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120 lines
14 KiB
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<document ID-CLB-Dataset="68943" ID-DOI="http://dx.doi.org/10.3897/zitteliana.96.83571" ID-GBIF-Dataset="f614d115-44b1-41ef-89ce-8651629bceb0" ID-Pensoft-Pub="2747-8106-96-103" ID-Pensoft-UUID="4E0188A4D3D7563C980CB7C5B9424F3C" ID-ZooBank="5B10FFF604074E09B986F931635E9BCC" ModsDocID="2747-8106-96-103" checkinTime="1654860115112" checkinUser="pensoft" docAuthor="Schwarzhans, Werner, Ohe, Fumio, Tsuchiya, Yuki & Ujihara, Atsushi" docDate="2022" docId="0CC9D2FF1EFF5D908C88A98B32242BE2" docLanguage="en" docName="Zitteliana 96: 103-134" docOrigin="Zitteliana 96" docPubDate="2022-06-10" docSource="http://dx.doi.org/10.3897/zitteliana.96.83571" docTitle="Diaphus epipedus Schwarzhans & Ohe & Tsuchiya & Ujihara 2022, sp. nov." docType="treatment" docUuid="FEEDFC9A-53E6-468B-B369-53B3BD7ED6B4" docUuidSource="ZooBank" docVersion="5" id="4E0188A4D3D7563C980CB7C5B9424F3C" lastPageNumber="103" masterDocId="4E0188A4D3D7563C980CB7C5B9424F3C" masterDocTitle="Lanternfish otoliths (Myctophidae, Teleostei) from the Miocene of Japan" masterLastPageNumber="134" masterPageNumber="103" pageNumber="103" updateTime="1732684795358" updateUser="ExternalLinkService">
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<mods:mods id="780958FB75480F2F64A406A7B93EE877" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="55FF6219415A2589B43B9A774E9D211A">
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<mods:title id="74A3C822B6253E7E2F0EABB120F4627D">Lanternfish otoliths (Myctophidae, Teleostei) from the Miocene of Japan</mods:title>
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<mods:name id="D07B166C1EC1627DDB2DC95A62D3470D" type="personal">
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<mods:role id="848ACC08126618953DE15306DD530B48">
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<mods:roleTerm id="038DF46982DE689EF7AD4E5B70D3D2FC">Author</mods:roleTerm>
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<mods:namePart id="6F875D40DA0521EE661C848F25BA6CC5">Schwarzhans, Werner</mods:namePart>
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<mods:affiliation id="C0F1FB341AFF1C8F9648A2B9049567F5">Zoological Museum, Natural History Museum of Denmark, Universitetsparken 15, 2100 Kobenhavn, Denmark & Ahrensburger Weg 103, 22359 Hamburg, Germany</mods:affiliation>
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<mods:nameIdentifier id="9BF2D3700B47293F9415E005B043F32C" type="email">wwschwarz@aol.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="4762CC2F1EDBEABCCB9B3D19A81DA465" type="personal">
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<mods:roleTerm id="FC1DA1CFA293863CEE6B7266F14E38A1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="49127674499D785DB9D7E81C509DB158">Ohe, Fumio</mods:namePart>
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<mods:affiliation id="D17AF000ADE856CA3C06947AF438FADB">Nara National Research Institute for Cultural Properties, Nara 630 - 8577, Japan</mods:affiliation>
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</mods:name>
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<mods:name id="8BEF193466B921E9253B68E8F8699891" type="personal">
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<mods:roleTerm id="01291CC6AD9EBD72F443F50CCF0C4085">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3433D1DA1442E348384B474DC342CCB7">Tsuchiya, Yuki</mods:namePart>
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<mods:affiliation id="A462A179FA4E11980C55DE3B94A2652E">Graduate School of Environmental Studies, Nagoya University, Nagoya, Japan</mods:affiliation>
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</mods:name>
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<mods:name id="BE4E4922BEAB7BF0FE8414FBD55B28F5" type="personal">
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</mods:role>
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<mods:namePart id="A3A22DA12043A1EB60413FBEF223F650">Ujihara, Atsushi</mods:namePart>
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<mods:affiliation id="D9CCACE1496213A1B38B73567D029135">Graduate School of Environmental Studies, Nagoya University, Nagoya, Japan</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="FAC456808CE885289119A9DFC7DB7D22">text</mods:typeOfResource>
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<mods:title id="C0E84361E6EC71C362F4D409889C2E2A">Zitteliana</mods:title>
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<mods:part id="9B79324715E661F7595EF2988A56E04A">
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<mods:date id="31408ECEC24DC7577C051BDEF00A1422">2022</mods:date>
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<mods:number id="DB23811576CCBCBA27F1A566FD4B0D19">2022-06-10</mods:number>
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<mods:detail id="31FFC77EA38C271A01B7197DA3EE1A22" type="volume">
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<mods:number id="2B3FF1374CC6282776C481044B045F8A">96</mods:number>
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<mods:start id="A92695B94CF833CBBF2AE7BA172A44F2">103</mods:start>
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<mods:end id="2DA266FEAE123147AA02FC900690B474">134</mods:end>
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<mods:url id="68933FBEACE7B4F2D8C4E2B848A46DB2">http://dx.doi.org/10.3897/zitteliana.96.83571</mods:url>
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<mods:classification id="E23F95410CDD6E62C1D2B3578AFAA7B4">journal article</mods:classification>
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<mods:identifier id="6D7B323DEEF4179263B9900850BDF158" type="DOI">http://dx.doi.org/10.3897/zitteliana.96.83571</mods:identifier>
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<mods:identifier id="742F7165BCB902430136ECBB7AF9239E" type="Pensoft-Pub">2747-8106-96-103</mods:identifier>
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<treatment id="0CC9D2FF1EFF5D908C88A98B32242BE2" ID-GBIF-Taxon="195832375" LSID="urn:lsid:zoobank.org:act:FEEDFC9A-53E6-468B-B369-53B3BD7ED6B4" httpUri="http://treatment.plazi.org/id/0CC9D2FF1EFF5D908C88A98B32242BE2" lastPageNumber="103" pageId="0" pageNumber="103" scope_family="Myctophidae" scope_order="Myctophiformes" scope_phylum="Teleostei">
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<subSubSection id="40B88CBE1A2C2DD0DF9C5C9FA863F363" pageId="0" pageNumber="103" type="nomenclature">
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<paragraph id="D8A145666F66D8FB1DED363CF517FAD4" pageId="0" pageNumber="103">
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<taxonomicName id="5169F6E59FA071FCC98AC833491171CF" LSID="http://zoobank.org/FEEDFC9A-53E6-468B-B369-53B3BD7ED6B4" authority="Schwarzhans & Ohe & Tsuchiya & Ujihara, 2022" authorityName="Schwarzhans & Ohe & Tsuchiya & Ujihara" authorityYear="2022" family="Myctophidae" genus="Diaphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diaphus epipedus" order="Myctophiformes" pageId="0" pageNumber="103" phylum="Chordata" rank="species" species="epipedus" status="sp. nov.">Diaphus epipedus</taxonomicName>
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<taxonomicNameLabel id="D97B2A201FE681E792F674A899D1636B" pageId="0" pageNumber="103">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="3EDF83E7340CE41F739A771C12A12978" pageId="0" pageNumber="103" type="description">
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<paragraph id="F7C1B269C4B001B26F155BF6A7EC67DA" pageId="0" pageNumber="103">
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<figureCitation id="1CFB3C14A30678795BCBDA8D94EBCEBA" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-F: Diaphus angulatus Ohe & Araki, 1973; A-C: SMF PO 101.130, Inabacho, Tsu City, Katada Formation, level ML 2305; D-E: SMF PO 101.131, Fuchu machi, Toyama City, Higashibessho Formation, level H 20; G-N: Diaphus epipedus sp. nov.; G-I: holotype, SMF PO 101.132, Tsuyama City, Takakura Formation, level MS 15; K-J: paratype, SMF PO 101.133, Tsuyama City, Takakura Formation, level MS 15; L-N: paratype, SMF PO 101.134, Yatsuo, Toyama City, Higashibessho Formation, level H 21; O-T: Diaphus metopoclampoides Steurbaut, 1983, SMF PO 101.135, Tsuyama City, Takakura Formation level MS 06, f; U-AB: Diaphus watatsumi sp. nov., Tsuyama City, Takakura Formation; U-X: holotype, SMF PO 101.136, level MS 52; Y-AB: paratypes, SMF PO 101.137, levels gh, j; AC-AE: Diaphus sp. 1, col. Tsuchiya, Ichishi Group." figureDoi="10.3897/zitteliana.96.83571.figure8" httpUri="https://binary.pensoft.net/fig/694121" pageId="0" pageNumber="103">Fig. 8G-N</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="9872DCB3DDA6907CAB0FAC8C97815024" pageId="0" pageNumber="103" type="etymology">
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<paragraph id="9F60188CB5B20A77796074F47BA24499" pageId="0" pageNumber="103">Etymology.</paragraph>
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<paragraph id="0DDF0818226ACB7B29E42DE1047D120C" pageId="0" pageNumber="103">
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From epipedos (Greek) = flat, referring to the relatively shallow ventral rim, the main distinguishing character from
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<taxonomicName id="C33F5D3072674F03A05D2AD7074ACCD0" lsidName="D. metopoclampoides" pageId="0" pageNumber="103" rank="species" species="metopoclampoides">
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<emphasis id="E16836C5653862AE8361D0438B6EFB2B" italics="true" pageId="0" pageNumber="103">D. metopoclampoides</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="D8C5E4940BBC74EFB0AF61393143355D" pageId="0" pageNumber="103" type="holotype">
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<paragraph id="250203E263B23E42A8CFBE748A5BE74A" pageId="0" pageNumber="103">Holotype.</paragraph>
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<paragraph id="C44E393BCEFD7174EB7D63AD51AB970C" pageId="0" pageNumber="103">
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SMF PO 101.132 (Fig.
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<figureCitation id="B421C7C75A294EBEBC0C386F821941E6" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-F: Diaphus angulatus Ohe & Araki, 1973; A-C: SMF PO 101.130, Inabacho, Tsu City, Katada Formation, level ML 2305; D-E: SMF PO 101.131, Fuchu machi, Toyama City, Higashibessho Formation, level H 20; G-N: Diaphus epipedus sp. nov.; G-I: holotype, SMF PO 101.132, Tsuyama City, Takakura Formation, level MS 15; K-J: paratype, SMF PO 101.133, Tsuyama City, Takakura Formation, level MS 15; L-N: paratype, SMF PO 101.134, Yatsuo, Toyama City, Higashibessho Formation, level H 21; O-T: Diaphus metopoclampoides Steurbaut, 1983, SMF PO 101.135, Tsuyama City, Takakura Formation level MS 06, f; U-AB: Diaphus watatsumi sp. nov., Tsuyama City, Takakura Formation; U-X: holotype, SMF PO 101.136, level MS 52; Y-AB: paratypes, SMF PO 101.137, levels gh, j; AC-AE: Diaphus sp. 1, col. Tsuchiya, Ichishi Group." figureDoi="10.3897/zitteliana.96.83571.figure8" httpUri="https://binary.pensoft.net/fig/694121" pageId="0" pageNumber="103">8G-I</figureCitation>
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), Miya River at Numa, Tsuyama City, Okayama Prefecture, Takakura FM, level MS 15, late Burdigalian, diatom zone 3A-3B.
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</paragraph>
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</subSubSection>
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<subSubSection id="DDC72971723A95975A378BDBA3ABB4DD" pageId="0" pageNumber="103" type="paratypes">
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<paragraph id="076447E56BBE9FA31804BE71E898D6A4" pageId="0" pageNumber="103">Paratypes.</paragraph>
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<paragraph id="BA53440D1E3CB057402A6DE4597CB6B4" pageId="0" pageNumber="103">8 specimens; 6 specimens (SMF PO 101.133), same data as holotype, levels MS 15, MS 27, MS 90, j; 2 specimens (SMF PO 101.134), Osedani, Toyama City, Toyama Prefecture, Higashibessho FM, level H 21, late Burdigalian to early Langhian, diatom zone 3A-4A.</paragraph>
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</subSubSection>
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<subSubSection id="3394BB37583E88CC336B93558F8BA72C" pageId="0" pageNumber="103" type="diagnosis">
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<paragraph id="904F6D5E6F92FD8AD465FC6EAEE4F965" pageId="0" pageNumber="103">Diagnosis.</paragraph>
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<paragraph id="ED8F643858798243F2C1484EAE834F4E" pageId="0" pageNumber="103">OL:OH = 1.2-1.25. Shape rounded triangular with relatively shallow ventral rim, inferior rostrum and posterior tip and broad mediodorsal bulge. Antirostrum and excisura minute. Ventral rim with 7-9 denticles. Postdorsal rim broadly concave. OCL:CCL = 1.5-1.7.</paragraph>
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</subSubSection>
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<subSubSection id="1FCC3F137DD95DD1E7DD19377E6DD600" pageId="0" pageNumber="103" type="description">
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<paragraph id="FBE813A00BBF7EE4003C7C7347FE2820" pageId="0" pageNumber="103">Description.</paragraph>
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<paragraph id="E89E68207CAEB8359E68A1E1D79508F9" pageId="0" pageNumber="103">Otoliths with rounded triangular outline, relatively robust, up to 2.8 mm in length (holotype 2.5 mm); OH:OT = 3.4-3.7. Rostrum far inferior, relatively short and pointed, 10-12% of OL. Antirostrum and excisura minute. Dorsal rim with broad, rounded, pronounced mediodorsal bulge, anteriorly steeply inclined, posteriorly inclined with long depressed or concave section. Ventral rim rather shallow, regularly curved, with 7-9 medium strong denticles. Posterior rim curved, slightly projecting behind termination of cauda, its tip centrally or inframedian.</paragraph>
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<paragraph id="94A427B3019281303D10DD1439EFA374" pageId="0" pageNumber="103">Inner face relatively flat, only slightly bent in horizontal direction. Sulcus terminating relatively far from posterior rim of otolith, straight, narrow. Ostial colliculum distinctly longer than caudal colliculum (OCL:CCL = 1.5-1.7) but only slightly wider (OCH:CCH = 1.2-1.4). Caudal pseudocolliculum distinct, about as long as caudal colliculum. Ventral furrow indistinct, sometimes not discernable, relatively close to ventral rim of otolith; dorsal depression small, ventrally marked by short crista superior. Outer face moderately convex throughout without distinct umbo, with distinct excisural furrow and few other weak and short radial furrows.</paragraph>
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</subSubSection>
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<subSubSection id="3AF515A9CB5C366497899209BAEF7A39" pageId="0" pageNumber="103" type="discussion">
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<paragraph id="D3FB310F548E3B3649031FCEBEB8C0C0" pageId="0" pageNumber="103">Discussion.</paragraph>
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<paragraph id="EF96ADF188424C93240F13A06138670E" pageId="0" pageNumber="103">
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<taxonomicName id="D4A26653FCDFBE03658E9685B25E2F06" authorityName="Schwarzhans & Ohe & Tsuchiya & Ujihara" authorityYear="2022" family="Myctophidae" genus="Diaphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diaphus epipedus" order="Myctophiformes" pageId="0" pageNumber="103" phylum="Chordata" rank="species" species="epipedus">
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<emphasis id="B93CA5F70B81CAAE5D57EF66639EBC5F" italics="true" pageId="0" pageNumber="103">Diaphus epipedus</emphasis>
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</taxonomicName>
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is an easily recognizable otolith that shares many features, such as the shape of the dorsal rim and the robust appearance without distinct umbo on the outer face, with
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<taxonomicName id="57FB20F8685A168FEB1385EBE2E60311" lsidName="D. metopoclampoides" pageId="0" pageNumber="103" rank="species" species="metopoclampoides">
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<emphasis id="21DA4960F792F0781475E29345F66CDC" italics="true" pageId="0" pageNumber="103">D. metopoclampoides</emphasis>
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</taxonomicName>
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(see below) but differs in the shallow ventral rim, which results in a higher ratio OL:OH of 1.2-1.25 (vs. 1.05-1.1); the relatively long ostium (OCL:CCL = 1.5-1.7 vs. 1.1-1.4); and the narrower sulcus.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |