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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.174.2668" ID-GBIF-Dataset="4bf0c20f-f2bd-4473-ae8b-75da30f9292f" ID-PMC="PMC3307351" ID-Pensoft-Pub="1313-2970-174-7" ID-PubMed="22451781" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-174-7" ModsDocOrigin="ZooKeys 174" ModsDocTitle="The Ochodaeidae of Argentina (Coleoptera, Scarabaeoidea)" checkinTime="1451249306841" checkinUser="pensoft" docAuthor="Paulsen, M. J. &amp; Ocampo, Federico C." docDate="2012" docId="2EC23AEDA1E5655181306F027A2362B4" docLanguage="en" docName="ZooKeys 174: 7-30" docOrigin="ZooKeys 174" docSource="http://dx.doi.org/10.3897/zookeys.174.2668" docTitle="Parochodaeus stupendus Paulsen &amp; Ocampo, sp. n." docType="treatment" docVersion="4" lastPageNumber="22" masterDocId="FFD2D2245A236B244A673E72FFB87A71" masterDocTitle="The Ochodaeidae of Argentina (Coleoptera, Scarabaeoidea)" masterLastPageNumber="30" masterPageNumber="7" pageNumber="22" updateTime="1668153319038" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The Ochodaeidae of Argentina (Coleoptera, Scarabaeoidea)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Paulsen, M. J.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ocampo, Federico C.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="volume">
<mods:number>174</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>7</mods:start>
<mods:end>30</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.174.2668</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.174.2668</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-174-7</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152033515" LSID="urn:lsid:zoobank.org:act:523C896D-1CFC-4281-824D-F2C2BAF3DD7D" httpUri="http://treatment.plazi.org/id/2EC23AEDA1E5655181306F027A2362B4" lastPageId="16" lastPageNumber="22" pageId="15" pageNumber="22">
<subSubSection pageId="15" pageNumber="22" type="nomenclature">
<paragraph pageId="15" pageNumber="22">
<taxonomicName LSID="urn:lsid:zoobank.org:act:523C896D-1CFC-4281-824D-F2C2BAF3DD7D" authority="Paulsen &amp; Ocampo" class="Insecta" family="Ochodaeidae" genus="Parochodaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parochodaeus stupendus" order="Coleoptera" pageId="15" pageNumber="22" phylum="Arthropoda" rank="species" species="stupendus">Parochodaeus stupendus Paulsen &amp; Ocampo</taxonomicName>
<taxonomicNameLabel pageId="15" pageNumber="22">sp. n.</taxonomicNameLabel>
Figs 716Map 3
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="22" type="type material">
<paragraph pageId="15" pageNumber="22">Type material.</paragraph>
<paragraph pageId="15" pageNumber="22">
Holotype male (CMNC), pinned. Original labels: a) &quot;ARGENTINA / SALTA / D° San
<normalizedToken originalValue="Martín">Martin</normalizedToken>
/ Hito 1 / Coll.
<normalizedToken originalValue="Martínez">Martinez</normalizedToken>
/ Dic 971&quot;; b) red label &quot;Parochodaeus / stupendus /Paulsen &amp; Ocampo / HOLOTYPE&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="22" type="type locality">
<paragraph pageId="15" pageNumber="22">Type locality.</paragraph>
<paragraph pageId="15" pageNumber="22">
Argentina: Salta: San
<normalizedToken originalValue="Martín">Martin</normalizedToken>
, Hito 1.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="22" type="diagnosis">
<paragraph pageId="15" pageNumber="22">Diagnosis.</paragraph>
<paragraph pageId="15" pageNumber="22">
<taxonomicName class="Insecta" family="Ochodaeidae" genus="Parochodaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parochodaeus stupendus" order="Coleoptera" pageId="15" pageNumber="22" phylum="Arthropoda" rank="species" species="stupendus">Parochodaeus stupendus</taxonomicName>
(Fig. 17) is the only species of
<taxonomicName class="Insecta" family="Ochodaeidae" genus="Parochodaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parochodaeus" order="Coleoptera" pageId="15" pageNumber="22" phylum="Arthropoda" rank="genus">Parochodaeus</taxonomicName>
to exhibit a doubly-toothed posterior margin on the metafemur. The tooth on the ventral surface is fairly common in ochodaeids, but the second tooth on the dorsal surface is unique. The toothed metatrochanter and strongly curved metatibiae (Fig. 7) are not found in any other species in
<taxonomicName class="Actinopterygii" family="Argentinidae" genus="Argentina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Argentina" order="Osmeriformes" pageId="15" pageNumber="22" phylum="Chordata" rank="genus">Argentina</taxonomicName>
, or elsewhere. The strongly produced
<normalizedToken originalValue="thumb">'thumb'</normalizedToken>
(internal apical tooth) is also species specific. The female is unknown.
</paragraph>
<caption pageId="15" pageNumber="22">
<paragraph pageId="15" pageNumber="22">
Figure 17. Dorsal habitus of
<taxonomicName class="Insecta" family="Ochodaeidae" genus="Gauchodaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gauchodaeus patagonicus" order="Coleoptera" pageId="15" pageNumber="22" phylum="Arthropoda" rank="species" species="patagonicus">Gauchodaeus patagonicus</taxonomicName>
gen. n., sp. n.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="15" pageNumber="22" type="description">
<paragraph pageId="15" pageNumber="22">Description, Holotype.</paragraph>
<paragraph pageId="15" pageNumber="22">Length:7.5 mm. Width:4.1 mm. Head: Surface tuberculate, tubercles moderately large, setose. Frons unarmed but indistinctly tumid medially. Clypeus evenly rounded, long (length equal to 1/2 width), margin thickened, produced anteriorly, declivous. Labrum emarginate. Mandibles rounded externally. Mentum tumid, not deeply furrowed. Antennal club strongly globose (Fig. 12). Pronotum: Form convex. Surface with moderately large, setose tubercles mixed with glabrous punctures; punctures moderate in size, sparse. Elytra: Setae of interstrial tubercles moderately long, erect, somewhat abraded. Legs: Protibia with apical spur strongly curved externally; internal apical tooth distinctly produced, subequal to basal protibial tooth in size. Metatrochanter with acute tooth. Metafemur with posterior margin acutely toothed on both dorsal and ventral surfaces. Metatibia strongly curved, slender, abruptly widening in distal third. Metatarsomere 1 not greatly enlarged. Abdomen:Stridulatory peg absent.</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="22" type="etymology">
<paragraph pageId="15" pageNumber="22">Etymology.</paragraph>
<paragraph pageId="15" pageNumber="22">
The specific epithet
<normalizedToken originalValue="stupendus">'stupendus'</normalizedToken>
is Latin meaning &quot;causing astonishment or wonder&quot;. With several autapomorphies not seen in other species of the genus,
<taxonomicName class="Insecta" family="Ochodaeidae" genus="Parochodaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parochodaeus stupendus" order="Coleoptera" pageId="15" pageNumber="22" phylum="Arthropoda" rank="species" species="stupendus">Parochodaeus stupendus</taxonomicName>
is arguably the most wondrous ochodaeid in the New World. The name is used a masculine adjective in the nominative singular.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="22" type="distribution">
<paragraph pageId="15" pageNumber="22">Distribution</paragraph>
<paragraph pageId="15" pageNumber="22">(Map 3).The species is known from a locality near the Bolivian border. 1 specimen examined.</paragraph>
<paragraph pageId="15" pageNumber="22">
ARGENTINA: SALTA: San
<normalizedToken originalValue="Martín">Martin</normalizedToken>
, Hito 1 (1).
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="22" type="temporal distribution">
<paragraph pageId="15" pageNumber="22">Temporal distribution</paragraph>
<paragraph pageId="15" pageNumber="22">.December (1).</paragraph>
</subSubSection>
<subSubSection lastPageId="16" lastPageNumber="23" pageId="15" pageNumber="22" type="remarks">
<paragraph pageId="15" pageNumber="22">Remarks.</paragraph>
<paragraph lastPageId="16" lastPageNumber="23" pageId="15" pageNumber="22">
This species is known only from the holotype. Like
<taxonomicName class="Insecta" family="Ochodaeidae" genus="Parochodaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parochodaeus jujuyus" order="Coleoptera" pageId="15" pageNumber="22" phylum="Arthropoda" rank="species" species="jujuyus">Parochodaeus jujuyus</taxonomicName>
, it is from the far north of Argentina. It is probable that
<taxonomicName class="Insecta" family="Ochodaeidae" genus="Parochodaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parochodaeus stupendus" order="Coleoptera" pageId="15" pageNumber="22" phylum="Arthropoda" rank="species" species="stupendus">Parochodaeus stupendus</taxonomicName>
will be discovered in Bolivia
<pageBreakToken pageId="16" pageNumber="23" start="start">as</pageBreakToken>
well, especially because the name of the locality &quot;Hito 1&quot; is in reference to the
<normalizedToken originalValue="“hito”">&quot;hito&quot;</normalizedToken>
or boundary post that indicates the border between Argentina and Bolivia.
</paragraph>
</subSubSection>
</treatment>
</document>