127 lines
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127 lines
13 KiB
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<document id="FA2ACF361486657EB5CD0E34AB2CD2E1" ID-CLB-Dataset="35529" ID-DOI="http://dx.doi.org/10.3897/zookeys.531.6097" ID-GBIF-Dataset="70702787-a6a8-4af7-9702-738f344a5cbe" ID-PMC="PMC4668911" ID-Pensoft-Pub="1313-2970-531-1" ID-PubMed="26692803" ID-ZooBank="C845838EC9124BD8AB4E07980F91959E" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-531-1" ModsDocOrigin="ZooKeys 531" ModsDocTitle="Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species" checkinTime="1451243834596" checkinUser="pensoft" docAuthor="Vermeulen, Jaap J., Liew, Thor-Seng & Schilthuizen, Menno" docDate="2015" docId="50D9CF5F68F512D0133D228C2B84BDF9" docLanguage="en" docName="ZooKeys 531: 1-139" docOrigin="ZooKeys 531" docSource="http://dx.doi.org/10.3897/zookeys.531.6097" docTitle="Microcystina Moerch 1872" docType="treatment" docVersion="11" lastPageNumber="27" masterDocId="1527DF5A6645C540FFF1FF85AF4AE358" masterDocTitle="Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species" masterLastPageNumber="139" masterPageNumber="1" pageNumber="27" updateTime="1701381844169" updateUser="ExternalLinkService">
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<mods:titleInfo id="C1CC7A10A5A87CB7E8FD5B5DF1371B87">
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<mods:title id="6BE655BD006625F9416B014A2D839016">Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species</mods:title>
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<mods:namePart id="944B844A6C6FBCA9161EE2BB502A950E">Vermeulen, Jaap J.</mods:namePart>
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<mods:namePart id="4BA61B28A37DCFDBC1ED15CB15E89F04">Liew, Thor-Seng</mods:namePart>
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<mods:namePart id="10355DB70054E88649B42609062FA2B9">Schilthuizen, Menno</mods:namePart>
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<mods:title id="2F1F2BD3721006649DB464359C9EBDB7">ZooKeys</mods:title>
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<mods:part id="B01899520FA81F507264C8E18B4AFC78">
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<mods:date id="7F0931C86E522733C6EF4D07C78E24F5">2015</mods:date>
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<mods:detail id="F52BA7CC177D8ADD4598D22B1B6685D5" type="volume">
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<mods:number id="19A9341C61AC2447CFE3B324C4A76644">531</mods:number>
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<mods:start id="D092BC3373CB77BE51654C7A0093C31C">1</mods:start>
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<mods:end id="EA7039667B2874D33AD755D57A1780D6">139</mods:end>
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<mods:url id="DB7CA64EB7F4FD79C05D9ED245295994">http://dx.doi.org/10.3897/zookeys.531.6097</mods:url>
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<mods:classification id="36AF152B900F9ED3355B2DBDB5B6C66B">journal article</mods:classification>
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<mods:identifier id="E7ED81F56D058F0610725D0F43D12274" type="DOI">http://dx.doi.org/10.3897/zookeys.531.6097</mods:identifier>
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<mods:identifier id="D178E043FFD440177C2561985B1B37D4" type="Pensoft-Pub">1313-2970-531-1</mods:identifier>
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<mods:identifier id="1F359CABCE0830D7B98F81ECFEBB5D35" type="ZooBank">C845838EC9124BD8AB4E07980F91959E</mods:identifier>
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</mods:mods>
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<treatment id="50D9CF5F68F512D0133D228C2B84BDF9" ID-GBIF-Taxon="144057565" LSID="urn:lsid:plazi:treatment:50D9CF5F68F512D0133D228C2B84BDF9" httpUri="http://treatment.plazi.org/id/50D9CF5F68F512D0133D228C2B84BDF9" lastPageNumber="27" pageId="26" pageNumber="27">
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<subSubSection id="A10213907FD671C73372591FC23C0A2D" pageId="26" pageNumber="27" type="multiple">
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<paragraph id="C45F33E3FD149C61CE25FB2673DE8AD7" pageId="26" pageNumber="27">Taxon classification Animalia Stylommatophora Ariophantidae</paragraph>
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</subSubSection>
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<subSubSection id="C3A47D2D6C23A983CAA4CFB5D9043A4E" pageId="26" pageNumber="27" type="nomenclature">
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<paragraph id="82029E1D81A8CF0D84915C1A55E7CB5B" pageId="26" pageNumber="27">
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Genus
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<taxonomicName id="A2BCCB4B91763B43E0C4B0E6E7072858" ID-CoL="7P8VS" authority="Moerch, 1872" authorityName="Moerch" authorityYear="1872" class="Gastropoda" family="Ariophantidae" genus="Microcystina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcystina" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="genus">
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Microcystina
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<normalizedToken id="1411394192B7D427488C113ECD29DCD1" originalValue="Mörch" pageId="26" pageNumber="27">Moerch</normalizedToken>
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, 1872
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="9B44A2C22121071B5EA4200FD868A7CB" pageId="26" pageNumber="27" type="reference_group">
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<paragraph id="B1F629BE74E909CA02D75506ACE715B0" pageId="26" pageNumber="27">
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<taxonomicName id="703573FB2604120A9990C431A66E54AD" class="Gastropoda" family="Ariophantidae" genus="Nanina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanina" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="genus">Nanina</taxonomicName>
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subgenus
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<taxonomicName id="3C7C07373F2ED722F7A164235F92E793" class="Gastropoda" family="Ariophantidae" genus="Microcystina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcystina" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="genus">Microcystina</taxonomicName>
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<normalizedToken id="7C9B8E9207D883FEFDC44D50AFF9FF08" originalValue="Mörch" pageId="26" pageNumber="27">Moerch</normalizedToken>
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, 1872: 311.
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<taxonomicName id="88CB5CF617801927F597708A0C74DF77" class="Gastropoda" family="Ariophantidae" genus="Microcystina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcystina" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="genus">Microcystina</taxonomicName>
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(
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<normalizedToken id="9A5EAEC78AD38AF82671426C3F7AD125" originalValue="Mörch" pageId="26" pageNumber="27">Moerch</normalizedToken>
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) Godwin Austen, 1882 (1882-1888): 11.
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</paragraph>
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</subSubSection>
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<subSubSection id="8E7DCB4CACC51E871E51CFDFB33F94A2" pageId="26" pageNumber="27" type="diagnosis for the sabah species">
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<paragraph id="45EE229A4F9A29EBC68DF5C54B7E5C96" pageId="26" pageNumber="27">Diagnosis for the Sabah species.</paragraph>
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<paragraph id="B2563EB66D18E5D054D5A083F84FC263" pageId="26" pageNumber="27">Shell minute or very small, thin, lenticular, inflated-lenticular to depressed-ovoid; spire almost flat to moderately elevated. Surface shiny or glossy. Sculpture very fine, hardly prominent. Umbilicus open but very narrow, to closed, often partly or entirely covered by a minute, triangular spur protruding from the columellar corner of the peristome; this spur lacking or inconspicuous in juveniles. Dimensions: Width of adult shells 1.7-3.6 mm.</paragraph>
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</subSubSection>
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<subSubSection id="A8BE21A8570476DD4D38F10D75357F37" pageId="26" pageNumber="27" type="remarks">
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<paragraph id="2C54EFFC514C1F2DEB5D0FDE5BDF75CD" pageId="26" pageNumber="27">Remarks.</paragraph>
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<paragraph id="CD22D3DF85AFCBE7E5756FD745B0926B" pageId="26" pageNumber="27">
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The genus
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<taxonomicName id="5C740F02745C6ADF075CC67594B1E16F" class="Gastropoda" family="Ariophantidae" genus="Microcystina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcystina" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="genus">Microcystina</taxonomicName>
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includes small (up to 3.6 mm wide) species with lenticular (depressed ovoid in
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<taxonomicName id="4C82E2ABA7D05CB5D28D690A87B6960E" class="Gastropoda" family="Ariophantidae" genus="Microcystina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcystina physotrochus" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="physotrochus">Microcystina physotrochus</taxonomicName>
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) shells. Invariably, the shell surface seems smooth at a first impression, any sculpture present on the shells is very fine and inconspicuous. This distinguishes the smaller species of
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<taxonomicName id="837719B6103EF54898FF3AFACC293A92" class="Gastropoda" family="Ariophantidae" genus="Microcystina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcystina" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="genus">Microcystina</taxonomicName>
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(particularly
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<taxonomicName id="01D48D1851CCF92E5067582BEE4F8B53" class="Gastropoda" family="Ariophantidae" genus="Microcystina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcystina sinica" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="sinica">Microcystina sinica</taxonomicName>
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) from sympatric, yet undescribed
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<taxonomicName id="AEC54922E8E3E118AD94A7AE7EAEFFF6" class="Gastropoda" family="Charopidae" genus="Charopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charopa" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="genus">Charopa</taxonomicName>
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species (
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<taxonomicName id="CB573AFFC5C7F0DE3AF8AC54CE23B75F" family="Charopidae" lsidName="" pageId="26" pageNumber="27" rank="family">Charopidae</taxonomicName>
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), which have lenticular, minute shells (width of adult shells 0.9-1.9 mm) with a much more conspicuous sculpture. In several
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<taxonomicName id="5A73FB009327D1266BD8BDD372B5F58B" class="Gastropoda" family="Ariophantidae" genus="Microcystina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcystina" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="genus">Microcystina</taxonomicName>
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species the umbilicus is partly or entirely covered by a minute, triangular spur protruding from the columellar corner of the peristome.
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</paragraph>
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<paragraph id="008A50B9600AC40728F369522F473711" pageId="26" pageNumber="27">
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Borneo
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<taxonomicName id="9AC76099FC62957AE6061FA8602EC691" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrochlamys" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="genus">Macrochlamys</taxonomicName>
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and
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<taxonomicName id="198A535650B5CB4D21CB5B5B673FC5FB" class="Gastropoda" family="Dyakiidae" genus="Everettia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Everettia" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="genus">Everettia</taxonomicName>
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(
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<taxonomicName id="0A07F72A1C11301F19F4458E1B7FD202" family="Ariophantidae" lsidName="" pageId="26" pageNumber="27" rank="family">Ariophantidae</taxonomicName>
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), and
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<taxonomicName id="E0339719E9A8E3DC4726C7AAE5EDE051" class="Gastropoda" family="Helicarionidae" genus="Helicarion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helicarion" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="genus">Helicarion</taxonomicName>
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(
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<taxonomicName id="48C2B4C8A65F895B809D115B2A3B4488" family="Helicarionidae" lsidName="" pageId="26" pageNumber="27" rank="family">Helicarionidae</taxonomicName>
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) may have similarly shaped shells, including the inconspicuous sculpture, but they are larger (width of adult shells 8 mm or more).
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</paragraph>
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<paragraph id="CD734EE0FFB558581FBB902CD8FC4771" pageId="26" pageNumber="27">
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Placement of
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<taxonomicName id="B1A76C49466D9A24082AF812D59C10F1" class="Gastropoda" family="Ariophantidae" genus="Microcystina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcystina" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="genus">Microcystina</taxonomicName>
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into the family
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<taxonomicName id="4B8C6DDF09EDB739DF8BE79B0314F461" family="Ariophantidae" lsidName="" pageId="26" pageNumber="27" rank="family">Ariophantidae</taxonomicName>
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follows
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<bibRefCitation id="74232FC915424F227F64962B7B96B7E3" author="Schileyko, AA" journalOrPublisher="Ruthenica" pageId="26" pageNumber="27" pagination="1309 - 1466" title="Treatise on Recent Terrestrial Pulmonate Molluscs; Part 10; Ariophantidae, Ostracolethidae, Ryssotidae, Milacidae, Dyakiidae, Staffordiidae, Gastrodontidae, Zonitidae, Daudebardiidae, Parmacellidae." volume="10" year="2003">Schileyko (2003)</bibRefCitation>
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.
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</paragraph>
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<paragraph id="BF39DE763CC8AA84D323CE703410B189" pageId="26" pageNumber="27">
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We provide a review of the Sabah species of
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<taxonomicName id="353C4460EB32F77BD320F1E3097AE274" class="Gastropoda" family="Ariophantidae" genus="Microcystina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcystina" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="genus">Microcystina</taxonomicName>
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. We divide the genus into two informal groups.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |