378 lines
43 KiB
XML
378 lines
43 KiB
XML
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<mods:title id="F7C946A01F99DD51200BA6135862F6DD">Revision of northeast Pacific Paleogene cypraeoidean gastropods, including recognition of three new species: implications for paleobiogeographic distribution and faunal turnover</mods:title>
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<mods:title id="57A38393643B04CB056C78313CDE1D8B">PaleoBios</mods:title>
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<mods:date id="9CC36A15AFA62E7D31806D7AED4E44DF">2023</mods:date>
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<paragraph id="1A09685AFFDEFFA5FC26FBEEFA16FB95" blockId="13.[900,1500,1109,1172]" box="[900,1500,1109,1137]" pageId="13" pageNumber="12">
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<taxonomicName id="DDB613D9FFDEFFA5FC26FBEEFA16FB95" ID-CoL="C4MRQ" authority="SQUIRES AND DEMETRION, 1992" authorityName="Squires and Demetrion" authorityYear="1992" box="[900,1500,1109,1137]" class="Gastropoda" family="Cypraeidae" genus="Bernaya" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="grovesi">
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<emphasis id="28C2B448FFDEFFA5FC26FBEEFB99FB95" box="[900,1107,1109,1137]" italics="true" pageId="13" pageNumber="12">BERNAYA GROVESI</emphasis>
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SQUIRES AND DEMETRION, 1992
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<figureCitation id="828D74DFFFDEFFA5FBD2FBC2FB3AFB71" box="[1136,1264,1145,1172]" captionStart="Figure 5" captionStartId="15.[113,200,200,226]" captionTargetId="figure-12@14.[141,1485,197,1966]" captionTargetPageId="14" captionText="Figure 5 (previous page) A–C. Bernaya grovesi Squires and Demetrion, 1992, Bateque Formation (Eocene, “Ca- pay Stage”), Mesa La Salina, Baja California Sur, México. Basal (A), dorsal (B), and right (labral) (C) views, all 2.6x, length 17 mm, width 12 mm, height 7.7 mm, of holotype IGM 5172, LACMIP Locality 401220a (ex CSUN Locality 1220b). D–F. Bernaya kaylinae n. sp., Llajas Formation, (Eocene, “Domengine Stage”), Las Llajas Canyon, north side Simi Valley, Ventura County, southern California. Basal (D), dorsal (E), and right (labral) side (F) views, 3.8x, length 25.6 mm, width 17.3 mm, height 12.9 mm, of holotype, LACMIP 40374.80, LACMIP Type 14938 (LACMIP Locality 40374). G–I. Bernaya squiresi Groves, 2011, unnamed Eocene strata (“Capay Stage”), southwest end of Discovery Bay, Jefferson County, Washington. Basal (G), dorsal (H), and right (labral) side (I) views, 0.97x, length 52.7 mm, width 40 mm, height 26.3 mm, of holotype LACMIP 22341.1, LACMIP Type 13644, LACMIP Locality 22341. J–L. Bernaya sp., unnamed Paleocene strata (Thanetian Stage), near Lower Lake, Lake County, northern California. Basal (J), dorsal (K), and right (labral) side (L) views, 1.7x, length 33.8 mm, width 23 mm, height 18.3 mm, hypotype LACMIP 7047.72, LACMIP Type 14923 (LACMIP Locality 7047). M–O. Protocypraea? simiensis (Nelson, 1925), Santa Susana Formation (Thanetian Stage), Simi Hills, Ventura County, Paleocene, southern Cali- fornia. Basal (M), dorsal (N), and right (labral) side (O) views, 1.6x, length 32.3 mm, width 24.6 mm, height 17.3 mm, of holotype UCMP 30498, UCMP Locality 3818. P.Protocypraea? sp. 1, Oyster Bay Formation, Paleocene, Appian Way, Central Island District, Vancouver Island, British Columbia. Basal view, 1.8x, length 48.1 mm, width 31.4 mm, height 25.3 mm, of CDM 997.91.3, informal locality AP3." figureDoi="http://doi.org/10.5281/zenodo.13869594" httpUri="https://zenodo.org/record/13869594/files/figure.png" pageId="13" pageNumber="12">Figs. 5A–C</figureCitation>
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<subSubSection id="52AC3BD1FFDEFFA5FCC2FB7BFC2BFA6F" pageId="13" pageNumber="12" type="reference_group">
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<paragraph id="1A09685AFFDEFFA5FCC2FB7BFC62FAA1" blockId="13.[832,1521,1216,1946]" pageId="13" pageNumber="12">
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<taxonomicName id="DDB613D9FFDEFFA5FCC2FB7BFA93FB3F" ID-CoL="C4MRQ" authority="Squires" authorityName="Squires and Demetrion" authorityYear="1992" box="[864,1369,1216,1244]" class="Gastropoda" family="Cypraeidae" genus="Bernaya" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="12" phylum="Mollusca" rank="species" species="grovesi" subGenus="Protocypraea">
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<emphasis id="28C2B448FFDEFFA5FCC2FB7BFC0DFB39" box="[864,967,1216,1244]" italics="true" pageId="13" pageNumber="12">Bernaya</emphasis>
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(
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<emphasis id="28C2B448FFDEFFA5FC7FFB7BFB4EFB39" box="[989,1156,1216,1244]" italics="true" pageId="13" pageNumber="12">Protocypraea</emphasis>
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)
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<emphasis id="28C2B448FFDEFFA5FB3BFB7BFB3BFB39" box="[1177,1265,1216,1244]" italics="true" pageId="13" pageNumber="12">grovesi</emphasis>
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Squires
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and Deme- trion, 1992. p. 31, figs. 74-76.
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<bibRefCitation id="7E2715ABFFDEFFA5FB06FB58FA8FFB18" author="Groves, L. T." box="[1188,1349,1251,1277]" pageId="13" pageNumber="12" pagination="11 - 14" refId="ref37858" refString="Groves, L. T. 1993. Fossil and recent species of eastern Pacific Cypraeacea (Pediculariidae, Cypraeidae, and Ovulidae). The Western Society of Malacologists Annual Report 25: 11 - 14." type="book chapter" year="1993">Groves, 1993</bibRefCitation>
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. p. 11.
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<bibRefCitation id="7E2715ABFFDEFFA5FA37FB58FC5BFAC5" author="Groves, L. T." pageId="13" pageNumber="12" pagination="5 - 16" refId="ref37963" refString="Groves, L. T. 1994 b. Catalog of fossil and recent Cypraei- dae and Eocypraeinae (Ovulidae) described since 1971. The Cowry, new series, 1 (1): 5 - 16." type="journal article" year="1994">Groves, 1994b</bibRefCitation>
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. p. 12.
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<bibRefCitation id="7E2715ABFFDEFFA5FC50FABDFB53FAC5" author="Groves, L. T." box="[1010,1177,1286,1312]" pageId="13" pageNumber="12" pagination="7 - 10" refId="ref38049" refString="Groves, L. T. 1997. Fossil and recent species of eastern Pacific Cypraeacea (Cypraeidae and Eocypraeinae [Ovulidae]): An update. The Western Society of Malacology Annual Report 29: 7 - 10." type="book chapter" year="1997">Groves, 1997</bibRefCitation>
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. p. 7 [as
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<taxonomicName id="DDB613D9FFDEFFA5FAB0FABEFC5DFAA3" ID-CoL="C4MRQ" authorityName="Squires and Demetrion" authorityYear="1992" class="Gastropoda" family="Cypraeidae" genus="Bernaya" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="12" phylum="Mollusca" rank="species" species="grovesi" subGenus="Protocypraea">
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<emphasis id="28C2B448FFDEFFA5FAB0FABEFAE0FAC4" box="[1298,1322,1285,1313]" italics="true" pageId="13" pageNumber="12">B.</emphasis>
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(
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<emphasis id="28C2B448FFDEFFA5FA9DFABDFA2FFAC7" box="[1343,1509,1286,1314]" italics="true" pageId="13" pageNumber="12">Protocypraea</emphasis>
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)
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<emphasis id="28C2B448FFDEFFA5FCE2FA91FC5DFAA3" box="[832,919,1322,1350]" italics="true" pageId="13" pageNumber="12">grovesi</emphasis>
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].
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<taxonomicName id="DDB613D9FFDEFFA5FCC2FAF6FA47FA82" ID-CoL="C4MRS" authority="(Squires and Demetrion)" baseAuthorityName="Squires and Demetrion" box="[864,1421,1356,1385]" class="Gastropoda" family="Cypraeidae" genus="Protocypraea" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="12" phylum="Mollusca" rank="species" species="grovesi">
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<emphasis id="28C2B448FFDEFFA5FCC2FAF6FB91FA8C" box="[864,1115,1357,1385]" italics="true" pageId="13" pageNumber="12">Protocypraea grovesi</emphasis>
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(Squires and Demetrion)
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</taxonomicName>
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.
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||
<bibRefCitation id="7E2715ABFFDEFFA5FA37FAF6FC4BFA6F" author="Lorenz, F." pageId="13" pageNumber="12" refId="ref39506" refString="Lorenz, F. 2017. Cowries. A guide to the gastropod family Cypraeidae. Volume 1. Biology and Systematics. Conch- Books, Harxheim, Germany. 644 pp., numerous figures." type="book" year="2017">Lorenz, 2017</bibRefCitation>
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. p. 205.
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</paragraph>
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and Type Locality—
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(
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), length
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, width
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, height
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Lo- cality 41220b [
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<emphasis id="28C2B448FFDEFFA5FC55FA63FBDBFA11" box="[1015,1041,1496,1524]" italics="true" pageId="13" pageNumber="12">ex</emphasis>
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Locality 1220b],
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Forma- tion, near
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,
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,
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|
||
<typeStatus id="C50DD6F8FFDEFFA5FCC2F99BFC13F9DF" box="[864,985,1568,1594]" pageId="13" pageNumber="12" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
—
|
||
<specimenCode id="4A10C021FFDEFFA5FC54F99BFBBAF9DF" box="[1014,1136,1567,1594]" collectionCode="IGM" country="Mongolia" httpUri="http://grbio.org/cool/0h5a-qjh4" name="Geological Institute, Mongolian Academy of Sciences" pageId="13" pageNumber="12">IGM 5173</specimenCode>
|
||
, length
|
||
<quantity id="DD4EC5BFFFDEFFA5FB6CF99BFAECF9DF" box="[1230,1318,1568,1594]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="13" pageNumber="12" unit="mm" value="16.0">16 mm</quantity>
|
||
, width
|
||
<quantity id="DD4EC5BFFFDEFFA5FADCF9A4FA21F9DF" box="[1406,1515,1567,1594]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="13" pageNumber="12" unit="mm" value="10.5">10.5 mm</quantity>
|
||
, height
|
||
<quantity id="DD4EC5BFFFDEFFA5FC37F9F8FC2AF9B8" box="[917,992,1603,1629]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="13" pageNumber="12" unit="mm" value="8.0">8 mm</quantity>
|
||
; same locality as the holotype
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="1A09685AFFDEFFA5FCC2F9DDFB21F893" blockId="13.[832,1521,1216,1946]" pageId="13" pageNumber="12">
|
||
<emphasis id="28C2B448FFDEFFA5FCC2F9DDFBB2F965" bold="true" box="[864,1144,1638,1664]" pageId="13" pageNumber="12">Referred Specimen—</emphasis>
|
||
<materialsCitation id="AADE6207FFDEFFA5FBDAF9DDFB2CF893" collectionCode="LACMIP, CSUN" county="Locality" location="lower Eocene" municipality="Crescent Formation" pageId="13" pageNumber="12" specimenCode="LACMIP 41653.6" specimenCount="1" stateProvince="Washington">
|
||
A single, slightly crushed speci- men (
|
||
<specimenCode id="4A10C021FFDEFFA5FC21F932FB9DF941" box="[899,1111,1673,1700]" collectionCode="LACMIP" pageId="13" pageNumber="12">LACMIP 41653.6</specimenCode>
|
||
,
|
||
<collectionCode id="7CA7F09FFFDEFFA5FBC2F932FB0EF946" box="[1120,1220,1673,1699]" pageId="13" pageNumber="12">LACMIP</collectionCode>
|
||
<typeStatus id="C50DD6F8FFDEFFA5FB6BF931FACDF941" box="[1225,1287,1674,1700]" pageId="13" pageNumber="12">Type</typeStatus>
|
||
14937) measuring length
|
||
<quantity id="DD4EC5BFFFDEFFA5FC3EF916FBDBF922" box="[924,1041,1709,1735]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="13" pageNumber="12" unit="mm" value="14.3">14.3 mm</quantity>
|
||
, width
|
||
<quantity id="DD4EC5BFFFDEFFA5FBD7F916FB10F922" box="[1141,1242,1709,1735]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="13" pageNumber="12" unit="mm" value="9.2">9.2 mm</quantity>
|
||
, height
|
||
<quantity id="DD4EC5BFFFDEFFA5FAE7F917FA6DF922" box="[1349,1447,1708,1735]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="13" pageNumber="12" unit="mm" value="5.8">5.8 mm</quantity>
|
||
from
|
||
<collectionCode id="7CA7F09FFFDEFFA5FCE2F96BFC6EF90F" box="[832,932,1744,1770]" pageId="13" pageNumber="12">LACMIP</collectionCode>
|
||
<collectingCounty id="F36810D6FFDEFFA5FC0CF96BFBDBF90F" box="[942,1041,1744,1770]" pageId="13" pageNumber="12">Locality</collectingCounty>
|
||
16655 [=
|
||
<collectionCode id="7CA7F09FFFDEFFA5FB3BF96BFB37F90F" box="[1177,1277,1744,1770]" pageId="13" pageNumber="12">LACMIP</collectionCode>
|
||
Locality 41653;
|
||
<emphasis id="28C2B448FFDEFFA5FA74F975FA3AF90F" box="[1494,1520,1742,1770]" italics="true" pageId="13" pageNumber="12">ex</emphasis>
|
||
<collectionCode id="7CA7F09FFFDEFFA5FCE2F948FC4CF8E8" box="[832,902,1779,1805]" country="USA" httpUri="http://grbio.org/cool/sgeh-wrgd" name="California State University, Northridge" pageId="13" pageNumber="12" type="University or college">CSUN</collectionCode>
|
||
Locality 1563],
|
||
<collectingMunicipality id="FA6DF220FFDEFFA5FBEBF948FA8AF8E8" box="[1097,1344,1779,1805]" pageId="13" pageNumber="12">Crescent Formation</collectingMunicipality>
|
||
,
|
||
<location id="1F693E81FFDEFFA5FAEBF948FA3AF8E8" LSID="urn:lsid:plazi:treatment:921FD94CFFDEFFA7FC26FBEEFD45FA88:1F693E81FFDEFFA5FAEBF948FA3AF8E8" box="[1353,1520,1779,1805]" county="Locality" municipality="Crescent Formation" name="lower Eocene" pageId="13" pageNumber="12" stateProvince="Washington">lower Eocene</location>
|
||
(“Capay” Stage),
|
||
<location id="1F693E81FFDEFFA5FBAFF8ADFADAF8D4" LSID="urn:lsid:plazi:treatment:921FD94CFFDEFFA7FC26FBEEFD45FA88:1F693E81FFDEFFA5FBAFF8ADFADAF8D4" box="[1037,1296,1814,1841]" county="Locality" municipality="Crescent Formation" name="Larch Mountain area" pageId="13" pageNumber="12" stateProvince="Washington">Larch Mountain area</location>
|
||
,
|
||
<location id="1F693E81FFDEFFA5FAB8F8ADFA54F8D5" LSID="urn:lsid:plazi:treatment:921FD94CFFDEFFA7FC26FBEEFD45FA88:1F693E81FFDEFFA5FAB8F8ADFA54F8D5" box="[1306,1438,1814,1840]" county="Locality" municipality="Crescent Formation" name="Black Hills" pageId="13" pageNumber="12" stateProvince="Washington">Black Hills</location>
|
||
, Thur- ston
|
||
<location id="1F693E81FFDEFFA5FC20F882FC17F8B6" LSID="urn:lsid:plazi:treatment:921FD94CFFDEFFA7FC26FBEEFD45FA88:1F693E81FFDEFFA5FC20F882FC17F8B6" box="[898,989,1849,1875]" county="Locality" municipality="Crescent Formation" name="County" pageId="13" pageNumber="12" stateProvince="Washington">County</location>
|
||
, southwestern
|
||
<collectingRegion id="D872A6B8FFDEFFA5FB04F882FAF5F8B6" box="[1190,1343,1849,1875]" country="United States of America" name="Washington" pageId="13" pageNumber="12">Washington</collectingRegion>
|
||
that strongly resembles
|
||
<taxonomicName id="DDB613D9FFDEFFA5FC65F8E0FBF1F89C" box="[967,1083,1883,1913]" class="Gastropoda" family="Cypraeidae" genus="Protocypraea" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="12" phylum="Mollusca" rank="species" species="grovesi">
|
||
<emphasis id="28C2B448FFDEFFA5FC65F8E0FBF1F89C" box="[967,1083,1883,1913]" italics="true" pageId="13" pageNumber="12">B. grovesi</emphasis>
|
||
</taxonomicName>
|
||
is here noted
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="1A09685AFFDEFFA5FCC2F8C4FA25F87F" blockId="13.[832,1521,1216,1946]" box="[864,1519,1919,1946]" pageId="13" pageNumber="12">
|
||
<emphasis id="28C2B448FFDEFFA5FCC2F8C4FBEEF87C" bold="true" box="[864,1060,1919,1945]" pageId="13" pageNumber="12">Occurrence—</emphasis>
|
||
Lower to middle Eocene (“Capay
|
||
</paragraph>
|
||
<caption id="4EC938D2FFDCFFA7FFD3FF73FC69FCF3" ID-DOI="http://doi.org/10.5281/zenodo.13869594" ID-Zenodo-Dep="13869594" httpUri="https://zenodo.org/record/13869594/files/figure.png" pageId="15" pageNumber="14" startId="15.[113,200,200,226]" targetBox="[165,1470,206,1937]" targetPageId="14">
|
||
<paragraph id="1A09685AFFDCFFA7FFD3FF73FC69FCF3" blockId="15.[112,1520,200,790]" pageId="15" pageNumber="14">
|
||
<emphasis id="28C2B448FFDCFFA7FFD3FF73FE30FF07" bold="true" box="[113,506,200,226]" pageId="15" pageNumber="14">Figure 5 (previous page) A–C.</emphasis>
|
||
<taxonomicName id="DDB613D9FFDCFFA7FE5DFF72FBF2FF07" authority="Squires and Demetrion, 1992" authorityName="Squires and Demetrion" authorityYear="1992" box="[511,1080,200,229]" class="Gastropoda" family="Cypraeidae" genus="Bernaya" kingdom="Animalia" order="Littorinimorpha" pageId="15" pageNumber="14" phylum="Mollusca" rank="species" species="grovesi">
|
||
<emphasis id="28C2B448FFDCFFA7FE5DFF72FD0EFF00" box="[511,708,201,229]" italics="true" pageId="15" pageNumber="14">Bernaya grovesi</emphasis>
|
||
<bibRefCitation id="7E2715ABFFDCFFA7FD68FF73FBF2FF07" author="Squires, R. L. & R. A. Demetrion" box="[714,1080,200,226]" pageId="15" pageNumber="14" pagination="1 - 55" refId="ref43379" refString="Squires, R. L. and R. A. Demetrion. 1992. Paleontology of the Eocene Bateque Formation, Baja California Sur, Mexico. Contributions in Science, Natural History Museum of Los Angeles County 434: 1 - 55, figs. 1 - 144." type="journal article" year="1992">Squires and Demetrion, 1992</bibRefCitation>
|
||
</taxonomicName>
|
||
, Bateque Formation (Eocene, “Ca- pay Stage”), Mesa La Salina,
|
||
<collectingRegion id="D872A6B8FFDCFFA7FE6CFF50FD7FFEE3" box="[462,693,235,262]" country="Mexico" name="Baja California Sur" pageId="15" pageNumber="14">Baja California Sur</collectingRegion>
|
||
,
|
||
<collectingCountry id="62A128CAFFDCFFA7FD62FF50FCD1FEE0" box="[704,795,235,261]" name="Mexico" pageId="15" pageNumber="14">México</collectingCountry>
|
||
. Basal (
|
||
<emphasis id="28C2B448FFDCFFA7FCD8FF50FC47FEE0" bold="true" box="[890,909,235,261]" pageId="15" pageNumber="14">A</emphasis>
|
||
), dorsal (
|
||
<emphasis id="28C2B448FFDCFFA7FBA7FF57FBD2FEE3" bold="true" box="[1029,1048,236,262]" pageId="15" pageNumber="14">B</emphasis>
|
||
), and right (labral) (
|
||
<emphasis id="28C2B448FFDCFFA7FABBFF50FAE0FEE0" bold="true" box="[1305,1322,235,261]" pageId="15" pageNumber="14">C</emphasis>
|
||
) views, all 2.6x, length
|
||
<quantity id="DD4EC5BFFFDCFFA7FF64FEB4FED5FECC" box="[198,287,271,297]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="15" pageNumber="14" unit="mm" value="17.0">17 mm</quantity>
|
||
, width
|
||
<quantity id="DD4EC5BFFFDCFFA7FED8FEB4FE19FECC" box="[378,467,271,297]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="15" pageNumber="14" unit="mm" value="12.0">12 mm</quantity>
|
||
, height
|
||
<quantity id="DD4EC5BFFFDCFFA7FD91FEB4FD58FECC" box="[563,658,271,297]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="15" pageNumber="14" unit="mm" value="7.7">7.7 mm</quantity>
|
||
, of
|
||
<typeStatus id="C50DD6F8FFDCFFA7FD1FFEB4FCE6FECC" box="[701,812,271,297]" pageId="15" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
<collectionCode id="7CA7F09FFFDCFFA7FC90FEB4FCAFFECC" box="[818,869,271,297]" country="Mongolia" httpUri="http://grbio.org/cool/0h5a-qjh4" name="Geological Institute, Mongolian Academy of Sciences" pageId="15" pageNumber="14">IGM</collectionCode>
|
||
5172,
|
||
<collectionCode id="7CA7F09FFFDCFFA7FC1BFEB4FBD7FECC" box="[953,1053,271,297]" pageId="15" pageNumber="14">LACMIP</collectionCode>
|
||
Locality 401220a (
|
||
<emphasis id="28C2B448FFDCFFA7FAADFEB6FAE3FECC" box="[1295,1321,269,297]" italics="true" pageId="15" pageNumber="14">ex</emphasis>
|
||
<collectionCode id="7CA7F09FFFDCFFA7FA92FEB4FABCFECC" box="[1328,1398,271,297]" country="USA" httpUri="http://grbio.org/cool/sgeh-wrgd" name="California State University, Northridge" pageId="15" pageNumber="14" type="University or college">CSUN</collectionCode>
|
||
Locality 1220b).
|
||
<emphasis id="28C2B448FFDCFFA7FF7BFE89FEDAFEA9" bold="true" box="[217,272,306,332]" pageId="15" pageNumber="14">D–F.</emphasis>
|
||
<taxonomicName id="DDB613D9FFDCFFA7FEB4FE89FE20FEAB" authority="Groves & Squires, 2023" authorityName="Groves & Squires" authorityYear="2023" box="[278,490,306,334]" class="Gastropoda" family="Cypraeidae" genus="Bernaya" kingdom="Animalia" order="Littorinimorpha" pageId="15" pageNumber="14" phylum="Mollusca" rank="species" species="kaylinae" status="sp. nov.">
|
||
<emphasis id="28C2B448FFDCFFA7FEB4FE89FE20FEAB" box="[278,490,306,334]" italics="true" pageId="15" pageNumber="14">Bernaya kaylinae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="33F10933FFDCFFA7FE53FE89FDFBFEA9" box="[497,561,306,332]" pageId="15" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
|
||
, Llajas Formation, (Eocene, “Domengine Stage”), Las Llajas Canyon, north side Simi Valley, Ventura County, southern California. Basal (
|
||
<emphasis id="28C2B448FFDCFFA7FCFEFEEEFCBBFE8A" bold="true" box="[860,881,341,367]" pageId="15" pageNumber="14">D</emphasis>
|
||
), dorsal (
|
||
<emphasis id="28C2B448FFDCFFA7FC4BFEEEFC30FE8A" bold="true" box="[1001,1018,341,367]" pageId="15" pageNumber="14">E</emphasis>
|
||
), and right (labral) side (
|
||
<emphasis id="28C2B448FFDCFFA7FA96FEEEFA8EFE8A" bold="true" box="[1332,1348,341,367]" pageId="15" pageNumber="14">F</emphasis>
|
||
) views, 3.8x, length
|
||
<quantity id="DD4EC5BFFFDCFFA7FF64FEC3FEFFFE76" box="[198,309,376,403]" metricMagnitude="-2" metricUnit="m" metricValue="2.56" pageId="15" pageNumber="14" unit="mm" value="25.6">25.6 mm</quantity>
|
||
, width
|
||
<quantity id="DD4EC5BFFFDCFFA7FE32FEC3FE35FE76" box="[400,511,376,403]" metricMagnitude="-2" metricUnit="m" metricValue="1.73" pageId="15" pageNumber="14" unit="mm" value="17.3">17.3 mm</quantity>
|
||
, height
|
||
<quantity id="DD4EC5BFFFDCFFA7FDC2FEC3FD05FE76" box="[608,719,376,403]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="15" pageNumber="14" unit="mm" value="12.9">12.9 mm</quantity>
|
||
, of
|
||
<typeStatus id="C50DD6F8FFDCFFA7FD5BFEC3FCA3FE77" box="[761,873,376,402]" pageId="15" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
,
|
||
<collectionCode id="7CA7F09FFFDCFFA7FCD6FEC3FC12FE77" box="[884,984,376,402]" pageId="15" pageNumber="14">LACMIP</collectionCode>
|
||
40374.80,
|
||
<collectionCode id="7CA7F09FFFDCFFA7FBC1FEC3FB0DFE77" box="[1123,1223,376,402]" pageId="15" pageNumber="14">LACMIP</collectionCode>
|
||
Type 14938 (
|
||
<collectionCode id="7CA7F09FFFDCFFA7FAD3FEC3FA13FE77" box="[1393,1497,376,402]" pageId="15" pageNumber="14">LACMIP</collectionCode>
|
||
Locality 40374).
|
||
<emphasis id="28C2B448FFDCFFA7FEE1FE20FEBFFE50" bold="true" box="[323,373,411,437]" pageId="15" pageNumber="14">G–I.</emphasis>
|
||
<taxonomicName id="DDB613D9FFDCFFA7FEDEFE27FD3BFE53" authority="Groves, 2011" authorityName="Groves" authorityYear="2011" box="[380,753,410,440]" class="Gastropoda" family="Cypraeidae" genus="Bernaya" kingdom="Animalia" order="Littorinimorpha" pageId="15" pageNumber="14" phylum="Mollusca" rank="species" species="squiresi">
|
||
<emphasis id="28C2B448FFDCFFA7FEDEFE27FD8DFE53" box="[380,583,410,440]" italics="true" pageId="15" pageNumber="14">Bernaya squiresi</emphasis>
|
||
Groves, 2011
|
||
</taxonomicName>
|
||
, unnamed Eocene strata (“Capay Stage”), southwest end of Discovery Bay, Jefferson County, Washington. Basal (
|
||
<emphasis id="28C2B448FFDCFFA7FD59FE05FCC4FE3D" bold="true" box="[763,782,446,472]" pageId="15" pageNumber="14">G</emphasis>
|
||
), dorsal (
|
||
<emphasis id="28C2B448FFDCFFA7FC24FE04FC51FE3C" bold="true" box="[902,923,447,473]" pageId="15" pageNumber="14">H</emphasis>
|
||
), and right (labral) side (
|
||
<emphasis id="28C2B448FFDCFFA7FB74FE04FB2AFE3C" bold="true" box="[1238,1248,447,473]" pageId="15" pageNumber="14">I</emphasis>
|
||
) views, 0.97x, length
|
||
<quantity id="DD4EC5BFFFDCFFA7FFD2FE59FF15FE19" box="[112,223,482,509]" metricMagnitude="-2" metricUnit="m" metricValue="5.27" pageId="15" pageNumber="14" unit="mm" value="52.7">52.7 mm</quantity>
|
||
, width
|
||
<quantity id="DD4EC5BFFFDCFFA7FE98FE59FE59FE19" box="[314,403,482,508]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="14" unit="mm" value="40.0">40 mm</quantity>
|
||
, height
|
||
<quantity id="DD4EC5BFFFDCFFA7FE56FE59FDA9FE19" box="[500,611,482,508]" metricMagnitude="-2" metricUnit="m" metricValue="2.63" pageId="15" pageNumber="14" unit="mm" value="26.3">26.3 mm</quantity>
|
||
, of
|
||
<typeStatus id="C50DD6F8FFDCFFA7FD2FFE59FD36FE19" box="[653,764,482,508]" pageId="15" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
<collectionCode id="7CA7F09FFFDCFFA7FCA0FE59FCACFE19" box="[770,870,482,508]" pageId="15" pageNumber="14">LACMIP</collectionCode>
|
||
22341.1,
|
||
<collectionCode id="7CA7F09FFFDCFFA7FC43FE59FB8FFE19" box="[993,1093,482,508]" pageId="15" pageNumber="14">LACMIP</collectionCode>
|
||
Type 13644,
|
||
<collectionCode id="7CA7F09FFFDCFFA7FB4CFE59FA98FE19" box="[1262,1362,482,508]" pageId="15" pageNumber="14">LACMIP</collectionCode>
|
||
Locality 22341.
|
||
<emphasis id="28C2B448FFDCFFA7FF6CFDBEFF34FDFA" bold="true" box="[206,254,517,543]" pageId="15" pageNumber="14">J–L.</emphasis>
|
||
<taxonomicName id="DDB613D9FFDCFFA7FEA6FDBEFE5EFDC5" box="[260,404,517,545]" class="Gastropoda" family="Cypraeidae" genus="Bernaya" kingdom="Animalia" order="Littorinimorpha" pageId="15" pageNumber="14" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="28C2B448FFDCFFA7FEA6FDBEFEA1FDC4" box="[260,363,517,545]" italics="true" pageId="15" pageNumber="14">Bernaya</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
, unnamed Paleocene strata (Thanetian Stage), near Lower Lake, Lake County, northern California. Basal (
|
||
<emphasis id="28C2B448FFDCFFA7FEE9FD93FE9FFDA7" bold="true" box="[331,341,552,578]" pageId="15" pageNumber="14">J</emphasis>
|
||
), dorsal (
|
||
<emphasis id="28C2B448FFDCFFA7FE6FFD93FE2BFDA7" bold="true" box="[461,481,552,578]" pageId="15" pageNumber="14">K</emphasis>
|
||
), and right (labral) side (
|
||
<emphasis id="28C2B448FFDCFFA7FCBEFD93FCE6FDA7" bold="true" box="[796,812,552,578]" pageId="15" pageNumber="14">L</emphasis>
|
||
) views, 1.7x, length
|
||
<quantity id="DD4EC5BFFFDCFFA7FB8BFD93FB52FDA6" box="[1065,1176,552,579]" metricMagnitude="-2" metricUnit="m" metricValue="3.38" pageId="15" pageNumber="14" unit="mm" value="33.8">33.8 mm</quantity>
|
||
, width
|
||
<quantity id="DD4EC5BFFFDCFFA7FB51FD93FA86FDA6" box="[1267,1356,552,579]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="15" pageNumber="14" unit="mm" value="23.0">23 mm</quantity>
|
||
, height
|
||
<quantity id="DD4EC5BFFFDCFFA7FA0FFD93FF68FD83" metricMagnitude="-2" metricUnit="m" metricValue="1.83" pageId="15" pageNumber="14" unit="mm" value="18.3">18.3 mm</quantity>
|
||
, hypotype
|
||
<collectionCode id="7CA7F09FFFDCFFA7FE8BFDF7FE47FD83" box="[297,397,588,614]" pageId="15" pageNumber="14">LACMIP</collectionCode>
|
||
7047.72,
|
||
<collectionCode id="7CA7F09FFFDCFFA7FDAAFDF7FDA6FD83" box="[520,620,588,614]" pageId="15" pageNumber="14">LACMIP</collectionCode>
|
||
Type 14923 (
|
||
<collectionCode id="7CA7F09FFFDCFFA7FCB4FDF7FCB4FD83" box="[790,894,588,614]" pageId="15" pageNumber="14">LACMIP</collectionCode>
|
||
Locality 7047).
|
||
<emphasis id="28C2B448FFDCFFA7FBE5FDF0FB43FD80" bold="true" box="[1095,1161,587,613]" pageId="15" pageNumber="14">M–O.</emphasis>
|
||
<taxonomicName id="DDB613D9FFDCFFA7FB2DFDF7FEE0FD6C" authority="(Nelson, 1925)" authorityName="Groves & Squires" authorityYear="2023" baseAuthorityName="Nelson" baseAuthorityYear="1925" class="Gastropoda" family="Cypraeidae" genus="Protocypraea" kingdom="Animalia" order="Littorinimorpha" pageId="15" pageNumber="14" phylum="Mollusca" rank="species" species="simiensis">
|
||
<emphasis id="28C2B448FFDCFFA7FB2DFDF7FAFEFD8D" box="[1167,1332,588,616]" italics="true" pageId="15" pageNumber="14">Protocypraea</emphasis>
|
||
?
|
||
<emphasis id="28C2B448FFDCFFA7FAEAFDF1FA7CFD83" box="[1352,1462,586,614]" italics="true" pageId="15" pageNumber="14">simiensis</emphasis>
|
||
(
|
||
<bibRefCitation id="7E2715ABFFDCFFA7FFD9FDD4FED5FD6C" author="Nelson, R. N." box="[123,287,622,649]" pageId="15" pageNumber="14" pagination="397 - 466" refId="ref39989" refString="Nelson, R. N. 1925. A contribution to the paleontology of the Martinez Eocene of California. University of California Publications, Bulletin of the Department of Geological Sciences 15 (11): 397 - 466, pls. 49 - 61." type="journal article" year="1925">Nelson, 1925</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, Santa Susana Formation (Thanetian Stage), Simi Hills, Ventura County, Paleocene, southern Cali- fornia. Basal (
|
||
<emphasis id="28C2B448FFDCFFA7FEBEFD29FEFFFD49" bold="true" box="[284,309,658,684]" pageId="15" pageNumber="14">M</emphasis>
|
||
), dorsal (
|
||
<emphasis id="28C2B448FFDCFFA7FE0FFD29FE0BFD49" bold="true" box="[429,449,658,684]" pageId="15" pageNumber="14">N</emphasis>
|
||
), and right (labral) side (
|
||
<emphasis id="28C2B448FFDCFFA7FD5EFD2AFCDAFD4E" bold="true" box="[764,784,657,683]" pageId="15" pageNumber="14">O</emphasis>
|
||
) views, 1.6x, length
|
||
<quantity id="DD4EC5BFFFDCFFA7FBAFFD29FBB7FD49" box="[1037,1149,658,684]" metricMagnitude="-2" metricUnit="m" metricValue="3.2299999999999995" pageId="15" pageNumber="14" unit="mm" value="32.3">32.3 mm</quantity>
|
||
, width
|
||
<quantity id="DD4EC5BFFFDCFFA7FB7AFD29FA8DFD49" box="[1240,1351,658,684]" metricMagnitude="-2" metricUnit="m" metricValue="2.46" pageId="15" pageNumber="14" unit="mm" value="24.6">24.6 mm</quantity>
|
||
, height
|
||
<quantity id="DD4EC5BFFFDCFFA7FA0AFD29FF68FD35" metricMagnitude="-2" metricUnit="m" metricValue="1.73" pageId="15" pageNumber="14" unit="mm" value="17.3">17.3 mm</quantity>
|
||
, of
|
||
<typeStatus id="C50DD6F8FFDCFFA7FF6EFD0EFEF1FD2A" box="[204,315,693,719]" pageId="15" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
<collectionCode id="7CA7F09FFFDCFFA7FEE3FD0EFE47FD2A" box="[321,397,693,719]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="15" pageNumber="14" type="University or college">UCMP</collectionCode>
|
||
30498,
|
||
<collectionCode id="7CA7F09FFFDCFFA7FE50FD0EFDF4FD2A" box="[498,574,693,719]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="15" pageNumber="14" type="University or college">UCMP</collectionCode>
|
||
Locality 3818.
|
||
<emphasis id="28C2B448FFDCFFA7FD59FD0EFCD8FD2A" bold="true" box="[763,786,693,719]" pageId="15" pageNumber="14">P.</emphasis>
|
||
<taxonomicName id="DDB613D9FFDCFFA7FCB1FD0EFC72FD34" authorityName="SCHILDER" authorityYear="1927" box="[787,952,693,721]" class="Gastropoda" family="Cypraeidae" genus="Protocypraea" kingdom="Animalia" order="Littorinimorpha" pageId="15" pageNumber="14" phylum="Mollusca" rank="genus">
|
||
<emphasis id="28C2B448FFDCFFA7FCB1FD0EFC72FD34" box="[787,952,693,721]" italics="true" pageId="15" pageNumber="14">Protocypraea</emphasis>
|
||
</taxonomicName>
|
||
? sp. 1, Oyster Bay Formation, Paleocene, Appian Way, Central Island District, Vancouver Island, British Columbia. Basal view, 1.8x, length
|
||
<quantity id="DD4EC5BFFFDCFFA7FABDFD63FA45FD16" box="[1311,1423,728,755]" metricMagnitude="-2" metricUnit="m" metricValue="4.81" pageId="15" pageNumber="14" unit="mm" value="48.1">48.1 mm</quantity>
|
||
, width
|
||
<quantity id="DD4EC5BFFFDCFFA7FFD2FD47FF15FCF3" box="[112,223,764,790]" metricMagnitude="-2" metricUnit="m" metricValue="3.1399999999999997" pageId="15" pageNumber="14" unit="mm" value="31.4">31.4 mm</quantity>
|
||
, height
|
||
<quantity id="DD4EC5BFFFDCFFA7FEE2FD40FE65FCF3" box="[320,431,763,790]" metricMagnitude="-2" metricUnit="m" metricValue="2.53" pageId="15" pageNumber="14" unit="mm" value="25.3">25.3 mm</quantity>
|
||
, of
|
||
<collectionCode id="7CA7F09FFFDCFFA7FE7BFD47FDDFFCF3" box="[473,533,764,790]" pageId="15" pageNumber="14">CDM</collectionCode>
|
||
997.91.3, informal locality AP3.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="1A09685AFFDCFFA7FFD2FCF8FDE2FCB8" blockId="15.[112,800,835,1391]" box="[112,552,835,861]" pageId="15" pageNumber="14">
|
||
Stage”),
|
||
<collectingRegion id="D872A6B8FFDCFFA7FF75FCF8FE74FCB8" box="[215,446,835,861]" country="Mexico" name="Baja California Sur" pageId="15" pageNumber="14">Baja California Sur</collectingRegion>
|
||
,
|
||
<collectingCountry id="62A128CAFFDCFFA7FE6BFCF8FDEEFCB8" box="[457,548,835,861]" name="Mexico" pageId="15" pageNumber="14">México</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="52AC3BD1FFDCFFA7FF32FCDDFEBBFC22" pageId="15" pageNumber="14" type="etymology">
|
||
<paragraph id="1A09685AFFDCFFA7FF32FCDDFEBBFC22" blockId="15.[112,800,835,1391]" pageId="15" pageNumber="14">
|
||
<emphasis id="28C2B448FFDCFFA7FF32FCDDFEF1FC65" bold="true" box="[144,315,870,896]" pageId="15" pageNumber="14">Etymology—</emphasis>
|
||
Originally named for Lindsey T. Groves (Natural History Museum of Los Angeles County, Mala- cology Department).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="52AC3BD1FFDCFFA7FF32FC74FE1BFAC2" pageId="15" pageNumber="14" type="description">
|
||
<paragraph id="1A09685AFFDCFFA7FF32FC74FE1BFAC2" blockId="15.[112,800,835,1391]" pageId="15" pageNumber="14">
|
||
<emphasis id="28C2B448FFDCFFA7FF32FC74FE86FC0C" bold="true" box="[144,332,975,1001]" pageId="15" pageNumber="14">Description—</emphasis>
|
||
Shell small (
|
||
<quantity id="DD4EC5BFFFDCFFA7FE42FC6BFDFDFC0F" box="[480,567,976,1002]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="15" pageNumber="14" unit="mm" value="17.0">17 mm</quantity>
|
||
length), oval, high- est at mid-point of dorsum. Base flattened with promi- nent concavity near anterior end. Spire covered, spire area coincident with small pit. Aperture straight, except for bending leftward in posterior region. Denticulation strong, outer lip with 16 teeth, inner lip with 12 teeth, sculpture extending halfway to shell margin, and ribs slightly stronger on inner lip. Outer lip callus prominent and with dorsal ring circumscribing entire shell (
|
||
<bibRefCitation id="7E2715ABFFDCFFA7FD61FB52FEB9FAC2" author="Squires, R. L. & R. A. Demetrion" pageId="15" pageNumber="14" pagination="1 - 55" refId="ref43379" refString="Squires, R. L. and R. A. Demetrion. 1992. Paleontology of the Eocene Bateque Formation, Baja California Sur, Mexico. Contributions in Science, Natural History Museum of Los Angeles County 434: 1 - 55, figs. 1 - 144." type="journal article" year="1992">Squires and Demetrion 1992</bibRefCitation>
|
||
: p. 31).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="52AC3BD1FFDCFFA7FF32FA8BFD45FA88" pageId="15" pageNumber="14" type="discussion">
|
||
<paragraph id="1A09685AFFDCFFA7FF32FA8BFD45FA88" blockId="15.[112,800,835,1391]" pageId="15" pageNumber="14">
|
||
<emphasis id="28C2B448FFDCFFA7FF32FA8BFEE3FAAF" bold="true" box="[144,297,1328,1354]" pageId="15" pageNumber="14">Remarks—</emphasis>
|
||
The two known specimens of
|
||
<taxonomicName id="DDB613D9FFDCFFA7FD15FA8BFF0DFA8A" authorityName="Squires and Demetrion" authorityYear="1992" class="Gastropoda" family="Cypraeidae" genus="Bernaya" kingdom="Animalia" order="Littorinimorpha" pageId="15" pageNumber="14" phylum="Mollusca" rank="species" species="grovesi">
|
||
<emphasis id="28C2B448FFDCFFA7FD15FA8BFF0DFA8A" italics="true" pageId="15" pageNumber="14">Bernaya grovesi</emphasis>
|
||
</taxonomicName>
|
||
have good to excellent preservation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |