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<mods:title id="39D9B0DB124A53184E6D938291440D4C">A revision of dragon millipedes I: genus Desmoxytes Chamberlin, 1923, with the description of eight new species (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:namePart id="419CC33702E8F45059DCAA2C876A10E7">Srisonchai, Ruttapon</mods:namePart>
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<mods:namePart id="9A0156AABC6A6C4CFAD04F4D69DBAD8A">Enghoff, Henrik</mods:namePart>
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<taxonomicName id="46C1D2BF9D03F76B8C21C261D4DDD21E" ID-CoL="354CG" authority="Jeekel, 1964" authorityName="Jeekel" authorityYear="1964" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes terae" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="terae">Desmoxytes terae (Jeekel, 1964)</taxonomicName>
Figs 88, 89, 90, 91, 92
</paragraph>
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<paragraph id="8D721EEE50E8DF2F66FBD29D66A5BEF4" pageId="0" pageNumber="1">
<taxonomicName id="ED72EBDBBFF6DF6E749B30F46AA0C48A" class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus terae" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="terae">Pratinus terae</taxonomicName>
Jeekel, 1964: 69; 1968: 51.
</paragraph>
<paragraph id="80A87AB58C0582B3712601715FD7D90F" pageId="0" pageNumber="1">
<taxonomicName id="B21BBDF97781945DDCC1DBAE61F1EEE4" class="Diplopoda" family="Paradoxosomatidae" genus="Pteroxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pteroxytes terae" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="terae">Pteroxytes terae</taxonomicName>
-
<bibRefCitation id="A9E52EC4D33205E61FEA2078A06C1DAB" author="Jeekel, CAW" journalOrPublisher="Revue suisse de Zoologie" pageId="0" pageNumber="1" pagination="651 - 670" title="The generic allocation of some little-known Paradoxosomatidae from South-East Asia (Diplopoda, Polydesmida)." url="https://biodiversitylibrary.org/page/41420961" volume="87" year="1980 a">Jeekel 1980a</bibRefCitation>
: 655.
</paragraph>
<paragraph id="6FE661EC66D94250DE40822989CAE15F" pageId="0" pageNumber="1">
<taxonomicName id="AED585FA52282AD30E3E28D1713DE3FF" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes terae" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="terae">Desmoxytes terae</taxonomicName>
-
<bibRefCitation id="3E65D5E84AA0B0CE6E34672505B69BFA" author="Golovatch, SI" journalOrPublisher="Steenstrupia" pageId="0" pageNumber="1" pagination="45 - 71" title="Review of the dragon millipedes, genus Desmoxytes Chamberlin, 1923 (Diplopoda, Polydesmida, Paradoxosomatidae)." volume="20" year="1994">Golovatch and Enghoff 1994</bibRefCitation>
: 59.
<bibRefCitation id="7818E4333CC28A119342D8BA5D13B5E8" author="Enghoff, H" journalOrPublisher="Steenstrupia" pageId="0" pageNumber="1" pagination="87 - 103" title="The millipedes of Thailand (Diplopoda)." url="http://www.zmuc.dk/commonweb/JOURNALS/PDF/Vol29-1/Enghoff.pdf" volume="29" year="2005">Enghoff 2005</bibRefCitation>
: 97.
<bibRefCitation id="535C36820648B21AAFA269B7890E2BF0" author="Nguyen, DA" journalOrPublisher="Check List" pageId="0" pageNumber="1" pagination="1132 - 1353" title="A worldwide catalog of the family Paradoxosomatidae Daday, 1889 (Diplopoda: Polydesmida)." url="https://doi.org/10.15560/9.6.1132" volume="9" year="2013">Nguyen and Sierwald 2013</bibRefCitation>
: 1243.
</paragraph>
</subSubSection>
<subSubSection id="8DD40B6A02DD91C8DBC82FEECB7DA1B7" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="1898AEDEE38B095A8CB2EADF1556ABF9" pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph id="56498F6983B6D881DDEA733774C3B4FB" pageId="0" pageNumber="1">Holotype. Male (NBC), MALAYSIA, Perlis, Kaki Bukit, near Kampong Wang Tangga, 19 December 1958, leg. W.S.S van der Feen-van Benthem Jutting.</paragraph>
<paragraph id="3D3E4E2540876973A02A398D6E182084" pageId="0" pageNumber="1">Paratypes. 1 female, 1 female fragment (NBC), same data as holotype.</paragraph>
</subSubSection>
<subSubSection id="AEE3027F33B0C8009DA82D5EC0AF9855" pageId="0" pageNumber="1" type="further specimens">
<paragraph id="95302AB4F29FCDB3C503A55558A8AA22" pageId="0" pageNumber="1">Further specimens, all from THAILAND, Satun Province:</paragraph>
<paragraph id="974C021AB30ACBEB2BD821D36E89201B" pageId="0" pageNumber="1">
1 male (ZMUC), Thale Ban National Park, in logs, litter, under stones,
<geoCoordinate id="C284D7BC7A5A273DF8B41689081A6670" direction="north" orientation="latitude" precision="925" value="6.7">6°42'N</geoCoordinate>
,
<geoCoordinate id="C2B79D6E83CE9975B3EF2C116562D589" direction="east" orientation="longitude" precision="925" value="100.166664">100°10'E</geoCoordinate>
, 8 November 1990, leg. M. Andersen and A. R. Rasmussen. 1 female (CUMZ), Khuan Don District, Thale Ban National Park, Tham Tone Din (Tone Din Cave),
<geoCoordinate id="9550B8F5E973230987E5E06370CED277" direction="north" orientation="latitude" precision="15" value="6.726389">6°43'35&quot;N</geoCoordinate>
,
<geoCoordinate id="B710E3C15F4AA1AB3A5853E713E3A2E4" direction="east" orientation="longitude" precision="15" value="100.1625">100°09'45&quot;E</geoCoordinate>
, ca. 154 m a.s.l., 31 August 2015, leg. S. Sutcharit, A. Pholyotha, T. Seesamut, and R. Srisonchai. 2 males, 6 females (CUMZ), 1 female (ZMUC), Khuan Don District, Thale Ban National Park, Tham Tone Din (Tone Din Cave),
<geoCoordinate id="3CB781502E319BAE7DB2951CF7A09DC4" direction="north" orientation="latitude" precision="15" value="6.726389">6°43'35&quot;N</geoCoordinate>
,
<geoCoordinate id="C6BDC6557111F077FDAB76F937B9FD2D" direction="east" orientation="longitude" precision="15" value="100.1625">100°09'45&quot;E</geoCoordinate>
, ca. 154 m a.s.l., 7 July 2017, leg. S. Sutcharit, R. Srisonchai, and ASRU members.
</paragraph>
</subSubSection>
<subSubSection id="F505F08FDC644025D03120A4404DF97F" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="E441782CACDD2C9E158D0E161C9923F0" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="81BB27F4CA166763BD5685920C5C6AD7" pageId="0" pageNumber="1">
Differs from all other
<taxonomicName id="EC2E7CF42278DF7B29D292B13598C32A" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Desmoxytes</taxonomicName>
species by the combination of the following characters; body black or brownish black contrasting with yellowish white paraterga with a triangular dorsal, dark spot; sternal lobe between male coxae 4 short and stout, broad at base, trapeziform or semicircular; male femora 5 and 6 without modification; lamina lateralis (ll) with big and long lobe-like structure projecting ventroanteriad.
</paragraph>
</subSubSection>
<subSubSection id="2D0A8C4EBAD98FF02F6435D88204D778" pageId="0" pageNumber="1" type="type locality">
<paragraph id="51D72F83C9CB925F810F0CB91B8A41E2" pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph id="F89B4046252411D527337896D734396C" pageId="0" pageNumber="1">MALAYSIA, Perlis, Kaki Bukit, near Kampong Wang Tangga.</paragraph>
<paragraph id="FB99404989733E1775E7999FBB446615" pageId="0" pageNumber="1">
The redescription hereunder is modified from
<bibRefCitation id="4F34CB982078ECC507F5DFD7CCB916D3" author="Jeekel, CAW" journalOrPublisher="Beaufortia" pageId="0" pageNumber="1" pagination="61 - 73" title="Two new species of Pratinus Attems, with taxonomic notes on the genus and a redescription of its type-species (Diplopoda, Polydesmida)." url="http://www.repository.naturalis.nl/document/548475" volume="11" year="1964">Jeekel (1964)</bibRefCitation>
; we
<normalizedToken id="B8B5F8CC787DB8240AFD2A6422C69356" originalValue="harmonised">'harmonised'</normalizedToken>
descriptions of all morphological characters and added some morphological characteristics from additional specimens.
</paragraph>
</subSubSection>
<subSubSection id="B37D60977A3218B86566BE501A68DE4F" pageId="0" pageNumber="1" type="redescription">
<paragraph id="3BF764C61A4A2656A0AAEFB404FC1123" pageId="0" pageNumber="1">Redescription.</paragraph>
<paragraph id="7E731F467AE135F6DB70250BC4D98E99" pageId="0" pageNumber="1">SIZE: Length 28-32 mm (male), 28-34 mm (female); width of midbody metazona ca. 2.4 mm (male), 3.1 mm (female). Width of head &lt;collum ≤ body ring 2 ≤ 3 = 4 &lt;5-17, thereafter body gradually tapering toward telson.</paragraph>
<paragraph id="7A16A225A5CB0D44E41837CA6EE27CF3" pageId="0" pageNumber="1">COLOUR (Fig. 88A, B): In life with body black or brownish black; paraterga yellowish white (dorsal side with triangular dark brown spot); metaterga, surface below paraterga and prozona black or brownish black; head black; antenna brownish black (except distal part of antennomere 7 and antennomere 8 whitish); legs brown; sterna brown to yellowish brown; a few basal podomeres and epiproct pale brown. Colour in alcohol: after 49 years changed to whitish brown, 2-5 years changed to pale blackish brown.</paragraph>
<caption id="8780DB6DC52260A65CD0381FF927E233" pageId="0" pageNumber="1">
<paragraph id="E31846F44C4985AA1D52F08A0070426A" pageId="0" pageNumber="1">
Figure 88. Photographs of live
<taxonomicName id="A27DF9F59A48D773C041B7B44FB2AC87" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes terae" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="terae">Desmoxytes terae</taxonomicName>
(Jeekel, 1964) (specimens from Tham Tone Din) and habitat. A male B female C habitat.
</paragraph>
</caption>
<paragraph id="480499C92F57F63AF0F888D944EB0663" pageId="0" pageNumber="1">ANTENNAE (Fig. 89D): Long and slender, reaching to body ring 6 (male), and 5 (female) when stretched dorsally.</paragraph>
<caption id="F95027484D3A9A5F56E1E4427FCA687C" pageId="0" pageNumber="1">
<paragraph id="6CEA12737E865D2571959B4E35256906" pageId="0" pageNumber="1">
Figure 89.
<taxonomicName id="3FCDAD184603D73D3032714149504040" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes terae" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="terae">Desmoxytes terae</taxonomicName>
(Jeekel, 1964), specimen from Tham Tone Din. A anterior body part B body rings 8-10 C posteriormost body rings and telson D head and antenna E paraterga of ring 10 F body ring 10.
</paragraph>
</caption>
<paragraph id="A1D12824C58BBEE2048A67F0DD37FF53" pageId="0" pageNumber="1">COLLUM (Fig. 89A): With 3 transverse rows of setae and tubercles, 3+3 anterior setae, 1+1 intermediate tubercles and 2+2 posterior tubercles (tubercles small, without setae), lateral tubercles of posterior row located more anteriorly, almost halfway to intermediate row; paraterga of collum with two distinct setiferous notches on lateral margin, directed caudolaterad, almost horizontal.</paragraph>
<paragraph id="145748ED886D0E33235480986A1440D9" pageId="0" pageNumber="1">TEGUMENT: Quite dull, sometimes shining; collum coarsely microgranulate; prozona, metaterga and surface below paraterga finely microgranulate; paraterga smooth (dorsal side finely microgranulate); sterna and epiproct smooth.</paragraph>
<paragraph id="A6B53B2354067EE1AD8D7EA111232585" pageId="0" pageNumber="1">
METATERGA (Fig. 89
<normalizedToken id="97279A5634962F1FE5581001754DC8BE" originalValue="AC">A-C</normalizedToken>
): With 2 transverse rows of tubercles; metaterga 2-19 with 2+2 anterior and 2+2 posterior tubercles.
</paragraph>
<paragraph id="2177DC425BF5A3CC8706904911B7274E" pageId="0" pageNumber="1">PARATERGA (Fig. 89E, F): Directed caudolaterad on body rings 2-17, elevated ca. 5°-10° above the horizontal plane in both sexes; directed increasingly caudad on body rings 18 and 19; anterior margin with 2 distinct notches, on lateral margin of body rings 9, 10, 12, 13, 15-18 with very small and tiny denticle near the tip.</paragraph>
<paragraph id="29DDF17C0BFB357E5EC599276D31D99B" pageId="0" pageNumber="1">
TELSON (Fig. 90
<normalizedToken id="9FF3361881849AE9F3A5E751F177BAFA" originalValue="CG">C-G</normalizedToken>
): Epiproct: tip usually subtruncate (in some specimens slightly emarginate); lateral setiferous tubercles usually conspicuous (in some specimens inconspicuous), quite short; apical tubercles inconspicuous. Hypoproct trapeziform; caudal margin usually subtruncate (in some specimens slightly round), with big and conspicuous setiferous tubercles.
</paragraph>
<caption id="CC42FD0896D223D7785B4CB171EF124B" pageId="0" pageNumber="1">
<paragraph id="B0AB44322D6AA0B7BFE84E1B1719A845" pageId="0" pageNumber="1">
Figure 90.
<taxonomicName id="CA76BF34BF8D6CAE19B82E682A56308A" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes terae" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="terae">Desmoxytes terae</taxonomicName>
(Jeekel, 1964), specimen from Tham Tone Din. A body rings 8-10 B sculpture of ring 10 C, E last ring and telson D hypoproct F, G epiproct H male leg 5 (right) I male leg 6 (right) J male leg 13 (right).
</paragraph>
</caption>
<paragraph id="C94D177B9FE293C93C9D8D90AB06C966" pageId="0" pageNumber="1">STERNA (Fig. 91): Cross-impressions quite deep. Sternal lobe between male coxae 4 swollen, short and stout, broad at base, usually trapeziform (in some specimens semicircular), tip usually truncate (in some specimens round).</paragraph>
<caption id="A617AFDD9ED128423111B838BE47C106" pageId="0" pageNumber="1">
<paragraph id="F22A150B9C3D21FD87F55819771256E1" pageId="0" pageNumber="1">
Figure 91.
<taxonomicName id="9C464E489A1642EB28D6D606D19371DB" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes terae" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="terae">Desmoxytes terae</taxonomicName>
(Jeekel, 1964), specimen from Tham Tone Din - sternal lobe between male coxae 4. A lateral view B mesal view
<normalizedToken id="35F615E3A5C0A574FD95A19D24AC2319" originalValue="CE">C-E</normalizedToken>
caudal view.
</paragraph>
</caption>
<paragraph id="6FD3DDF5FF3DE71DE5376FE6FAC3215C" pageId="0" pageNumber="1">
LEGS (Fig. 90
<normalizedToken id="28B5B98B5809D14C9E8B397EC2C1F934" originalValue="HJ">H-J</normalizedToken>
): Long and slender. Male femora 5 and 6 without modification.
</paragraph>
<paragraph id="759457B149826E0AD8A4F0C87708CA0D" pageId="0" pageNumber="1">GONOPODS (Fig. 92): Coxa (cx) longer than prefemur. Cannula (ca) slender. Prefemur (pfe) ca. 2/3 as long as femur, quite short and stout. Femur (fe) long and slender. Mesal sulcus (ms) and lateral sulcus (ls) deep. Postfemur (pof) conspicuous, ventrally wide. Solenophore (sph) well-developed: lamina lateralis (ll) swollen, with a big and long lobe-like structure projecting ventroanteriad: lamina medialis (lm) well-developed, longer than lamina lateralis; process (plm) short; distal lobe (dlm) quite broad, distally with two lamellae (mesal lamella shorter than lateral one); broad lobe (blm) long and thick, obviously demarcated from distal lobe (dlm) by a slightly deep and wide indentation. Solenomere (sl) very long.</paragraph>
<caption id="03F1F4FFD7997432DFC256300197ED27" pageId="0" pageNumber="1">
<paragraph id="9A42F4464BB0E28D9696D82627C164DE" pageId="0" pageNumber="1">
Figure 92.
<taxonomicName id="2BF15C6C8C844C1526D834EE02747B4B" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes terae" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="terae">Desmoxytes terae</taxonomicName>
(Jeekel, 1964), holotype - right gonopod. A mesal view B dorsal view C ventral view D submesal view (arrow = indentation) E lateral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="FE6AA592A22F08061E21D56C85668D38" pageId="0" pageNumber="1" type="distribution">
<paragraph id="505BBDA64667AB8AC3F5450A26DB49A8" pageId="0" pageNumber="1">Distribution and habitat.</paragraph>
<paragraph id="86E666CCF2B662B100E7683615147918" pageId="0" pageNumber="1">
This species is known only from the
<normalizedToken id="5F3CC3A1A438FA8F2EC5561F44F323C0" originalValue="MalaysiaThailand">Malaysia-Thailand</normalizedToken>
borderland (Kaki Bukit and Thale Ban National Park). We noticed that it prefers to live on humid mosses, logs or litter in limestone habitats (Fig. 88C). The localities are all on limestone mountains. The species is probably distributed in a narrow range in limestone areas near the
<normalizedToken id="B34808F47D88281FA4B73AF60E75E9B1" originalValue="ThaiMalay">Thai-Malay</normalizedToken>
border.
<taxonomicName id="95ADD95EF5009323BBF9A50E3981F42F" lsidName="D. terae" pageId="0" pageNumber="1" rank="species" species="terae">D. terae</taxonomicName>
has so far been recorded from three localities (near Kaki Bukit, Thale Ban National Park, and Tham Tone Din) which are located only 10-15 km apart. Despite several attempts by us,
<taxonomicName id="8A1485736AD8A43110DC23CABB56DFFF" lsidName="D. terae" pageId="0" pageNumber="1" rank="species" species="terae">D. terae</taxonomicName>
was not found in other areas. Hence, this species should be regarded as endemic to the Malaysia and Thailand faunas.
</paragraph>
</subSubSection>
<subSubSection id="0237BF764F4D090B8F8CB6073ACDA884" pageId="0" pageNumber="1" type="remarks">
<paragraph id="999311526AD32A152CCA3BB2D2D7F845" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="B06F51CCAF0B5F65E1A2BAB935107228" pageId="0" pageNumber="1">
In the recent field surveys we noticed that the colour of living specimens is black or brownish black with contrasting white paraterga as reported earlier by
<bibRefCitation id="40FC2AC921BD0B9A385A417BC3D15629" author="Jeekel, CAW" journalOrPublisher="Beaufortia" pageId="0" pageNumber="1" pagination="61 - 73" title="Two new species of Pratinus Attems, with taxonomic notes on the genus and a redescription of its type-species (Diplopoda, Polydesmida)." url="http://www.repository.naturalis.nl/document/548475" volume="11" year="1964">Jeekel (1964)</bibRefCitation>
and
<bibRefCitation id="74E0B3727DB5C3AFAF70797D1AAB2EAA" author="Golovatch, SI" journalOrPublisher="Steenstrupia" pageId="0" pageNumber="1" pagination="45 - 71" title="Review of the dragon millipedes, genus Desmoxytes Chamberlin, 1923 (Diplopoda, Polydesmida, Paradoxosomatidae)." volume="20" year="1994">Golovatch and Enghoff (1994)</bibRefCitation>
. This species blends so perfectly with its environment that it is difficult to collect specimens without a flashlight.
</paragraph>
<paragraph id="6422CE07BE19F126C771DA7AA5C719F4" pageId="0" pageNumber="1">
In the original description,
<bibRefCitation id="9E2FD670722C8D337B7502BA2A40C94E" author="Jeekel, CAW" journalOrPublisher="Beaufortia" pageId="0" pageNumber="1" pagination="61 - 73" title="Two new species of Pratinus Attems, with taxonomic notes on the genus and a redescription of its type-species (Diplopoda, Polydesmida)." url="http://www.repository.naturalis.nl/document/548475" volume="11" year="1964">Jeekel (1964)</bibRefCitation>
stated about the collum: &quot;near the anterior margin a transverse row of six hairs, which may be present partly rubbed off, the lateral pair placed on the low tubercles&quot;. This means collum with one row of 3+3 anterior setae/tubercles. Moreover, Jeekel also described paraterga without a tiny denticle near the tip. After we examined all specimens, it is clear that:
</paragraph>
<paragraph id="4E125A97BA663DED15470DA9404E9E86" pageId="0" pageNumber="1">- Collum with 3 rows of 3+3 anterior setae, 1+1 intermediate tubercles and 2+2 posterior tubercles (tubercles without setae, quite small but conspicuous).</paragraph>
<paragraph id="0AC2D57C10015E7D8E2701B10062910C" pageId="0" pageNumber="1">- Paraterga of body rings 9, 10, 12, 13, 15-18 with tiny denticle near ther tip, albeit all quite small.</paragraph>
<paragraph id="A49FA61937DFBB5BED36AE5F641EC389" pageId="0" pageNumber="1">The sternal lobe between male coxae 4 shows some variation within populations; the lobe of some specimens is trapeziform whereas in others it is semicircular. We also found some variability on the telson: tip of epiproct subtruncate in some individuals, in others slightly emarginate; lateral setiferous tubercles conspicuous in some specimens, inconspicuous in others; caudal margin of hypoproct truncate in some individuals, slightly round in others.</paragraph>
</subSubSection>
<subSubSection id="8109BB0401D60B86548293C71C35F2FF" pageId="0" pageNumber="1" type="coexisting species">
<paragraph id="5525ABD1B5646FC5ADB8AC28DC2F2866" pageId="0" pageNumber="1">Coexisting species.</paragraph>
<paragraph id="50BC39444CC180E9277E30DE5D0F55D8" pageId="0" pageNumber="1">
This species and
<taxonomicName id="0A486D39DF5AB618F468B38F97DFC835" lsidName="D. delfae" pageId="0" pageNumber="1" rank="species" species="delfae">D. delfae</taxonomicName>
are sympatric at Tam Ton Din.
</paragraph>
</subSubSection>
</treatment>
</document>