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<document id="218F580B9D5C75E7BE89544134AD7EF0" ID-DOI="10.3853/j.2201-4349.74.2022.1775" ID-ISSN="2201-4349" ID-Zenodo-Dep="7177922" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665305657841" checkinUser="felipe" docAuthor="Billingham, Zacariah D. &amp; Theischinger, Gunther" docDate="2022" docId="0392DE4AFC10FFEEFCA20BA2FD13A9DE" docLanguage="en" docName="RecAustMus.74.1.1940.pdf" docOrigin="Records of the Australian Museum (Rec. Aust. Mus.) 74 (1)" docSource="http://dx.doi.org/10.3853/j.2201-4349.74.2022.1775" docStyle="DocumentStyle:997BA24B490466A2F99594A367FA147F.3:RecAustMus.2019-.journal_article" docStyleId="997BA24B490466A2F99594A367FA147F" docStyleName="RecAustMus.2019-.journal_article" docStyleVersion="3" docTitle="Orimarga (Orimarga) joana Alexander 1926" docType="treatment" docVersion="4" lastPageNumber="38" masterDocId="FFABA632FC01FFFDFF990B35FFFEAA2C" masterDocTitle="New and Poorly Known Species of Crane Flies (Diptera: Limoniidae) from New South Wales, Australia" masterLastPageNumber="40" masterPageNumber="19" pageNumber="36" updateTime="1699383991684" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="82102FB862A212A2ED94DBD47D2EC83B">New and Poorly Known Species of Crane Flies (Diptera: Limoniidae) from New South Wales, Australia</mods:title>
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<mods:namePart id="55A59933A74ECE5C4DDD493FCAB0F7D7">Billingham, Zacariah D.</mods:namePart>
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<mods:namePart id="934FA81EE3497967C1A534F8830A5F5A">Theischinger, Gunther</mods:namePart>
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<mods:date id="C01355CCCB79144671568B5B5020EBC2">2022</mods:date>
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<taxonomicName id="4C3B14DFFC10FFECFCA20BA2FA93AA9D" ID-CoL="757BN" authority="Alexander, 1926" authorityName="Alexander" authorityYear="1926" box="[827,1389,151,177]" class="Insecta" family="Limoniidae" genus="Orimarga" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="36" phylum="Arthropoda" rank="species" species="joana" subGenus="Orimarga">
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<emphasis id="B94FB34EFC10FFECFCA20BA2FB69AA9D" bold="true" box="[827,1175,151,177]" italics="true" pageId="17" pageNumber="36">Orimarga (Orimarga) joana</emphasis>
<bibRefCitation id="EFAA12ADFC10FFECFB060BA2FA93AA9D" author="Alexander, C. P." box="[1183,1389,151,177]" pageId="17" pageNumber="36" pagination="169 - 193" refId="ref12144" refString="Alexander, C. P. 1926. New or little-known Tipulidae (Diptera). XXXII. Australasian species. Annals and Magazine of Natural History 9 (18): 169 - 193. https: // doi. org / 10.1080 / 00222932608633493" type="journal article" year="1926">Alexander, 1926</bibRefCitation>
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<figureCitation id="130073D9FC10FFECFB890BF9FB69AACB" box="[1040,1175,204,231]" captionStart-0="Figures 4143" captionStart-1="Figure 44" captionStartId-0="18.[166,243,1784,1806]" captionStartId-1="19.[121,189,684,706]" captionTargetBox-0="[167,1462,151,1762]" captionTargetBox-1="[154,1295,159,673]" captionTargetId-0="figure-10@18.[167,1467,151,1768]" captionTargetId-1="figure-340@19.[121,1421,151,683]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionText-0="Figures 4143. Orimarga joana, male.(41) thoracic colouration, lateral; (42) wing. (43) hypopygium, dorsal view, with aedeagal complex and sternite 9 inset." captionText-1="Figure 44. Orimarga joana female, ovipositor, lateral." figureDoi-0="http://doi.org/10.5281/zenodo.7177960" figureDoi-1="http://doi.org/10.5281/zenodo.7177962" httpUri-0="https://zenodo.org/record/7177960/files/figure.png" httpUri-1="https://zenodo.org/record/7177962/files/figure.png" pageId="17" pageNumber="36">Figs 4144</figureCitation>
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, GHD (
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<paragraph id="8B846F5CFC10FFECFC830923FA70A8AA" blockId="17.[794,1422,534,764]" pageId="17" pageNumber="36">
<emphasis id="B94FB34EFC10FFECFC830923FC7CA802" bold="true" box="[794,898,534,558]" pageId="17" pageNumber="36">Remarks</emphasis>
.
<bibRefCitation id="EFAA12ADFC10FFECFC150923FBB9A802" author="Alexander, C. P." box="[908,1095,534,558]" pageId="17" pageNumber="36" pagination="169 - 193" refId="ref12144" refString="Alexander, C. P. 1926. New or little-known Tipulidae (Diptera). XXXII. Australasian species. Annals and Magazine of Natural History 9 (18): 169 - 193. https: // doi. org / 10.1080 / 00222932608633493" type="journal article" year="1926">Alexander (1926)</bibRefCitation>
described
<taxonomicName id="4C3B14DFFC10FFECFB2E0922FABAA802" authorityName="Alexander" authorityYear="1926" box="[1207,1348,535,558]" class="Insecta" family="Limoniidae" genus="Orimarga" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="36" phylum="Arthropoda" rank="species" species="joana" subGenus="Orimarga">
<emphasis id="B94FB34EFC10FFECFB2E0922FABAA802" box="[1207,1348,535,558]" italics="true" pageId="17" pageNumber="36">O. (O.) joana</emphasis>
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from a unique female, leaving the male of the species undescribed. With the availability of fresh material general notes on the male and a description of the hypopygium are now possible.
</paragraph>
<paragraph id="8B846F5CFC10FFECFCA109B9FC34A8D0" blockId="17.[794,1422,534,764]" pageId="17" pageNumber="36">
Photographs are included to show a lateral view of the thoracic colouration (
<figureCitation id="130073D9FC10FFECFB9B099CFBADA8ED" box="[1026,1107,681,705]" captionStart="Figures 4143" captionStartId="18.[166,243,1784,1806]" captionTargetBox="[167,1462,151,1762]" captionTargetId="figure-10@18.[167,1467,151,1768]" captionTargetPageId="18" captionText="Figures 4143. Orimarga joana, male.(41) thoracic colouration, lateral; (42) wing. (43) hypopygium, dorsal view, with aedeagal complex and sternite 9 inset." figureDoi="http://doi.org/10.5281/zenodo.7177960" httpUri="https://zenodo.org/record/7177960/files/figure.png" pageId="17" pageNumber="36">Fig. 41</figureCitation>
), the wing (
<figureCitation id="130073D9FC10FFECFB4E099CFAD7A8ED" box="[1239,1321,681,705]" captionStart="Figures 4143" captionStartId="18.[166,243,1784,1806]" captionTargetBox="[167,1462,151,1762]" captionTargetId="figure-10@18.[167,1467,151,1768]" captionTargetPageId="18" captionText="Figures 4143. Orimarga joana, male.(41) thoracic colouration, lateral; (42) wing. (43) hypopygium, dorsal view, with aedeagal complex and sternite 9 inset." figureDoi="http://doi.org/10.5281/zenodo.7177960" httpUri="https://zenodo.org/record/7177960/files/figure.png" pageId="17" pageNumber="36">Fig. 42</figureCitation>
), and the ovipositor (
<figureCitation id="130073D9FC10FFECFC0F09F3FC19A8F2" box="[918,999,710,734]" captionStart="Figure 44" captionStartId="19.[121,189,684,706]" captionTargetBox="[154,1295,159,673]" captionTargetId="figure-340@19.[121,1421,151,683]" captionTargetPageId="19" captionText="Figure 44. Orimarga joana female, ovipositor, lateral." figureDoi="http://doi.org/10.5281/zenodo.7177962" httpUri="https://zenodo.org/record/7177962/files/figure.png" pageId="17" pageNumber="36">Fig. 44</figureCitation>
). A figure of the hypopygium (
<figureCitation id="130073D9FC10FFECFAAE09F3FA78A8F2" box="[1335,1414,710,734]" captionStart="Figures 4143" captionStartId="18.[166,243,1784,1806]" captionTargetBox="[167,1462,151,1762]" captionTargetId="figure-10@18.[167,1467,151,1768]" captionTargetPageId="18" captionText="Figures 4143. Orimarga joana, male.(41) thoracic colouration, lateral; (42) wing. (43) hypopygium, dorsal view, with aedeagal complex and sternite 9 inset." figureDoi="http://doi.org/10.5281/zenodo.7177960" httpUri="https://zenodo.org/record/7177960/files/figure.png" pageId="17" pageNumber="36">Fig. 43</figureCitation>
) is also provided.
</paragraph>
<paragraph id="8B846F5CFC10FFECFC830824FACDAE1D" blockId="17.[793,1422,785,1073]" pageId="17" pageNumber="36">
<emphasis id="B94FB34EFC10FFECFC830824FCAAA905" bold="true" box="[794,852,785,809]" pageId="17" pageNumber="36">Male</emphasis>
. Size and colouration much as described for female. Hypopygium largely yellowish brown. Distal margin of T9 broadly concave, sternite with triangular medial membranous area, narrowest at the distal margin. Gonostyli narrow and blade-like, the outer gonostylus faintly dilated towards base, weakly hooked at tip, inner gonostylus more simple, styli about ⅔ the length of the elongate gonocoxites. Aedeagus short and curving ventrad, posterior bridge of parameres narrow, dorsal arm of parameres with two short, bluntly tipped branches, the terminal branch a little larger.
</paragraph>
<paragraph id="8B846F5CFC10FFECFC830F73FB58AE9A" blockId="17.[793,1422,1094,1206]" pageId="17" pageNumber="36">
<emphasis id="B94FB34EFC10FFECFC830F73FC93AE72" bold="true" box="[794,877,1094,1118]" pageId="17" pageNumber="36">Female</emphasis>
. Size and colouration much the same as for male. Ovipositor with tergite yellowish brown, sternite dark brown. Cercus quite short, lightly curved. Tip of hypogynial valve reaching to about ¾ length of cercus.
</paragraph>
<caption id="DF443FD4FC10FFECFFE00C08FEB1AD41" ID-DOI="http://doi.org/10.5281/zenodo.7177958" ID-Zenodo-Dep="7177958" httpUri="https://zenodo.org/record/7177958/files/figure.png" pageId="17" pageNumber="36" startId="17.[121,198,1853,1875]" targetBox="[180,1324,1249,1834]" targetPageId="17">
<paragraph id="8B846F5CFC10FFECFFE00C08FEB1AD41" blockId="17.[121,1422,1853,1901]" pageId="17" pageNumber="36">
<emphasis id="B94FB34EFC10FFECFFE00C08FF1BAD7E" bold="true" box="[121,229,1853,1875]" pageId="17" pageNumber="36">Figures 40</emphasis>
.
<taxonomicName id="4C3B14DFFC10FFECFF6B0C08FD57AD7E" authorityName="Alexander" authorityYear="1922" box="[242,681,1853,1874]" class="Insecta" family="Limoniidae" genus="Elephantomyia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="36" phylum="Arthropoda" rank="species" species="fumicosta" subGenus="Elephantomyodes">
<emphasis id="B94FB34EFC10FFECFF6B0C08FD57AD7E" box="[242,681,1853,1874]" italics="true" pageId="17" pageNumber="36">Elephantomyia (Elephantomyodes) fumicosta</emphasis>
</taxonomicName>
male, hypopygium, dorsal view, with aedeagal complex inset, coiled extent of aedeagus excluded.
</paragraph>
</caption>
<caption id="DF443FD4FC13FFEFFF3F0DCDFE9CAD05" ID-DOI="http://doi.org/10.5281/zenodo.7177960" ID-Zenodo-Dep="7177960" httpUri="https://zenodo.org/record/7177960/files/figure.png" pageId="18" pageNumber="37" startId="18.[166,243,1784,1806]" targetBox="[167,1462,151,1762]" targetPageId="18">
<paragraph id="8B846F5CFC13FFEFFF3F0DCDFE9CAD05" blockId="18.[166,1466,1784,1833]" pageId="18" pageNumber="37">
<emphasis id="B94FB34EFC13FFEFFF3F0DCDFECDAD21" bold="true" box="[166,307,1784,1806]" pageId="18" pageNumber="37">Figures 4143</emphasis>
.
<taxonomicName id="4C3B14DFFC13FFEFFEA40DCDFE28AD22" authorityName="Alexander" authorityYear="1926" box="[317,470,1784,1806]" class="Insecta" family="Limoniidae" genus="Orimarga" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="37" phylum="Arthropoda" rank="species" species="joana">
<emphasis id="B94FB34EFC13FFEFFEA40DCDFE28AD22" box="[317,470,1784,1806]" italics="true" pageId="18" pageNumber="37">Orimarga joana</emphasis>
</taxonomicName>
, male. (
<emphasis id="B94FB34EFC13FFEFFDBB0DCDFDC4AD21" box="[546,570,1784,1805]" italics="true" pageId="18" pageNumber="37">41</emphasis>
) thoracic colouration, lateral; (
<emphasis id="B94FB34EFC13FFEFFCC40DCDFC8BAD21" box="[861,885,1784,1805]" italics="true" pageId="18" pageNumber="37">42</emphasis>
) wing. (
<emphasis id="B94FB34EFC13FFEFFC5D0DCDFC22AD21" box="[964,988,1784,1805]" italics="true" pageId="18" pageNumber="37">43</emphasis>
) hypopygium, dorsal view, with aedeagal complex and sternite 9 inset.
</paragraph>
</caption>
<caption id="DF443FD4FC12FFEEFFE00999FD7BA8EE" ID-DOI="http://doi.org/10.5281/zenodo.7177962" ID-Zenodo-Dep="7177962" box="[121,645,684,706]" httpUri="https://zenodo.org/record/7177962/files/figure.png" pageId="19" pageNumber="38" startId="19.[121,189,684,706]" targetBox="[154,1295,159,673]" targetPageId="19">
<paragraph id="8B846F5CFC12FFEEFFE00999FD7BA8EE" blockId="19.[121,645,684,706]" box="[121,645,684,706]" pageId="19" pageNumber="38">
<emphasis id="B94FB34EFC12FFEEFFE00999FF25A8ED" bold="true" box="[121,219,684,706]" pageId="19" pageNumber="38">Figure 44</emphasis>
.
<taxonomicName id="4C3B14DFFC12FFEEFF7E0999FE7DA8EE" authorityName="Alexander" authorityYear="1926" box="[231,387,684,706]" class="Insecta" family="Limoniidae" genus="Orimarga" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="38" phylum="Arthropoda" rank="species" species="joana">
<emphasis id="B94FB34EFC12FFEEFF7E0999FE7DA8EE" box="[231,387,684,706]" italics="true" pageId="19" pageNumber="38">Orimarga joana</emphasis>
</taxonomicName>
female, ovipositor, lateral.
</paragraph>
</caption>
<paragraph id="8B846F5CFC12FFEEFFE009C5FD13A9DE" blockId="19.[121,749,752,1011]" pageId="19" pageNumber="38">
<emphasis id="B94FB34EFC12FFEEFFE009C5FF0EA924" bold="true" box="[121,240,752,776]" pageId="19" pageNumber="38">Discussion</emphasis>
. Only three species of
<taxonomicName id="4C3B14DFFC12FFEEFE4309C4FDBDA924" authorityName="Osten Sacken" authorityYear="1869" box="[474,579,753,776]" class="Insecta" family="Limoniidae" genus="Orimarga" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="B94FB34EFC12FFEEFE4309C4FDBDA924" box="[474,579,753,776]" italics="true" pageId="19" pageNumber="38">Orimarga</emphasis>
</taxonomicName>
are known from
<collectingCountry id="F32C2FCCFC12FFEEFFE0083BFF20A90A" box="[121,222,782,806]" name="Australia" pageId="19" pageNumber="38">Australia</collectingCountry>
, seemingly occurring only in
<collectingRegion id="49FFA1BEFC12FFEEFDB0083BFD13A90A" box="[553,749,782,806]" country="Australia" name="New South Wales" pageId="19" pageNumber="38">New South Wales</collectingRegion>
from the Sydney-Blue Mountains area to the area around Wollongong.
<taxonomicName id="4C3B14DFFC12FFEEFE93087CFE12A94C" authorityName="Alexander" authorityYear="1926" box="[266,492,841,864]" class="Insecta" family="Limoniidae" genus="Orimarga" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="38" phylum="Arthropoda" rank="species" species="joana" subGenus="Orimarga">
<emphasis id="B94FB34EFC12FFEEFE93087CFE12A94C" box="[266,492,841,864]" italics="true" pageId="19" pageNumber="38">Orimarga (O.) joana</emphasis>
</taxonomicName>
is readily distinguished from
<taxonomicName id="4C3B14DFFC12FFEEFF2F0853FE02A952" authority="Skuse, 1890" authorityName="Skuse" authorityYear="1890" box="[182,508,870,894]" class="Insecta" family="Limoniidae" genus="Orimarga" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="38" phylum="Arthropoda" rank="species" species="australis" subGenus="Orimarga">
<emphasis id="B94FB34EFC12FFEEFF2F0853FE95A952" box="[182,363,870,894]" italics="true" pageId="19" pageNumber="38">O. (O.) australis</emphasis>
<bibRefCitation id="EFAA12ADFC12FFEEFEED0853FE02A952" author="Skuse, F. A. A." box="[372,508,870,894]" pageId="19" pageNumber="38" pagination="757 - 892" refId="ref12726" refString="Skuse, F. A. A. 1890. Diptera of Australia. Part VII. The Tipulidae Brevipalpi. Proceedings of the Linnean Society of New South Wales 4: 757 - 892. https: // doi. org / 10.5962 / bhl. part. 15067" type="journal article" year="1890">Skuse, 1890</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C3B14DFFC12FFEEFDA00853FEF4A9B7" authority="Skuse, 1890" authorityName="Skuse" authorityYear="1890" class="Insecta" family="Limoniidae" genus="Orimarga" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="38" phylum="Arthropoda" rank="species" species="inornata" subGenus="Orimarga">
<emphasis id="B94FB34EFC12FFEEFDA00853FD13A951" box="[569,749,870,893]" italics="true" pageId="19" pageNumber="38">O. (O.) inornata</emphasis>
<bibRefCitation id="EFAA12ADFC12FFEEFFE008B6FEF4A9B7" author="Skuse, F. A. A." box="[121,266,899,923]" pageId="19" pageNumber="38" pagination="757 - 892" refId="ref12726" refString="Skuse, F. A. A. 1890. Diptera of Australia. Part VII. The Tipulidae Brevipalpi. Proceedings of the Linnean Society of New South Wales 4: 757 - 892. https: // doi. org / 10.5962 / bhl. part. 15067" type="journal article" year="1890">Skuse, 1890</bibRefCitation>
</taxonomicName>
, by the patterned wings, particularly the coloured stigma, and by the unusual placement of crossvein r-m, at nearly ½ the length of R
<subScript id="17BF6D19FC12FFEEFE2C08FFFE31A9F4" attach="left" box="[437,463,970,984]" fontSize="6" pageId="19" pageNumber="38">4+5</subScript>
, the vein occupying a much more basal position in
<taxonomicName id="4C3B14DFFC12FFEEFEFB08EEFDF2A9DF" authorityName="Skuse" authorityYear="1890" box="[354,524,987,1011]" class="Insecta" family="Limoniidae" genus="Orimarga" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="38" phylum="Arthropoda" rank="species" species="australis" subGenus="Orimarga">
<emphasis id="B94FB34EFC12FFEEFEFB08EEFDF2A9DF" box="[354,524,987,1011]" italics="true" pageId="19" pageNumber="38">O. (O.) australis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3B14DFFC12FFEEFDA508EEFD18A9DF" authorityName="Skuse" authorityYear="1890" box="[572,742,987,1011]" class="Insecta" family="Limoniidae" genus="Orimarga" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="38" phylum="Arthropoda" rank="species" species="inornata" subGenus="Orimarga">
<emphasis id="B94FB34EFC12FFEEFDA508EEFD18A9DF" box="[572,742,987,1011]" italics="true" pageId="19" pageNumber="38">O. (O.) inornata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>