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<document ID-CLB-Dataset="6050" ID-DOI="http://dx.doi.org/10.3897/zookeys.1044.60072" ID-GBIF-Dataset="a8375122-971c-4804-92a6-fe16e10e3224" ID-PMC="PMC8222295" ID-Pensoft-Pub="1313-2970-1044-427" ID-Pensoft-UUID="8320077CEFA5571D971EFEA66B0C6268" ID-PubMed="34183882" ID-ZooBank="9AA72654B6CC4081B557CD0BAE3939A3" ModsDocID="1313-2970-1044-427" checkinTime="1623875398087" checkinUser="pensoft" docAuthor="Allegro, Gianni &amp; Giachino, Pier Mauro" docDate="2021" docId="3C1B7F5006C75DE790EAECA39CF1CC62" docLanguage="en" docName="ZooKeys 1044: 427-448" docOrigin="ZooKeys 1044" docPubDate="2021-06-16" docSource="http://dx.doi.org/10.3897/zookeys.1044.60072" docTitle="Diplocheila (Submera) laevis" docType="treatment" docVersion="6" id="8320077CEFA5571D971EFEA66B0C6268" lastPageNumber="427" masterDocId="8320077CEFA5571D971EFEA66B0C6268" masterDocTitle="The genus Diplocheila Brulle, 1834 in Cambodia, with descriptions of two new species (Coleoptera, Carabidae, Licinini)" masterLastPageNumber="448" masterPageNumber="427" pageNumber="427" updateTime="1732756944497" updateUser="ExternalLinkService">
<mods:mods id="9951940DCEA831BECD46E0E30EFD3247" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="CBB32BE386EED3959581E7C0E9E4B297">The genus Diplocheila Brulle, 1834 in Cambodia, with descriptions of two new species (Coleoptera, Carabidae, Licinini)</mods:title>
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<mods:namePart id="F7BE4B96DC6D781D0F6E3F343684B9A4">Allegro, Gianni</mods:namePart>
<mods:nameIdentifier id="9305E7B706A39FF1CB4F50B649136A7E" type="ORCID">https://orcid.org/0000-0002-5079-6671</mods:nameIdentifier>
<mods:affiliation id="56073BD00893D516AB3D7DD9F4CE1D1C">Strada Patro 11, I- 14036 Moncalvo (AT), Italy</mods:affiliation>
<mods:nameIdentifier id="95268F046CFE11351BEACB6C8CB5BED7" type="email">gianni.allegro54@gmail.com</mods:nameIdentifier>
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<mods:namePart id="A1B8F903A25977CE33FAFBF747DAA822">Giachino, Pier Mauro</mods:namePart>
<mods:nameIdentifier id="78FED299288C57D9D1E91115DE44500F" type="ORCID">https://orcid.org/0000-0002-1167-5447</mods:nameIdentifier>
<mods:affiliation id="1AF30CE670163493D1A9BEC124F6D01D">Via della Trinita 13, I- 10010 San Martino Canavese (TO), Italy</mods:affiliation>
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<mods:typeOfResource id="0D9203649552EB5FABB59E5301BB1A14">text</mods:typeOfResource>
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<mods:date id="CE9F7FDEF40FF2FF9CC69401AF952C3D">2021</mods:date>
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<mods:number id="C8AB1FB2F44A9120FBA6020055C04339">2021-06-16</mods:number>
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<mods:detail id="47DB5F4B418A3936AF7ACB6ABF8BDB5E" type="volume">
<mods:number id="AAFB8D2E301DA1457FBF7443DCD54DA6">1044</mods:number>
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<mods:classification id="E55915AF2FD03C32D3301217E132342E">journal article</mods:classification>
<mods:identifier id="F46B430C7372E5B4747AAE73ACC19BF7" type="DOI">http://dx.doi.org/10.3897/zookeys.1044.60072</mods:identifier>
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<treatment id="3C1B7F5006C75DE790EAECA39CF1CC62" ID-GBIF-Taxon="182754012" LSID="urn:lsid:plazi:treatment:3C1B7F5006C75DE790EAECA39CF1CC62" httpUri="http://treatment.plazi.org/id/3C1B7F5006C75DE790EAECA39CF1CC62" lastPageNumber="427" pageId="0" pageNumber="427" scope_family="Carabidae" scope_order="Coleoptera" scope_tribe="Licinini">
<subSubSection id="54D856ECC4D43D52F7C294A9459D913E" pageId="0" pageNumber="427" type="nomenclature">
<paragraph id="7B5F3BFCA047720BB7AE6A0A5C8B160E" pageId="0" pageNumber="427">
<taxonomicName id="FC4D5EADB1A3B6366A870A6300B53392" ID-CoL="36JS5" LSID="3C1B7F50-06C7-5DE7-90EA-ECA39CF1CC62" authority="(Lesne, 1896)" baseAuthorityName="Lesne" baseAuthorityYear="1896" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila (Submera) laevis" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="laevis" subGenus="Submera">Diplocheila (Submera) laevis (Lesne, 1896)</taxonomicName>
<figureCitation id="03DC87621D480704FB3CB294C04036E6" captionStart="Figures 20–25" captionStartId="F9" captionText="Figures 20 - 25. Aedeagus in lateral view of Diplocheila species 20 D. laevigata from Cambodia 21 Diplocheila sp. from Thailand 22 D. walterrossii sp. nov. HT 23 D. erwini sp. nov. HT 24 D. laevis from Cambodia 25 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures20-25" httpUri="https://binary.pensoft.net/fig/554100" pageId="0" pageNumber="427">Figures 24</figureCitation>
<figureCitation id="0E5917EBDA9BDE6272EA83963A05EBA0" captionStart="Figures 26–32" captionStartId="F10" captionText="Figures 26 - 32. Aedeagus in dorsal view of Diplocheila species 26 D. laevigata HT 27 D. laevigata from Cambodia 28 Diplocheila sp. from Thailand 29 D. walterrossii sp. nov. HT 30 D. erwini sp. nov. HT 31 D. laevis from Cambodia 32 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures26-32" httpUri="https://binary.pensoft.net/fig/554101" pageId="0" pageNumber="427">, 31</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="5367CB608BFFE5FF6DF3F22C8CCAC076" pageId="0" pageNumber="427" type="reference_group">
<paragraph id="05BFCE444C561ACA0057F8B25405998A" pageId="0" pageNumber="427">
<taxonomicName id="35245E6CF9AAC09DBBC9583D4E810B95" class="Insecta" family="Carabidae" genus="Rhembus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhembus laevis" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="laevis">Rhembus laevis</taxonomicName>
<bibRefCitation id="6358A550F34645200D09153F18F8EF80" author="Lesne, MP" journalOrPublisher="Entomological Review" pageId="0" pageNumber="427" refId="B20" refString="Lesne, MP, 1896. Cicindelides et Carabides Indo-Chinois Recueillis par M. Pavie. Diagnoses des Especes Nouvelles et d'un Genre Nouveau. Bulletin du Museum d'Histoire Naturelle, Paris, 238-245." title="Cicindelides et Carabides Indo-Chinois Recueillis par M. Pavie. Diagnoses des Especes Nouvelles et d'un Genre Nouveau. Bulletin du Museum d'Histoire Naturelle, Paris, 238 - 245." year="1896">Lesne 1896</bibRefCitation>
: 243
</paragraph>
<paragraph id="AA2542C4AA43B56684B3DEC7A1711DAF" pageId="0" pageNumber="427">
<taxonomicName id="9DC8F695E3D2D07E4A204695468929F7" class="Insecta" family="Carabidae" genus="Rhembus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhembus laevis" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="laevis">Rhembus laevis</taxonomicName>
Lesne, 1896:
<bibRefCitation id="C5F3829F79FDCFF8B9BCA8EF475C9035" author="Bouchard, J" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="427" pagination="169 - 176" refId="B11" refString="Bouchard, J, 1903. Insectes recueillis par M. le Professeur Dr. Forster a Borneo, Java et Sumatra (Palembang). Coleopteres, carabiques. Annales de la Societe Entomologique de France 72: 169 - 176" title="Insectes recueillis par M. le Professeur Dr. Forster a Borneo, Java et Sumatra (Palembang). Coleopteres, carabiques." volume="72" year="1903">Bouchard 1903</bibRefCitation>
: 171
</paragraph>
<paragraph id="4F3D2663F7E6224D3314FC22E8A23688" pageId="0" pageNumber="427">
<taxonomicName id="FF89ACAD50C261CB4167C39F64A4235F" class="Insecta" family="Carabidae" genus="Diplochila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplochila laevis" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="laevis">Diplochila laevis</taxonomicName>
(Lesne, 1896):
<bibRefCitation id="5B994F244D19E7A3B23B28D79FC44BAC" author="Lesne, MP" journalOrPublisher="Mission Pavie d'Indo Chine" pageId="0" pageNumber="427" pagination="62 - 75" refId="B21" refString="Lesne, MP, 1904. Famille des carabides. Mission Pavie d'Indo Chine 3: 62 - 75" title="Famille des carabides." volume="3" year="1904">Lesne 1904</bibRefCitation>
: 72
</paragraph>
<paragraph id="5ACF35AEED8A645BFFCCE89A466478AC" pageId="0" pageNumber="427">
<taxonomicName id="56DCDBDAD50D285FD4CF7CD511EF7B95" class="Insecta" family="Carabidae" genus="Diplochila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplochila laevis" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="laevis">Diplochila laevis</taxonomicName>
(Lesne, 1896):
<bibRefCitation id="07A2058957ABF78EC008EF689AABAFE6" DOI="https://doi.org/10.1080/00222932208632677" author="Andrewes, HE" pageId="0" pageNumber="427" refId="B2" refString="Andrewes, HE, 1922. Papers on Oriental Carabidae. VII. The Annals and Magazine of Natural History (Series 9) 52: 281-295. https://doi.org/10.1080/00222932208632677" title="Papers on Oriental Carabidae. VII. The Annals and Magazine of Natural History (Series 9) 52: 281 - 295." url="https://doi.org/10.1080/00222932208632677" year="1922">Andrewes 1922</bibRefCitation>
: 284
</paragraph>
<paragraph id="49D1DCD1DAC602970648AE1F78E1F9B1" pageId="0" pageNumber="427">
<taxonomicName id="9F8B9B38A9CD2F53F59BE0AA3491D750" baseAuthorityName="Lesne" baseAuthorityYear="1896" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila laevis" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="laevis">Diplocheila laevis</taxonomicName>
(Lesne, 1896):
<bibRefCitation id="D5605599C620E38F00C2259D9696D97F" author="Andrewes, HE" journalOrPublisher="Government of India Central Publication Branch, Calcutta" pageId="0" pageNumber="427" refId="B3" refString="Andrewes, HE, 1930. Catalogue of Indian insects. Part 18 - Carabidae. Government of India Central Publication Branch, Calcutta" title="Catalogue of Indian insects. Part 18 - Carabidae." year="1930">Andrewes 1930</bibRefCitation>
: 150
</paragraph>
<paragraph id="7CEA90CE87FCB948A6F8CC0231317C2E" pageId="0" pageNumber="427">
<taxonomicName id="B1E04CD1B2A84FC057FBC4638D636650" baseAuthorityName="Lesne" baseAuthorityYear="1896" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila laevis" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="laevis">Diplocheila laevis</taxonomicName>
(Lesne, 1896):
<bibRefCitation id="96A854512B9932482B058106CD99F979" author="Ball, GE" journalOrPublisher="Memoirs of the American Entomological Society" pageId="0" pageNumber="427" pagination="1 - 258" refId="B5" refString="Ball, GE, 1959. A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society 16: 1 - 258" title="A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera)." volume="16" year="1959">Ball 1959</bibRefCitation>
: 52
</paragraph>
<paragraph id="6CC35759ADA02F4262C45211B0DF0566" pageId="0" pageNumber="427">
<taxonomicName id="2E22736A1F7138E87321528CA3859DFC" baseAuthorityName="Lesne" baseAuthorityYear="1896" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila laevis" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="laevis">Diplocheila laevis</taxonomicName>
(Lesne, 1896):
<bibRefCitation id="A5EAFECD12062023411ED858B44507A7" author="Lorenz, W" journalOrPublisher="Published by the author, Tutzing" pageId="0" pageNumber="427" refId="B22" refString="Lorenz, W, 2005a. Systematic List of Extant Ground Beetles of the World (InsectaColeoptera Geadephaga: Trachypachidae and Carabidae Incl. Paussinae, Cicindelinae, Rhysodinae). Published by the author, Tutzing" title="Systematic List of Extant Ground Beetles of the World (InsectaColeoptera Geadephaga: Trachypachidae and Carabidae Incl. Paussinae, Cicindelinae, Rhysodinae)." year="2005 a">Lorenz 2005a</bibRefCitation>
: 343
</paragraph>
<paragraph id="597696B975174612231030093BCEFB67" pageId="0" pageNumber="427">
<taxonomicName id="DB1300C6F08F12BF50D7BACC8D9FB991" baseAuthorityName="Lesne" baseAuthorityYear="1896" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila laevis" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="laevis">Diplocheila laevis</taxonomicName>
(Lesne, 1896):
<bibRefCitation id="943ABA5DA5FA2AA36592CBF2578A4965" author="Lorenz, W" journalOrPublisher="Published by the author, Tutzing" pageId="0" pageNumber="427" refId="B23" refString="Lorenz, W, 2005b. Nomina Carabidarum. A Directory of the Scientific Names of Ground Beetles (InsectaColeoptera 'Geadephaga': Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). (2nd Edn.). Published by the author, Tutzing" title="Nomina Carabidarum. A Directory of the Scientific Names of Ground Beetles (InsectaColeoptera ' Geadephaga': Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). (2 nd Edn.)." year="2005 b">Lorenz 2005b</bibRefCitation>
: 571
</paragraph>
<paragraph id="2ECD9047277226EA677C73085B61E9A1" pageId="0" pageNumber="427">
<taxonomicName id="0A407F191D9776F3D3002100A4DF1C1B" baseAuthorityName="Lesne" baseAuthorityYear="1896" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila laevis" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="laevis">Diplocheila laevis</taxonomicName>
(Lesne, 1896):
<bibRefCitation id="51E2E556FF44CECC6FB3FD0AE35FF3BF" DOI="https://doi.org/10.1134/S0013873808060079" author="Lafer, GS" journalOrPublisher="Entomological Review" pageId="0" pageNumber="427" pagination="679 - 695" refId="B19" refString="Lafer, GS, Kataev, BM, 2008. On the species of the genus Diplocheila Brulle (Coleoptera: Carabidae) of the Far East of Russia, with a brief review of the East Asian species. Entomological Review 88 (6): 679 - 695, DOI: https://doi.org/10.1134/S0013873808060079" title="On the species of the genus Diplocheila Brulle (Coleoptera: Carabidae) of the Far East of Russia, with a brief review of the East Asian species." url="https://doi.org/10.1134/S0013873808060079" volume="88" year="2008">Lafer and Kataev 2008</bibRefCitation>
: 690
</paragraph>
<paragraph id="A50DB942C6598174C4DFCFEACBE25131" pageId="0" pageNumber="427">
<taxonomicName id="6B3831D67DC2F7C80FD256D851FE9FC7" baseAuthorityName="Lesne" baseAuthorityYear="1896" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila laevis" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="laevis">Diplocheila laevis</taxonomicName>
(Lesne, 1896):
<bibRefCitation id="CE58FD6E3FD728E7F0AF937678331727" author="Huber, C" editor="Loebl, I" journalOrPublisher="Archostemata - Myxophaga - Adephaga (Vol. 1). Revised and Updated Edition. Koninklijke Brill NV, Leiden" pageId="0" pageNumber="427" pagination="621 - 577" refId="B16" refString="Huber, C, Marggi, W, 2017. Licinini. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera. Archostemata - Myxophaga - Adephaga (Vol. 1). Revised and Updated Edition. Koninklijke Brill NV, Leiden: 621 - 577" title="Licinini." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2017">Huber and Marggi 2017</bibRefCitation>
: 626
</paragraph>
</subSubSection>
<subSubSection id="9C817F27CBE18CE23B3E86D63173796C" pageId="0" pageNumber="427" type="type locality">
<paragraph id="35148B1CDA25AF7387B00058B0CC1542" pageId="0" pageNumber="427">Type locality.</paragraph>
<paragraph id="76D705E156449892CE2D253B5B857A47" pageId="0" pageNumber="427">
Bangkok (Le P. Larnaudie); Chantaboun
<normalizedToken id="C6EA4D071BE7154DFA162D7767FD245F" originalValue="à">a</normalizedToken>
Battambang (Siam Cambodgien); Meuwen Bay (Java).
</paragraph>
</subSubSection>
<subSubSection id="39A9278AC59D508A5C75BE7EDEE4B53C" pageId="0" pageNumber="427" type="materials_examined">
<paragraph id="22B2D9BE629BDEE393E61F45D96F3257" pageId="0" pageNumber="427">Material examined.</paragraph>
<paragraph id="5010E5CEB6273EE72AAD615C5E1B0C5A" pageId="0" pageNumber="427">
<materialsCitation id="73E96F12622308C1DED6CC3B8E61A595" collectingDate="2019-05-17" collectorName="W. Rossi, V. Kong" country="Cambodia" location="Tonle Sap Lake" specimenCount="1" specimenCount-male="1" stateProvince="Kampong Chhnang">
<collectingCountry id="E1E8A8DF64E7022C7251FF8B90EE8044" name="Cambodia">Cambodia</collectingCountry>
:
<specimenCount id="9926D763A1835CA07E9E4441E3A66574" type="male">1 ♂</specimenCount>
<collectingRegion id="6F7B4A8D0728058644A4905D12744DD8" country="Cambodia" name="Kampong Chhnang">Kampong Chhnang</collectingRegion>
, banks of
<location id="FCB8F51D884ABE3CEBE62D8CE366227C" LSID="urn:lsid:plazi:treatment:3C1B7F5006C75DE790EAECA39CF1CC62:FCB8F51D884ABE3CEBE62D8CE366227C" country="Cambodia" name="Tonle Sap Lake" stateProvince="Kampong Chhnang">Tonle Sap Lake</location>
,
<collectingDate id="DC41F8A34DC2F39FD854C95F4E49A4AE" value="2019-05-17">17.V.2019</collectingDate>
,
<collectorName id="5261955DB3D8B7202CDD4498A9DC125A">W. Rossi</collectorName>
and
<collectorName id="46FC8184AE4F6198DB2852785092C143">V. Kong</collectorName>
leg. (CAl)
</materialsCitation>
;
<materialsCitation id="0D1BAA569196D40FE09EB0EDCF0877D6" collectingDate="2018-11-04" collectorName="W. Rossi, V. Kong" country="Cambodia" county="Sankat" latitude="12.233334" location="Tonle Sap Lake" longLatPrecision="1293" longitude="104.683334" municipality="Phum Toul Ompel" specimenCount="1" specimenCount-male="1" stateProvince="Kampong Chhnang Province">
<specimenCount id="46E0B13C0CB5D78C0542A45D0E346AD8" type="male">1 ♂</specimenCount>
<collectingRegion id="3485517E1A27861E4ECDD6085CE0DA18" country="Cambodia" name="Kampong Chhnang">Kampong Chhnang Province</collectingRegion>
,
<collectingCounty id="B7D951D769B46493C07AD997C4F25C3A">Sankat</collectingCounty>
<collectingRegion id="33194AF9796D3DD5A1158B5F3D06CAF4" country="Cambodia" name="Kampong Chhnang">Kampong Chhnang</collectingRegion>
,
<collectingMunicipality id="E99AFE47C30C5F32BD13A33E868E3E3B">Phum Toul Ompel</collectingMunicipality>
, banks of a branch of
<location id="0C54ED2927E917D602F6F0DABBB725FE" LSID="urn:lsid:plazi:treatment:3C1B7F5006C75DE790EAECA39CF1CC62:0C54ED2927E917D602F6F0DABBB725FE" country="Cambodia" county="Sankat" latitude="12.233334" longLatPrecision="1293" longitude="104.683334" municipality="Phum Toul Ompel" name="Tonle Sap Lake" stateProvince="Kampong Chhnang Province">Tonle Sap Lake</location>
,
<geoCoordinate id="FED86E89712362134269FA38B4703262" degrees="12" direction="north" minutes="14" orientation="latitude" precision="925" value="12.233334">12°14'N</geoCoordinate>
,
<geoCoordinate id="D7D1B8C5212B6288CD48FE57F902C2D9" degrees="104" direction="east" minutes="41" orientation="longitude" precision="925" value="104.683334">104°41'E</geoCoordinate>
,
<collectingDate id="E9A80D25D5E922CD9DAF6EB2B9B8AC69" value="2018-11-04">4.XI.2018</collectingDate>
,
<collectorName id="04F37A06A3F5CE73126339ABEF94676B">W. Rossi</collectorName>
and
<collectorName id="2F86E7D46CA5C172D141EC437357BACF">V. Kong</collectorName>
leg. (CGi)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="6C4C7933CDE4D2603CE60EC7B243F412" pageId="0" pageNumber="427" type="remarks">
<paragraph id="65A60846B532BF97E80242565D2FEC95" pageId="0" pageNumber="427">Remarks.</paragraph>
<paragraph id="17E8F7B31C10420D277738924B9C4111" pageId="0" pageNumber="427">
<taxonomicName id="159379CDD09F6DDD4FC934BA080D0CFB" baseAuthorityName="Lesne" baseAuthorityYear="1896" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila laevis" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="laevis">
<emphasis id="4179D6E753ECD77C195F49D906AD6A3B" italics="true" pageId="0" pageNumber="427">Diplocheila laevis</emphasis>
</taxonomicName>
is widely distributed across South-East Asia, China, Indonesia, and the Philippines (
<bibRefCitation id="4099E37D9EF5DFFB24369D492CE9DAC8" DOI="https://doi.org/10.1080/00222932208632677" author="Andrewes, HE" pageId="0" pageNumber="427" refId="B2" refString="Andrewes, HE, 1922. Papers on Oriental Carabidae. VII. The Annals and Magazine of Natural History (Series 9) 52: 281-295. https://doi.org/10.1080/00222932208632677" title="Papers on Oriental Carabidae. VII. The Annals and Magazine of Natural History (Series 9) 52: 281 - 295." url="https://doi.org/10.1080/00222932208632677" year="1922">Andrewes 1922</bibRefCitation>
;
<bibRefCitation id="B019EB9BFF8F538C0F56C01D40A10640" author="Ball, GE" journalOrPublisher="Memoirs of the American Entomological Society" pageId="0" pageNumber="427" pagination="1 - 258" refId="B5" refString="Ball, GE, 1959. A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society 16: 1 - 258" title="A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera)." volume="16" year="1959">Ball 1959</bibRefCitation>
;
<bibRefCitation id="C95162911820D665CFE223E7F10AFF78" DOI="https://doi.org/10.1134/S0013873808060079" author="Lafer, GS" journalOrPublisher="Entomological Review" pageId="0" pageNumber="427" pagination="679 - 695" refId="B19" refString="Lafer, GS, Kataev, BM, 2008. On the species of the genus Diplocheila Brulle (Coleoptera: Carabidae) of the Far East of Russia, with a brief review of the East Asian species. Entomological Review 88 (6): 679 - 695, DOI: https://doi.org/10.1134/S0013873808060079" title="On the species of the genus Diplocheila Brulle (Coleoptera: Carabidae) of the Far East of Russia, with a brief review of the East Asian species." url="https://doi.org/10.1134/S0013873808060079" volume="88" year="2008">Lafer and Kataev 2008</bibRefCitation>
). The aedeagus is also illustrated by
<bibRefCitation id="8A7255E364EF2C28C9FFB9F6F22CC125" DOI="https://doi.org/10.1134/S0013873808060079" author="Lafer, GS" journalOrPublisher="Entomological Review" pageId="0" pageNumber="427" pagination="679 - 695" refId="B19" refString="Lafer, GS, Kataev, BM, 2008. On the species of the genus Diplocheila Brulle (Coleoptera: Carabidae) of the Far East of Russia, with a brief review of the East Asian species. Entomological Review 88 (6): 679 - 695, DOI: https://doi.org/10.1134/S0013873808060079" title="On the species of the genus Diplocheila Brulle (Coleoptera: Carabidae) of the Far East of Russia, with a brief review of the East Asian species." url="https://doi.org/10.1134/S0013873808060079" volume="88" year="2008">Lafer and Kataev (2008)</bibRefCitation>
. As far as we know, this is the first record of the species from Cambodia.
</paragraph>
</subSubSection>
</treatment>
</document>
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