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<document id="D0B2E65F6D0C0684BF1DAC623940B16C" ID-DOI="http://dx.doi.org/10.3897/subtbiol.29.34123" ID-GBIF-Dataset="888bf9da-679c-414e-bbe8-addf0c5b34f4" ID-Pensoft-Pub="1314-2615--89" ID-ZooBank="4691E5225A074CA4A45ED4D5F9182BCD" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1314-2615--89" ModsDocOrigin="Subterranean Biology " ModsDocTitle="New genus, two new species and new records of subterranean freshwater snails (Caenogastropoda; Cochliopidae and Lithoglyphidae) from Coahuila and Durango, Northern Mexico" checkinTime="1553126946128" checkinUser="pensoft" docAuthor="Czaja, Alexander, Cardoza-Martinez, Gabriel Fernando, Meza-Sanchez, Iris Gabriela, Estrada-Rodriguez, Jose Luis, Saenz-Mata, Jorge, Becerra-Lopez, Jorge Luis, Romero-Mendez, Ulises, Estrada-Arellano, Josue Raymundo, Garza-Martinez, Miguel Angel &amp; Paulin, Jose Antonio Davila" docDate="2019" docId="634D4BF72F57474E3775ADAD036B414B" docLanguage="en" docName="SubterBiol 29: 89-102" docOrigin="Subterranean Biology 29" docSource="http://dx.doi.org/10.3897/subtbiol.29.34123" docTitle="Balconorbis sabinasense Czaja, Cardoza-Martinez &amp; Estrada-Rodriguez, sp. n." docType="treatment" docUuid="BAEA864F-38F1-4251-854E-D6AAD57D1B86" docUuidSource="ZooBank" docVersion="5" lastPageNumber="95" masterDocId="FF841B3C9A129C05394EFFB1FFA5FFAD" masterDocTitle="New genus, two new species and new records of subterranean freshwater snails (Caenogastropoda; Cochliopidae and Lithoglyphidae) from Coahuila and Durango, Northern Mexico" masterLastPageNumber="102" masterPageNumber="89" pageNumber="94" updateTime="1701317714002" updateUser="plazi">
<mods:mods id="35052C96C97F7B0AEA5D0F9063DFC5FB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3F41DB7965FF3873F6125877580B303B">
<mods:title id="BE6D5B855525F38FE259A9F026FE00FB">New genus, two new species and new records of subterranean freshwater snails (Caenogastropoda; Cochliopidae and Lithoglyphidae) from Coahuila and Durango, Northern Mexico</mods:title>
</mods:titleInfo>
<mods:name id="4B61F906AA7C58504E84F2D81D70E071" type="personal">
<mods:role id="7FAC31D74FED69AD803294AE81015055">
<mods:roleTerm id="DC1B5933F4AE9819A75184143A33FC34">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7B7B1A0E345FF317909C2A5C73D7EC2B">Czaja, Alexander</mods:namePart>
</mods:name>
<mods:name id="0A1F27DE6B7413E1D8574A9FB31AFB90" type="personal">
<mods:role id="1F40DCBB8C18802157714FED6D82EA3D">
<mods:roleTerm id="014E9B3FAD124D26F54EB590AC90BC6E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="12BA899CFA73BD2740CBAD8EE0D3E287">Cardoza-Martinez, Gabriel Fernando</mods:namePart>
</mods:name>
<mods:name id="A0D1A2BD260469773362C4C8A0A9CF17" type="personal">
<mods:role id="92B9297373C74DBA495D076DB0F736F3">
<mods:roleTerm id="82A7410204F6B179AA90139E608A3451">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1FD2CD8E12597EB177139B7AF0FBEB1E">Meza-Sanchez, Iris Gabriela</mods:namePart>
</mods:name>
<mods:name id="53C7C63B5CE20D366BCD12F1823A1A12" type="personal">
<mods:role id="8A1DD20239E3CC00FAFBA913B8263932">
<mods:roleTerm id="A5B65F8C464D7AC5A4E637B51BCD63C2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A2F4679AFACAD76CD56C87AD0C3D50D6">Estrada-Rodriguez, Jose Luis</mods:namePart>
</mods:name>
<mods:name id="A2AEB0BF10ADBC45CB2F22F3ECFF0B10" type="personal">
<mods:role id="1DB6BA81E5F4AD0E1570F16D22AAED2B">
<mods:roleTerm id="4D416D1FD4F1BA2B05F208395827A663">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="688E4A92C24172AB0C7666B37989AEE7">Saenz-Mata, Jorge</mods:namePart>
</mods:name>
<mods:name id="34842489FA2E56E6519DD788CB9CA1D6" type="personal">
<mods:role id="86404A6DD4A9DEE40F48CF9F8993677C">
<mods:roleTerm id="C5BAB9B87D6CB0D414BFEFA77C6DF7CD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="31F81A2FABFE15369D42C08411F7C256">Becerra-Lopez, Jorge Luis</mods:namePart>
</mods:name>
<mods:name id="A48736BA0D9C0F901FBE5DFA938E761D" type="personal">
<mods:role id="D61D32EFA152C3CB06D0064F3F0AF92D">
<mods:roleTerm id="B2A7F49E4CB3F98B5056F0BE2E5CDCC6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1F2A17DD7E05A027EC2C5FEB2CE7806C">Romero-Mendez, Ulises</mods:namePart>
</mods:name>
<mods:name id="B0274B5DC6DE2F0851DDA0C8D38C6348" type="personal">
<mods:role id="33872226F86B9FA42D65898B86EED8A8">
<mods:roleTerm id="9D203EE870EB8757334917A93D7C717F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D96C3C80F82588C224A6FEF44525F232">Estrada-Arellano, Josue Raymundo</mods:namePart>
</mods:name>
<mods:name id="45FCDE76BEC9ED7DCBF957539AE69922" type="personal">
<mods:role id="13C1E372D315BA4B43C419B4C87FCFB4">
<mods:roleTerm id="A3394B05A32EF9162849C829A6FE1CCB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FF0AFF47F3F745506E0D7051D70854EC">Garza-Martinez, Miguel Angel</mods:namePart>
</mods:name>
<mods:name id="F5B49A080967E0A176DB328739EB9196" type="personal">
<mods:role id="F74C6CBE9B9FB6EEA75F9B6A38F56E70">
<mods:roleTerm id="C875094F9A45C2142FB868BF32A6685A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5004496AADDF2EF8F17EC5CAF796FF92">Paulin, Jose Antonio Davila</mods:namePart>
</mods:name>
<mods:typeOfResource id="41053920DD357EC46E23A686DB476B6A">text</mods:typeOfResource>
<mods:relatedItem id="271558C7BB06A06470B0AC747F2D434B" type="host">
<mods:titleInfo id="673C61880674A7A8B63BEAD828A57DF5">
<mods:title id="1BC11700B710589A30A1F3A024A78D47">Subterranean Biology</mods:title>
</mods:titleInfo>
<mods:part id="4DA2B7DD036B36EB9F5DD151766929F7">
<mods:date id="F2A84E6D0F732A8C2286F5ACD04197FD">2019</mods:date>
<mods:detail id="43487971963213D01C4BAFF4A07DC9B6" type="volume">
<mods:number id="6FC144C1108FDAAA2D8370FECE04CCDB">29</mods:number>
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<mods:start id="EF64D4F3CBC5A8A36A5BB9E505323387">89</mods:start>
<mods:end id="BD19084A83F4BCC36EC84BD1E8E94142">102</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:location id="874E0AD2EF311405E360555E178BF53A">
<mods:url id="481EC56C69FB308F50E347A394CB9AE3">http://dx.doi.org/10.3897/subtbiol.29.34123</mods:url>
</mods:location>
<mods:classification id="B8AC1552400A38A171A721DB2984A125">journal article</mods:classification>
<mods:identifier id="A66A37622CE82193D08AA39488B4D002" type="DOI">http://dx.doi.org/10.3897/subtbiol.29.34123</mods:identifier>
<mods:identifier id="FC2EA490A8A15FD1907D72F554519EF0" type="Pensoft-Pub">1314-2615--89</mods:identifier>
<mods:identifier id="44CB783E0AA52A968B82D2816A1F8E65" type="ZooBank">4691E5225A074CA4A45ED4D5F9182BCD</mods:identifier>
</mods:mods>
<treatment id="634D4BF72F57474E3775ADAD036B414B" ID-GBIF-Taxon="154474162" LSID="urn:lsid:zoobank.org:act:BAEA864F-38F1-4251-854E-D6AAD57D1B86" httpUri="http://treatment.plazi.org/id/634D4BF72F57474E3775ADAD036B414B" lastPageId="6" lastPageNumber="95" pageId="5" pageNumber="94">
<subSubSection id="AE63EFA8D803DE5BE92760ED61291B2B" pageId="5" pageNumber="94" type="nomenclature">
<paragraph id="4B24A87E12D5C98440252177040A3354" pageId="5" pageNumber="94">
<taxonomicName id="2B7CE40B5A10B10026EF810B087FB953" LSID="http://zoobank.org/BAEA864F-38F1-4251-854E-D6AAD57D1B86" authority="Czaja, Cardoza-Martinez &amp; Estrada-Rodriguez" class="Gastropoda" family="Lithoglyphidae" genus="Balconorbis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balconorbis sabinasense" order="Littorinimorpha" pageId="5" pageNumber="94" phylum="Mollusca" rank="species" species="sabinasense">
<pageBreakToken id="A02927A0815949B3B35127B940E515D6" pageId="5" pageNumber="94" start="start">Balconorbis</pageBreakToken>
sabinasense Czaja,
<normalizedToken id="CC48CBDE3BFE398974C235F50777686E" originalValue="Cardoza-Martínez">Cardoza-Martinez</normalizedToken>
&amp;
<normalizedToken id="AC8442209FB75F9A69B078755D41C3A9" originalValue="Estrada-Rodríguez">Estrada-Rodriguez</normalizedToken>
</taxonomicName>
<taxonomicNameLabel id="87B542F9322A177550F41C50B2477C41" pageId="5" pageNumber="94">sp. n.</taxonomicNameLabel>
Figures 12-18
</paragraph>
</subSubSection>
<subSubSection id="FE5B3297FB29791BBF4383A339966BE1" pageId="5" pageNumber="94" type="type locality">
<paragraph id="04EE0F9BA4C83EBA0EE03965271BE738" pageId="5" pageNumber="94">Type locality.</paragraph>
<paragraph id="8317D39B2D673ED60EFE13714D735AD8" pageId="5" pageNumber="94">
MEXICO, Coahuila state, Don
<normalizedToken id="B4AE44E4876260785396163FD815637D" originalValue="Martín">Martin</normalizedToken>
Basin, Sabinas River, Las Adjuntas (Rancho San Carlos):
<geoCoordinate id="5559133F1A7AD173FEF9A09E01CC7E21" direction="west" orientation="longitude" precision="15" value="-101.194725">101°11'41&quot;W</geoCoordinate>
,
<geoCoordinate id="E043CC6A9E3D1B401D8982701BFBE0FF" direction="north" orientation="latitude" precision="15" value="27.8925">27°53'33&quot;N</geoCoordinate>
, 354 m a.s.l. (Fig. 1).
</paragraph>
</subSubSection>
<subSubSection id="75B4D8C6149D2208FAC45F3A9F8399B0" pageId="5" pageNumber="94" type="type material">
<paragraph id="083A1F94B6B6E40993431E42B3747B58" pageId="5" pageNumber="94">Type material.</paragraph>
<paragraph id="51A76E9B633B191D1537BE527E69185E" pageId="5" pageNumber="94">Holotype (Figs 12-14), UJMC 410, from type locality, leg. Cecilio Arreola Chapa, 30/xi/2018. Paratypes, UJMC 411, CNMO 7901, from the same lot, 11 specimens.</paragraph>
<caption id="0BED1DCD6684E3EEB606675C7930C73D" pageId="5" pageNumber="94">
<paragraph id="74B53A7229F8FF6D8D483B44627519CC" pageId="5" pageNumber="94">
Figures 12-23. Shells of
<taxonomicName id="5DDF2A825D8A83E64A05740F4FA3B0CB" class="Gastropoda" family="Cochliopidae" genus="Balconorbis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balconorbis sabinasense" order="Littorinimorpha" pageId="5" pageNumber="94" phylum="Mollusca" rank="species" species="sabinasense">Balconorbis sabinasense</taxonomicName>
sp. n. and
<taxonomicName id="36EAED2AA3DB82FE0B37B3F834FDFE27" class="Gastropoda" family="Cochliopidae" genus="Coahuilix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coahuilix parrasense" order="Littorinimorpha" pageId="5" pageNumber="94" phylum="Mollusca" rank="species" species="parrasense">Coahuilix parrasense</taxonomicName>
Czaja,
<normalizedToken id="CFCB9AE433C46DA3955607A0CC44DA03" originalValue="Estrada-Rodríguez">Estrada-Rodriguez</normalizedToken>
,
<normalizedToken id="25F90D3644B9C5080A98B78A466297A5" originalValue="Romero-Méndez">Romero-Mendez</normalizedToken>
,
<normalizedToken id="08D72BA1F3A8A801D2E82B9311FC916F" originalValue="Ávila-Rodríguez">Avila-Rodriguez</normalizedToken>
,
<normalizedToken id="604A7C2C34B0BA21FADB583D0FCAC73A" originalValue="Meza-Sánchez">Meza-Sanchez</normalizedToken>
&amp; Covich, 2017. 12-14
<taxonomicName id="E367F6F9B1932CA3BB8DE3A94053EA2C" lsidName="B. sabinasense" pageId="5" pageNumber="94" rank="species" species="sabinasense">B. sabinasense</taxonomicName>
n. sp., holotype, UJMC 410. 15-18 Paratype 1, UJMC 411. 19-21
<taxonomicName id="63F3E522CF67501C496C17AF5B9C4844" lsidName="C. parrasense" pageId="5" pageNumber="94" rank="species" species="parrasense">C. parrasense</taxonomicName>
, UJMC 418, from Nazas River, Durango. 22, 23
<taxonomicName id="46432F1904DAB16E9E3A5266DC81BB8C" lsidName="C. parrasense" pageId="5" pageNumber="94" rank="species" species="parrasense">C. parrasense</taxonomicName>
, fossil specimen from Parras de la Fuente, Coahuila, holotype, UJMC-320, from
<bibRefCitation id="53F908D9ACBEA3216BEC6E32A57E2962" author="Czaja, A" journalOrPublisher="Revista Mexicana de Biodiversidad" pageId="9" pageNumber="98" pagination="912 - 917" title="Primer registro fosil del gasteropodo Cincinnatia (Hydrobiidae: Nymphophilinae) en Mexico." url="https://doi.org/10.1016/j.rmb.2017.10.025" volume="88" year="2017 c">Czaja et al. 2017c</bibRefCitation>
, Fig. 4A, C. Scale bar: 0.5 mm (12-14, 19-23).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="3D29D83454D82EAAF8367C52BB7E8A96" pageId="5" pageNumber="94" type="etymology">
<paragraph id="0732C54A3865FF96EF5456B189BCA6EB" pageId="5" pageNumber="94">Etymology.</paragraph>
<paragraph id="1C2600FE5B9AC51BF7C9EA0463635F91" pageId="5" pageNumber="94">Derived from the name of Sabinas River, Coahuila, where the type locality is situated.</paragraph>
</subSubSection>
<subSubSection id="04E700F92C1CB309E824D074452119F1" pageId="5" pageNumber="94" type="materials_examined">
<paragraph id="7032E4187A0708F4B4D742CE1794874B" pageId="5" pageNumber="94">Referred material.</paragraph>
<paragraph id="5687F1B5070DEF8BC1660F3BB659D50C" pageId="5" pageNumber="94">
COAHUILA. Don
<normalizedToken id="7BE69BD61FD018FC63E4BDA7CBE086E6" originalValue="Martín">Martin</normalizedToken>
Basin, Sabinas River, UJMC 412, Ejido Santa
<normalizedToken id="46334B6204A1896F35F8482656F77370" originalValue="María">Maria</normalizedToken>
, Y. A.
<normalizedToken id="F9EFD3F58E1F32E3824249CF16922F8C" originalValue="Sanchez-Montañez">Sanchez-Montanez</normalizedToken>
, 03/vii/2018. UJMC 413, Ejido Paso del Coyote, Cecilio Arreola Chapa, 30/xi/2018. UJMC 414, Sabinas (Agua Prieta), Y. A.
<normalizedToken id="E2E61740E2BC76653C623BCABEEF5CD4" originalValue="Sanchez-Montañez">Sanchez-Montanez</normalizedToken>
, 08/vii/2018. UJMC 415, La Vega, Y. A.
<normalizedToken id="9E0801D277625DEB76114E04102A3353" originalValue="Sanchez-Montañez">Sanchez-Montanez</normalizedToken>
, 08/vii/2018.
</paragraph>
</subSubSection>
<subSubSection id="17D5D8FB58A9A12611DDDA5F08A6DAD7" pageId="5" pageNumber="94" type="diagnosis">
<paragraph id="B00B951BBA61128E19AC2FF3B8761B40" pageId="5" pageNumber="94">Diagnosis.</paragraph>
<paragraph id="5F6E1B829D2490EC24CA27CE5551AC3D" pageId="5" pageNumber="94">
Shell minute, less than 2 mm in diameter (width), planispiral, apertural plane not or only slightly inclined, with
<normalizedToken id="9525D54A9468128E242AA4C6AF2986B8" originalValue="3¾">33/4</normalizedToken>
or less whorls, bodywhorl keeled, with spiral lines and axial growth lines, protoconch smooth.
</paragraph>
</subSubSection>
<subSubSection id="63CF38FDC0B609F7982CA465EFA45C4A" pageId="5" pageNumber="94" type="differential diagnosis">
<paragraph id="9F8B42D2D431F352869C6FBF79AAA943" pageId="5" pageNumber="94">Differential diagnosis.</paragraph>
<paragraph id="4B6980CF7A9E061E56D4BBDC7AEC5DEB" pageId="5" pageNumber="94">
According to
<bibRefCitation id="10447549E09943C277F9CB8EE359EB35" author="Hershler, R" journalOrPublisher="Proceedings of the Biological Society of Washington" pageId="9" pageNumber="98" pagination="121 - 136" title="Hadocerastaylori, a new genus and species of phreatic Hydrobiidae (Gastropoda: Rissoacea) from South-central Texas." volume="99" year="1986">Hershler and Longley (1986)</bibRefCitation>
, the characteristic spiral structure (Figs 16-18) of the shells distinguish the genus
<taxonomicName id="258968115016F8E16167C8CE0AE7D22F" class="Gastropoda" family="Cochliopidae" genus="Balconorbis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balconorbis" order="Littorinimorpha" pageId="5" pageNumber="94" phylum="Mollusca" rank="genus">Balconorbis</taxonomicName>
clearly from other subterranean littoridinines. Shells of the new species differs from
<taxonomicName id="E8CEC1B6170161A6C99A638BDD79DAB7" lsidName="B. uvaldensis" pageId="5" pageNumber="94" rank="species" species="uvaldensis">B. uvaldensis</taxonomicName>
by larger shells (1.51-1.81 mm by
<taxonomicName id="F27321819320A0364F518D016582D014" genus="B." infraspecific-rank="sp. n." lsidName="B. sabinasense" pageId="5" pageNumber="94" rank="infraspecific-rank" species="sabinasense">B. sabinasense sp. n.</taxonomicName>
versus 0.90-1.22 mm by
<taxonomicName id="68291744E1605F5C047733335F40CAB8" lsidName="B. uvaldensis" pageId="5" pageNumber="94" rank="species" species="uvaldensis">B. uvaldensis</taxonomicName>
) with weaker suture, more whorls, smooth to slightly pitted protoconch without spiral lines, and characteristic keels on the bodywhorl. Similar planispiral shells of the genus
<taxonomicName id="2710370C74470E1F788E3D72BA0BDA3B" class="Gastropoda" family="Cochliopidae" genus="Phreatodrobia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phreatodrobia" order="Littorinimorpha" pageId="5" pageNumber="94" phylum="Mollusca" rank="genus">Phreatodrobia</taxonomicName>
(
<taxonomicName id="CC2270A9D5D7DE0C63434C2C6A40720F" lsidName="P. micra" pageId="5" pageNumber="94" rank="species" species="micra">P. micra</taxonomicName>
Hershler &amp; Longley, 1986,
<taxonomicName id="B4EE5FADF495FB4122269F674EB8531D" lsidName="P. plana" pageId="5" pageNumber="94" rank="species" species="plana">P. plana</taxonomicName>
Hershler &amp; Longley, 1986 and
<taxonomicName id="DD1DF233F79424252FBBCA915FF75170" lsidName="P. rotunda" pageId="5" pageNumber="94" rank="species" species="rotunda">P. rotunda</taxonomicName>
Hershler &amp; Longley, 1986) are considerable smaller, lack the keels on the bodywhorl and do not have the characteristic regularly-spaced spiral lines and axial growth lines.
</paragraph>
</subSubSection>
<subSubSection id="3A5B1D370A2304F52012A463DA953378" pageId="5" pageNumber="94" type="description">
<paragraph id="748304C60F383ED015A5FEF201BA1677" pageId="5" pageNumber="94">Description.</paragraph>
<paragraph id="10CD9B5B52F180DE253004244D630D85" pageId="5" pageNumber="94">
Shell minute, planispiral, width 1.51-1.81 mm, height 0.42-0.61 mm, with
<normalizedToken id="A8BE97C921C3AE23E01FB7FAC5F8FF9F" originalValue="3¼–3¾">31/4-33/4</normalizedToken>
whorls; protoconch smooth to slightly pitted (Fig. 16), hidden in ventral view, has
<normalizedToken id="577D811FB0B11EBF2C97062A4D348C33" originalValue="1¼">11/4</normalizedToken>
whorls, first teleoconch whorl with strong and elevated axial growth lines which cross the spiral lines producing a square pattern (Figs 17, 18), about 80 elevated spiral lines are present on the bodywhorl, bodywhorl with one or two keels, one keel usually stronger; aperture rounded to ovate, apertural plane only slightly inclined from the columellar axis (Fig. 13); inner peristome is fused to the penultimate whorl, outer lip strong and advanced relative to the remaining peristome; shells transparent or whitish.
</paragraph>
</subSubSection>
<subSubSection id="E8578B4D4E0105C5346E39B83EA6D54B" pageId="5" pageNumber="94" type="description">
<paragraph id="E975A69462FEBA9B53C1623755E18F1F" pageId="5" pageNumber="94">Shell measurements</paragraph>
<paragraph id="BB5CF2741A4D1D8CB92139F1908BCCA8" pageId="5" pageNumber="94">
(mean
<normalizedToken id="74310673FB7B38A7D46C2243F0E726D5" originalValue="±">+/-</normalizedToken>
standard deviation in parentheses; n = 11): SH 0.53 (0.07) mm, SW 1.68 (0.10) mm, AH 0.59 (0.05) mm, AW 0.49 (0.08) mm, WN 3.5 (0.19), aperture plane inclination relative to shell axis = 22°-31°. Paratypes from the type locality.
</paragraph>
</subSubSection>
<subSubSection id="EFAA271375225BBAAF74B2744CA80D33" pageId="5" pageNumber="94" type="description">
<paragraph id="DEC1880A449AE34F372C316D1EC481C4" pageId="5" pageNumber="94">Measurements of holotype.</paragraph>
<paragraph id="8BEE3AE0B1E2DEC45541980A67872CD4" pageId="5" pageNumber="94">
SH 0.55 mm, SW 1.58 mm, AH 0.59 mm, AW 0.58 mm, HBW 0.50 mm, WN
<normalizedToken id="789B724733ACEF70671D716C17907F50" originalValue="3½">31/2</normalizedToken>
, aperture plane inclination relative to shell axis = 25°.
</paragraph>
</subSubSection>
<subSubSection id="2E6F808864744E594D7617C6B5DFB2C4" lastPageId="6" lastPageNumber="95" pageId="5" pageNumber="94" type="habitat">
<paragraph id="3C4576C2F36BB4C8450119AB42BB7008" pageId="5" pageNumber="94">Habitat.</paragraph>
<paragraph id="13DE467D8EE98C210A74BD0D1358AA01" pageId="5" pageNumber="94">
Similar to
<taxonomicName id="A192E4290E8FF7E7000AB2B48F2AEF90" class="Gastropoda" family="Lithoglyphidae" genus="Phreatomascogos" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phreatomascogos gregoi" order="Littorinimorpha" pageId="5" pageNumber="94" phylum="Mollusca" rank="species" species="gregoi">Phreatomascogos gregoi</taxonomicName>
sp. n.
</paragraph>
<paragraph id="8DCF3AEBF56407E24BC0FA04D5474067" pageId="6" pageNumber="95">
<pageBreakToken id="8003DF1562F5858FEF5651CFBF018E03" pageId="6" pageNumber="95" start="start">Distribution</pageBreakToken>
. Known from the type locality and various sites among the Sabinas River between the localities Ejido Santa
<normalizedToken id="CFA8A4021D8F64CA279BCC51CECD4686" originalValue="María">Maria</normalizedToken>
and La Vega. Within the type locality and most other sites the new species occurs sympatrically to
<taxonomicName id="D8BDEDA2FF706200C5ACAA2698DA14ED" class="Gastropoda" family="Lithoglyphidae" genus="Phreatomascogos" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phreatomascogos gregoi" order="Littorinimorpha" pageId="6" pageNumber="95" phylum="Mollusca" rank="species" species="gregoi">Phreatomascogos gregoi</taxonomicName>
sp. n.
</paragraph>
</subSubSection>
<subSubSection id="EDFFC82A7A70C192415FDCF713183BD5" pageId="6" pageNumber="95" type="remarks">
<paragraph id="97992996A03B187579B5C43F6E220562" pageId="6" pageNumber="95">Remarks.</paragraph>
<paragraph id="CA3E21F5FC77033A70EB908EA038EB29" pageId="6" pageNumber="95">
Almost all the members of
<taxonomicName id="F15574BBF6027408A6870A62621DB1C4" family="Cochliopidae" lsidName="" pageId="6" pageNumber="95" rank="family">Cochliopidae</taxonomicName>
family with planispiral shells are subterranean forms. Freshwater gastropods with similar shell characteristics can be found only in the genus
<taxonomicName id="A872FA491E832AB3256956C757B14114" class="Gastropoda" family="Cochliopidae" genus="Balconorbis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balconorbis" order="Littorinimorpha" pageId="6" pageNumber="95" phylum="Mollusca" rank="genus">Balconorbis</taxonomicName>
which is monotypic with
<taxonomicName id="5197985C1C2424B66CF0752091E62DCA" lsidName="B. uvaldensis" pageId="6" pageNumber="95" rank="species" species="uvaldensis">B. uvaldensis</taxonomicName>
in the Edwards (Balcones Fault Zone) Aquifer in Uvalde County, Texas, United State. Our new species resembles
<taxonomicName id="730E2D5B965F53C4797D06D58B3FD455" lsidName="B. uvaldensis" pageId="6" pageNumber="95" rank="species" species="uvaldensis">B. uvaldensis</taxonomicName>
but differs from this by having larger shells, smooth protoconch, one more whorl and a bodywhorl with one or two more or less strong keels and by much closer, compressed set of the whorls resulting in weaker suture and larger columellar peristome. The later feature is unique in the genus
<taxonomicName id="2527FB6467D73756A4E30790ADC163D0" class="Gastropoda" family="Cochliopidae" genus="Balconorbis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balconorbis" order="Littorinimorpha" pageId="6" pageNumber="95" phylum="Mollusca" rank="genus">Balconorbis</taxonomicName>
and could indicate the possible position of the new species in a new geographically isolated genus. These conchological differences confirm that the new species is morphologically diagnosable and distinct to shells of the snails from Texas. It is the first record of the genus
<taxonomicName id="202F00013F61083A47C82392B9CF944E" class="Gastropoda" family="Cochliopidae" genus="Balconorbis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balconorbis" order="Littorinimorpha" pageId="6" pageNumber="95" phylum="Mollusca" rank="genus">Balconorbis</taxonomicName>
in Mexico.
</paragraph>
</subSubSection>
</treatment>
</document>
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