8FFAF0C03E414FA1CFAA4" bold="true" box="[1373,1429,1327,1353]" pageId="4" pageNumber="87">Legs</emphasis>
. Metatibial spur spatulate, with lateral edges weakly sinuous; apex rounded, with inconspicuous inner denticle (
<figureCitation id="130887E5FFD8FFAF0C2BE469FF35FA7C" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="4" pageNumber="87">Fig 1H</figureCitation>
). First meso- and metatarsomeres triangular; tarsomeres 2–4 trapezoidal, apical tarsomere subrectangular (
<figureCitation id="130887E5FFD8FFAF0C2BE44DFF35FA58" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="4" pageNumber="87">Fig 1H</figureCitation>
). Tarsal claws reduced (
<figureCitation id="130887E5FFD8FFAF0889E4A1FDD9FA58" box="[471,592,1434,1461]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="4" pageNumber="87">Figs 1G–I</figureCitation>
). 
<emphasis id="B9474772FFD8FFAF0B39E4A1FCDAFA59" bold="true" box="[615,851,1434,1460]" pageId="4" pageNumber="87">Sexual dimorphism</emphasis>
. Females can be distinguished from males by last abdominal ventrite evenly wide. Last abdominal ventrite of males subtly narrower medially. Females with protibial spur slightly emarginate apically (bifurcate). Protibial spur of males wide and rounded, lacking denticle apically (
<figureCitation id="130887E5FFD8FFAF09FEE73DFF71F9CC" box="[160,248,1542,1569]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="4" pageNumber="87">Fig. 1G</figureCitation>
). Females with metatibial spur apically sinuous and bifurcate, with denticle (spiniform process) on internal and external edges (
<figureCitation id="130887E5FFD8FFAF0828E711FE48F9A8" box="[374,449,1578,1605]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="4" pageNumber="87">Fig. 1I</figureCitation>
). Males with metatibial spur rounded apically, inner denticle almost inconspicuous (
<figureCitation id="130887E5FFD8FFAF0C31E711FF35F984" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="4" pageNumber="87">Fig. 1H</figureCitation>
). Apical third of metatibiae almost straight in females (
<figureCitation id="130887E5FFD8FFAF0A65E775FC00F984" box="[827,905,1614,1641]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="4" pageNumber="87">Fig. 1I</figureCitation>
), distinctly curved inward in males (
<figureCitation id="130887E5FFD8FFAF0C6DE775FA07F984" box="[1331,1422,1614,1641]" captionStart="FIGURE 1" captionStartId="5.[151,250,1798,1822]" captionTargetBox="[164,1425,491,1772]" captionTargetId="figure-157@5.[151,1436,489,1775]" captionTargetPageId="5" captionText="FIGURE 1. Details of the external morphology of Deltepilissus diabolicus. A, Body, male, dorsal view; B, body, male, ventral view; C, head, male; D, hypomeron, male; E, abdominal ventrites, male; F, pygidium, male; G, detail of protibial spur, male; H–I, left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur)." figureDoi="http://doi.org/10.5281/zenodo.6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="4" pageNumber="87">Fig. 1H</figureCitation>
).
</paragraph>
<paragraph id="8B8C9B60FFD8FFAF09C9E749FBEAF8F0" blockId="4.[151,1437,642,2037]" pageId="4" pageNumber="87">
<emphasis id="B9474772FFD8FFAF09C9E749FE83F961" bold="true" box="[151,266,1650,1676]" pageId="4" pageNumber="87">Aedeagus</emphasis>
. Parameres symmetrical, subparallel in lateral view, with abruptly pointing downward apically (
<figureCitation id="130887E5FFD8FFAF0C31E749FF32F95C" captionStart="FIGURE 3" captionStartId="9.[151,250,1773,1797]" captionTargetBox="[231,1367,208,1740]" captionTargetId="figure-19@9.[214,1374,181,1750]" captionTargetPageId="9" captionText="FIGURE 3. Details of the male genitalia in Deltepilissus. A–C, Deltepilissus diabolicus: A, parameres, dorsal view; B, parameres, ventral view; C, aedeagus, lateral view. D–F. Deltepilissus infernalis: D, parameres, dorsal view; E, parameres, ventral view; F, aedeagus, lateral view. G, Deltepilissus diabolicus superior right peripheral (SRP) endophallite. H, Deltepilissus infernalis superior right peripheral (SRP) endophallite. I, Deltepilissus diabolicus frontolateral peripheral (FLP) endophallite. J, Deltepilissus infernalis frontolateral peripheral (FLP) endophallite. K, Deltepilissus diabolicus additional endophallites (AE). L, Deltepilissus infernalis additional endophallites (AE). M, Deltepilissus diabolicus complex of axial and subaxial (A+SA) endophallites. N, Deltepilissus infernalis complex of axial and subaxial (A+SA) endophallites." figureDoi="http://doi.org/10.5281/zenodo.6388915" httpUri="https://zenodo.org/record/6388915/files/figure.png" pageId="4" pageNumber="87">Fig. 3C</figureCitation>
). Apex of parameres curved inward in dorsal and ventral views (
<figureCitation id="130887E5FFD8FFAF0AD1E7ADFB86F95C" box="[911,1039,1686,1713]" captionStart="FIGURE 3" captionStartId="9.[151,250,1773,1797]" captionTargetBox="[231,1367,208,1740]" captionTargetId="figure-19@9.[214,1374,181,1750]" captionTargetPageId="9" captionText="FIGURE 3. Details of the male genitalia in Deltepilissus. A–C, Deltepilissus diabolicus: A, parameres, dorsal view; B, parameres, ventral view; C, aedeagus, lateral view. D–F. Deltepilissus infernalis: D, parameres, dorsal view; E, parameres, ventral view; F, aedeagus, lateral view. G, Deltepilissus diabolicus superior right peripheral (SRP) endophallite. H, Deltepilissus infernalis superior right peripheral (SRP) endophallite. I, Deltepilissus diabolicus frontolateral peripheral (FLP) endophallite. J, Deltepilissus infernalis frontolateral peripheral (FLP) endophallite. K, Deltepilissus diabolicus additional endophallites (AE). L, Deltepilissus infernalis additional endophallites (AE). M, Deltepilissus diabolicus complex of axial and subaxial (A+SA) endophallites. N, Deltepilissus infernalis complex of axial and subaxial (A+SA) endophallites." figureDoi="http://doi.org/10.5281/zenodo.6388915" httpUri="https://zenodo.org/record/6388915/files/figure.png" pageId="4" pageNumber="87">Figs 3A–B</figureCitation>
). 
<emphasis id="B9474772FFD8FFAF0D7DE7ACFB48F95C" bold="true" box="[1059,1217,1687,1713]" pageId="4" pageNumber="87">Endophallus.</emphasis>
Lamella copulatrix absent. Superior right peripheral endophallite ring shaped; lateral projection arched, rounded apically (
<figureCitation id="130887E5FFD8FFAF0C6DE781FA06F938" box="[1331,1423,1722,1749]" captionStart="FIGURE 3" captionStartId="9.[151,250,1773,1797]" captionTargetBox="[231,1367,208,1740]" captionTargetId="figure-19@9.[214,1374,181,1750]" captionTargetPageId="9" captionText="FIGURE 3. Details of the male genitalia in Deltepilissus. A–C, Deltepilissus diabolicus: A, parameres, dorsal view; B, parameres, ventral view; C, aedeagus, lateral view. D–F. Deltepilissus infernalis: D, parameres, dorsal view; E, parameres, ventral view; F, aedeagus, lateral view. G, Deltepilissus diabolicus superior right peripheral (SRP) endophallite. H, Deltepilissus infernalis superior right peripheral (SRP) endophallite. I, Deltepilissus diabolicus frontolateral peripheral (FLP) endophallite. J, Deltepilissus infernalis frontolateral peripheral (FLP) endophallite. K, Deltepilissus diabolicus additional endophallites (AE). L, Deltepilissus infernalis additional endophallites (AE). M, Deltepilissus diabolicus complex of axial and subaxial (A+SA) endophallites. N, Deltepilissus infernalis complex of axial and subaxial (A+SA) endophallites." figureDoi="http://doi.org/10.5281/zenodo.6388915" httpUri="https://zenodo.org/record/6388915/files/figure.png" pageId="4" pageNumber="87">Fig. 3G</figureCitation>
). frontolateral peripheral endophallite elongate, irregular (
<figureCitation id="130887E5FFD8FFAF0A78E7E5FCF0F914" box="[806,889,1758,1785]" captionStart="FIGURE 3" captionStartId="9.[151,250,1773,1797]" captionTargetBox="[231,1367,208,1740]" captionTargetId="figure-19@9.[214,1374,181,1750]" captionTargetPageId="9" captionText="FIGURE 3. Details of the male genitalia in Deltepilissus. A–C, Deltepilissus diabolicus: A, parameres, dorsal view; B, parameres, ventral view; C, aedeagus, lateral view. D–F. Deltepilissus infernalis: D, parameres, dorsal view; E, parameres, ventral view; F, aedeagus, lateral view. G, Deltepilissus diabolicus superior right peripheral (SRP) endophallite. H, Deltepilissus infernalis superior right peripheral (SRP) endophallite. I, Deltepilissus diabolicus frontolateral peripheral (FLP) endophallite. J, Deltepilissus infernalis frontolateral peripheral (FLP) endophallite. K, Deltepilissus diabolicus additional endophallites (AE). L, Deltepilissus infernalis additional endophallites (AE). M, Deltepilissus diabolicus complex of axial and subaxial (A+SA) endophallites. N, Deltepilissus infernalis complex of axial and subaxial (A+SA) endophallites." figureDoi="http://doi.org/10.5281/zenodo.6388915" httpUri="https://zenodo.org/record/6388915/files/figure.png" pageId="4" pageNumber="87">Fig. 3I</figureCitation>
). Complex of axial and subaxial endophallites elongate, irregular (
<figureCitation id="130887E5FFD8FFAF082AE639FE5AF8F0" box="[372,467,1794,1821]" captionStart="FIGURE 3" captionStartId="9.[151,250,1773,1797]" captionTargetBox="[231,1367,208,1740]" captionTargetId="figure-19@9.[214,1374,181,1750]" captionTargetPageId="9" captionText="FIGURE 3. Details of the male genitalia in Deltepilissus. A–C, Deltepilissus diabolicus: A, parameres, dorsal view; B, parameres, ventral view; C, aedeagus, lateral view. D–F. Deltepilissus infernalis: D, parameres, dorsal view; E, parameres, ventral view; F, aedeagus, lateral view. G, Deltepilissus diabolicus superior right peripheral (SRP) endophallite. H, Deltepilissus infernalis superior right peripheral (SRP) endophallite. I, Deltepilissus diabolicus frontolateral peripheral (FLP) endophallite. J, Deltepilissus infernalis frontolateral peripheral (FLP) endophallite. K, Deltepilissus diabolicus additional endophallites (AE). L, Deltepilissus infernalis additional endophallites (AE). M, Deltepilissus diabolicus complex of axial and subaxial (A+SA) endophallites. N, Deltepilissus infernalis complex of axial and subaxial (A+SA) endophallites." figureDoi="http://doi.org/10.5281/zenodo.6388915" httpUri="https://zenodo.org/record/6388915/files/figure.png" pageId="4" pageNumber="87">Fig. 3M</figureCitation>
). Additional endophallites short, comma shaped (
<figureCitation id="130887E5FFD8FFAF0AA2E639FBDFF8F0" box="[1020,1110,1794,1821]" captionStart="FIGURE 3" captionStartId="9.[151,250,1773,1797]" captionTargetBox="[231,1367,208,1740]" captionTargetId="figure-19@9.[214,1374,181,1750]" captionTargetPageId="9" captionText="FIGURE 3. Details of the male genitalia in Deltepilissus. A–C, Deltepilissus diabolicus: A, parameres, dorsal view; B, parameres, ventral view; C, aedeagus, lateral view. D–F. Deltepilissus infernalis: D, parameres, dorsal view; E, parameres, ventral view; F, aedeagus, lateral view. G, Deltepilissus diabolicus superior right peripheral (SRP) endophallite. H, Deltepilissus infernalis superior right peripheral (SRP) endophallite. I, Deltepilissus diabolicus frontolateral peripheral (FLP) endophallite. J, Deltepilissus infernalis frontolateral peripheral (FLP) endophallite. K, Deltepilissus diabolicus additional endophallites (AE). L, Deltepilissus infernalis additional endophallites (AE). M, Deltepilissus diabolicus complex of axial and subaxial (A+SA) endophallites. N, Deltepilissus infernalis complex of axial and subaxial (A+SA) endophallites." figureDoi="http://doi.org/10.5281/zenodo.6388915" httpUri="https://zenodo.org/record/6388915/files/figure.png" pageId="4" pageNumber="87">Fig. 3K</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C329C8EBFFD8FFAE0999E61CFE14FE3C" lastPageId="5" lastPageNumber="88" pageId="4" pageNumber="87" type="materials_examined">
<paragraph id="8B8C9B60FFD8FFAF0999E61CFD92F864" blockId="4.[151,1437,642,2037]" pageId="4" pageNumber="87">
<emphasis id="B9474772FFD8FFAF0999E61CFEF6F8AD" bold="true" box="[199,383,1830,1857]" pageId="4" pageNumber="87">Type material.</emphasis>
<taxonomicName id="4C33E0E3FFD8FFAF08CFE61DFD62F8AC" authority="Harold." authorityName="Harold." box="[401,747,1830,1857]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="diabolicus">
<emphasis id="B9474772FFD8FFAF08CFE61DFD6DF8AC" bold="true" box="[401,740,1830,1857]" pageId="4" pageNumber="87">
<emphasis id="B9474772FFD8FFAF08CFE61DFDF4F8AC" bold="true" box="[401,637,1830,1857]" italics="true" pageId="4" pageNumber="87">Canthon diabolicus</emphasis>
Harold
</emphasis>
.
</taxonomicName>
<materialsCitation id="3B5B913DFFD8FFAF0BA3E61CFD92F864" ID-GBIF-Occurrence="3738408303" collectionCode="MFNB" county="Sello" location="Har" municipality="Type" pageId="4" pageNumber="87" specimenCount="1" specimenCount-female="1" stateProvince="Bahia" typeStatus="lectotype">
<emphasis id="B9474772FFD8FFAF0BA3E61CFCFAF8AC" bold="true" box="[765,883,1831,1857]" pageId="4" pageNumber="87">
<typeStatus id="548825C2FFD8FFAF0BA3E61CFCFAF8AC" box="[765,883,1831,1857]" pageId="4" pageNumber="87" type="lectotype">Lectotype</typeStatus>
</emphasis>
<specimenCount id="9D3550E9FFD8FFAF0ADAE61CFC10F8AC" box="[900,921,1831,1857]" pageId="4" pageNumber="87" type="female">♀</specimenCount>
(
<emphasis id="B9474772FFD8FFAF0AEAE61CFBF3F8AD" bold="true" box="[948,1146,1830,1857]" pageId="4" pageNumber="87">here designated</emphasis>
; 
<collectionCode id="ED2203A5FFD8FFAF0DCDE61CFB6DF8AC" box="[1171,1252,1831,1857]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="4" pageNumber="87">MFNB</collectionCode>
) (
<figureCitation id="130887E5FFD8FFAF0C5BE61DFA18F8AC" box="[1285,1425,1830,1857]" captionStart="FIGURE 4" captionStartId="10.[151,250,1910,1934]" captionTargetBox="[278,1321,181,1886]" captionTargetId="figure-30@10.[267,1321,181,1886]" captionTargetPageId="10" captionText="FIGURE 4. Type specimens in Deltepilissus and respective label data. A–B, Lectotype of Canthon diabolicus Harold, 1880: A, body, dorsal view; B, labels. C–D, Lectotype of Canthon infernalis Harold, 1880: C, body, dorsal view; D, labels. E–H, holotype of Deltepilissus travassosi Pereira, 1949: E, body, dorsal view; F, head; G, left protibia, dorsal view; H, protibial and tarsal claws. Photographs A–D provided by Fernando Z. Vaz-de-Mello." figureDoi="http://doi.org/10.5281/zenodo.6388917" httpUri="https://zenodo.org/record/6388917/files/figure.png" pageId="4" pageNumber="87">Figs 4A–B</figureCitation>
): “ 
<typeStatus id="548825C2FFD8FFAF09FEE670FEBEF888" box="[160,311,1867,1893]" pageId="4" pageNumber="87" type="holotype">HOLOTYPE</typeStatus>
// 
<emphasis id="B9474772FFD8FFAF0807E671FE2AF889" box="[345,419,1866,1892]" italics="true" pageId="4" pageNumber="87">
<collectingRegion id="49F75582FFD8FFAF0807E671FE16F889" box="[345,415,1866,1892]" country="Brazil" name="Bahia" pageId="4" pageNumber="87">Bahia</collectingRegion>
,
</emphasis>
/ 
<emphasis id="B9474772FFD8FFAF08E3E671FE7CF889" box="[445,501,1866,1892]" italics="true" pageId="4" pageNumber="87">
<collectingCounty id="62EDE3ECFFD8FFAF08E3E671FE7CF889" box="[445,501,1866,1892]" pageId="4" pageNumber="87">Sello</collectingCounty>
</emphasis>
// 9004 // 
<collectingMunicipality id="6BE8011AFFD8FFAF0B2EE670FD21F888" box="[624,680,1867,1893]" pageId="4" pageNumber="87">Type</collectingMunicipality>
// 
<taxonomicName id="4C33E0E3FFD8FFAF0B94E671FCB6F889" box="[714,831,1866,1892]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="diabolicus">
<emphasis id="B9474772FFD8FFAF0B94E671FCB6F889" box="[714,831,1866,1892]" italics="true" pageId="4" pageNumber="87">diabolicus</emphasis>
</taxonomicName>
/ 
<emphasis id="B9474772FFD8FFAF0A07E670FC0FF889" box="[857,902,1867,1892]" italics="true" pageId="4" pageNumber="87">
<location id="8EECCDBBFFD8FFAF0A07E670FC0FF889" LSID="urn:lsid:plazi:treatment:039A2A76FFD8FFAD09C9E001FB4BFC6C:8EECCDBBFFD8FFAF0A07E670FC0FF889" box="[857,902,1867,1892]" county="Sello" municipality="Type" name="Har" pageId="4" pageNumber="87" stateProvince="Bahia">Har</location>
</emphasis>
// 
<typeStatus id="548825C2FFD8FFAF0AF6E670FBB0F888" box="[936,1081,1867,1893]" pageId="4" pageNumber="87" type="syntype">SYNTYPUS</typeStatus>
/ 
<taxonomicName id="4C33E0E3FFD8FFAF0D0CE671FB38F889" authorityName="Hoffmannsegg" authorityYear="1817" box="[1106,1201,1866,1892]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="genus">Canthon</taxonomicName>
/ 
<taxonomicName id="4C33E0E3FFD8FFAF0D95E671FF46F864" authority="Harold, 1880" authorityName="Harold" authorityYear="1880" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="diabolicus">
diabolicus 
<bibRefCitation id="EFA2E691FFD8FFAF0C19E671FF46F864" author="Harold, E." pageId="4" pageNumber="87" pagination="148 - 171" refId="ref9058" refString="Harold, E. (1880) Einige Neue Coleopteren. Mittheilungen des Munchener Entomologischen Vereins, 4, 148 - 171." type="journal article" year="1880">Harold, 1880</bibRefCitation>
</taxonomicName>
/ labelled by MNHUB 2013”
</materialsCitation>
</paragraph>
<paragraph id="8B8C9B60FFD8FFAF0999E6A9FCDBF83C" blockId="4.[151,1437,642,2037]" pageId="4" pageNumber="87">
<emphasis id="B9474772FFD8FFAF0999E6A9FE36F841" bold="true" box="[199,447,1938,1964]" pageId="4" pageNumber="87">Additional material.</emphasis>
<materialsCitation id="3B5B913DFFD8FFAF0892E6A9FCC7F83C" ID-GBIF-Occurrence="3738408307" collectingDate="1968-09-11" collectionCode="BMNH, CEMT" collectorName="Guy V. Santos" country="Brazil" location="Ilheus" pageId="4" pageNumber="87" specimenCount="1" specimenCount-male="1" stateProvince="Bahia">
<emphasis id="B9474772FFD8FFAF0892E6A9FDBEF841" bold="true" box="[460,567,1938,1964]" pageId="4" pageNumber="87">
<collectingCountry id="F324DBF0FFD8FFAF0892E6A9FDBEF841" box="[460,567,1938,1964]" name="Brazil" pageId="4" pageNumber="87">BRAZIL</collectingCountry>
</emphasis>
: 
<collectingRegion id="49F75582FFD8FFAF0B15E6A8FD2EF840" box="[587,679,1939,1965]" country="Brazil" name="Bahia" pageId="4" pageNumber="87">BAHIA</collectingRegion>
, no date and collector – 1 sex undetermined (
<collectionCode id="ED2203A5FFD8FFAF0D8CE6A8FAA1F840" box="[1234,1320,1939,1965]" pageId="4" pageNumber="87">BMNH</collectionCode>
– ex Fry collection); 
<location id="8EECCDBBFFD8FFAF0840E68DFEECF83D" LSID="urn:lsid:plazi:treatment:039A2A76FFD8FFAD09C9E001FB4BFC6C:8EECCDBBFFD8FFAF0840E68DFEECF83D" box="[286,357,1974,2000]" country="Brazil" name="Ilheus" pageId="4" pageNumber="87" stateProvince="Bahia">Ilhéus</location>
, 
<date id="FF8DBDA0FFD8FFAF082EE68DFE78F83D" box="[368,497,1974,2000]" pageId="4" pageNumber="87" value="1968-09-11">
<collectingDate id="EFC94448FFD8FFAF082EE68DFE78F83D" box="[368,497,1974,2000]" pageId="4" pageNumber="87" value="1968-09-11">11.IX.1968</collectingDate>
</date>
, 
<collectorName id="26C6FEB6FFD8FFAF08A3E68CFD14F83C" box="[509,669,1975,2001]" pageId="4" pageNumber="87">Guy V. Santos</collectorName>
– 
<specimenCount id="9D3550E9FFD8FFAF0BE7E68CFD6AF83C" box="[697,739,1975,2001]" pageId="4" pageNumber="87" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED2203A5FFD8FFAF0BACE68CFCCFF83C" box="[754,838,1975,2001]" pageId="4" pageNumber="87">CEMT</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8C9B60FFD8FFAE0999E6E1FE14FE3C" blockId="4.[151,1437,642,2037]" lastBlockId="5.[151,1437,151,465]" lastPageId="5" lastPageNumber="88" pageId="4" pageNumber="87">
<emphasis id="B9474772FFD8FFAF0999E6E1FEAAF819" bold="true" box="[199,291,2010,2036]" pageId="4" pageNumber="87">Habitat</emphasis>
. 
<materialsCitation id="3B5B913DFFD8FFAE086FE6E0FE13FE3C" ID-GBIF-Occurrence="3738408304" country="Brazil, Uruguay" county="The" lastPageId="5" lastPageNumber="88" location="Prussian" municipality="Atlantic" pageId="4" pageNumber="87" specimenCount="1">
Most likely from Atlantic coastal rainforest. The 
<typeStatus id="548825C2FFD8FFAF0A0BE6E0FC0CF818" box="[853,901,2011,2037]" pageId="4" pageNumber="87">type</typeStatus>
specimen of 
<taxonomicName id="4C33E0E3FFD8FFAF0D45E6E0FB3BF819" box="[1051,1202,2010,2036]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="diabolicus">
<emphasis id="B9474772FFD8FFAF0D45E6E0FB3BF819" box="[1051,1202,2010,2036]" italics="true" pageId="4" pageNumber="87">D. diabolicus</emphasis>
</taxonomicName>
was collected by the 
<location id="8EECCDBBFFD9FFAE09C9E1ACFF7FFF5C" LSID="urn:lsid:plazi:treatment:039A2A76FFD8FFAD09C9E001FB4BFC6C:8EECCDBBFFD9FFAE09C9E1ACFF7FFF5C" box="[151,246,151,177]" country="Brazil, Uruguay" county="The" municipality="Atlantic" name="Prussian" pageId="5" pageNumber="88">Prussian</location>
naturalist 
<location id="8EECCDBBFFD9FFAE082CE1ACFDB8FF5C" LSID="urn:lsid:plazi:treatment:039A2A76FFD8FFAD09C9E001FB4BFC6C:8EECCDBBFFD9FFAE082CE1ACFDB8FF5C" box="[370,561,151,177]" country="Brazil, Uruguay" county="The" municipality="Atlantic" name="Friedrich Sellow" pageId="5" pageNumber="88">Friedrich Sellow</location>
(1789–1831). 
<location id="8EECCDBBFFD9FFAE0B82E1ACFD74FF5C" LSID="urn:lsid:plazi:treatment:039A2A76FFD8FFAD09C9E001FB4BFC6C:8EECCDBBFFD9FFAE0B82E1ACFD74FF5C" box="[732,765,151,177]" country="Brazil, Uruguay" county="The" municipality="Atlantic" name="He" pageId="5" pageNumber="88">He</location>
travelled through 
<collectingCountry id="F324DBF0FFD9FFAE0A8AE1ACFB02FF5C" box="[980,1163,151,177]" isEnumeration="true" pageId="5" pageNumber="88">Brazil, Uruguay</collectingCountry>
and 
<collectingCountry id="F324DBF0FFD9FFAE0D98E1ACFABCFF5C" box="[1222,1333,151,177]" name="Argentina" pageId="5" pageNumber="88">Argentina</collectingCountry>
between 1814 and 1831. 
<location id="8EECCDBBFFD9FFAE0809E180FEF5FF38" LSID="urn:lsid:plazi:treatment:039A2A76FFD8FFAD09C9E001FB4BFC6C:8EECCDBBFFD9FFAE0809E180FEF5FF38" box="[343,380,187,213]" country="Brazil, Uruguay" county="The" municipality="Atlantic" name="We" pageId="5" pageNumber="88">We</location>
were able to check 
<location id="8EECCDBBFFD9FFAE0B31E180FD46FF38" LSID="urn:lsid:plazi:treatment:039A2A76FFD8FFAD09C9E001FB4BFC6C:8EECCDBBFFD9FFAE0B31E180FD46FF38" box="[623,719,187,213]" country="Brazil, Uruguay" county="The" municipality="Atlantic" name="Sellow's" pageId="5" pageNumber="88">Sellow’s</location>
itinerary in 
<location id="8EECCDBBFFD9FFAE0A3BE180FC2FFF38" LSID="urn:lsid:plazi:treatment:039A2A76FFD8FFAD09C9E001FB4BFC6C:8EECCDBBFFD9FFAE0A3BE180FC2FFF38" box="[869,934,187,213]" country="Brazil, Uruguay" county="The" municipality="Atlantic" name="Bahia" pageId="5" pageNumber="88">Bahia</location>
(
<bibRefCitation id="EFA2E691FFD9FFAE0AE7E180FBE1FF38" author="Papavero, N." box="[953,1128,187,213]" pageId="5" pageNumber="88" refId="ref9218" refString="Papavero, N. (1971) Essays on the history of Neotropical dipterology: with special reference to collectors: 1750 - 1905: Volume I. Museu de Zoologia, Universidade de Sao Paulo, Sao Paulo, Brazil, 216 pp. https: // doi. org / 10.5962 / bhl. title. 101715" type="book" year="1971">Papavero 1971</bibRefCitation>
, map 8). 
<location id="8EECCDBBFFD9FFAE0D82E180FABFFF38" LSID="urn:lsid:plazi:treatment:039A2A76FFD8FFAD09C9E001FB4BFC6C:8EECCDBBFFD9FFAE0D82E180FABFFF38" box="[1244,1334,187,213]" country="Brazil, Uruguay" county="The" municipality="Atlantic" name="Therein" pageId="5" pageNumber="88">Therein</location>
, he only visited areas in the 
<location id="8EECCDBBFFD9FFAE082AE1E4FE47FF14" LSID="urn:lsid:plazi:treatment:039A2A76FFD8FFAD09C9E001FB4BFC6C:8EECCDBBFFD9FFAE082AE1E4FE47FF14" box="[372,462,223,249]" country="Brazil, Uruguay" county="The" municipality="Atlantic" name="Atlantic" pageId="5" pageNumber="88">Atlantic</location>
coastal forest between Salvador and 
<location id="8EECCDBBFFD9FFAE0A26E1E4FB48FF14" LSID="urn:lsid:plazi:treatment:039A2A76FFD8FFAD09C9E001FB4BFC6C:8EECCDBBFFD9FFAE0A26E1E4FB48FF14" box="[888,1217,223,249]" country="Brazil, Uruguay" county="The" municipality="Atlantic" name="southern Bahia. Furthermore" pageId="5" pageNumber="88">southern Bahia. Furthermore</location>
, the only specimen studied here with a precise locality was collected from 
<location id="8EECCDBBFFD9FFAE0BA5E038FCCBFEF0" LSID="urn:lsid:plazi:treatment:039A2A76FFD8FFAD09C9E001FB4BFC6C:8EECCDBBFFD9FFAE0BA5E038FCCBFEF0" box="[763,834,259,285]" country="Brazil, Uruguay" county="The" municipality="Atlantic" name="Ilheus" pageId="5" pageNumber="88">Ilhéus</location>
, which is a coastal city located in 
<location id="8EECCDBBFFD9FFAE0DE2E038FA14FEF0" LSID="urn:lsid:plazi:treatment:039A2A76FFD8FFAD09C9E001FB4BFC6C:8EECCDBBFFD9FFAE0DE2E038FA14FEF0" box="[1212,1437,259,285]" country="Brazil, Uruguay" county="The" municipality="Atlantic" name="southern Bahia. The" pageId="5" pageNumber="88">southern Bahia. The</location>
predominant ecosystem surrounding the area is the 
<location id="8EECCDBBFFD9FFAE0B83E01CFCBEFEAC" LSID="urn:lsid:plazi:treatment:039A2A76FFD8FFAD09C9E001FB4BFC6C:8EECCDBBFFD9FFAE0B83E01CFCBEFEAC" box="[733,823,295,321]" country="Brazil, Uruguay" county="The" municipality="Atlantic" name="Atlantic" pageId="5" pageNumber="88">Atlantic</location>
coastal forest, which hosts a high number of endemic species. 
<location id="8EECCDBBFFD9FFAE09A9E070FE87FE88" LSID="urn:lsid:plazi:treatment:039A2A76FFD8FFAD09C9E001FB4BFC6C:8EECCDBBFFD9FFAE09A9E070FE87FE88" box="[247,270,331,357]" country="Brazil, Uruguay" county="The" municipality="Atlantic" name="In" pageId="5" pageNumber="88">In</location>
addition to this vegetation, there are areas of restinga (tropical rainforest close to coastal, sandy areas) and mangroves on the coastal line, and large cocoa-growing areas planted among the natural 
<collectingMunicipality id="6BE8011AFFD9FFAE0D93E054FAAEFE64" box="[1229,1319,367,393]" pageId="5" pageNumber="88">Atlantic</collectingMunicipality>
rainforest (Cabruca = ecological system of cultivation agroforestry). 
<collectingCounty id="62EDE3ECFFD9FFAE0A69E0A8FCEAFE40" box="[823,867,403,429]" pageId="5" pageNumber="88">The</collectingCounty>
specimen might have been collected from any of these different habitats
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF4CCBE8FFD9FFAE09C9E63DFB4BF88B" ID-DOI="http://doi.org/10.5281/zenodo.6388911" ID-Zenodo-Dep="6388911" httpUri="https://zenodo.org/record/6388911/files/figure.png" pageId="5" pageNumber="88" startId="5.[151,250,1798,1822]" targetBox="[164,1425,491,1772]" targetPageId="5">
<paragraph id="8B8C9B60FFD9FFAE09C9E63DFB4BF88B" blockId="5.[151,1436,1798,1894]" pageId="5" pageNumber="88">
<emphasis id="B9474772FFD9FFAE09C9E63DFE84F8F3" bold="true" box="[151,269,1798,1822]" pageId="5" pageNumber="88">FIGURE 1</emphasis>
. Details of the external morphology of 
<emphasis id="B9474772FFD9FFAE0BC3E63DFC1EF8F3" box="[669,919,1798,1822]" italics="true" pageId="5" pageNumber="88">
<taxonomicName id="4C33E0E3FFD9FFAE0BC3E63DFC1AF8F3" authorityName=": Halffter &amp; Martinez" authorityYear="1977" box="[669,915,1798,1822]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="species" species="diabolicus">Deltepilissus diabolicus</taxonomicName>
.
</emphasis>
<emphasis id="B9474772FFD9FFAE0AC2E63DFC3DF8F3" bold="true" box="[924,948,1798,1822]" pageId="5" pageNumber="88">A,</emphasis>
Body, male, dorsal view; 
<emphasis id="B9474772FFD9FFAE0DE3E63DFB5DF8F3" bold="true" box="[1213,1236,1798,1822]" pageId="5" pageNumber="88">B,</emphasis>
body, male, ventral view; 
<emphasis id="B9474772FFD9FFAE0989E611FF79F8AF" bold="true" box="[215,240,1834,1858]" pageId="5" pageNumber="88">C,</emphasis>
head, male; 
<emphasis id="B9474772FFD9FFAE082DE611FE05F8AF" bold="true" box="[371,396,1834,1858]" pageId="5" pageNumber="88">D,</emphasis>
hypomeron, male; 
<emphasis id="B9474772FFD9FFAE0B0AE611FDE2F8AF" bold="true" box="[596,619,1834,1858]" pageId="5" pageNumber="88">E,</emphasis>
abdominal ventrites, male; 
<emphasis id="B9474772FFD9FFAE0AD7E611FC15F8AF" bold="true" box="[905,924,1834,1858]" pageId="5" pageNumber="88">F,</emphasis>
pygidium, male; 
<emphasis id="B9474772FFD9FFAE0D0CE611FBE2F8AF" bold="true" box="[1106,1131,1834,1858]" pageId="5" pageNumber="88">G,</emphasis>
detail of protibial spur, male; 
<emphasis id="B9474772FFD9FFAE09C9E675FF41F88B" bold="true" box="[151,200,1870,1894]" pageId="5" pageNumber="88">H–I,</emphasis>
left metatibia, ventral view, male and female, respectively (arrows indicate apex of metatibial spur).
</paragraph>
</caption>
<subSubSection id="C329C8EBFFD9FFAD0999E6AFFB4BFC6C" lastPageId="6" lastPageNumber="89" pageId="5" pageNumber="88" type="distribution">
<paragraph id="8B8C9B60FFD9FFAE0999E6AFFA1AF842" blockId="5.[151,1437,1940,2039]" box="[199,1427,1940,1967]" pageId="5" pageNumber="88">
<emphasis id="B9474772FFD9FFAE0999E6AFFEE8F843" bold="true" box="[199,353,1940,1966]" pageId="5" pageNumber="88">Distribution.</emphasis>
Known only from Brazil (
<collectingRegion id="49F75582FFD9FFAE0BD5E6AFFD5BF842" box="[651,722,1940,1967]" country="Brazil" name="Bahia" pageId="5" pageNumber="88">Bahia</collectingRegion>
). 
<emphasis id="B9474772FFD9FFAE0BBAE6AFFBB3F843" bold="true" box="[740,1082,1940,1967]" pageId="5" pageNumber="88">
Literature records. 
<collectingCountry id="F324DBF0FFD9FFAE0A91E6AFFBB3F843" box="[975,1082,1940,1966]" name="Brazil" pageId="5" pageNumber="88">BRAZIL</collectingCountry>
</emphasis>
: 
<collectingRegion id="49F75582FFD9FFAE0D17E6AFFB03F843" box="[1097,1162,1940,1966]" country="Brazil" name="Bahia" pageId="5" pageNumber="88">Bahia</collectingRegion>
(
<bibRefCitation id="EFA2E691FFD9FFAE0DC7E6AFFAA0F842" author="Harold, E." box="[1177,1321,1940,1967]" pageId="5" pageNumber="88" pagination="148 - 171" refId="ref9058" refString="Harold, E. (1880) Einige Neue Coleopteren. Mittheilungen des Munchener Entomologischen Vereins, 4, 148 - 171." type="journal article" year="1880">Harold 1880</bibRefCitation>
) (
<figureCitation id="130887E5FFD9FFAE0C1EE6AFFA0CF842" box="[1344,1413,1940,1967]" captionStart="FIGURE 5" captionStartId="11.[151,250,1103,1127]" captionTargetBox="[151,1436,181,1066]" captionTargetId="figure-19@11.[151,1436,181,1080]" captionTargetPageId="11" captionText="FIGURE 5. Known distribution of species of Deltepilissus. A, Records plotted on terrestrial biomes layer; B, records plotted on elevational range layer." figureDoi="http://doi.org/10.5281/zenodo.6388921" httpUri="https://zenodo.org/record/6388921/files/figure.png" pageId="5" pageNumber="88">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="8B8C9B60FFD9FFAD0999E683FC22FDF4" blockId="5.[151,1437,1940,2039]" lastBlockId="6.[151,1437,151,897]" lastPageId="6" lastPageNumber="89" pageId="5" pageNumber="88">
<emphasis id="B9474772FFD9FFAE0999E683FE34F83E" bold="true" box="[199,445,1976,2003]" pageId="5" pageNumber="88">Conservation status.</emphasis>
Based on available data, this species seems to be endemic to the state of 
<collectingRegion id="49F75582FFD9FFAE0C5EE683FACDF83F" box="[1280,1348,1976,2002]" country="Brazil" name="Bahia" pageId="5" pageNumber="88">Bahia</collectingRegion>
, 
<collectingCountry id="F324DBF0FFD9FFAE0C0FE683FA10F83F" box="[1361,1433,1976,2002]" name="Brazil" pageId="5" pageNumber="88">Brazil</collectingCountry>
. The last known record for this species was from Ilhéus in 1968. Therefore, we do not have information about the conservation status of populations of this species. We know at least two cases of apparently rare species along the range of the two species of 
<taxonomicName id="4C33E0E3FFDAFFAD0895E180FDD5FF38" authorityName="Pereira" authorityYear="1949" box="[459,604,187,213]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="B9474772FFDAFFAD0895E180FDD5FF38" box="[459,604,187,213]" italics="true" pageId="6" pageNumber="89">Deltepilissus</emphasis>
</taxonomicName>
(southern 
<collectingRegion id="49F75582FFDAFFAD0B8DE180FC9DFF38" box="[723,788,187,213]" country="Brazil" name="Bahia" pageId="6" pageNumber="89">Bahia</collectingRegion>
and 
<collectingRegion id="49F75582FFDAFFAD0A15E180FC62FF38" box="[843,1003,187,213]" country="Brazil" name="Espirito Santo" pageId="6" pageNumber="89">Espírito Santo</collectingRegion>
) that when the number of collectors in the area increased, a great number of specimens have been collected. These cases can be verified with 
<taxonomicName id="4C33E0E3FFDAFFAD0C7BE1E4FD85FEF0" authority="Van Lansberge, 1874" authorityName="Van Lansberge" authorityYear="1874" class="Insecta" family="Scarabaeidae" genus="Streblopus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="89" phylum="Arthropoda" rank="species" species="opatroides">
<emphasis id="B9474772FFDAFFAD0C7BE1E4FE87FEF0" italics="true" pageId="6" pageNumber="89">Streblopus opatroides</emphasis>
Van Lansberge, 1874
</taxonomicName>
(
<bibRefCitation id="EFA2E691FFDAFFAD0B40E038FC8DFEF0" author="Cupello, M. &amp; Ribeiro-Costa, C. S. &amp; Vaz-de-Mello, F. Z." box="[542,772,259,285]" pageId="6" pageNumber="89" pagination="1 - 85" refId="ref8683" refString="Cupello, M., Ribeiro-Costa, C. S. &amp; Vaz-de-Mello, F. Z. (2020) Systematics of the enigmatic South American Streblopus Van Lansberge, 1874 dung beetles and their transatlantic origin: a case study on the role of dispersal events in the biogeographical history of the Scarabaeinae (Coleoptera: Scarabaeidae). European Journal of Taxonomy, 603, 1 - 85. https: // doi. org / 10.5852 / ejt. 2020.603" type="journal article" year="2020">
Cupello 
<emphasis id="B9474772FFDAFFAD0BDAE038FD31FEF0" box="[644,696,259,285]" italics="true" pageId="6" pageNumber="89">et al</emphasis>
. 2020
</bibRefCitation>
) and some species of 
<taxonomicName id="4C33E0E3FFDAFFAD0D52E038FAA7FEF0" authority="Harold, 1868" authorityName="Harold" authorityYear="1868" box="[1036,1326,259,285]" class="Insecta" family="Scarabaeidae" genus="Aphengium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="B9474772FFDAFFAD0D52E038FB05FEF0" box="[1036,1164,259,285]" italics="true" pageId="6" pageNumber="89">Aphengium</emphasis>
Harold, 1868
</taxonomicName>
(
<bibRefCitation id="EFA2E691FFDAFFAD0C1FE038FEF3FEAC" author="Silva, F. A. &amp; Vaz-de-Mello, F." pageId="6" pageNumber="89" pagination="505 - 520" refId="ref9575" refString="Silva, F. A. B &amp; Vaz-de-Mello, F. (2015) A revision of the genus Aphengium Harold, 1868 (Coleoptera: Scarabaeidae: Scarabaeinae: Ateuchini). Zootaxa, 3955 (4), 505 - 520. https: // doi. org / 10.11646 / zootaxa. 3955.4.3" type="journal article" year="2015">Silva &amp; Vaz-de-Mello 2015</bibRefCitation>
). However, despite many collecting efforts with pitfalls and/or flight interception traps in Atlantic rainforest remnants throughout that region and neighboring areas (F. Silva and F. Vaz-de-Mello, personal communication) no specimens of 
<taxonomicName id="4C33E0E3FFDAFFAD0B4DE054FD22FE64" authorityName=": Halffter &amp; Martinez" authorityYear="1977" box="[531,683,367,393]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="89" phylum="Arthropoda" rank="species" species="diabolicus">
<emphasis id="B9474772FFDAFFAD0B4DE054FD22FE64" box="[531,683,367,393]" italics="true" pageId="6" pageNumber="89">D. diabolicus</emphasis>
</taxonomicName>
have been recently collected. The rarity of 
<taxonomicName id="4C33E0E3FFDAFFAD0DCDE054FAA2FE64" authorityName=": Halffter &amp; Martinez" authorityYear="1977" box="[1171,1323,367,393]" class="Insecta" family="Scarabaeidae" genus="Deltepilissus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="89" phylum="Arthropoda" rank="species" species="diabolicus">
<emphasis id="B9474772FFDAFFAD0DCDE054FAA2FE64" box="[1171,1323,367,393]" italics="true" pageId="6" pageNumber="89">D. diabolicus</emphasis>
</taxonomicName>
in natural history collections can be due to its unknown natural history and/or some unusual ecological specialization. The baits used for collecting ‒ usually, human feces ‒ might not be attractive for this species. Even so, flight interception traps would have likely collected specimens were the populations substantial, suggesting that population densities are indeed quite low or that the specimens flying higher in the canopy.
</paragraph>
<paragraph id="8B8C9B60FFDAFFAD0999E318FB4BFC6C" blockId="6.[151,1437,151,897]" pageId="6" pageNumber="89">
Based on its supposed endemicity, the extent of occurrence of this species is being estimated as up to 
<quantity id="4CCB3685FFDAFFAD0C66E318FA04FDD0" box="[1336,1421,547,573]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="89" unit="km" value="100.0">100 km</quantity>
<superScript id="7C463628FFDAFFAD0CD3E319FA1CFDDD" attach="left" box="[1421,1429,546,560]" fontSize="6" pageId="6" pageNumber="89">2</superScript>
, with large surrounding areas of unsuitable habitat. The Atlantic rainforest is probably the most extinction-prone ecosystem of 
<collectingCountry id="F324DBF0FFDAFFAD0873E350FEFCFD68" box="[301,373,619,645]" name="Brazil" pageId="6" pageNumber="89">Brazil</collectingCountry>
. Its ecosystems have been suffering intense degradation early during European colonization. The major threats for the southern 
<collectingRegion id="49F75582FFDAFFAD08B2E3B4FDF4FD44" box="[492,637,655,681]" country="Brazil" name="Bahia" pageId="6" pageNumber="89">Bahia region</collectingRegion>
are urbanization (residential and industrial development), illegal wood exploitation, and impacts associated with tourism (
<bibRefCitation id="EFA2E691FFDAFFAD0B84E388FC2FFD20" author="Vieira, L. &amp; Louzada, J. &amp; Vaz-de-Mello, F. Z. &amp; Lopes, P. P. &amp; Silva, F. A. B." box="[730,934,691,717]" pageId="6" pageNumber="89" pagination="282 - 284" refId="ref10224" refString="Vieira, L., Louzada, J., Vaz-de-Mello, F. Z., Lopes, P. P. &amp; Silva, F. A. B. (2011) New records, threatens and conservation status for Dichotomius schiffleri Vaz-de-Mello, Louzada &amp; Gavino (Coleoptera: Scarabaeidae): an endangered dung beetle species from Brazilian atlantic forest ecosystems. Neotropical Entomology, 40, 282 - 284. https: // doi. org / 10.1590 / S 1519 - 566 X 2011000200020" type="journal article" year="2011">
Vieira 
<emphasis id="B9474772FFDAFFAD0A77E388FCEAFD20" box="[809,867,691,717]" italics="true" pageId="6" pageNumber="89">et al.</emphasis>
2011
</bibRefCitation>
; 
<bibRefCitation id="EFA2E691FFDAFFAD0AEAE388FBFFFD20" author="Silva, F. A. B. &amp; Moura, A. B. G. &amp; Araujo, J. F. &amp; Moura, R. C." box="[948,1142,691,717]" pageId="6" pageNumber="89" pagination="434 - 442" refId="ref9632" refString="Silva, F. A. B., Moura, A. B. G., Araujo, J. F. &amp; Moura, R. C. (2020) Brazilian Atlantic rainforest endangered biodiversity: a new species of the Dichotomius sericeus (Harold, 1867) species group (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa, 4834 (3), 434 - 442. https: // doi. org / 10.11646 / zootaxa. 4834.3.6" type="journal article" year="2020">
Silva 
<emphasis id="B9474772FFDAFFAD0AA8E388FBA0FD20" box="[1014,1065,691,717]" italics="true" pageId="6" pageNumber="89">et al</emphasis>
. 2020
</bibRefCitation>
). Therefore, we evaluated this species using the International Union for Conservation of Nature (2012) criteria and found that it could be assessed as Critically Endangered (CR) under criterion B1ab(iii)+B2ab(iii), mainly due to high fragmentation and destruction of the Atlantic rainforest, and its apparent restricted distributional range, and also due to the low number of individuals historically collected. Sellow died in 1831 (
<bibRefCitation id="EFA2E691FFDAFFAD0A48E278FC48FCB0" author="Papavero, N." box="[790,961,835,861]" pageId="6" pageNumber="89" refId="ref9218" refString="Papavero, N. (1971) Essays on the history of Neotropical dipterology: with special reference to collectors: 1750 - 1905: Volume I. Museu de Zoologia, Universidade de Sao Paulo, Sao Paulo, Brazil, 216 pp. https: // doi. org / 10.5962 / bhl. title. 101715" type="book" year="1971">Papavero 1971</bibRefCitation>
), therefore the specimen studied by Harold was certainly collected at least 190 years ago, and after that date few specimens were recorded.
</paragraph>
</subSubSection>
</treatment>
</document>