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<document id="300BCAE1719C6A01474CDA4328F805B3" ID-DOI="10.5281/zenodo.209799" ID-GBIF-Dataset="e2c293ee-749c-4e93-87b9-e7cf68d7dc20" ID-ISSN="1175-5326" ID-Zenodo-Dep="209799" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1460492091377" checkinUser="plazi" docAuthor="Olmi, Massimo &amp; Guglielmino, Adalgisa" docDate="2012" docId="CA5A4823FFBBFFB2FF4CF99EFEE8FEE4" docLanguage="en" docName="zt03351p026.pdf" docOrigin="Zootaxa 3351" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Protodryinus eocenicus Olmi &amp; Guglielmino, sp. nov." docType="treatment" docVersion="14" lastPageNumber="20" masterDocId="3663305BFFB8FFB7FFDBFF8DFF8BFFDB" masterDocTitle="A contribution to the knowledge of Dryinidae from Late Eocene Baltic amber (Hymenoptera: Chrysidoidea), with description of new subfamily, Protodryininae subfam. nov." masterLastPageNumber="26" masterPageNumber="15" pageNumber="18" updateTime="1698310245601" updateUser="plazi">
<mods:mods id="8973EE7516FFCD3AACE1A7D5E454BCCA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CEE698893109F4ACCCDC77DFBE249A25">
<mods:title id="7A6D2521DD34831674A4406B716CDD55">A contribution to the knowledge of Dryinidae from Late Eocene Baltic amber (Hymenoptera: Chrysidoidea), with description of new subfamily, Protodryininae subfam. nov.</mods:title>
</mods:titleInfo>
<mods:name id="C52B3B2A876DB8A20D887F607163461A" type="personal">
<mods:role id="40A40CBEE802BBF716DFD14CDD187707">
<mods:roleTerm id="074A3921156C2E251CFC6194DC7ABE14">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="02BBF8E600D24730F6F6CF90024478E3">Olmi, Massimo</mods:namePart>
</mods:name>
<mods:name id="773B34C3E77A0AF29A70602475A980BD" type="personal">
<mods:role id="9D49E7DA9E8C80A5C1DFD7EC75B9C88B">
<mods:roleTerm id="936CB6AA21D43947F71226163FDD9D47">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5B8BF3CE796ADBA450E81C5B81D7C7A3">Guglielmino, Adalgisa</mods:namePart>
</mods:name>
<mods:typeOfResource id="E8746445DD6DDECC5877CCDD472F23FF">text</mods:typeOfResource>
<mods:relatedItem id="7962F4FAFA3B1622171ACAC5C10023CA" type="host">
<mods:titleInfo id="A2A33BAB5ACE3467442576E25E7F11F9">
<mods:title id="6BB00D794A26F55CE62DB5B36BB7E23A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="DC60ADB8EFEF2FEA8133E896D355C8E8">
<mods:date id="2BFC17D707D1ACDC1810C801AF79D814">2012</mods:date>
<mods:detail id="8660AF6255FE40ABDCFAC9090CEE640F" type="volume">
<mods:number id="0477317CE70A50E16A030C95A3400046">3351</mods:number>
</mods:detail>
<mods:extent id="06B01B623C1AA004BFDA73034B7F7B66" unit="page">
<mods:start id="994BD9C6BFDEF52F1647531CAEC1601A">15</mods:start>
<mods:end id="EB8DFD134409F6ACBF147FA1FCA625D3">26</mods:end>
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<mods:classification id="2C425EF2D221593EF966BAC879606B68">journal article</mods:classification>
<mods:identifier id="4F1A9D9CCDDB57FA98B973765A1D4B91" type="DOI">10.5281/zenodo.209799</mods:identifier>
<mods:identifier id="57C491C9426F42CDD003AC13B5CD196E" type="GBIF-Dataset">e2c293ee-749c-4e93-87b9-e7cf68d7dc20</mods:identifier>
<mods:identifier id="C8E06C5C3782D41246AA3734479299B9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="BE20F90B15A0E7E1DA4F0A8B5F641B46" type="Zenodo-Dep">209799</mods:identifier>
</mods:mods>
<treatment id="CA5A4823FFBBFFB2FF4CF99EFEE8FEE4" ID-DOI="http://doi.org/10.5281/zenodo.6180390" ID-GBIF-Taxon="119519845" ID-Zenodo-Dep="6180390" LSID="urn:lsid:plazi:treatment:CA5A4823FFBBFFB2FF4CF99EFEE8FEE4" httpUri="http://treatment.plazi.org/id/CA5A4823FFBBFFB2FF4CF99EFEE8FEE4" lastPageId="5" lastPageNumber="20" pageId="3" pageNumber="18">
<subSubSection id="0AE9AABEFFBBFFB4FF4CF99EFE80F995" pageId="3" pageNumber="18" type="nomenclature">
<paragraph id="424CF935FFBBFFB4FF4CF99EFCACF9F7" blockId="3.[151,807,1554,1614]" box="[151,807,1554,1581]" pageId="3" pageNumber="18">
<heading id="19044E59FFBBFFB4FF4CF99EFCACF9F7" bold="true" box="[151,807,1554,1581]" fontSize="11" level="1" pageId="3" pageNumber="18" reason="1">
<emphasis id="70872527FFBBFFB4FF4CF99EFCACF9F7" bold="true" box="[151,807,1554,1581]" pageId="3" pageNumber="18">
<taxonomicName id="85F382B6FFBBFFB4FF4CF99EFD36F9F7" authority="Olmi &amp; Guglielmino" authorityName="Olmi &amp; Guglielmino" box="[151,701,1554,1581]" class="Insecta" family="Dryinidae" genus="Protodryinus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="eocenicus" status="sp. nov.">
<emphasis id="70872527FFBBFFB4FF4CF99EFE38F9F6" bold="true" box="[151,435,1555,1581]" italics="true" pageId="3" pageNumber="18">Protodryinus eocenicus</emphasis>
Olmi &amp; Guglielmino
</taxonomicName>
,
<taxonomicNameLabel id="6BB4985CFFBBFFB4FD11F99FFCACF9F7" box="[714,807,1554,1580]" pageId="3" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="424CF935FFBBFFB4FF4CF9BBFE80F995" blockId="3.[151,807,1554,1614]" box="[151,267,1590,1614]" pageId="3" pageNumber="18">
(
<figureCitation id="DAC8E5B0FFBBFFB4FF44F9BBFE88F995" box="[159,259,1590,1614]" captionStart-0="FIGURES 1, 2. 1" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="2.[151,264,1206,1229]" captionStartId-1="3.[151,250,1401,1424]" captionStartId-2="5.[151,250,942,965]" captionStartId-3="5.[151,250,1957,1980]" captionTargetBox-0="[286,1305,689,1178]" captionTargetBox-1="[176,1417,322,1356]" captionTargetBox-2="[183,1407,393,901]" captionTargetBox-3="[376,1204,1049,1928]" captionTargetId-0="figure@2.[286,1306,689,1178]" captionTargetId-1="figure@3.[151,1436,303,1377]" captionTargetId-2="figure@5.[166,1421,375,916]" captionTargetId-3="figure@5.[358,1215,1045,1930]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="5" captionText-0="FIGURES 1, 2. 1. Lonchodryinus balticus sp. nov., holotype, female, length 4.0 mm; 2. Protodryinus eocenicus gen. et sp. nov., female holotype, length 5.0 mm" captionText-1="FIGURE 3. Protodryinus eocenicus gen. et sp. nov., female holotype. A. Scutum, pronotum and head, lateral view (pronotum indicated by an arrow); B. Mesosoma, dorsal view (scutum indicated by an arrow); C. Propodeum, lateral view (indicated by an arrow); D. Fore wing (pterostigma and stigmal vein indicated by an arrow)." captionText-2="FIGURE 4. Protodryinus eocenicus gen. et sp. nov., female holotype. A. Protrochanter (indicated by an arrow); B. Chela (rudimentary claw indicated by an arrow)." captionText-3="FIGURE 5. Protodryinus eocenicus gen. et sp. nov., female holotype. Protarsus and chela (rudimentary claw indicated by an arrow)." httpUri-0="https://zenodo.org/record/209800/files/figure.png" httpUri-1="https://zenodo.org/record/209801/files/figure.png" httpUri-2="https://zenodo.org/record/209802/files/figure.png" httpUri-3="https://zenodo.org/record/209803/files/figure.png" pageId="3" pageNumber="18">Figs 2–5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0AE9AABEFFBBFFB4FF4CF9F0FE24F94C" box="[151,431,1661,1687]" pageId="3" pageNumber="18" type="diagnosis">
<paragraph id="424CF935FFBBFFB4FF4CF9F0FE24F94C" blockId="3.[151,1436,1661,2012]" box="[151,431,1661,1687]" pageId="3" pageNumber="18">
<emphasis id="70872527FFBBFFB4FF4CF9F0FE81F94D" bold="true" box="[151,266,1661,1686]" pageId="3" pageNumber="18">Diagnosis</emphasis>
. As for genus.
</paragraph>
</subSubSection>
<subSubSection id="0AE9AABEFFBBFFB4FF12F92CFBF0F8DA" pageId="3" pageNumber="18" type="materials_examined">
<paragraph id="424CF935FFBBFFB4FF12F92CFBF0F8DA" blockId="3.[151,1436,1661,2012]" pageId="3" pageNumber="18">
<emphasis id="70872527FFBBFFB4FF12F92CFE27F961" bold="true" box="[201,428,1697,1722]" pageId="3" pageNumber="18">Material examined</emphasis>
.
<typeStatus id="9D484797FFBBFFB4FE61F92CFDD9F962" box="[442,594,1697,1721]" pageId="3" pageNumber="18" type="holotype">HOLOTYPE</typeStatus>
: female, in Baltic amber.
<typeStatus id="9D484797FFBBFFB4FCA8F92FFC20F961" box="[883,939,1698,1722]" pageId="3" pageNumber="18">Type</typeStatus>
horizon: Late Eocene (40–50 mybp), Baltic amber (Succinite). The locality where the piece was originally found is unknown. The
<typeStatus id="9D484797FFBBFFB4FB41F94AFB41F904" box="[1178,1226,1735,1759]" pageId="3" pageNumber="18">type</typeStatus>
specimen will be deposited in
<collectionCode id="24E261F0FFBBFFB4FEFDF964FEFEF8DA" box="[294,373,1769,1793]" pageId="3" pageNumber="18">SNMS</collectionCode>
, though actually it is in JVC (the buying process is being dealt with).
</paragraph>
</subSubSection>
<subSubSection id="0AE9AABEFFBBFFB3FF12F880FE1BFAEB" lastPageId="4" lastPageNumber="19" pageId="3" pageNumber="18" type="description">
<paragraph id="424CF935FFBBFFB3FF12F880FD95FB33" blockId="3.[151,1436,1661,2012]" lastBlockId="4.[151,1437,151,2013]" lastPageId="4" lastPageNumber="19" pageId="3" pageNumber="18">
<emphasis id="70872527FFBBFFB4FF12F880FED1F8FD" bold="true" box="[201,346,1805,1830]" pageId="3" pageNumber="18">Description.</emphasis>
<emphasis id="70872527FFBBFFB4FEBAF883FE3EF8FD" box="[353,437,1806,1830]" italics="true" pageId="3" pageNumber="18">Female</emphasis>
. Fully winged (
<figureCitation id="DAC8E5B0FFBBFFB4FDB3F883FD3BF8FD" box="[616,688,1806,1830]" captionStart="FIGURES 1, 2. 1" captionStartId="2.[151,264,1206,1229]" captionTargetBox="[286,1305,689,1178]" captionTargetId="figure@2.[286,1306,689,1178]" captionTargetPageId="2" captionText="FIGURES 1, 2. 1. Lonchodryinus balticus sp. nov., holotype, female, length 4.0 mm; 2. Protodryinus eocenicus gen. et sp. nov., female holotype, length 5.0 mm" httpUri="https://zenodo.org/record/209800/files/figure.png" pageId="3" pageNumber="18">Figs 2</figureCitation>
,
<figureCitation id="DAC8E5B0FFBBFFB4FD66F883FD40F8FD" box="[701,715,1806,1830]" captionStart="FIGURE 3" captionStartId="3.[151,250,1401,1424]" captionTargetBox="[176,1417,322,1356]" captionTargetId="figure@3.[151,1436,303,1377]" captionTargetPageId="3" captionText="FIGURE 3. Protodryinus eocenicus gen. et sp. nov., female holotype. A. Scutum, pronotum and head, lateral view (pronotum indicated by an arrow); B. Mesosoma, dorsal view (scutum indicated by an arrow); C. Propodeum, lateral view (indicated by an arrow); D. Fore wing (pterostigma and stigmal vein indicated by an arrow)." httpUri="https://zenodo.org/record/209801/files/figure.png" pageId="3" pageNumber="18">3</figureCitation>
D); length 5.0 mm. Colour apparently black, except tarsi brown. Antenna 10-segmented, long and slender, apparently covered with dense short hairs, slightly thickened distally, without tufts of long hairs on segments 5–10; antennal rhinaria apparently present from 5th to 10th segment; in 10th segment two rhinaria are present; one rhinarium per segment is present in segments 5-9; antennal segments in following proportions: 9:6:28:26:24:19:14:13:12:14; antenna shorter than body, more than four times as long as head (approximately 180:39 = 4.62) (length of head dorsally measured from occipital carina behind ocelli to distal apex of mandible). Head laterally and dorsally visible; dorsal surface of head completely reticulate rugose; clypeus apparently with anterior margin rounded, not emarginated; occipital carina incomplete, present behind vertex, laterally do not reaching eyes, not present on temple and ventral side of head; occiput excavated; eye normally bulging, much shorter than head (22:39); ocellar triangle distinctly visible; POL = 4; OL = 3; OOL = 10; OPL = 4; TL = 9; greatest breadth of posterior ocellus about as long as OL; frontal line absent. Maxillary palpus 6-segmented. Labial palpus 3-segmented. Pronotum (
<figureCitation id="DAC8E5B0FFBCFFB3FD8BFEC1FD1AFEBF" box="[592,657,332,356]" captionStart="FIGURE 3" captionStartId="3.[151,250,1401,1424]" captionTargetBox="[176,1417,322,1356]" captionTargetId="figure@3.[151,1436,303,1377]" captionTargetPageId="3" captionText="FIGURE 3. Protodryinus eocenicus gen. et sp. nov., female holotype. A. Scutum, pronotum and head, lateral view (pronotum indicated by an arrow); B. Mesosoma, dorsal view (scutum indicated by an arrow); C. Propodeum, lateral view (indicated by an arrow); D. Fore wing (pterostigma and stigmal vein indicated by an arrow)." httpUri="https://zenodo.org/record/209801/files/figure.png" pageId="4" pageNumber="19">Fig. 3</figureCitation>
A) hairless, dull, completely granulated, excavated, crossed by strong transverse impression, without disc; dorsal region of pronotum much shorter than head (7:39); pronotal tubercle reaching tegula. Epicnemium present. Scutum (
<figureCitation id="DAC8E5B0FFBCFFB3FD66FE19FC89FE77" box="[701,770,404,428]" captionStart="FIGURE 3" captionStartId="3.[151,250,1401,1424]" captionTargetBox="[176,1417,322,1356]" captionTargetId="figure@3.[151,1436,303,1377]" captionTargetPageId="3" captionText="FIGURE 3. Protodryinus eocenicus gen. et sp. nov., female holotype. A. Scutum, pronotum and head, lateral view (pronotum indicated by an arrow); B. Mesosoma, dorsal view (scutum indicated by an arrow); C. Propodeum, lateral view (indicated by an arrow); D. Fore wing (pterostigma and stigmal vein indicated by an arrow)." httpUri="https://zenodo.org/record/209801/files/figure.png" pageId="4" pageNumber="19">Fig. 3</figureCitation>
B) shiny, hairless, strongly punctate, unsculptured among punctures, much longer than dorsal pronotal region (25:7). Notauli complete (
<figureCitation id="DAC8E5B0FFBCFFB3FBDAFE3AFBC8FE14" box="[1025,1091,439,463]" captionStart="FIGURE 3" captionStartId="3.[151,250,1401,1424]" captionTargetBox="[176,1417,322,1356]" captionTargetId="figure@3.[151,1436,303,1377]" captionTargetPageId="3" captionText="FIGURE 3. Protodryinus eocenicus gen. et sp. nov., female holotype. A. Scutum, pronotum and head, lateral view (pronotum indicated by an arrow); B. Mesosoma, dorsal view (scutum indicated by an arrow); C. Propodeum, lateral view (indicated by an arrow); D. Fore wing (pterostigma and stigmal vein indicated by an arrow)." httpUri="https://zenodo.org/record/209801/files/figure.png" pageId="4" pageNumber="19">Fig. 3</figureCitation>
B), posteriorly separated; minimum distance between notauli longer than antennal segment 2 (9:6). Scutellum shorter than scutum (9:25), apparently strongly punctate. Metanotum convex, smooth, apparently unsculptured, shorter than scutellum (5:9). Propodeum (
<figureCitation id="DAC8E5B0FFBCFFB3FEFDFDA9FEE6FDE7" box="[294,365,548,572]" captionStart="FIGURE 3" captionStartId="3.[151,250,1401,1424]" captionTargetBox="[176,1417,322,1356]" captionTargetId="figure@3.[151,1436,303,1377]" captionTargetPageId="3" captionText="FIGURE 3. Protodryinus eocenicus gen. et sp. nov., female holotype. A. Scutum, pronotum and head, lateral view (pronotum indicated by an arrow); B. Mesosoma, dorsal view (scutum indicated by an arrow); C. Propodeum, lateral view (indicated by an arrow); D. Fore wing (pterostigma and stigmal vein indicated by an arrow)." httpUri="https://zenodo.org/record/209801/files/figure.png" pageId="4" pageNumber="19">Figs 3</figureCitation>
B, C) completely sculptured by many transverse keels, longer than scutellum (24:9), with dorsal surface much shorter than posterior surface (9:15); posterior surface without longitudinal keels. Mesopleuron and metapleuron reticulate rugose. Meso-metapleural suture complete. Fore wing (
<figureCitation id="DAC8E5B0FFBCFFB3FBF6FDE1FBF8FD5F" box="[1069,1139,620,644]" captionStart="FIGURE 3" captionStartId="3.[151,250,1401,1424]" captionTargetBox="[176,1417,322,1356]" captionTargetId="figure@3.[151,1436,303,1377]" captionTargetPageId="3" captionText="FIGURE 3. Protodryinus eocenicus gen. et sp. nov., female holotype. A. Scutum, pronotum and head, lateral view (pronotum indicated by an arrow); B. Mesosoma, dorsal view (scutum indicated by an arrow); C. Propodeum, lateral view (indicated by an arrow); D. Fore wing (pterostigma and stigmal vein indicated by an arrow)." httpUri="https://zenodo.org/record/209801/files/figure.png" pageId="4" pageNumber="19">Fig. 3</figureCitation>
D) apparently completely darkened, with three basal cells completely enclosed by pigmented veins (costa, median and submedian); pterostigma (
<figureCitation id="DAC8E5B0FFBCFFB3FED9FD39FECDFD17" box="[258,326,692,716]" captionStart="FIGURE 3" captionStartId="3.[151,250,1401,1424]" captionTargetBox="[176,1417,322,1356]" captionTargetId="figure@3.[151,1436,303,1377]" captionTargetPageId="3" captionText="FIGURE 3. Protodryinus eocenicus gen. et sp. nov., female holotype. A. Scutum, pronotum and head, lateral view (pronotum indicated by an arrow); B. Mesosoma, dorsal view (scutum indicated by an arrow); C. Propodeum, lateral view (indicated by an arrow); D. Fore wing (pterostigma and stigmal vein indicated by an arrow)." httpUri="https://zenodo.org/record/209801/files/figure.png" pageId="4" pageNumber="19">Fig. 3</figureCitation>
D) much longer than broad (32:10); marginal cell open; stigmal vein composed of strongly pigmented straight and short proximal part and less pigmented long distal part; stigmal vein (
<figureCitation id="DAC8E5B0FFBCFFB3FB6BFD5AFB7EFD34" box="[1200,1269,727,751]" captionStart="FIGURE 3" captionStartId="3.[151,250,1401,1424]" captionTargetBox="[176,1417,322,1356]" captionTargetId="figure@3.[151,1436,303,1377]" captionTargetPageId="3" captionText="FIGURE 3. Protodryinus eocenicus gen. et sp. nov., female holotype. A. Scutum, pronotum and head, lateral view (pronotum indicated by an arrow); B. Mesosoma, dorsal view (scutum indicated by an arrow); C. Propodeum, lateral view (indicated by an arrow); D. Fore wing (pterostigma and stigmal vein indicated by an arrow)." httpUri="https://zenodo.org/record/209801/files/figure.png" pageId="4" pageNumber="19">Fig. 3</figureCitation>
D) forming an angle between proximal and distal less sclerotized part; distal less sclerotized part much longer than proximal part (38:9), not reaching margin of fore wing.
<collectingCountry id="3AE4B9A5FFBCFFB3FDA8FC92FD20FCEC" box="[627,683,799,823]" name="India" pageId="4" pageNumber="19">Hind</collectingCountry>
wing not visible, hidden under fore wing. Mesosoma shorter than metasoma (70:94), longer than head (70:39). Petiole very short, much shorter than metasoma (2:94). Foreleg segments in following proportions: 16 (coxa): 7 (trochanter): 45 (femur): 39 (tibia): 13 (tarsal segment 1): 10 (tarsal segment 2): 7 (tarsal segment 3): 5 (tarsal segment 4): 12 (tarsal segment 5). Enlarged claw slightly shorter than protarsal segment 5 (9:12). Protrochanter (
<figureCitation id="DAC8E5B0FFBCFFB3FDADFC22FD32FC1C" box="[630,697,943,967]" captionStart="FIGURE 4" captionStartId="5.[151,250,942,965]" captionTargetBox="[183,1407,393,901]" captionTargetId="figure@5.[166,1421,375,916]" captionTargetPageId="5" captionText="FIGURE 4. Protodryinus eocenicus gen. et sp. nov., female holotype. A. Protrochanter (indicated by an arrow); B. Chela (rudimentary claw indicated by an arrow)." httpUri="https://zenodo.org/record/209802/files/figure.png" pageId="4" pageNumber="19">Fig. 4</figureCitation>
A) large, not slender, not curved, longer than broad (7:4), without long proximal stalk and distal broadened apex. Segment 3 of protarsus apparently produced into hook. Rudimentary claw (
<figureCitation id="DAC8E5B0FFBCFFB3FED6FC7AFEDDFBD4" box="[269,342,1015,1039]" captionStart="FIGURE 4" captionStartId="5.[151,250,942,965]" captionTargetBox="[183,1407,393,901]" captionTargetId="figure@5.[166,1421,375,916]" captionTargetPageId="5" captionText="FIGURE 4. Protodryinus eocenicus gen. et sp. nov., female holotype. A. Protrochanter (indicated by an arrow); B. Chela (rudimentary claw indicated by an arrow)." httpUri="https://zenodo.org/record/209802/files/figure.png" pageId="4" pageNumber="19">Figs 4</figureCitation>
B, 5) present. Arolium much shorter than enlarged claw (2:9); inner side of enlarged claw not visible, because of closed chela. Segment 5 of protarsus with inner side not visible, with basal part about as long as distal part. Segment 5 of protarsus less than twice as broad as enlarged claw. Midleg segments in following proportions: 15 (coxa): 5 (trochanter): 38 (femur): 41(tibia): 18 (tarsal segment 1): 7 (tarsal segment 2): 6 (tarsal segment 3): 5 (tarsal segment 4): 8 (tarsal segment 5).
<collectingCountry id="3AE4B9A5FFBCFFB3FD54FB05FD4CFB7B" box="[655,711,1160,1184]" name="India" pageId="4" pageNumber="19">Hind</collectingCountry>
leg segments in following proportions: 27 (coxa): 8 (trochanter): 40 (femur): 50 (tibia): 21 (tarsal segment 1): 12 (tarsal segment 2): 9 (tarsal segment 3): 5 (tarsal segment 4): 5 (tarsal segment 5). Tibial spurs 1, 1, 2.
</paragraph>
<paragraph id="424CF935FFBCFFB3FF12FB79FE01FAD7" blockId="4.[151,1437,151,2013]" box="[201,394,1268,1292]" pageId="4" pageNumber="19">
<emphasis id="70872527FFBCFFB3FF12FB79FE8DFAD7" bold="true" box="[201,262,1268,1292]" pageId="4" pageNumber="19">Male</emphasis>
. Unknown.
</paragraph>
<paragraph id="424CF935FFBCFFB3FF12FA95FE1BFAEB" blockId="4.[151,1437,151,2013]" box="[201,400,1304,1328]" pageId="4" pageNumber="19">
<emphasis id="70872527FFBCFFB3FF12FA95FE87FAEB" bold="true" box="[201,268,1304,1328]" pageId="4" pageNumber="19">Hosts</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="0AE9AABEFFBCFFB3FF12FAB1FDBBFA8F" box="[201,560,1340,1364]" pageId="4" pageNumber="19" type="etymology">
<paragraph id="424CF935FFBCFFB3FF12FAB1FDBBFA8F" blockId="4.[151,1437,151,2013]" box="[201,560,1340,1364]" pageId="4" pageNumber="19">
<emphasis id="70872527FFBCFFB3FF12FAB1FEC3FA8F" bold="true" box="[201,328,1340,1364]" pageId="4" pageNumber="19">Etymology</emphasis>
. From Baltic amber.
</paragraph>
</subSubSection>
<subSubSection id="0AE9AABEFFBCFFB2FF12FAEDFEE8FEE4" lastPageId="5" lastPageNumber="20" pageId="4" pageNumber="19" type="discussion">
<paragraph id="424CF935FFBCFFB2FF12FAEDFDA1FF2C" blockId="4.[151,1437,151,2013]" lastBlockId="5.[151,1436,151,319]" lastPageId="5" lastPageNumber="20" pageId="4" pageNumber="19">
<emphasis id="70872527FFBCFFB3FF12FAEDFEBDFAA3" bold="true" box="[201,310,1376,1400]" pageId="4" pageNumber="19">Remarks</emphasis>
.
<taxonomicName id="85F382B6FFBCFFB3FE9EFAEDFDC5FAA3" box="[325,590,1376,1400]" class="Insecta" family="Dryinidae" genus="Protodryinus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="eocenicus" status="sp. nov.">
<emphasis id="70872527FFBCFFB3FE9EFAEDFDC5FAA3" box="[325,590,1376,1400]" italics="true" pageId="4" pageNumber="19">Protodryinus eocenicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6BB4985CFFBCFFB3FD8CFAEDFD39FAA3" box="[599,690,1376,1400]" pageId="4" pageNumber="19" rank="species">
<emphasis id="70872527FFBCFFB3FD8CFAEDFD39FAA3" bold="true" box="[599,690,1376,1400]" pageId="4" pageNumber="19">sp. nov.</emphasis>
</taxonomicNameLabel>
, at first sight, for the general aspect of the body, the shape of the pronotum (with dorsal surface excavated) (
<figureCitation id="DAC8E5B0FFBCFFB3FDA5FA09FD4AFA47" box="[638,705,1412,1436]" captionStart="FIGURE 3" captionStartId="3.[151,250,1401,1424]" captionTargetBox="[176,1417,322,1356]" captionTargetId="figure@3.[151,1436,303,1377]" captionTargetPageId="3" captionText="FIGURE 3. Protodryinus eocenicus gen. et sp. nov., female holotype. A. Scutum, pronotum and head, lateral view (pronotum indicated by an arrow); B. Mesosoma, dorsal view (scutum indicated by an arrow); C. Propodeum, lateral view (indicated by an arrow); D. Fore wing (pterostigma and stigmal vein indicated by an arrow)." httpUri="https://zenodo.org/record/209801/files/figure.png" pageId="4" pageNumber="19">Fig. 3</figureCitation>
A) and propodeum (with dorsal surface much longer than posterior surface) (
<figureCitation id="DAC8E5B0FFBCFFB3FEEFFA25FEFCFA1B" box="[308,375,1448,1472]" captionStart="FIGURE 3" captionStartId="3.[151,250,1401,1424]" captionTargetBox="[176,1417,322,1356]" captionTargetId="figure@3.[151,1436,303,1377]" captionTargetPageId="3" captionText="FIGURE 3. Protodryinus eocenicus gen. et sp. nov., female holotype. A. Scutum, pronotum and head, lateral view (pronotum indicated by an arrow); B. Mesosoma, dorsal view (scutum indicated by an arrow); C. Propodeum, lateral view (indicated by an arrow); D. Fore wing (pterostigma and stigmal vein indicated by an arrow)." httpUri="https://zenodo.org/record/209801/files/figure.png" pageId="4" pageNumber="19">Fig. 3</figureCitation>
C) and the presence of a rudimentary claw (
<figureCitation id="DAC8E5B0FFBCFFB3FCA9FA25FC36FA1B" box="[882,957,1448,1472]" captionStart="FIGURE 4" captionStartId="5.[151,250,942,965]" captionTargetBox="[183,1407,393,901]" captionTargetId="figure@5.[166,1421,375,916]" captionTargetPageId="5" captionText="FIGURE 4. Protodryinus eocenicus gen. et sp. nov., female holotype. A. Protrochanter (indicated by an arrow); B. Chela (rudimentary claw indicated by an arrow)." httpUri="https://zenodo.org/record/209802/files/figure.png" pageId="4" pageNumber="19">Figs 4</figureCitation>
B, 5), seems to be a species of
<taxonomicName id="85F382B6FFBCFFB3FAFEFA25FA12FA1B" box="[1317,1433,1448,1472]" class="Insecta" family="Dryinidae" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="subFamily" subFamily="Dryininae">Dryininae</taxonomicName>
. However, the following characters contradict the first impression: 1) the shape of the protrochanter (protrochanter large, less than twice as long as broad (7:4) (i.e. plesiomorphic) (
<figureCitation id="DAC8E5B0FFBCFFB3FC52FA7DFC46F9D3" box="[905,973,1520,1544]" captionStart="FIGURE 4" captionStartId="5.[151,250,942,965]" captionTargetBox="[183,1407,393,901]" captionTargetId="figure@5.[166,1421,375,916]" captionTargetPageId="5" captionText="FIGURE 4. Protodryinus eocenicus gen. et sp. nov., female holotype. A. Protrochanter (indicated by an arrow); B. Chela (rudimentary claw indicated by an arrow)." httpUri="https://zenodo.org/record/209802/files/figure.png" pageId="4" pageNumber="19">Fig. 4</figureCitation>
A), not slender, not curved, without long proximal stalk and distal broadened apex); 2) the shape of the stigmal vein (stigmal vein composed of strongly pigmented straight and short proximal part and less pigmented long distal part) (
<figureCitation id="DAC8E5B0FFBCFFB3FB9AF9B5FB0CF98B" box="[1089,1159,1592,1616]" captionStart="FIGURE 3" captionStartId="3.[151,250,1401,1424]" captionTargetBox="[176,1417,322,1356]" captionTargetId="figure@3.[151,1436,303,1377]" captionTargetPageId="3" captionText="FIGURE 3. Protodryinus eocenicus gen. et sp. nov., female holotype. A. Scutum, pronotum and head, lateral view (pronotum indicated by an arrow); B. Mesosoma, dorsal view (scutum indicated by an arrow); C. Propodeum, lateral view (indicated by an arrow); D. Fore wing (pterostigma and stigmal vein indicated by an arrow)." httpUri="https://zenodo.org/record/209801/files/figure.png" pageId="4" pageNumber="19">Fig. 3</figureCitation>
D); 3) the shape of the pterostigma (pterostigma very large and broad, less than four times as long as broad: 32:10) (
<figureCitation id="DAC8E5B0FFBCFFB3FB72F9D1FB61F9AF" box="[1193,1258,1628,1652]" captionStart="FIGURE 3" captionStartId="3.[151,250,1401,1424]" captionTargetBox="[176,1417,322,1356]" captionTargetId="figure@3.[151,1436,303,1377]" captionTargetPageId="3" captionText="FIGURE 3. Protodryinus eocenicus gen. et sp. nov., female holotype. A. Scutum, pronotum and head, lateral view (pronotum indicated by an arrow); B. Mesosoma, dorsal view (scutum indicated by an arrow); C. Propodeum, lateral view (indicated by an arrow); D. Fore wing (pterostigma and stigmal vein indicated by an arrow)." httpUri="https://zenodo.org/record/209801/files/figure.png" pageId="4" pageNumber="19">Fig. 3</figureCitation>
D); 4) the shape of the protarsomere 5 (with basal part about as long as distal part) (
<figureCitation id="DAC8E5B0FFBCFFB3FC72F90DFC7FF943" box="[937,1012,1664,1688]" captionStart="FIGURE 4" captionStartId="5.[151,250,942,965]" captionTargetBox="[183,1407,393,901]" captionTargetId="figure@5.[166,1421,375,916]" captionTargetPageId="5" captionText="FIGURE 4. Protodryinus eocenicus gen. et sp. nov., female holotype. A. Protrochanter (indicated by an arrow); B. Chela (rudimentary claw indicated by an arrow)." httpUri="https://zenodo.org/record/209802/files/figure.png" pageId="4" pageNumber="19">Figs 4</figureCitation>
B, 5). The above four characters and mainly the shape of protrochanter are not present in
<taxonomicName id="85F382B6FFBCFFB3FD2CF929FCE1F967" box="[759,874,1700,1724]" class="Insecta" family="Dryinidae" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="subFamily" subFamily="Dryininae">Dryininae</taxonomicName>
. The typical shape of protrochanter in
<taxonomicName id="85F382B6FFBCFFB3FAF7F929FA17F967" box="[1324,1436,1700,1724]" class="Insecta" family="Dryinidae" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="subFamily" subFamily="Dryininae">Dryininae</taxonomicName>
(very long and slender protrochanter, with a long stalk and a rounded and broad distal apex) (
<figureCitation id="DAC8E5B0FFBCFFB3FB1BF945FA88F93B" box="[1216,1283,1736,1760]" captionStart="FIGURE 6" captionStartId="7.[151,250,1030,1053]" captionTargetBox="[301,1294,210,1005]" captionTargetId="figure@7.[285,1303,197,1009]" captionTargetPageId="7" captionText="FIGURE 6. Megadryinus magnificus Richards, female holotype, lateral view (from Olmi, 1984), length 6.0 mm. Protrochanter (indicated by an arrow), pterostigma and stigmal vein." httpUri="https://zenodo.org/record/209804/files/figure.png" pageId="4" pageNumber="19">Fig. 6</figureCitation>
) is a peculiar synapomorphy common also to
<taxonomicName id="85F382B6FFBCFFB3FDCBF961FD35F8DF" box="[528,702,1772,1796]" class="Insecta" family="Dryinidae" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="subFamily" subFamily="Gonatopodinae">Gonatopodinae</taxonomicName>
, Transdryininae and Palaeoanteoninae. This character, together with the extraordinary length of the foreleg, increases the raptorial ability of
<taxonomicName id="85F382B6FFBCFFB3FBE5F89DFB25F8F3" box="[1086,1198,1808,1832]" class="Insecta" family="Dryinidae" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="subFamily" subFamily="Dryininae">Dryininae</taxonomicName>
and
<taxonomicName id="85F382B6FFBCFFB3FB2AF89DFA17F8F3" box="[1265,1436,1808,1832]" class="Insecta" family="Dryinidae" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="subFamily" subFamily="Gonatopodinae">Gonatopodinae</taxonomicName>
females. In
<taxonomicName id="85F382B6FFBCFFB3FEC2F8B8FE29F897" box="[281,418,1845,1868]" class="Insecta" family="Dryinidae" genus="Protodryinus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="eocenicus">
<emphasis id="70872527FFBCFFB3FEC2F8B8FE29F897" box="[281,418,1845,1868]" italics="true" pageId="4" pageNumber="19">P. eocenicus</emphasis>
</taxonomicName>
the foreleg is very long, as in
<taxonomicName id="85F382B6FFBCFFB3FD29F8B9FCE9F897" box="[754,866,1844,1868]" class="Insecta" family="Dryinidae" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="subFamily" subFamily="Dryininae">Dryininae</taxonomicName>
and
<taxonomicName id="85F382B6FFBCFFB3FC43F8B9FBCDF897" box="[920,1094,1844,1868]" class="Insecta" family="Dryinidae" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="subFamily" subFamily="Gonatopodinae">Gonatopodinae</taxonomicName>
, but the trochanter is short and squat. In addition, in
<taxonomicName id="85F382B6FFBCFFB3FE59F8D5FE79F8AB" box="[386,498,1880,1904]" class="Insecta" family="Dryinidae" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="subFamily" subFamily="Dryininae">Dryininae</taxonomicName>
the pterostigma (
<figureCitation id="DAC8E5B0FFBCFFB3FD6FF8D5FD7DF8AB" box="[692,758,1880,1904]" captionStart="FIGURE 6" captionStartId="7.[151,250,1030,1053]" captionTargetBox="[301,1294,210,1005]" captionTargetId="figure@7.[285,1303,197,1009]" captionTargetPageId="7" captionText="FIGURE 6. Megadryinus magnificus Richards, female holotype, lateral view (from Olmi, 1984), length 6.0 mm. Protrochanter (indicated by an arrow), pterostigma and stigmal vein." httpUri="https://zenodo.org/record/209804/files/figure.png" pageId="4" pageNumber="19">Fig. 6</figureCitation>
) is always more than four times as long as broad, the stigmal vein (
<figureCitation id="DAC8E5B0FFBCFFB3FF0CF8F1FE90F84F" box="[215,283,1916,1940]" captionStart="FIGURE 6" captionStartId="7.[151,250,1030,1053]" captionTargetBox="[301,1294,210,1005]" captionTargetId="figure@7.[285,1303,197,1009]" captionTargetPageId="7" captionText="FIGURE 6. Megadryinus magnificus Richards, female holotype, lateral view (from Olmi, 1984), length 6.0 mm. Protrochanter (indicated by an arrow), pterostigma and stigmal vein." httpUri="https://zenodo.org/record/209804/files/figure.png" pageId="4" pageNumber="19">Fig. 6</figureCitation>
) is always composed of a proximal and distal part, both at least partly well pigmented, and the protarsomere 5 has always a basal part much shorter than distal part. The shapes of pterostigma, stigmal vein, protrochanter and segment 5 of protarsus characterize
<taxonomicName id="85F382B6FFBCFFB3FD21F849FCF0F807" box="[762,891,1988,2012]" class="Insecta" family="Dryinidae" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="subFamily" subFamily="Anteoninae">Anteoninae</taxonomicName>
and never can be found in
<taxonomicName id="85F382B6FFBCFFB3FB29F849FAE9F807" box="[1266,1378,1988,2012]" class="Insecta" family="Dryinidae" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="subFamily" subFamily="Dryininae">Dryininae</taxonomicName>
and
<taxonomicName id="85F382B6FFBDFFB2FF4CFF1AFECDFF74" box="[151,326,151,175]" class="Insecta" family="Dryinidae" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="subFamily" subFamily="Gonatopodinae">Gonatopodinae</taxonomicName>
. The combination of the above characters cannot be found in any subfamilies of
<taxonomicName id="85F382B6FFBDFFB2FB15FF1AFAC9FF74" box="[1230,1346,151,175]" class="Insecta" family="Dryinidae" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="family">Dryinidae</taxonomicName>
. For the above reasons,
<taxonomicName id="85F382B6FFBDFFB2FE97FF31FE54FF0F" box="[332,479,188,212]" class="Insecta" family="Dryinidae" genus="Protodryinus" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="70872527FFBDFFB2FE97FF31FE54FF0F" box="[332,479,188,212]" italics="true" pageId="5" pageNumber="20">Protodryinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6BB4985CFFBDFFB2FE37FF31FDD2FF0F" box="[492,601,188,212]" pageId="5" pageNumber="20" rank="genus">
<emphasis id="70872527FFBDFFB2FE37FF31FDD2FF0F" bold="true" box="[492,601,188,212]" pageId="5" pageNumber="20">gen. nov.</emphasis>
</taxonomicNameLabel>
, belongs to Protodryininae
<emphasis id="70872527FFBDFFB2FC7FFF36FBCBFF0F" bold="true" box="[932,1088,187,212]" pageId="5" pageNumber="20">subfam. nov.</emphasis>
, with characters of transition between
<taxonomicName id="85F382B6FFBDFFB2FF20FF52FEF7FF2C" box="[251,380,223,247]" class="Insecta" family="Dryinidae" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="subFamily" subFamily="Anteoninae">Anteoninae</taxonomicName>
and
<taxonomicName id="85F382B6FFBDFFB2FE68FF52FDACFF2C" box="[435,551,223,247]" class="Insecta" family="Dryinidae" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="subFamily" subFamily="Dryininae">Dryininae</taxonomicName>
.
</paragraph>
<paragraph id="424CF935FFBDFFB2FF12FE89FEE8FEE4" blockId="5.[151,1436,151,319]" pageId="5" pageNumber="20">
Protodryininae can be included in the key to
<taxonomicName id="85F382B6FFBDFFB2FD64FE89FCA4FEC7" box="[703,815,260,284]" class="Insecta" family="Dryinidae" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="family">Dryinidae</taxonomicName>
subfamilies females presented by Olmi &amp; Guglielmino (2010) as follows:
</paragraph>
</subSubSection>
</treatment>
</document>
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