343 lines
52 KiB
XML
343 lines
52 KiB
XML
<document id="07D6EA8177385CBC090B859F4891AFCD" ID-DOI="10.11646/zootaxa.5497.2.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="13618317" ID-ZooBank="B3C66D95-3585-4920-BE93-A44D33FB2FBB" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="admin" IM.treatments_approvedBy="tatiana" checkinTime="1725025887147" checkinUser="plazi" docAuthor="Orr, Albert G. W." docDate="2024" docId="937387ADE034D750FF79EA82FA56F88C" docLanguage="en" docName="zootaxa.5497.2.3.pdf" docOrigin="Zootaxa 5497 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5497.2.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calopterygoidea Sélys 1850" docType="key" docVersion="2" lastPageNumber="211" masterDocId="6F4AFFD5E035D752FFEEEC20FFCAFFDC" masterDocTitle="A review of present knowledge of larvae of the Calopterygoidea (Zygoptera) of the Oriental realm, including keys to families and known genera" masterLastPageNumber="243" masterPageNumber="209" pageNumber="210" updateTime="1731029118091" updateUser="admin" zenodo-license-document="CLOSED">
|
||
<mods:mods id="306DB6B80E0B88FBF6C4CAB1D01CD54C" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="3A89E3992F21CCE90B171C0530EC73D3">
|
||
<mods:title id="657CC22718B7A3DEEF506722E102354E">A review of present knowledge of larvae of the Calopterygoidea (Zygoptera) of the Oriental realm, including keys to families and known genera</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="E2681BD3B37C59FEF12A451A490986FB" type="personal">
|
||
<mods:role id="B50FDF374331863D21E80FFAE5991BE4">
|
||
<mods:roleTerm id="CED45B27B121641F9BDA50CE1FC6F76B">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="26EFDBF7806175AD9F6A7DCFD7EA44E8">Orr, Albert G. W.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="3108600AE3D3B9387B10B55249D918B2">text</mods:typeOfResource>
|
||
<mods:relatedItem id="39CAE2119E49C9DF7BD13F8FE5F5B60D" type="host">
|
||
<mods:titleInfo id="DF2C68804D52750376312668F6D11CD6">
|
||
<mods:title id="106DA3267F91AA04D1AB1FB46F5770E6">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="C00C60FC7F9E2D0763245BC4099F448E">
|
||
<mods:date id="B836917A0CD5209CCCEC8717F2F1EE9A">2024</mods:date>
|
||
<mods:detail id="D074D61F538F4D7D8D3973849D9CDBB9" type="pubDate">
|
||
<mods:number id="1999E1F756B70AC05E326A48F5963A52">2024-08-23</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="732A063FA4A5CC380CFD1DB7211A7701" type="volume">
|
||
<mods:number id="63D15485A1B7A7DD77628E882AB5BF5C">5497</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="FC91AFCDEB209677F1EBFDFB6E7227B6" type="issue">
|
||
<mods:number id="04B9FFB454CCCEE3F2AC334C68131FFA">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="AAF17986CD625EAF7F02F0318C9B9ADB" unit="page">
|
||
<mods:start id="7DBE9CBE03C40DE884FA6D5E2D7EA9EF">209</mods:start>
|
||
<mods:end id="7C61117D018BAF49FB411D420046586E">243</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="A6FE10D30E66393F5CB6C086C98A3DB9">
|
||
<mods:url id="93535FF8185D6753D7080CF5D67797D6">http://dx.doi.org/10.11646/zootaxa.5497.2.3</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="318730C78D562F054E00C4D56452BD82">journal article</mods:classification>
|
||
<mods:identifier id="1AC76B220A670623275821E4D1AB5E94" type="DOI">10.11646/zootaxa.5497.2.3</mods:identifier>
|
||
<mods:identifier id="79BEFC0985C6D41EAD41E479793858C4" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="DB22C88DA0726AF4B7FC2C78627ADE1D" type="Zenodo-Dep">13618317</mods:identifier>
|
||
<mods:identifier id="FAE2CBF09CDD8FAA39247F2788B149A1" type="ZooBank">B3C66D95-3585-4920-BE93-A44D33FB2FBB</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="937387ADE034D750FF79EA82FA56F88C" LSID="urn:lsid:plazi:treatment:937387ADE034D750FF79EA82FA56F88C" httpUri="http://treatment.plazi.org/id/937387ADE034D750FF79EA82FA56F88C" lastPageId="2" lastPageNumber="211" pageId="1" pageNumber="210">
|
||
<subSubSection id="53C06530E034D753FF79EA82FDC2F960" box="[151,520,1698,1724]" pageId="1" pageNumber="210" type="nomenclature">
|
||
<paragraph id="1B6536BBE034D753FF79EA82FDC2F960" blockId="1.[151,520,1698,1724]" box="[151,520,1698,1724]" pageId="1" pageNumber="210">
|
||
<heading id="402D81D7E034D753FF79EA82FDC2F960" bold="true" box="[151,520,1698,1724]" fontSize="11" level="1" pageId="1" pageNumber="210" reason="1">
|
||
<emphasis id="29AEEAA9E034D753FF79EA82FDC2F960" bold="true" box="[151,520,1698,1724]" pageId="1" pageNumber="210">
|
||
Superfamily
|
||
<taxonomicName id="DCDA4D38E034D753FED7EA82FDC2F960" authorityName="Sélys" authorityYear="1850" box="[313,520,1698,1724]" class="Insecta" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="210" phylum="Arthropoda" rank="superFamily" superFamily="Calopterygoidea">Calopterygoidea</taxonomicName>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="53C06530E034D750FF79EACBFC68FDE1" lastPageId="2" lastPageNumber="211" pageId="1" pageNumber="210" type="description">
|
||
<paragraph id="1B6536BBE034D750FF79EACBFD7EFE71" blockId="1.[151,1436,1771,2013]" lastBlockId="2.[151,1436,151,429]" lastPageId="2" lastPageNumber="211" pageId="1" pageNumber="210">
|
||
The superfamily
|
||
<taxonomicName id="DCDA4D38E034D753FEB7EACBFDDFF8D9" box="[345,533,1771,1797]" pageId="1" pageNumber="210" rank="superFamily" superFamily="Calopterygoidea">Calopterygoidea</taxonomicName>
|
||
, as it is presently understood, is a relatively new assemblage of taxa which has been recognised in recent years through the morphological studies of several authors, such as
|
||
<bibRefCitation id="7F4B4B4AE034D753FB34EB2FFAA0F8F5" author="Rehn, A. C." box="[1242,1386,1807,1833]" pageId="1" pageNumber="210" pagination="181 - 239" refId="ref26517" refString="Rehn, A. C. (2003) Phylogenetic analysis of higher-level relationships of Odonata. Systematic Entomology, 28, 181 - 239. https: // doi. org / 10.1046 / j. 1365 - 3113.2003.00210. x" type="journal article" year="2003">Rehn (2003)</bibRefCitation>
|
||
and molecular studies, notably by
|
||
<bibRefCitation id="7F4B4B4AE034D753FE0BEB13FD04F891" author="Dijkstra, K-D. B. & Kalkman, V. J. & Dow, R. A. & Stokvis, F. R. & van Tol, J." box="[485,718,1843,1869]" pageId="1" pageNumber="210" pagination="68 - 96" refId="ref22480" refString="Dijkstra, K-D. B., Kalkman, V. J., Dow, R. A., Stokvis, F. R. & van Tol, J. (2014) Redefining the damselfly families: a comprehensive molecular phylogeny of Zygoptera (Odonata). Systematic entomology, 39, 68 - 96. https: // doi. org / 10.1111 / syen. 12035" type="journal article" year="2014">
|
||
Dijkstra
|
||
<emphasis id="29AEEAA9E034D753FDA8EB13FDBCF891" box="[582,630,1843,1869]" italics="true" pageId="1" pageNumber="210">et al</emphasis>
|
||
. (2014)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="7F4B4B4AE034D753FCEDEB13FC16F891" author="Bybee, S. M. & Kalkman, V. J. & Erickson, R. J. & Breinholt, J. W. & Suvorov, A. & Dijkstra, K-D. B. & Cordero-Riviera, A. & Skevington, J. H. & Abbott, J. C. & Sanchez Herrera, M. & Lemmon, A. R. & Moriaty Lemmon, E. & Ware, J. L." box="[771,988,1843,1869]" pageId="1" pageNumber="210" pagination="1 - 15" refId="ref22012" refString="Bybee, S. M., Kalkman, V. J., Erickson, R. J., Breinholt, J. W., Suvorov, A., Dijkstra, K-D. B., Cordero-Riviera, A., Skevington, J. H., Abbott, J. C., Sanchez Herrera, M., Lemmon, A. R., Moriaty Lemmon, E. & Ware, J. L. (2021) Phylogeny and classification of Odonata using targeted genomics. Molecular Phylogenetics and Evolution, 160, 1 - 15. https: // doi. org / 10.1016 / j. ympev. 2021.107115" type="journal article" year="2021">
|
||
Bybee
|
||
<emphasis id="29AEEAA9E034D753FCBFEB13FC48F891" box="[849,898,1843,1869]" italics="true" pageId="1" pageNumber="210">et al</emphasis>
|
||
. (2021)
|
||
</bibRefCitation>
|
||
. One unexpected result of the molecular studies has been the inclusion within the group of the family
|
||
<taxonomicName id="DCDA4D38E034D753FCBBEB77FC24F8AD" authorityName="Fraser" authorityYear="1960" box="[853,1006,1879,1905]" class="Insecta" family="Argiolestidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="210" phylum="Arthropoda" rank="family">Argiolestidae</taxonomicName>
|
||
, defined by
|
||
<bibRefCitation id="7F4B4B4AE034D753FB93EB77FF2FF849" author="Kalkman, V. J. & Theischinger, G." pageId="1" pageNumber="210" pagination="1 - 52" refId="ref23895" refString="Kalkman, V. J. & Theischinger, G. (2013) Generic revision of Argiolestidae (Odonata), with four new genera. International Journal of Odonatology, 16, 1 - 52. https: // doi. org / 10.1080 / 13887890.2012.749450" type="journal article" year="2013">Kalkman & Theischinger (2013)</bibRefCitation>
|
||
. However both molecular studies assess the
|
||
<taxonomicName id="DCDA4D38E034D753FD35EB5BFC5EF849" box="[731,916,1915,1941]" pageId="1" pageNumber="210" rank="superFamily" superFamily="Calopterygoidea">Calopterygoidea</taxonomicName>
|
||
as paraphyletic (Djikstra
|
||
<emphasis id="29AEEAA9E034D753FB56EB5BFB23F849" box="[1208,1257,1915,1941]" italics="true" pageId="1" pageNumber="210">et al</emphasis>
|
||
. 2014) or ‘nonmonophyletic’ (
|
||
<bibRefCitation id="7F4B4B4AE034D753FEABEBBFFDC1F865" author="Bybee, S. M. & Kalkman, V. J. & Erickson, R. J. & Breinholt, J. W. & Suvorov, A. & Dijkstra, K-D. B. & Cordero-Riviera, A. & Skevington, J. H. & Abbott, J. C. & Sanchez Herrera, M. & Lemmon, A. R. & Moriaty Lemmon, E. & Ware, J. L." box="[325,523,1951,1977]" pageId="1" pageNumber="210" pagination="1 - 15" refId="ref22012" refString="Bybee, S. M., Kalkman, V. J., Erickson, R. J., Breinholt, J. W., Suvorov, A., Dijkstra, K-D. B., Cordero-Riviera, A., Skevington, J. H., Abbott, J. C., Sanchez Herrera, M., Lemmon, A. R., Moriaty Lemmon, E. & Ware, J. L. (2021) Phylogeny and classification of Odonata using targeted genomics. Molecular Phylogenetics and Evolution, 160, 1 - 15. https: // doi. org / 10.1016 / j. ympev. 2021.107115" type="journal article" year="2021">
|
||
Bybee
|
||
<emphasis id="29AEEAA9E034D753FE7DEBBFFE09F865" box="[403,451,1951,1977]" italics="true" pageId="1" pageNumber="210">et al</emphasis>
|
||
. 2021
|
||
</bibRefCitation>
|
||
), but while the former authors left many minor taxa in the category ‘
|
||
<emphasis id="29AEEAA9E034D753FB1DEBBFFA5FF865" box="[1267,1429,1951,1977]" italics="true" pageId="1" pageNumber="210">incertae sedis’</emphasis>
|
||
, the latter did assign taxa to named families within
|
||
<taxonomicName id="DCDA4D38E034D753FD35EBE3FC5EF801" box="[731,916,1987,2013]" pageId="1" pageNumber="210" rank="superFamily" superFamily="Calopterygoidea">Calopterygoidea</taxonomicName>
|
||
(as ‘Calopterygoidea’ see above). Worldwide it includes 26 families and one
|
||
<emphasis id="29AEEAA9E037D750FE12ECB7FD52FF6D" box="[508,664,151,177]" italics="true" pageId="2" pageNumber="211">incertae sedis</emphasis>
|
||
group (
|
||
<bibRefCitation id="7F4B4B4AE037D750FD1CECB7FC0AFF6D" author="Bybee, S. M. & Kalkman, V. J. & Erickson, R. J. & Breinholt, J. W. & Suvorov, A. & Dijkstra, K-D. B. & Cordero-Riviera, A. & Skevington, J. H. & Abbott, J. C. & Sanchez Herrera, M. & Lemmon, A. R. & Moriaty Lemmon, E. & Ware, J. L." box="[754,960,151,177]" pageId="2" pageNumber="211" pagination="1 - 15" refId="ref22012" refString="Bybee, S. M., Kalkman, V. J., Erickson, R. J., Breinholt, J. W., Suvorov, A., Dijkstra, K-D. B., Cordero-Riviera, A., Skevington, J. H., Abbott, J. C., Sanchez Herrera, M., Lemmon, A. R., Moriaty Lemmon, E. & Ware, J. L. (2021) Phylogeny and classification of Odonata using targeted genomics. Molecular Phylogenetics and Evolution, 160, 1 - 15. https: // doi. org / 10.1016 / j. ympev. 2021.107115" type="journal article" year="2021">
|
||
Bybee
|
||
<emphasis id="29AEEAA9E037D750FCADECB7FCBFFF6D" box="[835,885,151,177]" italics="true" pageId="2" pageNumber="211">et al</emphasis>
|
||
. 2021
|
||
</bibRefCitation>
|
||
). It is the family arrangement of
|
||
<bibRefCitation id="7F4B4B4AE037D750FAD6ECB7FECDFF09" author="Bybee, S. M. & Kalkman, V. J. & Erickson, R. J. & Breinholt, J. W. & Suvorov, A. & Dijkstra, K-D. B. & Cordero-Riviera, A. & Skevington, J. H. & Abbott, J. C. & Sanchez Herrera, M. & Lemmon, A. R. & Moriaty Lemmon, E. & Ware, J. L." pageId="2" pageNumber="211" pagination="1 - 15" refId="ref22012" refString="Bybee, S. M., Kalkman, V. J., Erickson, R. J., Breinholt, J. W., Suvorov, A., Dijkstra, K-D. B., Cordero-Riviera, A., Skevington, J. H., Abbott, J. C., Sanchez Herrera, M., Lemmon, A. R., Moriaty Lemmon, E. & Ware, J. L. (2021) Phylogeny and classification of Odonata using targeted genomics. Molecular Phylogenetics and Evolution, 160, 1 - 15. https: // doi. org / 10.1016 / j. ympev. 2021.107115" type="journal article" year="2021">
|
||
Bybee
|
||
<emphasis id="29AEEAA9E037D750FA66ECB7FF7EFF09" italics="true" pageId="2" pageNumber="211">et al.</emphasis>
|
||
(2021)
|
||
</bibRefCitation>
|
||
that is followed here. This comprises 11 families in the Oriental realm, and for 10 of them, at least
|
||
<specimenCount id="0DDCFD32E037D750FA9AEC9BFF05FF25" count="1" pageId="2" pageNumber="211" type="larva">one larva</specimenCount>
|
||
is known. The key is expected to generally be reliable at the family level, with the possible exception of some genera of the weakly defined
|
||
<taxonomicName id="DCDA4D38E037D750FE0AED23FD52FEC1" box="[484,664,259,285]" family="Rhipidolestidae" pageId="2" pageNumber="237" rank="family">Rhipidolestidae</taxonomicName>
|
||
. Larvae of the newly erected family
|
||
<taxonomicName id="DCDA4D38E037D750FBDCED23FB29FEC1" box="[1074,1251,259,285]" family="Priscagrionidae" pageId="2" pageNumber="211" rank="family">Priscagrionidae</taxonomicName>
|
||
, first recognised in
|
||
<bibRefCitation id="7F4B4B4AE037D750FF56ED07FE54FE9D" author="Bybee, S. M. & Kalkman, V. J. & Erickson, R. J. & Breinholt, J. W. & Suvorov, A. & Dijkstra, K-D. B. & Cordero-Riviera, A. & Skevington, J. H. & Abbott, J. C. & Sanchez Herrera, M. & Lemmon, A. R. & Moriaty Lemmon, E. & Ware, J. L." box="[184,414,295,321]" pageId="2" pageNumber="211" pagination="1 - 15" refId="ref22012" refString="Bybee, S. M., Kalkman, V. J., Erickson, R. J., Breinholt, J. W., Suvorov, A., Dijkstra, K-D. B., Cordero-Riviera, A., Skevington, J. H., Abbott, J. C., Sanchez Herrera, M., Lemmon, A. R., Moriaty Lemmon, E. & Ware, J. L. (2021) Phylogeny and classification of Odonata using targeted genomics. Molecular Phylogenetics and Evolution, 160, 1 - 15. https: // doi. org / 10.1016 / j. ympev. 2021.107115" type="journal article" year="2021">
|
||
Bybee
|
||
<emphasis id="29AEEAA9E037D750FEE4ED07FE8AFE9D" box="[266,320,295,321]" italics="true" pageId="2" pageNumber="211">et al</emphasis>
|
||
. (2021)
|
||
</bibRefCitation>
|
||
, remain unknown. Within the region larvae of the large families
|
||
<taxonomicName id="DCDA4D38E037D750FB70ED07FA83FE9D" box="[1182,1353,295,321]" class="Insecta" family="Calopterygidae" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="211" phylum="Arthropoda" rank="family">Calopterygidae</taxonomicName>
|
||
(11/12 genera),
|
||
<taxonomicName id="DCDA4D38E037D750FF17ED6BFEB6FEB9" baseAuthorityName="Jacobson & Bianchi" baseAuthorityYear="1905" box="[249,380,331,357]" class="Insecta" family="Euphaeidae" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="211" phylum="Arthropoda" rank="family">Euphaeidae</taxonomicName>
|
||
(5/8 genera) and
|
||
<taxonomicName id="DCDA4D38E037D750FDADED6BFD3CFEB9" authorityName="Cowley" authorityYear="1937" box="[579,758,331,357]" class="Insecta" family="Chlorocyphidae" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="211" phylum="Arthropoda" rank="family">Chlorocyphidae</taxonomicName>
|
||
(5/16 genera) are unevenly known. The generic divisions of the
|
||
<taxonomicName id="DCDA4D38E037D750FF2FED4FFEBEFE55" authorityName="Cowley" authorityYear="1937" box="[193,372,367,393]" class="Insecta" family="Chlorocyphidae" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="211" phylum="Arthropoda" rank="family">Chlorocyphidae</taxonomicName>
|
||
are not fully resolved, and only small morphological differences have been found between the larvae of even quite distant genera in the family.
|
||
</paragraph>
|
||
<paragraph id="1B6536BBE037D750FF79EDDEFDBEFDC4" blockId="2.[151,930,510,573]" box="[151,628,510,537]" pageId="2" pageNumber="211">
|
||
<heading id="402D81D7E037D750FF79EDDEFDBEFDC4" bold="true" box="[151,628,510,537]" fontSize="11" level="1" pageId="2" pageNumber="211" reason="1">
|
||
<emphasis id="29AEEAA9E037D750FF79EDDEFDBEFDC4" bold="true" box="[151,628,510,537]" pageId="2" pageNumber="211">
|
||
Family key Oriental
|
||
<taxonomicName id="DCDA4D38E037D750FE75EDDEFDA5FDC4" box="[411,623,510,536]" pageId="2" pageNumber="210" rank="superFamily" superFamily="Calopterygoidea">Calopterygoidea</taxonomicName>
|
||
:
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="1B6536BBE037D750FF79EE03FC68FDE1" blockId="2.[151,930,510,573]" box="[151,930,547,573]" pageId="2" pageNumber="211">
|
||
larva unknown:
|
||
<taxonomicName id="DCDA4D38E037D750FEA3EE03FE31FDE1" box="[333,507,547,573]" family="Priscagrionidae" pageId="2" pageNumber="211" rank="family">Priscagrionidae</taxonomicName>
|
||
(genera
|
||
<taxonomicName id="DCDA4D38E037D750FDB3EE03FD29FDE0" authorityName="Zhou & Wilson" authorityYear="2001" box="[605,739,547,572]" class="Insecta" family="Megapodagrionidae" genus="Priscagrion" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="211" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="29AEEAA9E037D750FDB3EE03FD29FDE0" box="[605,739,547,572]" italics="true" pageId="2" pageNumber="211">Priscagrion</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="DCDA4D38E037D750FCF7EE03FC53FDE1" authorityName="Wilson & Zhou" authorityYear="2000" box="[793,921,547,573]" class="Insecta" family="Megapodagrionidae" genus="Sinocnemis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="211" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="29AEEAA9E037D750FCF7EE03FC53FDE1" box="[793,921,547,573]" italics="true" pageId="2" pageNumber="211">Sinocnemis</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="53C06530E037D750FF79EE48FA56F88C" pageId="2" pageNumber="211" type="key">
|
||
<key id="6F4B618AE037D750FF79EE48FA56F88C" pageId="2" pageNumber="211">
|
||
<keyStep id="A02E2E1EE037D750FF79EE48FA56FD29" pageId="2" pageNumber="211">
|
||
<paragraph id="1B6536BBE037D750FF79EE48FA56FD67" blockId="2.[151,1437,616,1872]" pageId="2" pageNumber="211">
|
||
<keyLead id="A02B958EE037D750FF79EE48FA56FD67" pageId="2" pageNumber="211">
|
||
1 Abdomen with two ventral rows of long fleshy, filamentous gills on S2–8; build generally robust; caudal gills saccoid with terminal filament or blunt process (
|
||
<figureCitation id="83E12A3EE037D750FDDAEEA5FDA6FD46" box="[564,620,645,667]" captionStart="FIGURES 1–7. 1" captionStartId="3.[151,275,1843,1869]" captionTargetBox="[252,1341,160,1825]" captionTargetId="figure-15@3.[243,1344,149,1836]" captionTargetPageId="3" captionText="FIGURES 1–7. 1. Ventral view of abdomen of Euphaea sp. showing abdominal gills and saccoid gills (original drawing). 2. Head and antennae of Calopterygidae (Neurobasis chinensis) (original drawing). 3. Head and antennae of Chlorocyphidae (Heliocypha biseriata) (original drawing). 4. Antenna of Podolestes orientalis Selys, 1852, scape arrowed (from Choong & Orr 2010). 5. Mask of Calopterygidae (Calopteryx japonica Selys redrawn from Miyakawa 1983). 6. Mask of Chlorocyphidae (Libellago hyalina, from Orr et al. 2024). 7. Habitus of L. hyalina, with detail of right caudal spike (from Orr et al. 2024)." figureDoi="http://doi.org/10.5281/zenodo.13618321" httpUri="https://zenodo.org/record/13618321/files/figure.png" pageId="2" pageNumber="211">Fig. 1</figureCitation>
|
||
).........................................................
|
||
<taxonomicName id="DCDA4D38E037D750FACDEEA5FA56FD47" baseAuthorityName="Jacobson & Bianchi" baseAuthorityYear="1905" box="[1315,1436,645,667]" class="Insecta" family="Euphaeidae" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="211" phylum="Arthropoda" rank="family">
|
||
<emphasis id="29AEEAA9E037D750FACDEEA5FA56FD47" bold="true" box="[1315,1436,645,667]" pageId="2" pageNumber="211">Euphaeidae</emphasis>
|
||
</taxonomicName>
|
||
[absent from
|
||
<collectingRegion id="D91EF859E037D750FADDEE83FA5CFD67" box="[1331,1430,675,699]" country="Indonesia" name="Sulawesi Utara" pageId="2" pageNumber="211">Sulawesi</collectingRegion>
|
||
]
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="1B6536BBE037D750FF79EEE2FA56FD29" blockId="2.[151,1437,616,1872]" pageId="2" pageNumber="211">
|
||
<keyLead id="A02B958EE037D750FF79EEE2FA56FD29" pageId="2" pageNumber="211">1’ Abdomen lacking long fleshy gills on venter of abdomen, build very slender to robust, caudal gills of variable form, including saccoid............................................................................................. 2</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="A02E2E1EE037D750FF79EEDDFA56FC91" pageId="2" pageNumber="211">
|
||
<paragraph id="1B6536BBE037D750FF79EEDDFA56FCF3" blockId="2.[151,1437,616,1872]" pageId="2" pageNumber="211">
|
||
<keyLead id="A02B958EE037D750FF79EEDDFA56FCF3" pageId="2" pageNumber="211">
|
||
2 (1’) Antennal scape at least one third total antennal length; antennal sockets set in raised prominence; (
|
||
<figureCitation id="83E12A3EE037D750FB6BEEDDFB13FCCE" box="[1157,1241,765,787]" captionStart="FIGURES 1–7. 1" captionStartId="3.[151,275,1843,1869]" captionTargetBox="[252,1341,160,1825]" captionTargetId="figure-15@3.[243,1344,149,1836]" captionTargetPageId="3" captionText="FIGURES 1–7. 1. Ventral view of abdomen of Euphaea sp. showing abdominal gills and saccoid gills (original drawing). 2. Head and antennae of Calopterygidae (Neurobasis chinensis) (original drawing). 3. Head and antennae of Chlorocyphidae (Heliocypha biseriata) (original drawing). 4. Antenna of Podolestes orientalis Selys, 1852, scape arrowed (from Choong & Orr 2010). 5. Mask of Calopterygidae (Calopteryx japonica Selys redrawn from Miyakawa 1983). 6. Mask of Chlorocyphidae (Libellago hyalina, from Orr et al. 2024). 7. Habitus of L. hyalina, with detail of right caudal spike (from Orr et al. 2024)." figureDoi="http://doi.org/10.5281/zenodo.13618321" httpUri="https://zenodo.org/record/13618321/files/figure.png" pageId="2" pageNumber="211">Figs 2, 3</figureCitation>
|
||
); very small to large species with long legs; moderately robust to very slender build.................................................. 3
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="1B6536BBE037D750FF79EF18FA56FC91" blockId="2.[151,1437,616,1872]" box="[151,1436,824,846]" pageId="2" pageNumber="211">
|
||
<keyLead id="A02B958EE037D750FF79EF18FA56FC91" box="[151,1436,824,846]" pageId="2" pageNumber="211">
|
||
2’ Antennal scape less than one third total antennal length (
|
||
<figureCitation id="83E12A3EE037D750FCECEF18FCF6FC91" box="[770,828,824,846]" captionStart="FIGURES 1–7. 1" captionStartId="3.[151,275,1843,1869]" captionTargetBox="[252,1341,160,1825]" captionTargetId="figure-15@3.[243,1344,149,1836]" captionTargetPageId="3" captionText="FIGURES 1–7. 1. Ventral view of abdomen of Euphaea sp. showing abdominal gills and saccoid gills (original drawing). 2. Head and antennae of Calopterygidae (Neurobasis chinensis) (original drawing). 3. Head and antennae of Chlorocyphidae (Heliocypha biseriata) (original drawing). 4. Antenna of Podolestes orientalis Selys, 1852, scape arrowed (from Choong & Orr 2010). 5. Mask of Calopterygidae (Calopteryx japonica Selys redrawn from Miyakawa 1983). 6. Mask of Chlorocyphidae (Libellago hyalina, from Orr et al. 2024). 7. Habitus of L. hyalina, with detail of right caudal spike (from Orr et al. 2024)." figureDoi="http://doi.org/10.5281/zenodo.13618321" httpUri="https://zenodo.org/record/13618321/files/figure.png" pageId="2" pageNumber="211">Fig. 4</figureCitation>
|
||
),................................................ 4
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="A02E2E1EE037D750FF79EF75FA57FC3C" pageId="2" pageNumber="211">
|
||
<paragraph id="1B6536BBE037D750FF79EF75FA56FC54" blockId="2.[151,1437,616,1872]" pageId="2" pageNumber="211">
|
||
<keyLead id="A02B958EE037D750FF79EF75FA56FC54" pageId="2" pageNumber="211">
|
||
3(2) Prementum with long lateral lobes and often deep median cleft (
|
||
<figureCitation id="83E12A3EE037D750FCD3EF75FCBDFCB7" box="[829,887,853,875]" captionStart="FIGURES 1–7. 1" captionStartId="3.[151,275,1843,1869]" captionTargetBox="[252,1341,160,1825]" captionTargetId="figure-15@3.[243,1344,149,1836]" captionTargetPageId="3" captionText="FIGURES 1–7. 1. Ventral view of abdomen of Euphaea sp. showing abdominal gills and saccoid gills (original drawing). 2. Head and antennae of Calopterygidae (Neurobasis chinensis) (original drawing). 3. Head and antennae of Chlorocyphidae (Heliocypha biseriata) (original drawing). 4. Antenna of Podolestes orientalis Selys, 1852, scape arrowed (from Choong & Orr 2010). 5. Mask of Calopterygidae (Calopteryx japonica Selys redrawn from Miyakawa 1983). 6. Mask of Chlorocyphidae (Libellago hyalina, from Orr et al. 2024). 7. Habitus of L. hyalina, with detail of right caudal spike (from Orr et al. 2024)." figureDoi="http://doi.org/10.5281/zenodo.13618321" httpUri="https://zenodo.org/record/13618321/files/figure.png" pageId="2" pageNumber="211">Fig. 5</figureCitation>
|
||
); antennal scape mostly longer than pedicel and flagellum together (
|
||
<figureCitation id="83E12A3EE037D750FED0EF52FEB3FC5B" box="[318,377,882,904]" captionStart="FIGURES 1–7. 1" captionStartId="3.[151,275,1843,1869]" captionTargetBox="[252,1341,160,1825]" captionTargetId="figure-15@3.[243,1344,149,1836]" captionTargetPageId="3" captionText="FIGURES 1–7. 1. Ventral view of abdomen of Euphaea sp. showing abdominal gills and saccoid gills (original drawing). 2. Head and antennae of Calopterygidae (Neurobasis chinensis) (original drawing). 3. Head and antennae of Chlorocyphidae (Heliocypha biseriata) (original drawing). 4. Antenna of Podolestes orientalis Selys, 1852, scape arrowed (from Choong & Orr 2010). 5. Mask of Calopterygidae (Calopteryx japonica Selys redrawn from Miyakawa 1983). 6. Mask of Chlorocyphidae (Libellago hyalina, from Orr et al. 2024). 7. Habitus of L. hyalina, with detail of right caudal spike (from Orr et al. 2024)." figureDoi="http://doi.org/10.5281/zenodo.13618321" httpUri="https://zenodo.org/record/13618321/files/figure.png" pageId="2" pageNumber="211">Fig. 2</figureCitation>
|
||
); three well developed caudal gills of variable form; medium to large size...............
|
||
<taxonomicName id="DCDA4D38E037D750FB11EF52FA56FC54" box="[1279,1436,882,904]" class="Insecta" family="Calopterygidae" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="211" phylum="Arthropoda" rank="family">
|
||
<emphasis id="29AEEAA9E037D750FB11EF52FA56FC54" bold="true" box="[1279,1436,882,904]" pageId="2" pageNumber="211">Calopterygidae</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="1B6536BBE037D750FF79EFB0FA57FC3C" blockId="2.[151,1437,616,1872]" pageId="2" pageNumber="211">
|
||
<keyLead id="A02B958EE037D750FF79EFB0FA57FC3C" pageId="2" pageNumber="211">
|
||
3’ Prementum without long lateral lobes or deep median cleft (
|
||
<figureCitation id="83E12A3EE037D750FCAEEFB0FCB4FC7A" box="[832,894,912,934]" captionStart="FIGURES 1–7. 1" captionStartId="3.[151,275,1843,1869]" captionTargetBox="[252,1341,160,1825]" captionTargetId="figure-15@3.[243,1344,149,1836]" captionTargetPageId="3" captionText="FIGURES 1–7. 1. Ventral view of abdomen of Euphaea sp. showing abdominal gills and saccoid gills (original drawing). 2. Head and antennae of Calopterygidae (Neurobasis chinensis) (original drawing). 3. Head and antennae of Chlorocyphidae (Heliocypha biseriata) (original drawing). 4. Antenna of Podolestes orientalis Selys, 1852, scape arrowed (from Choong & Orr 2010). 5. Mask of Calopterygidae (Calopteryx japonica Selys redrawn from Miyakawa 1983). 6. Mask of Chlorocyphidae (Libellago hyalina, from Orr et al. 2024). 7. Habitus of L. hyalina, with detail of right caudal spike (from Orr et al. 2024)." figureDoi="http://doi.org/10.5281/zenodo.13618321" httpUri="https://zenodo.org/record/13618321/files/figure.png" pageId="2" pageNumber="211">Fig. 6</figureCitation>
|
||
); caudal gills reduced to two long, thin lateral spikes, triangular in section, between 0.5–0.9 times length of abdomen, (
|
||
<figureCitation id="83E12A3EE037D750FC84EF8DFC6DFC1E" box="[874,935,941,963]" captionStart="FIGURES 1–7. 1" captionStartId="3.[151,275,1843,1869]" captionTargetBox="[252,1341,160,1825]" captionTargetId="figure-15@3.[243,1344,149,1836]" captionTargetPageId="3" captionText="FIGURES 1–7. 1. Ventral view of abdomen of Euphaea sp. showing abdominal gills and saccoid gills (original drawing). 2. Head and antennae of Calopterygidae (Neurobasis chinensis) (original drawing). 3. Head and antennae of Chlorocyphidae (Heliocypha biseriata) (original drawing). 4. Antenna of Podolestes orientalis Selys, 1852, scape arrowed (from Choong & Orr 2010). 5. Mask of Calopterygidae (Calopteryx japonica Selys redrawn from Miyakawa 1983). 6. Mask of Chlorocyphidae (Libellago hyalina, from Orr et al. 2024). 7. Habitus of L. hyalina, with detail of right caudal spike (from Orr et al. 2024)." figureDoi="http://doi.org/10.5281/zenodo.13618321" httpUri="https://zenodo.org/record/13618321/files/figure.png" pageId="2" pageNumber="211">Fig. 7</figureCitation>
|
||
), vestigial central gill barely apparent as a conical epiproct; build moderately stout, small to very small............................................
|
||
<taxonomicName id="DCDA4D38E037D750FB17EFEAFA57FC3C" authorityName="Cowley" authorityYear="1937" box="[1273,1437,970,992]" class="Insecta" family="Chlorocyphidae" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="211" phylum="Arthropoda" rank="family">
|
||
<emphasis id="29AEEAA9E037D750FB17EFEAFA57FC3C" bold="true" box="[1273,1437,970,992]" pageId="2" pageNumber="211">Chlorocyphidae</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="A02E2E1EE037D750FF79EFC8FA56FBE4" pageId="2" pageNumber="211">
|
||
<paragraph id="1B6536BBE037D750FF79EFC8FA56FBC7" blockId="2.[151,1437,616,1872]" pageId="2" pageNumber="211">
|
||
<keyLead id="A02B958EE037D750FF79EFC8FA56FBC7" pageId="2" pageNumber="211">
|
||
4 (2’). Caudal gills bulbous and sclerotised, either ovate or conical with blunt trifid tips; venter of S10 with paired retractable gill tufts (
|
||
<figureCitation id="83E12A3EE037D750FF07E825FEF5FBC6" box="[233,319,1029,1051]" captionStart="FIGURES 8–11. 8" captionStartId="4.[151,264,1914,1938]" captionTargetBox="[200,1382,177,1900]" captionTargetId="figure-21@4.[193,1394,169,1908]" captionTargetPageId="4" captionText="FIGURES 8–11. 8. Pseudolestes mirabilis, ventral view of terminal abdominal segments and caudal gills of showing gill tufts on S10 (after Yu & Bu 2011). 9. Devadatta argyoides, detail of extruded right gill tuft, posterior view (redrawn from Watson 1966). 10. Pseudolestes mirabilis, habitus with detail of mask (after Yu & Bu 2011). 11. Devadatta argyoides, habitus with details of right antenna and mask (original drawing)." figureDoi="http://doi.org/10.5281/zenodo.13618323" httpUri="https://zenodo.org/record/13618323/files/figure.png" pageId="2" pageNumber="211">Figs 8, 9</figureCitation>
|
||
), sometimes just evident as tiny tumescent orifices when retracted; stoutly built............................ 5
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="1B6536BBE037D750FF79E802FA56FBE4" blockId="2.[151,1437,616,1872]" box="[151,1436,1058,1080]" pageId="2" pageNumber="211">
|
||
<keyLead id="A02B958EE037D750FF79E802FA56FBE4" box="[151,1436,1058,1080]" pageId="2" pageNumber="211">4’ Caudal gills not as above, either soft and saccoid, or else laterally or ventrally flattened. Ventral gill tufts on S10 absent.... 6</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="A02E2E1EE037D750FF79E81FFA56FB34" pageId="2" pageNumber="211">
|
||
<paragraph id="1B6536BBE037D750FF79E81FFA56FB4C" blockId="2.[151,1437,616,1872]" pageId="2" pageNumber="211">
|
||
<keyLead id="A02B958EE037D750FF79E81FFA56FB4C" pageId="2" pageNumber="211">
|
||
5(4) Caudal gills subspherical hardened balloons with slight acuminate tip (
|
||
<figureCitation id="83E12A3EE037D750FC67E860FC1AFB89" box="[905,976,1088,1110]" captionStart="FIGURES 8–11. 8" captionStartId="4.[151,264,1914,1938]" captionTargetBox="[200,1382,177,1900]" captionTargetId="figure-21@4.[193,1394,169,1908]" captionTargetPageId="4" captionText="FIGURES 8–11. 8. Pseudolestes mirabilis, ventral view of terminal abdominal segments and caudal gills of showing gill tufts on S10 (after Yu & Bu 2011). 9. Devadatta argyoides, detail of extruded right gill tuft, posterior view (redrawn from Watson 1966). 10. Pseudolestes mirabilis, habitus with detail of mask (after Yu & Bu 2011). 11. Devadatta argyoides, habitus with details of right antenna and mask (original drawing)." figureDoi="http://doi.org/10.5281/zenodo.13618323" httpUri="https://zenodo.org/record/13618323/files/figure.png" pageId="2" pageNumber="211">Fig. 10</figureCitation>
|
||
), antennae normally proportioned with scape ca half length of pedicel; prementum long, narrow and gently tapered...................................
|
||
<taxonomicName id="DCDA4D38E037D750FAE8E87DFA57FBAF" authorityName="Fraser" authorityYear="1957" box="[1286,1437,1117,1139]" class="Insecta" family="Pseudolestidae" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="211" phylum="Arthropoda" rank="family">
|
||
<emphasis id="29AEEAA9E037D750FAE8E87DFA57FBAF" bold="true" box="[1286,1437,1117,1139]" pageId="2" pageNumber="211">Pseudolestidae</emphasis>
|
||
</taxonomicName>
|
||
[endemic to
|
||
<collectingRegion id="D91EF859E037D750FAA1E85AFA5DFB4C" box="[1359,1431,1146,1168]" country="China" name="Hainan" pageId="2" pageNumber="211">Hainan</collectingRegion>
|
||
]
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="1B6536BBE037D750FF79E8B7FA56FB34" blockId="2.[151,1437,616,1872]" pageId="2" pageNumber="211">
|
||
<keyLead id="A02B958EE037D750FF79E8B7FA56FB34" pageId="2" pageNumber="211">
|
||
5’ Caudal gills hardened, conical, median tricorn (
|
||
<figureCitation id="83E12A3EE037D750FD5CE8B8FD30FB71" box="[690,762,1176,1198]" captionStart="FIGURES 8–11. 8" captionStartId="4.[151,264,1914,1938]" captionTargetBox="[200,1382,177,1900]" captionTargetId="figure-21@4.[193,1394,169,1908]" captionTargetPageId="4" captionText="FIGURES 8–11. 8. Pseudolestes mirabilis, ventral view of terminal abdominal segments and caudal gills of showing gill tufts on S10 (after Yu & Bu 2011). 9. Devadatta argyoides, detail of extruded right gill tuft, posterior view (redrawn from Watson 1966). 10. Pseudolestes mirabilis, habitus with detail of mask (after Yu & Bu 2011). 11. Devadatta argyoides, habitus with details of right antenna and mask (original drawing)." figureDoi="http://doi.org/10.5281/zenodo.13618323" httpUri="https://zenodo.org/record/13618323/files/figure.png" pageId="2" pageNumber="211">Fig. 11</figureCitation>
|
||
); first three segments of antennae abnormally thick with scape about twice length of pedicel; prementum broad, squared................................................
|
||
<taxonomicName id="DCDA4D38E037D750FAFAE895FA56FB17" authorityName="Dijkstra, Kalkman, Dow, Stokvis & van Tol" authorityYear="2014" box="[1300,1436,1205,1227]" class="Insecta" family="Devadattidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="211" phylum="Arthropoda" rank="family">
|
||
<emphasis id="29AEEAA9E037D750FAFAE895FA56FB17" bold="true" box="[1300,1436,1205,1227]" pageId="2" pageNumber="211">Devadattidae</emphasis>
|
||
</taxonomicName>
|
||
[mainland south-east Asia from southern
|
||
<collectingRegion id="D91EF859E037D750FB59E8F3FACAFB34" box="[1207,1280,1235,1256]" country="China" name="Yunnan" pageId="2" pageNumber="211">Yunnan</collectingRegion>
|
||
and Sundaland]
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="A02E2E1EE037D750FF79E8D0FA56FAFE" pageId="2" pageNumber="211">
|
||
<paragraph id="1B6536BBE037D750FF79E8D0FA56FAD9" blockId="2.[151,1437,616,1872]" box="[151,1436,1264,1286]" pageId="2" pageNumber="211">
|
||
<keyLead id="A02B958EE037D750FF79E8D0FA56FAD9" box="[151,1436,1264,1286]" pageId="2" pageNumber="211">6 (4’) Caudal gills flattened, foliate and arranged horizontally in a broad fan........................................... 7</keyLead>
|
||
</paragraph>
|
||
<paragraph id="1B6536BBE037D750FF79E92DFA56FAFE" blockId="2.[151,1437,616,1872]" box="[151,1436,1293,1315]" pageId="2" pageNumber="211">
|
||
<keyLead id="A02B958EE037D750FF79E92DFA56FAFE" box="[151,1436,1293,1315]" pageId="2" pageNumber="211">6’ Caudal gills either saccoid or broadly flattened and arranged vertically........................................... 8</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="A02E2E1EE037D750FF79E90AFA56FA2C" pageId="2" pageNumber="211">
|
||
<paragraph id="1B6536BBE037D750FF79E90AFA56FA44" blockId="2.[151,1437,616,1872]" pageId="2" pageNumber="211">
|
||
<keyLead id="A02B958EE037D750FF79E90AFA56FA44" pageId="2" pageNumber="211">
|
||
7(6) Stoutly built with broad, acutely tipped, foliate caudal gills ca 0.6 length of abdomen (
|
||
<figureCitation id="83E12A3EE037D750FBF9E90AFB94FAE3" box="[1047,1118,1322,1344]" captionStart="FIGURES 12–17. 12" captionStartId="5.[151,264,1903,1927]" captionTargetBox="[190,1400,160,1874]" captionTargetId="figure-15@5.[178,1409,151,1889]" captionTargetPageId="5" captionText="FIGURES 12–17. 12. Mesopodagrion tibetanum, habitus with detail of right profile of head (based on Yu 2016). 13. M. tibetanum, detail of mask (redrawn from Yu 2016). 14. Podolestes orientalis, habitus (from Choong & Orr 2010). 15. Podolestes orientalis, detail of mask (from Choong & Orr 2010). 16. Rhipidolestes aculeatus, habitus and detail of lateral and median gill (modified from Asahina 1994). 17. Rhipidolestes aculeatus, detail of mask (modified from Ishida 1996)." figureDoi="http://doi.org/10.5281/zenodo.13618327" httpUri="https://zenodo.org/record/13618327/files/figure.png" pageId="2" pageNumber="211">Fig. 12</figureCitation>
|
||
); head robust with distinct lateral cephalic lobes; antennae short, no longer than long axis of head from occipital rim to base of labrum; prementum broad and short with short labial palps (
|
||
<figureCitation id="83E12A3EE037D750FE14E945FDF5FAA6" box="[506,575,1381,1403]" captionStart="FIGURES 12–17. 12" captionStartId="5.[151,264,1903,1927]" captionTargetBox="[190,1400,160,1874]" captionTargetId="figure-15@5.[178,1409,151,1889]" captionTargetPageId="5" captionText="FIGURES 12–17. 12. Mesopodagrion tibetanum, habitus with detail of right profile of head (based on Yu 2016). 13. M. tibetanum, detail of mask (redrawn from Yu 2016). 14. Podolestes orientalis, habitus (from Choong & Orr 2010). 15. Podolestes orientalis, detail of mask (from Choong & Orr 2010). 16. Rhipidolestes aculeatus, habitus and detail of lateral and median gill (modified from Asahina 1994). 17. Rhipidolestes aculeatus, detail of mask (modified from Ishida 1996)." figureDoi="http://doi.org/10.5281/zenodo.13618327" httpUri="https://zenodo.org/record/13618327/files/figure.png" pageId="2" pageNumber="211">Fig. 13</figureCitation>
|
||
)......................................................
|
||
<taxonomicName id="DCDA4D38E037D750FB3CE945FA57FAA7" authorityName="Kalkman & Abbott" authorityYear="2021" box="[1234,1437,1381,1403]" class="Insecta" family="Mesopodagrionidae" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="211" phylum="Arthropoda" rank="family">
|
||
<emphasis id="29AEEAA9E037D750FB3CE945FA57FAA7" bold="true" box="[1234,1437,1381,1403]" pageId="2" pageNumber="211">Mesopodagrionidae</emphasis>
|
||
</taxonomicName>
|
||
[southern
|
||
<collectingCountry id="63CD762BE037D750FBF2E9A2FB92FA44" box="[1052,1112,1410,1432]" name="China" pageId="2" pageNumber="211">China</collectingCountry>
|
||
, northern Indochina &
|
||
<collectingCountry id="63CD762BE037D750FAD9E9A3FA5DFA44" box="[1335,1431,1411,1432]" name="Myanmar" pageId="2" pageNumber="211">Myanmar</collectingCountry>
|
||
]
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="1B6536BBE037D750FF79E980FA56FA2C" blockId="2.[151,1437,616,1872]" pageId="2" pageNumber="211">
|
||
<keyLead id="A02B958EE037D750FF79E980FA56FA2C" pageId="2" pageNumber="211">
|
||
7’ Variable in build but head never with lateral cephalic lobes; antenna moderately long or short, caudal gills always broad and arranged in a horizontal fan, but vary considerably in relative size;
|
||
<taxonomicName id="DCDA4D38E037D750FC8EE99DFC02FA0E" authorityName="Selys" authorityYear="1862" box="[864,968,1469,1490]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="211" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="29AEEAA9E037D750FC8EE99DFC02FA0E" box="[864,968,1469,1490]" italics="true" pageId="2" pageNumber="211">Podolestes</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="83E12A3EE037D750FC3AE99DFBF5FA0F" box="[980,1087,1469,1491]" captionStart="FIGURES 12–17. 12" captionStartId="5.[151,264,1903,1927]" captionTargetBox="[190,1400,160,1874]" captionTargetId="figure-15@5.[178,1409,151,1889]" captionTargetPageId="5" captionText="FIGURES 12–17. 12. Mesopodagrion tibetanum, habitus with detail of right profile of head (based on Yu 2016). 13. M. tibetanum, detail of mask (redrawn from Yu 2016). 14. Podolestes orientalis, habitus (from Choong & Orr 2010). 15. Podolestes orientalis, detail of mask (from Choong & Orr 2010). 16. Rhipidolestes aculeatus, habitus and detail of lateral and median gill (modified from Asahina 1994). 17. Rhipidolestes aculeatus, detail of mask (modified from Ishida 1996)." figureDoi="http://doi.org/10.5281/zenodo.13618327" httpUri="https://zenodo.org/record/13618327/files/figure.png" pageId="2" pageNumber="211">Figs 14, 15</figureCitation>
|
||
) may be broadly sympatric with last family but has antennae longer than the long axis of head; caudal lamellae relatively much larger...........
|
||
<taxonomicName id="DCDA4D38E037D750FAFAE9FAFA56FA2C" authorityName="Fraser" authorityYear="1960" box="[1300,1436,1498,1520]" class="Insecta" family="Argiolestidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="211" phylum="Arthropoda" rank="family">
|
||
<emphasis id="29AEEAA9E037D750FAFAE9FAFA56FA2C" bold="true" box="[1300,1436,1498,1520]" pageId="2" pageNumber="211">Argiolestidae</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="A02E2E1EE037D750FF79E9D8FA56F9B9" pageId="2" pageNumber="211">
|
||
<paragraph id="1B6536BBE037D750FF79E9D8FA57F994" blockId="2.[151,1437,616,1872]" pageId="2" pageNumber="211">
|
||
<keyLead id="A02B958EE037D750FF79E9D8FA57F994" pageId="2" pageNumber="211">
|
||
8(6’) Caudal gills flattened, fleshy short and broad, arranged vertically, median gill thin walled with complex tracheation, (
|
||
<figureCitation id="83E12A3EE037D750FA9CE9D8FED5F9F6" captionStart="FIGURES 18–26. 18" captionStartId="6.[151,264,1811,1835]" captionTargetBox="[265,1336,167,1790]" captionTargetId="figure-21@6.[248,1339,153,1804]" captionTargetPageId="6" captionText="FIGURES 18–26. 18. Rhinagrion borneense, habitus (original drawing). 19. Philosina alba, habitus (modified from Zhang et al. 2011). 20. Rhinagrion mima, caudal gills: above heavily tracheated median gill; below right lateral gill with basal spines along mid-rib (modified from Lieftinck 1956). 21. Philosina alba caudal gills: above heavily tracheated median gill; below right lateral gill with basal spines along mid-rib (modified from Zhang et al. 2011) 22. Rhinagrion borneense, detail of mask (original drawing). 23. Philosina buchi, detail of mask (redrawn from Zhang et al. 2011). 24. Philoganga vetusta, habitus (adapted from Xu 2016). 25. Philoganga vetusta, details of caudal gills and antenna (redrawn from Xu 2016). 26. Philoganga montana, detail of mask (original drawing)." figureDoi="http://doi.org/10.5281/zenodo.13618331" httpUri="https://zenodo.org/record/13618331/files/figure.png" pageId="2" pageNumber="211">Figs 18–21</figureCitation>
|
||
)....................................................................................
|
||
<taxonomicName id="DCDA4D38E037D750FAF1EA35FA56F9F7" authorityName="Kennedy" authorityYear="1925" box="[1311,1436,1557,1579]" class="Insecta" family="Philosinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="211" phylum="Arthropoda" rank="family">
|
||
<emphasis id="29AEEAA9E037D750FAF1EA35FA56F9F7" bold="true" box="[1311,1436,1557,1579]" pageId="2" pageNumber="211">Philosinidae</emphasis>
|
||
</taxonomicName>
|
||
[West of Wallace’s line]
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="1B6536BBE037D750FF79EA70FA56F9B9" blockId="2.[151,1437,616,1872]" box="[151,1436,1616,1638]" pageId="2" pageNumber="211">
|
||
<keyLead id="A02B958EE037D750FF79EA70FA56F9B9" box="[151,1436,1616,1638]" pageId="2" pageNumber="211">8’ Caudal gills saccoid, terminating in filament with median gill placed over two lower, lateral gills...................... 9</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="A02E2E1EE037D750FF79EA4DFA56F88C" pageId="2" pageNumber="211">
|
||
<paragraph id="1B6536BBE037D750FF79EA4DFA56F907" blockId="2.[151,1437,616,1872]" pageId="2" pageNumber="211">
|
||
<keyLead id="A02B958EE037D750FF79EA4DFA56F907" pageId="2" pageNumber="211">
|
||
9 (8’) Head nearly straight sided behind eyes producing an overall squared profile (
|
||
<figureCitation id="83E12A3EE037D750FC53EA4DFBCEF95E" box="[957,1028,1645,1667]" captionStart="FIGURES 18–26. 18" captionStartId="6.[151,264,1811,1835]" captionTargetBox="[265,1336,167,1790]" captionTargetId="figure-21@6.[248,1339,153,1804]" captionTargetPageId="6" captionText="FIGURES 18–26. 18. Rhinagrion borneense, habitus (original drawing). 19. Philosina alba, habitus (modified from Zhang et al. 2011). 20. Rhinagrion mima, caudal gills: above heavily tracheated median gill; below right lateral gill with basal spines along mid-rib (modified from Lieftinck 1956). 21. Philosina alba caudal gills: above heavily tracheated median gill; below right lateral gill with basal spines along mid-rib (modified from Zhang et al. 2011) 22. Rhinagrion borneense, detail of mask (original drawing). 23. Philosina buchi, detail of mask (redrawn from Zhang et al. 2011). 24. Philoganga vetusta, habitus (adapted from Xu 2016). 25. Philoganga vetusta, details of caudal gills and antenna (redrawn from Xu 2016). 26. Philoganga montana, detail of mask (original drawing)." figureDoi="http://doi.org/10.5281/zenodo.13618331" httpUri="https://zenodo.org/record/13618331/files/figure.png" pageId="2" pageNumber="211">Fig. 24</figureCitation>
|
||
); antennae mostly 8 segmented and clearly longer than head from rear occipital margin to base of frons; caudal gills elongate and terminating uniformly in a point (
|
||
<figureCitation id="83E12A3EE037D750FA9BEAAAFF37F961" captionStart="FIGURES 18–26. 18" captionStartId="6.[151,264,1811,1835]" captionTargetBox="[265,1336,167,1790]" captionTargetId="figure-21@6.[248,1339,153,1804]" captionTargetPageId="6" captionText="FIGURES 18–26. 18. Rhinagrion borneense, habitus (original drawing). 19. Philosina alba, habitus (modified from Zhang et al. 2011). 20. Rhinagrion mima, caudal gills: above heavily tracheated median gill; below right lateral gill with basal spines along mid-rib (modified from Lieftinck 1956). 21. Philosina alba caudal gills: above heavily tracheated median gill; below right lateral gill with basal spines along mid-rib (modified from Zhang et al. 2011) 22. Rhinagrion borneense, detail of mask (original drawing). 23. Philosina buchi, detail of mask (redrawn from Zhang et al. 2011). 24. Philoganga vetusta, habitus (adapted from Xu 2016). 25. Philoganga vetusta, details of caudal gills and antenna (redrawn from Xu 2016). 26. Philoganga montana, detail of mask (original drawing)." figureDoi="http://doi.org/10.5281/zenodo.13618331" httpUri="https://zenodo.org/record/13618331/files/figure.png" pageId="2" pageNumber="211">Fig. 25</figureCitation>
|
||
); mature larvae and exuviae huge,
|
||
<quantity id="DC229B5EE037D750FDA2EA88FD79F961" box="[588,691,1704,1725]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="211" unit="mm" value="35.0" valueMax="40.0" valueMin="30.0">30–40 mm</quantity>
|
||
in total length including gills; general build robust but elongate..................................................................................................
|
||
<taxonomicName id="DCDA4D38E037D750FAE5EAE5FA56F907" authorityName="Kennedy" authorityYear="1920" box="[1291,1436,1733,1755]" class="Insecta" family="Philogangidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="211" phylum="Arthropoda" rank="family">
|
||
<emphasis id="29AEEAA9E037D750FAE5EAE5FA56F907" bold="true" box="[1291,1436,1733,1755]" pageId="2" pageNumber="211">Philogangidae</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="1B6536BBE037D750FF79EAC2FA56F88C" blockId="2.[151,1437,616,1872]" pageId="2" pageNumber="211">
|
||
<keyLead id="A02B958EE037D750FF79EAC2FA56F88C" pageId="2" pageNumber="211">
|
||
9’ Head with postocular lobes rounded (
|
||
<figureCitation id="83E12A3EE037D750FDA6EAC2FD45F924" box="[584,655,1762,1784]" captionStart="FIGURES 12–17. 12" captionStartId="5.[151,264,1903,1927]" captionTargetBox="[190,1400,160,1874]" captionTargetId="figure-15@5.[178,1409,151,1889]" captionTargetPageId="5" captionText="FIGURES 12–17. 12. Mesopodagrion tibetanum, habitus with detail of right profile of head (based on Yu 2016). 13. M. tibetanum, detail of mask (redrawn from Yu 2016). 14. Podolestes orientalis, habitus (from Choong & Orr 2010). 15. Podolestes orientalis, detail of mask (from Choong & Orr 2010). 16. Rhipidolestes aculeatus, habitus and detail of lateral and median gill (modified from Asahina 1994). 17. Rhipidolestes aculeatus, detail of mask (modified from Ishida 1996)." figureDoi="http://doi.org/10.5281/zenodo.13618327" httpUri="https://zenodo.org/record/13618327/files/figure.png" pageId="2" pageNumber="211">Fig. 16</figureCitation>
|
||
); antennae short and robust, about same length as head from rear margin to frons; caudal gills ovate saccoid with distinct dorsal ridge and terminating in short, well defined filament; prementum broad and short with short labial palps (
|
||
<figureCitation id="83E12A3EE037D750FE27EB3DFDDBF8EE" box="[457,529,1821,1843]" captionStart="FIGURES 12–17. 12" captionStartId="5.[151,264,1903,1927]" captionTargetBox="[190,1400,160,1874]" captionTargetId="figure-15@5.[178,1409,151,1889]" captionTargetPageId="5" captionText="FIGURES 12–17. 12. Mesopodagrion tibetanum, habitus with detail of right profile of head (based on Yu 2016). 13. M. tibetanum, detail of mask (redrawn from Yu 2016). 14. Podolestes orientalis, habitus (from Choong & Orr 2010). 15. Podolestes orientalis, detail of mask (from Choong & Orr 2010). 16. Rhipidolestes aculeatus, habitus and detail of lateral and median gill (modified from Asahina 1994). 17. Rhipidolestes aculeatus, detail of mask (modified from Ishida 1996)." figureDoi="http://doi.org/10.5281/zenodo.13618327" httpUri="https://zenodo.org/record/13618327/files/figure.png" pageId="2" pageNumber="211">Fig. 17</figureCitation>
|
||
); general build stout, tapering caudad and less than
|
||
<quantity id="DC229B5EE037D750FBE0EB3DFB98F8EE" box="[1038,1106,1821,1843]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="211" unit="mm" value="15.0">15 mm</quantity>
|
||
in length (
|
||
<quantity id="DC229B5EE037D750FB53EB3DFAEDF8EE" box="[1213,1319,1821,1842]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="2" pageNumber="211" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10–12 mm</quantity>
|
||
in recorded specimens)..............................................................................
|
||
<taxonomicName id="DCDA4D38E037D750FB12EB1AFA56F88C" box="[1276,1436,1850,1872]" family="Rhipidolestidae" pageId="2" pageNumber="237" rank="family">
|
||
<emphasis id="29AEEAA9E037D750FB12EB1AFA56F88C" bold="true" box="[1276,1436,1850,1872]" pageId="2" pageNumber="211">Rhipidolestidae</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |