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<document id="E9347B3E2EA4339DA2EC5785165A420A" ID-CLB-Dataset="21058" ID-DOI="10.1080/00222933.2014.974703" ID-GBIF-Dataset="88042df7-dae6-4b66-93eb-9fcbe4706bdd" ID-ISSN="1464-5262" ID-Zenodo-Dep="4006314" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1598626814697" checkinUser="carolina" docAuthor="Disney, R. H. L. &amp; Russell-Smith, A." docDate="2014" docId="1B4687EDFFB1FFF544FE449DFB0AFAA2" docLanguage="en" docName="JNATHIST.49.25-26.1599-1626.pdf" docOrigin="Journal of Natural History 49 (25)" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.6:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="6" docTitle="Megaselia crassipes" docType="treatment" docVersion="7" lastPageNumber="1602" masterDocId="E77FFF95FFB0FFF1452C4059FF99FF9E" masterDocTitle="Additions to the British list of Megaselia Rondani (Diptera: Phoridae), including two new species, from the crowns of ancient pollarded trees" masterLastPageNumber="1626" masterPageNumber="1599" pageNumber="1600" updateTime="1698845724323" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="4725BE1C86EC04E545748905C3CEAB3D">Additions to the British list of Megaselia Rondani (Diptera: Phoridae), including two new species, from the crowns of ancient pollarded trees</mods:title>
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<mods:namePart id="2D5C05FCA803583463C11A5E190E49C5">Disney, R. H. L.</mods:namePart>
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<treatment id="1B4687EDFFB1FFF544FE449DFB0AFAA2" ID-DOI="http://doi.org/10.5281/zenodo.4329360" ID-GBIF-Taxon="167371454" ID-Zenodo-Dep="4329360" LSID="urn:lsid:plazi:treatment:1B4687EDFFB1FFF544FE449DFB0AFAA2" httpUri="http://treatment.plazi.org/id/1B4687EDFFB1FFF544FE449DFB0AFAA2" lastPageId="4" lastPageNumber="1602" pageId="1" pageNumber="1600">
<subSubSection id="DBF56570FFB1FFF044FE449DFC94FB43" box="[466,781,1220,1245]" pageId="1" pageNumber="1600" type="nomenclature">
<paragraph id="935036FBFFB1FFF044FE449DFC94FB43" blockId="1.[466,781,1220,1277]" box="[466,781,1220,1245]" pageId="1" pageNumber="1600">
<heading id="C8188197FFB1FFF044FE449DFC94FB43" box="[466,781,1220,1245]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="1600" reason="2">
<taxonomicName id="54EF4D78FFB1FFF044FE449DFC94FB43" ID-CoL="8CQPT" authority="(Wood)" baseAuthorityName="Wood" baseAuthorityYear="1909" box="[466,781,1220,1245]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="1600" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="A19BEAE9FFB1FFF044FE449DFD37FB42" bold="true" box="[466,686,1220,1244]" italics="true" pageId="1" pageNumber="1600">Megaselia crassipes</emphasis>
(Wood)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DBF56570FFB1FFF0470944BCFD22FB63" box="[549,699,1253,1277]" pageId="1" pageNumber="1600" type="description">
<paragraph id="935036FBFFB1FFF0470944BCFD22FB63" blockId="1.[466,781,1220,1277]" box="[549,699,1253,1277]" pageId="1" pageNumber="1600">
(
<figureCitation id="0BD42A7EFFB1FFF0470144BCFD2BFB63" box="[557,690,1253,1277]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="2.[212,279,1298,1320]" captionStartId-1="3.[144,211,1067,1089]" captionStartId-2="4.[212,279,1198,1220]" captionTargetBox-0="[213,1171,169,1265]" captionTargetBox-1="[145,1103,169,1034]" captionTargetBox-2="[212,1172,168,1166]" captionTargetId-0="figure@2.[212,1172,168,1266]" captionTargetId-1="figure@3.[144,1104,168,1035]" captionTargetId-2="figure@4.[212,1172,168,1166]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Figure 1. Megaselia crassipes male. (A) Posterior face of front tarsus; (B) left face of hypopytgium. Scale bar: 0.1 mm." captionText-1="Figure 2. Megaselia crassipes male (synonym of M. basiturgida). (A) Left face of hypopygium; (B) posterior face of front tibia and basitarsus. Scale bar: 0.1 mm." captionText-2="Figure 3. Megaselia crassipes male (synonym of M. gargarans), left face of anal tube." figureDoi-0="http://doi.org/10.5281/zenodo.4006316" figureDoi-1="http://doi.org/10.5281/zenodo.4006318" figureDoi-2="http://doi.org/10.5281/zenodo.4006322" httpUri-0="https://zenodo.org/record/4006316/files/figure.png" httpUri-1="https://zenodo.org/record/4006318/files/figure.png" httpUri-2="https://zenodo.org/record/4006322/files/figure.png" pageId="1" pageNumber="1600">Figures 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DBF56570FFB1FFF045BC457CFD8AFAA2" box="[144,531,1316,1341]" pageId="1" pageNumber="1600" type="nomenclature">
<paragraph id="935036FBFFB1FFF045BC457CFD8AFAA2" blockId="1.[144,1104,1316,1691]" box="[144,531,1316,1341]" pageId="1" pageNumber="1600">
<heading id="C8188197FFB1FFF045BC457CFD8AFAA2" box="[144,531,1316,1341]" fontSize="10" level="3" pageId="1" pageNumber="1600" reason="5">
<taxonomicName id="54EF4D78FFB1FFF045BC457CFD8AFAA2" ID-CoL="88XMD" authority="Wood 1909, p. 24" authorityName="Wood" authorityPageNumber="24" authorityYear="1909" box="[144,531,1316,1341]" class="Insecta" family="Phoridae" genus="Phora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="1600" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="A19BEAE9FFB1FFF045BC457CFEA4FAA2" box="[144,317,1317,1341]" italics="true" pageId="1" pageNumber="1600">Phora crassipes</emphasis>
<bibRefCitation id="F77E4B0AFFB1FFF0446A457DFE50FAA2" author="Wood JH" box="[326,457,1316,1340]" pageId="1" pageNumber="1600" pagination="240 - 244" refId="ref8804" refString="Wood JH. 1909. On the British species of Phora. Part II contd. Entomol Mon Mag. 45: 24 - 29, 59 - 63, 113 - 120, 143 - 149, 191 - 195, 240 - 244." type="journal article" year="1909">Wood 1909</bibRefCitation>
, p. 24
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DBF56570FFB1FFF045BC451CFE50F982" pageId="1" pageNumber="1600" type="reference_group">
<paragraph id="935036FBFFB1FFF045BC451CFC1BFAC2" blockId="1.[144,1104,1316,1691]" box="[144,898,1348,1373]" pageId="1" pageNumber="1600">
<taxonomicName id="54EF4D78FFB1FFF045BC451CFDE6FAC2" ID-CoL="87SRD" authority="Lundbeck 1920, p. 9" authorityName="Lundbeck" authorityPageNumber="9" authorityYear="1920" box="[144,639,1348,1373]" class="Insecta" family="Phoridae" genus="Aphiochaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="1600" phylum="Arthropoda" rank="species" species="dactylata">
<emphasis id="A19BEAE9FFB1FFF045BC451CFE10FAC3" box="[144,393,1349,1373]" italics="true" pageId="1" pageNumber="1600">Aphiochaeta dactylata</emphasis>
<bibRefCitation id="F77E4B0AFFB1FFF044BE451DFDD8FAC2" author="Lundbeck W" box="[402,577,1348,1372]" pageId="1" pageNumber="1600" pagination="1 - 34" refId="ref8425" refString="Lundbeck W. 1920. New species of Aphiochaeta from Denmark. Videnskabelige Meddelelser fra Dansk naturhistorisk Forening i Kjobenhavn. 71: 1 - 34." type="journal article" year="1920">Lundbeck 1920</bibRefCitation>
, p. 9
</taxonomicName>
.
<bibRefCitation id="F77E4B0AFFB1FFF047A1451DFCB2FAC2" author="Schmitz H" box="[653,811,1348,1372]" pageId="1" pageNumber="1600" pagination="350 - 379" refId="ref8553" refString="Schmitz H. 1952. Uber W. Lundbecks Sammlung und Beschreibung Danischer Phoriden. Entomologiske Meddelelser. 26: 350 - 379." type="journal article" year="1952">Schmitz, 1952</bibRefCitation>
, p. 359
</paragraph>
<paragraph id="935036FBFFB1FFF045BC453DFC11FAE2" blockId="1.[144,1104,1316,1691]" box="[144,904,1380,1404]" pageId="1" pageNumber="1600">
<taxonomicName id="54EF4D78FFB1FFF045BC453DFD1CFAE2" ID-CoL="87SRJ" authority="Lundbeck 1921, p. 134" authorityName="Lundbeck" authorityPageNumber="134" authorityYear="1921" box="[144,645,1380,1404]" class="Insecta" family="Phoridae" genus="Aphiochaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="1600" phylum="Arthropoda" rank="species" species="exclusa">
<emphasis id="A19BEAE9FFB1FFF045BC453DFEEDFAE2" box="[144,372,1380,1404]" italics="true" pageId="1" pageNumber="1600">Aphiochaeta exclusa</emphasis>
<bibRefCitation id="F77E4B0AFFB1FFF04451453DFDB5FAE2" author="Lundbeck W" box="[381,556,1380,1404]" pageId="1" pageNumber="1600" pagination="129 - 143" refId="ref8452" refString="Lundbeck W. 1921. New species of Phoridae from Denmark, together with remarks on Aphiochaeta groenlandica Lundb. Videnskabelige Meddelelser fra Dansk naturhistorisk Forening i Kjobenhavn. 72: 129 - 143." type="journal article" year="1921">Lundbeck 1921</bibRefCitation>
, p. 134
</taxonomicName>
.
<bibRefCitation id="F77E4B0AFFB1FFF047BE453DFCA8FAE2" author="Schmitz H" box="[658,817,1380,1404]" pageId="1" pageNumber="1600" pagination="350 - 379" refId="ref8553" refString="Schmitz H. 1952. Uber W. Lundbecks Sammlung und Beschreibung Danischer Phoriden. Entomologiske Meddelelser. 26: 350 - 379." type="journal article" year="1952">Schmitz, 1952</bibRefCitation>
, p. 361
</paragraph>
<paragraph id="935036FBFFB1FFF045BC45DDFD53FA02" blockId="1.[144,1104,1316,1691]" box="[144,714,1412,1437]" pageId="1" pageNumber="1600">
<taxonomicName id="54EF4D78FFB1FFF045BC45DDFDEBFA02" ID-CoL="8CQT4" authority="Schmitz 1948, p. 393" authorityName="Schmitz" authorityPageNumber="393" authorityYear="1948" box="[144,626,1412,1436]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="1600" phylum="Arthropoda" rank="species" species="gargarans">
<emphasis id="A19BEAE9FFB1FFF045BC45DDFEE1FA02" box="[144,376,1412,1436]" italics="true" pageId="1" pageNumber="1600">Megaselia gargarans</emphasis>
<bibRefCitation id="F77E4B0AFFB1FFF044AD45DDFD80FA02" author="Schmitz H" box="[385,537,1412,1436]" pageId="1" pageNumber="1600" pagination="375 - 399" refId="ref8524" refString="Schmitz H. 1948. Zweiter Beitrag zur Kenntnis der Phoriden Osterreichs (Diptera). Annalen des Naturhistorischen Museums, Wien. 56: 375 - 399." type="journal article" year="1948">Schmitz 1948</bibRefCitation>
, p. 393
</taxonomicName>
,
<taxonomicNameLabel id="BAA85792FFB1FFF0475345DCFD53FA02" box="[639,714,1413,1437]" pageId="1" pageNumber="1600">syn. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="935036FBFFB1FFF045BC45FDFCC2FA22" blockId="1.[144,1104,1316,1691]" box="[144,859,1444,1469]" pageId="1" pageNumber="1600">
<taxonomicName id="54EF4D78FFB1FFF045BC45FDFC9DFA22" ID-CoL="9HV6Y" authority="Disney and Durska 2011, p. 528" authorityName="Disney and Durska" authorityPageNumber="528" authorityYear="2011" box="[144,772,1444,1468]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="1600" phylum="Arthropoda" rank="species" species="basiturgida">
<emphasis id="A19BEAE9FFB1FFF045BC45FDFE1CFA22" box="[144,389,1444,1468]" italics="true" pageId="1" pageNumber="1600">Megaselia basiturgida</emphasis>
<bibRefCitation id="F77E4B0AFFB1FFF044A245FDFD34FA22" author="Disney RHL &amp; Durska E" box="[398,685,1444,1468]" pageId="1" pageNumber="1600" pagination="527 - 534" refId="ref8254" refString="Disney RHL, Durska E. 2011. Five new species and new records of Megaselia Rondani (Diptera: Phoridae) from Poland. Annales Zoologici (Warszawa). 61: 527 - 534." type="journal article" year="2011">Disney and Durska 2011</bibRefCitation>
, p. 528
</taxonomicName>
,
<taxonomicNameLabel id="BAA85792FFB1FFF0463C45FCFCC2FA22" box="[784,859,1445,1469]" pageId="1" pageNumber="1600">syn. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="935036FBFFB1FFF045BC459DFE50F982" blockId="1.[144,1104,1316,1691]" pageId="1" pageNumber="1600">
<taxonomicName id="54EF4D78FFB1FFF045BC459DFEF4FA42" ID-CoL="8CQPT" baseAuthorityName="Wood" baseAuthorityYear="1909" box="[144,365,1476,1500]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="1600" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="A19BEAE9FFB1FFF045BC459DFEF4FA42" box="[144,365,1476,1500]" italics="true" pageId="1" pageNumber="1600">Megaselia crassipes</emphasis>
</taxonomicName>
was not obtained in the present study but it needs to be distinguished from a new species (described below) that was obtained, and from the addition to the British List.
</paragraph>
</subSubSection>
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<paragraph id="935036FBFFB1FFF34594467AFDB3F904" blockId="1.[144,1104,1316,1691]" lastBlockId="2.[212,1172,1411,1690]" lastPageId="2" lastPageNumber="1601" pageId="1" pageNumber="1600">
The recognition of
<taxonomicName id="54EF4D78FFB1FFF0448F467CFDA8F9A5" baseAuthorityName="Wood" baseAuthorityYear="1909" box="[419,561,1572,1596]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="1600" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="A19BEAE9FFB1FFF0448F467CFDA8F9A5" box="[419,561,1572,1596]" italics="true" pageId="1" pageNumber="1600">M. crassipes</emphasis>
</taxonomicName>
has proved troublesome from the beginning.
<bibRefCitation id="F77E4B0AFFB1FFF045BC461AFED5F9C5" author="Lundbeck W" box="[144,332,1603,1627]" pageId="1" pageNumber="1600" refId="ref8487" refString="Lundbeck W. 1922. Diptera Danica Part VI Pipunculidae, Phoridae. London: Wesley." type="book" year="1922">Lundbeck (1922)</bibRefCitation>
only included Wood s species in his key to the species of his Group IV. However, he included his
<emphasis id="A19BEAE9FFB1FFF044FB463DFC88F9E2" box="[471,785,1636,1660]" italics="true" pageId="1" pageNumber="1600">
<taxonomicName id="54EF4D78FFB1FFF044FB463DFDFAF9E2" authorityName="Lundbeck" authorityYear="1920" box="[471,611,1636,1660]" class="Insecta" family="Phoridae" genus="Aphiochaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="1600" phylum="Arthropoda" rank="species" species="dactylata">M. dactylata</taxonomicName>
and
<taxonomicName id="54EF4D78FFB1FFF047B5463DFC88F9E2" authorityName="Lundbeck" authorityYear="1921" box="[665,785,1636,1660]" class="Insecta" family="Phoridae" genus="Aphiochaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="1600" phylum="Arthropoda" rank="species" species="exclusa">M. exclusa</taxonomicName>
</emphasis>
in both the key and the text. Subsequently,
<bibRefCitation id="F77E4B0AFFB1FFF0441E46DAFE40F905" author="Schmitz H" box="[306,473,1667,1691]" pageId="1" pageNumber="1600" pagination="350 - 379" refId="ref8553" refString="Schmitz H. 1952. Uber W. Lundbecks Sammlung und Beschreibung Danischer Phoriden. Entomologiske Meddelelser. 26: 350 - 379." type="journal article" year="1952">Schmitz (1952)</bibRefCitation>
synonymised both these species with
<emphasis id="A19BEAE9FFB1FFF046A546DDFB82F905" box="[905,1051,1668,1691]" italics="true" pageId="1" pageNumber="1600">
<taxonomicName id="54EF4D78FFB1FFF046A546DDFB8EF905" baseAuthorityName="Wood" baseAuthorityYear="1909" box="[905,1047,1668,1691]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="1600" phylum="Arthropoda" rank="species" species="crassipes">M. crassipes</taxonomicName>
.
</emphasis>
The effect of this was to play havoc with Lundbeck s key to his Group IV. For example, his first couplet separates those species with long costal cilia from those in which they are short. It is evident, however, that these cilia vary across accepted conventions regarding the boundary between short and long. The failure to appreciate this caused
<bibRefCitation id="F77E4B0AFFB2FFF3411045BBFE81F984" author="Schmitz H" pageId="2" pageNumber="1601" pagination="465 - 512" refId="ref8577" refString="Schmitz H. 1958. Phoridae. In: Lindner E, editor. Die Fliegen der palaearktischen Region 4 (33) (Lieferung 202). Stuttgart: E. Schweizerbart' sche Verlagsbuchhandlung; p. 465 - 512." type="book chapter" year="1958">Schmitz (1958)</bibRefCitation>
to key out this species in his Abteilung IV, Erste Reuhe.
<bibRefCitation id="F77E4B0AFFB2FFF34652465BFB0DF984" author="Schmitz H &amp; Beyer E" box="[894,1172,1538,1563]" pageId="2" pageNumber="1601" pagination="513 - 608" refId="ref8619" refString="Schmitz H, Beyer E. 1965. Phoridae. In: Lindner E editor. Die Fliegen der palaearktischen Region 4 (33) (Lieferung 258, 260). Stuttgart: E. Schweizerbart' sche Verlagsbuchhandlung; p. 513 - 608." type="book chapter" year="1965">Schmitz and Beyer (1965)</bibRefCitation>
then keyed out
<taxonomicName id="54EF4D78FFB2FFF344AD467AFD31F9A4" authority="in Abteilung" authorityName="in Abteilung" box="[385,680,1570,1594]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="1601" phylum="Arthropoda" rank="species" species="gargarans">
<emphasis id="A19BEAE9FFB2FFF344AD467AFD8FF9A4" box="[385,534,1571,1594]" italics="true" pageId="2" pageNumber="1601">M. gargarans</emphasis>
in Abteilung
</taxonomicName>
IV, Zweite Reihe. To add to the confusion,
<bibRefCitation id="F77E4B0AFFB2FFF345F8461BFE09F9C4" author="Borgmeier T" box="[212,400,1602,1626]" pageId="2" pageNumber="1601" pagination="1 - 367" refId="ref7815" refString="Borgmeier T. 1968. A catalogue of the Phoridae of the world (Diptera, Phoridae). Studia Entomologica, Petropolis. 11: 1 - 367." type="journal article" year="1968">Borgmeier (1968)</bibRefCitation>
cited the wrong paper of Schmitz for the synonymy of
<taxonomicName id="54EF4D78FFB2FFF346F5461AFBFAF9C4" authorityName="Lundbeck" authorityYear="1920" box="[985,1123,1602,1626]" class="Insecta" family="Phoridae" genus="Aphiochaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="1601" phylum="Arthropoda" rank="species" species="dactylata">
<emphasis id="A19BEAE9FFB2FFF346F5461AFBFAF9C4" box="[985,1123,1602,1626]" italics="true" pageId="2" pageNumber="1601">M. dactylata</emphasis>
</taxonomicName>
, but correctly observed that
<bibRefCitation id="F77E4B0AFFB2FFF344F4463BFDE5F9E4" author="Schmitz H" box="[472,636,1634,1658]" pageId="2" pageNumber="1601" pagination="465 - 512" refId="ref8577" refString="Schmitz H. 1958. Phoridae. In: Lindner E, editor. Die Fliegen der palaearktischen Region 4 (33) (Lieferung 202). Stuttgart: E. Schweizerbart' sche Verlagsbuchhandlung; p. 465 - 512." type="book chapter" year="1958">Schmitz (1958)</bibRefCitation>
had erroneously referred a photo of the wing of another species to
<emphasis id="A19BEAE9FFB2FFF344B146DAFDB3F904" box="[413,554,1667,1690]" italics="true" pageId="2" pageNumber="1601">
<taxonomicName id="54EF4D78FFB2FFF344B146DAFDBFF904" baseAuthorityName="Wood" baseAuthorityYear="1909" box="[413,550,1667,1690]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="1601" phylum="Arthropoda" rank="species" species="crassipes">M. crassipes</taxonomicName>
.
</emphasis>
</paragraph>
<caption id="C7906673FFB2FFF345F8454BFE6EFADB" ID-DOI="http://doi.org/10.5281/zenodo.4006316" ID-Zenodo-Dep="4006316" httpUri="https://zenodo.org/record/4006316/files/figure.png" pageId="2" pageNumber="1601" startId="2.[212,279,1298,1320]" targetBox="[213,1171,169,1265]" targetPageId="2">
<paragraph id="935036FBFFB2FFF345F8454BFE6EFADB" blockId="2.[212,1171,1298,1349]" pageId="2" pageNumber="1601">
Figure 1.
<taxonomicName id="54EF4D78FFB2FFF34412454BFD9CFAB9" baseAuthorityName="Wood" baseAuthorityYear="1909" box="[318,517,1298,1319]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="1601" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="A19BEAE9FFB2FFF34412454BFD9CFAB9" box="[318,517,1298,1319]" italics="true" pageId="2" pageNumber="1601">Megaselia crassipes</emphasis>
</taxonomicName>
male. (A) Posterior face of front tarsus; (B) left face of hypopytgium. Scale bar: 0.1 mm.
</paragraph>
</caption>
<caption id="C7906673FFB3FFF245BC4472FCAEFBC0" ID-DOI="http://doi.org/10.5281/zenodo.4006318" ID-Zenodo-Dep="4006318" httpUri="https://zenodo.org/record/4006318/files/figure.png" pageId="3" pageNumber="1602" startId="3.[144,211,1067,1089]" targetBox="[145,1103,169,1034]" targetPageId="3">
<paragraph id="935036FBFFB3FFF245BC4472FCAEFBC0" blockId="3.[144,1104,1067,1118]" pageId="3" pageNumber="1602">
Figure 2.
<taxonomicName id="54EF4D78FFB3FFF245DB4472FE22FBDE" baseAuthorityName="Wood" baseAuthorityYear="1909" box="[247,443,1067,1088]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="1602" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="A19BEAE9FFB3FFF245DB4472FE22FBDE" box="[247,443,1067,1088]" italics="true" pageId="3" pageNumber="1602">Megaselia crassipes</emphasis>
</taxonomicName>
male (synonym of
<taxonomicName id="54EF4D78FFB3FFF247534475FC88FBDE" authorityName="Disney and Durska" authorityYear="2011" box="[639,785,1067,1089]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="1602" phylum="Arthropoda" rank="species" species="basiturgida">
<emphasis id="A19BEAE9FFB3FFF247534475FC88FBDE" box="[639,785,1067,1089]" italics="true" pageId="3" pageNumber="1602">M. basiturgida</emphasis>
</taxonomicName>
). (A) Left face of hypopygium; (B) posterior face of front tibia and basitarsus. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="935036FBFFB3FFF5459444FBFB0AFAA2" blockId="3.[144,1104,1186,1689]" lastBlockId="4.[212,1172,1284,1340]" lastPageId="4" lastPageNumber="1603" pageId="3" pageNumber="1602">
The recognition of
<taxonomicName id="54EF4D78FFB3FFF244BF44FAFDB3FB24" box="[403,554,1187,1210]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="1602" phylum="Arthropoda" rank="species" species="gargarans">
<emphasis id="A19BEAE9FFB3FFF244BF44FAFDB3FB24" box="[403,554,1187,1210]" italics="true" pageId="3" pageNumber="1602">M. gargarans</emphasis>
</taxonomicName>
has also proved troublesome. In the keys to the males of the species of the British Isles (
<bibRefCitation id="F77E4B0AFFB3FFF2471F449BFD22FB44" author="Disney RHL" box="[563,699,1218,1242]" pageId="3" pageNumber="1602" pagination="1 - 155" refId="ref8043" refString="Disney RHL. 1989. Scuttle Flies - Diptera Phoridae Genus Megaselia. Handbk Ident Br Insects. 10: 1 - 155." type="journal article" year="1989">Disney 1989</bibRefCitation>
), it seemingly runs to couplet 81, lead 2, to
<taxonomicName id="54EF4D78FFB3FFF245EB44BAFED4FB64" baseAuthorityName="Wood" baseAuthorityYear="1909" box="[199,333,1251,1274]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="1602" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="A19BEAE9FFB3FFF245EB44BAFED4FB64" box="[199,333,1251,1274]" italics="true" pageId="3" pageNumber="1602">M. crassipes</emphasis>
</taxonomicName>
. Schmitz did not provide a figure of the hypopygium but described the anal tube as being kurz, kaum länger als hoch, which would immediately exclude
<emphasis id="A19BEAE9FFB3FFF24102455AFF6EFAA4" italics="true" pageId="3" pageNumber="1602">
<taxonomicName id="54EF4D78FFB3FFF24102455AFF6AFAA4" baseAuthorityName="Wood" baseAuthorityYear="1909" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="1602" phylum="Arthropoda" rank="species" species="crassipes">M. crassipes</taxonomicName>
.
</emphasis>
In the keys of
<bibRefCitation id="F77E4B0AFFB3FFF24489457BFD51FAA4" author="Schmitz H &amp; Beyer E" box="[421,712,1314,1339]" pageId="3" pageNumber="1602" pagination="513 - 608" refId="ref8619" refString="Schmitz H, Beyer E. 1965. Phoridae. In: Lindner E editor. Die Fliegen der palaearktischen Region 4 (33) (Lieferung 258, 260). Stuttgart: E. Schweizerbart' sche Verlagsbuchhandlung; p. 513 - 608." type="book chapter" year="1965">Schmitz and Beyer (1965)</bibRefCitation>
, the males of
<taxonomicName id="54EF4D78FFB3FFF24646457AFB98FAA4" box="[874,1025,1315,1338]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="1602" phylum="Arthropoda" rank="species" species="gargarans">
<emphasis id="A19BEAE9FFB3FFF24646457AFB98FAA4" box="[874,1025,1315,1338]" italics="true" pageId="3" pageNumber="1602">M. gargarans</emphasis>
</taxonomicName>
run to couplet 52 on p. 523. Furthermore, in the text Beyer pointed out that Schmitz s description of the anal tube was incorrect, as he reported his measurement of its length-tobreadth ratio as being 8:4. Examination of Schmitz s slide mount of a
<typeStatus id="4C548859FFB3FFF246B945DBFC6FFA04" box="[917,1014,1410,1434]" pageId="3" pageNumber="1602" type="paratype">paratype</typeStatus>
male of
<taxonomicName id="54EF4D78FFB3FFF245BC45FBFEBAFA27" box="[144,291,1442,1465]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="1602" phylum="Arthropoda" rank="species" species="gargarans">
<emphasis id="A19BEAE9FFB3FFF245BC45FBFEBAFA27" box="[144,291,1442,1465]" italics="true" pageId="3" pageNumber="1602">M. gargarans</emphasis>
</taxonomicName>
(in the Museum Koenig, Bonn) endorses this (
<figureCitation id="0BD42A7EFFB3FFF2463145F8FCE0FA27" box="[797,889,1441,1465]" captionStart="Figure 3" captionStartId="4.[212,279,1198,1220]" captionTargetBox="[212,1172,168,1166]" captionTargetId="figure@4.[212,1172,168,1166]" captionTargetPageId="4" captionText="Figure 3. Megaselia crassipes male (synonym of M. gargarans), left face of anal tube." figureDoi="http://doi.org/10.5281/zenodo.4006322" httpUri="https://zenodo.org/record/4006322/files/figure.png" pageId="3" pageNumber="1602">Figure 3</figureCitation>
). Furthermore, it is evident that it is a specimen of
<taxonomicName id="54EF4D78FFB3FFF244CD459BFDF1FA47" baseAuthorityName="Wood" baseAuthorityYear="1909" box="[481,616,1474,1497]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="1602" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="A19BEAE9FFB3FFF244CD459BFDF1FA47" box="[481,616,1474,1497]" italics="true" pageId="3" pageNumber="1602">M. crassipes</emphasis>
</taxonomicName>
at the larger and darker end of the spectrum of variation in this species. Therefore, the synonymy of
<taxonomicName id="54EF4D78FFB3FFF247CA45BBFCE0FA67" box="[742,889,1506,1529]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="1602" phylum="Arthropoda" rank="species" species="gargarans">
<emphasis id="A19BEAE9FFB3FFF247CA45BBFCE0FA67" box="[742,889,1506,1529]" italics="true" pageId="3" pageNumber="1602">M. gargarans</emphasis>
</taxonomicName>
with
<taxonomicName id="54EF4D78FFB3FFF2469845BBFBA0FA67" baseAuthorityName="Wood" baseAuthorityYear="1909" box="[948,1081,1506,1529]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="1602" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="A19BEAE9FFB3FFF2469845BBFBA0FA67" box="[948,1081,1506,1529]" italics="true" pageId="3" pageNumber="1602">M. crassipes</emphasis>
</taxonomicName>
is herewith formally proposed.
<taxonomicName id="54EF4D78FFB3FFF244F4465BFDE2F987" authorityName="Disney and Durska" authorityYear="2011" box="[472,635,1537,1561]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="1602" phylum="Arthropoda" rank="species" species="basiturgida">
<emphasis id="A19BEAE9FFB3FFF244F4465BFDE2F987" box="[472,635,1537,1561]" italics="true" pageId="3" pageNumber="1602">M. basiturgida</emphasis>
</taxonomicName>
was described from two similarly darker males, but which are not quite as large as the
<typeStatus id="4C548859FFB3FFF24758467BFD4CF9A4" box="[628,725,1570,1594]" pageId="3" pageNumber="1602" type="paratype">paratype</typeStatus>
of
<emphasis id="A19BEAE9FFB3FFF247DA467BFC16F9A7" box="[758,911,1570,1593]" italics="true" pageId="3" pageNumber="1602">
<taxonomicName id="54EF4D78FFB3FFF247DA467BFC15F9A7" box="[758,908,1570,1593]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="1602" phylum="Arthropoda" rank="species" species="gargarans">M. gargarans</taxonomicName>
.
</emphasis>
Furthermore, the left hypandrial lobes of these two males are paler than usual. However, in every other respect, they lie within the ranges of variation for
<emphasis id="A19BEAE9FFB3FFF24787463BFCA0F9E7" box="[683,825,1634,1657]" italics="true" pageId="3" pageNumber="1602">
<taxonomicName id="54EF4D78FFB3FFF24787463BFCACF9E7" baseAuthorityName="Wood" baseAuthorityYear="1909" box="[683,821,1634,1657]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="1602" phylum="Arthropoda" rank="species" species="crassipes">M. crassipes</taxonomicName>
.
</emphasis>
Therefore, the synonymy of
<taxonomicName id="54EF4D78FFB3FFF2458046DBFED2F907" authorityName="Disney and Durska" authorityYear="2011" box="[172,331,1665,1689]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="1602" phylum="Arthropoda" rank="species" species="basiturgida">
<emphasis id="A19BEAE9FFB3FFF2458046DBFED2F907" box="[172,331,1665,1689]" italics="true" pageId="3" pageNumber="1602">M. basiturgida</emphasis>
</taxonomicName>
with
<taxonomicName id="54EF4D78FFB3FFF244AA46DBFD95F907" baseAuthorityName="Wood" baseAuthorityYear="1909" box="[390,524,1666,1689]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="1602" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="A19BEAE9FFB3FFF244AA46DBFD95F907" box="[390,524,1666,1689]" italics="true" pageId="3" pageNumber="1602">M. crassipes</emphasis>
</taxonomicName>
is also herewith formally proposed. Critical details of
<taxonomicName id="54EF4D78FFB4FFF545F8455CFEC2FA82" baseAuthorityName="Wood" baseAuthorityYear="1909" box="[212,347,1285,1308]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="1603" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="A19BEAE9FFB4FFF545F8455CFEC2FA82" box="[212,347,1285,1308]" italics="true" pageId="4" pageNumber="1603">M. crassipes</emphasis>
</taxonomicName>
are given in
<figureCitation id="0BD42A7EFFB4FFF544CB455DFDF1FA82" box="[487,616,1284,1308]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStartId-0="2.[212,279,1298,1320]" captionStartId-1="3.[144,211,1067,1089]" captionTargetBox-0="[213,1171,169,1265]" captionTargetBox-1="[145,1103,169,1034]" captionTargetId-0="figure@2.[212,1172,168,1266]" captionTargetId-1="figure@3.[144,1104,168,1035]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Figure 1. Megaselia crassipes male. (A) Posterior face of front tarsus; (B) left face of hypopytgium. Scale bar: 0.1 mm." captionText-1="Figure 2. Megaselia crassipes male (synonym of M. basiturgida). (A) Left face of hypopygium; (B) posterior face of front tibia and basitarsus. Scale bar: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4006316" figureDoi-1="http://doi.org/10.5281/zenodo.4006318" httpUri-0="https://zenodo.org/record/4006316/files/figure.png" httpUri-1="https://zenodo.org/record/4006318/files/figure.png" pageId="4" pageNumber="1603">Figures 12</figureCitation>
and its recognition clarified in the revised couplets 7981 of the key to the males recorded from the British Isles (
<bibRefCitation id="F77E4B0AFFB4FFF54655457DFB9BFAA2" author="Disney RHL" box="[889,1026,1316,1340]" pageId="4" pageNumber="1603" pagination="1 - 155" refId="ref8043" refString="Disney RHL. 1989. Scuttle Flies - Diptera Phoridae Genus Megaselia. Handbk Ident Br Insects. 10: 1 - 155." type="journal article" year="1989">Disney 1989</bibRefCitation>
) (see below).
</paragraph>
</subSubSection>
</treatment>
</document>