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<document id="E66443813F3F7896D89AF457E179819D" ID-DOI="10.5281/zenodo.3678492" ID-GBIF-Dataset="8e18452b-1d9d-462e-bac8-8c86eeb4ddc3" ID-ISSN="1942-1354" ID-Zenodo-Dep="3678492" ID-ZooBank="930EAB11-37FA-41B0-980A-1A4736527842" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1582295086673" checkinUser="felipe" docAuthor="Hielkema, Auke J. &amp; Hielkema, Meindert A." docDate="2019" docId="03D7FF1CFF90FFB1FF48FDA1351CFD4A" docLanguage="en" docName="InsectaMundi.0732.1-306.pdf.imf" docOrigin="Insecta Mundi 732 (732)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Onthophagus clypeatus Blanchard 1846" docType="treatment" docVersion="12" lastPageNumber="91" masterDocId="FFEE8764FFCBFFEDFF88FB6E317CFFF0" masterDocTitle="An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas" masterLastPageNumber="306" masterPageNumber="1" pageNumber="90" updateTime="1698721431862" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="3D6596880D4D4F411776B1CB1B271E4B">An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas</mods:title>
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<mods:title id="013FB852F35A93FC83AD3B9D8317FB04">Insecta Mundi</mods:title>
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<mods:date id="20BD7E87C42E1E9F3E697D2D2C1285D3">2019</mods:date>
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<treatment id="03D7FF1CFF90FFB1FF48FDA1351CFD4A" ID-DOI="http://doi.org/10.5281/zenodo.3683548" ID-GBIF-Taxon="162246505" ID-Zenodo-Dep="3683548" LSID="urn:lsid:plazi:treatment:03D7FF1CFF90FFB1FF48FDA1351CFD4A" httpUri="http://treatment.plazi.org/id/03D7FF1CFF90FFB1FF48FDA1351CFD4A" lastPageId="92" lastPageNumber="91" pageId="91" pageNumber="90">
<subSubSection id="C3641D81FF90FFB6FF48FDA13399F918" box="[192,741,1743,1768]" pageId="91" pageNumber="90" type="nomenclature">
<paragraph id="8BC14E0AFF90FFB6FF48FDA13399F918" blockId="91.[192,741,1743,1768]" box="[192,741,1743,1768]" pageId="91" pageNumber="90">
<heading id="D089F966FF90FFB6FF48FDA13399F918" bold="true" box="[192,741,1743,1768]" fontSize="10" level="2" pageId="91" pageNumber="90" reason="3">
<taxonomicName id="4C7E3589FF90FFB6FF48FDA13399F918" ID-CoL="7XMM5" authority="Blanchard, 1846" authorityName="Blanchard" authorityYear="1846" box="[192,741,1743,1768]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="90" phylum="Arthropoda" rank="species" species="clypeatus">
<emphasis id="B90A9218FF90FFB6FF48FDA13399F918" bold="true" box="[192,741,1743,1768]" pageId="91" pageNumber="90">
<emphasis id="B90A9218FF90FFB6FF48FDA1337DF917" bold="true" box="[192,513,1743,1767]" italics="true" pageId="91" pageNumber="90">Onthophagus clypeatus</emphasis>
<bibRefCitation id="EFEF33FBFF90FFB6FD81FDBE3399F918" author="Blanchard, C. E." bookContentInfo="222 p." box="[521,741,1744,1768]" editor="C. E. Blanchard &amp; G. A. Brulle" journalOrPublisher="Execute pendant les annees 1826" pageId="91" pageNumber="90" part="1830" refId="ref194002" refString="Blanchard, C. E. 1846. Tribu des lamellicornes. p. 155 - 194. In: C. E. Blanchard and G. A. Brulle. Voyage dans l'Amerique meridionale (…). Execute pendant les annees 1826, 1827, 1828, 1829, 1830, 1831, 1832 et 1833 par Alcide d'Orbigny. Tome sixieme, 2. e Partie: Insectes. Bertrand; Paris. 222 p." title="Tribu des lamellicornes" type="book" volumeTitle="Voyage dans l'Amerique meridionale" year="1846">Blanchard, 1846</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3641D81FF90FFB6FF60FD983242F8FC" box="[232,830,1782,1804]" pageId="91" pageNumber="90" type="reference_group">
<paragraph id="8BC14E0AFF90FFB6FF60FD983242F8FC" blockId="91.[232,830,1782,1804]" box="[232,830,1782,1804]" pageId="91" pageNumber="90">
<treatmentCitationGroup id="AB6E6924FF90FFB6FF60FD983242F8FC" box="[232,830,1782,1804]" pageId="91" pageNumber="90">
<taxonomicName id="4C7E3589FF90FFB6FF60FD983242F8FC" ID-CoL="7XMM5" authority="Blanchard 1846: 182" authorityName="Blanchard" authorityPageNumber="182" authorityYear="1846" box="[232,830,1782,1804]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="90" phylum="Arthropoda" rank="species" species="clypeatus">
<treatmentCitation id="0ADF681BFF90FFB6FF60FD9833A4F8FB" author="Blanchard, C. E." box="[232,728,1782,1804]" page="182" pageId="91" pageNumber="90" year="1846">
<emphasis id="B90A9218FF90FFB6FF60FD983090F8FB" box="[232,492,1782,1803]" italics="true" pageId="91" pageNumber="90">Onthophagus clypeatus</emphasis>
<bibRefCitation id="EFEF33FBFF90FFB6FE7AFD9833A4F8FB" author="Blanchard, C. E." bookContentInfo="222 p." box="[498,728,1782,1804]" editor="C. E. Blanchard &amp; G. A. Brulle" journalOrPublisher="Execute pendant les annees 1826" pageId="91" pageNumber="90" part="1830" refId="ref194002" refString="Blanchard, C. E. 1846. Tribu des lamellicornes. p. 155 - 194. In: C. E. Blanchard and G. A. Brulle. Voyage dans l'Amerique meridionale (…). Execute pendant les annees 1826, 1827, 1828, 1829, 1830, 1831, 1832 et 1833 par Alcide d'Orbigny. Tome sixieme, 2. e Partie: Insectes. Bertrand; Paris. 222 p." title="Tribu des lamellicornes" type="book" volumeTitle="Voyage dans l'Amerique meridionale" year="1846">Blanchard 1846: 182</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F3690E9AFF90FFB6FD6EFD98324BF8FC" box="[742,823,1782,1804]" name="Bolivia" pageId="91" pageNumber="90">Bolivia</collectingCountry>
)
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3641D81FF90FFB1FF48FC72351CFD4A" lastPageId="92" lastPageNumber="91" pageId="91" pageNumber="90" type="distribution">
<paragraph id="8BC14E0AFF90FFB6FF48FC7234DCF884" blockId="91.[192,1440,1820,1909]" pageId="91" pageNumber="90">
<emphasis id="B90A9218FF90FFB6FF48FC723090F8C4" bold="true" box="[192,492,1820,1844]" pageId="91" pageNumber="90">
Distribution.
<collectingCountry id="F3690E9AFF90FFB6FEF1FC72309AF8C4" box="[377,486,1820,1844]" name="Guyana" pageId="91" pageNumber="90">Guyana</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF90FFB6FE7AFC733250F8C5" author="Forsyth, A. B. &amp; B. D. Gill" bookContentInfo="70 p." box="[498,812,1821,1845]" editor="T. A. Parker &amp; III &amp; R. B. Foster &amp; L. H. Emmons &amp; P. Freed &amp; A. B. Forsyth &amp; B. Hoffman &amp; B. D. Gill" journalOrPublisher="Conservation International" pageId="91" pageNumber="90" refId="ref202721" refString="Forsyth, A. B., and B. D. Gill. 1993. Invertebrate indicators. p. 32 - 33, 70. In: T. A. Parker III, R. B. Foster, L. H. Emmons, P. Freed, A. B. Forsyth, B. Hoffman, and B. D. Gill. RAP Working Papers 5. A biological assessment of the Kanuku Mountain region of southwestern Guyana. Conservation International. 70 p." title="Invertebrate indicators" type="book" volumeTitle="RAP Working Papers 5. A biological assessment of the Kanuku Mountain region of southwestern Guyana" year="1993">Forsyth and Gill 1993: 70</bibRefCitation>
.
<emphasis id="B90A9218FF90FFB6FCBFFC7232B9F8C4" bold="true" box="[823,965,1820,1844]" pageId="91" pageNumber="90">
<collectingCountry id="F3690E9AFF90FFB6FCBFFC7232C3F8C4" box="[823,959,1820,1844]" name="Suriname" pageId="91" pageNumber="90">Suriname</collectingCountry>
:
</emphasis>
MHCN -
<emphasis id="B90A9218FF90FFB6FBB2FC7334DCF8C4" bold="true" box="[1082,1440,1820,1845]" pageId="91" pageNumber="90">NEW COUNTRY RECORD</emphasis>
(Suriname,
<collectingRegion id="49BA80E8FF90FFB6FED8FC5330ACF8A5" box="[336,464,1853,1877]" country="Suriname" name="Sipaliwini" pageId="91" pageNumber="90">Sipaliwini</collectingRegion>
, eastern bank Corantijn opposite Arapahu Resort, (
<geoCoordinate id="EE4A28CDFF90FFB6FBCCFC5535E5F8A4" box="[1092,1177,1851,1876]" degrees="3" direction="north" minutes="27" orientation="latitude" pageId="91" pageNumber="90" precision="925" value="3.45">3°27′N</geoCoordinate>
,
<geoCoordinate id="EE4A28CDFF90FFB6FB2CFC553458F8A4" box="[1188,1316,1851,1876]" degrees="57" direction="west" minutes="37.5" orientation="longitude" pageId="91" pageNumber="90" precision="92" value="-57.625">57°37.5′W</geoCoordinate>
). 15/
<date id="FFC068CAFF90FFB6FAD4FC533183F885" pageId="91" pageNumber="90" value="2011-02-21">21-II- 2011</date>
. A. J. Hielkema. In pitfall with human excrements in forest.); NZCS; RMNH.
<emphasis id="B90A9218FF90FFB6FB40FC3334DCF884" bold="true" box="[1224,1440,1884,1909]" pageId="91" pageNumber="90">
<collectingCountry id="F3690E9AFF90FFB6FB40FC3334E7F884" box="[1224,1435,1884,1909]" name="French Guiana" pageId="91" pageNumber="90">French Guiana</collectingCountry>
:
</emphasis>
</paragraph>
<paragraph id="8BC14E0AFF97FFB1FF48FBB4354EFE61" blockId="92.[192,1440,217,402]" pageId="92" pageNumber="91">
<bibRefCitation id="EFEF33FBFF97FFB1FF48FBB430B3FF02" author="Boucomont, A." box="[192,463,218,242]" journalOrPublisher="Annales de la Societe entomologique de France" pageId="92" pageNumber="91" pagination="293 - 332" part="101" refId="ref195096" refString="Boucomont, A. 1932. Synopsis des Onthophagus d'Amerique du Sud [Col. Scarab.]. Annales de la Societe entomologique de France 101: 293 - 332." title="Synopsis des Onthophagus d'Amerique du Sud [Col. Scarab.]" type="journal article" year="1932">Boucomont 1932: 322</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF97FFB1FE57FBB73397FF01" author="Balthasar, V." bookContentInfo="358 p." box="[479,747,217,242]" editor="E. Titschack" journalOrPublisher="Behre; Hamburg, Germany" pageId="92" pageNumber="91" pagination="337 - 358" refId="ref192769" refString="Balthasar, V. 1941 c. Scarabaeidae laparostictae (Col.). 68. Beitrag zur Kenntnis der Scarabaeiden Col. p. 337 - 358. In: E. Titschack. Beitrage zur Fauna Perus: nach der Ausbeute der Hamburger Sudperu-Expedition 1936, anderer Sammlungen, wie auch auf Grund von Literaturangaben. Bd I. Behre; Hamburg, Germany. 358 p. [Note. Reprinted 1951 by Gustav Fischer; Jena, Germany. 344 p.]" title="Scarabaeidae laparostictae (Col.). 68. Beitrag zur Kenntnis der Scarabaeiden Col" type="book chapter" volumeTitle="Beitrage zur Fauna Perus: nach der Ausbeute der Hamburger Sudperu-Expedition 1936, anderer Sammlungen, wie auch auf Grund von Literaturangaben. Bd I" year="1941">Balthasar 1941c: 352</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF97FFB1FD72FBB73546FF02" author="Price and Feer" box="[762,1082,217,242]" firstAuthor="Price" pageId="92" pageNumber="91" pagination="325 - 331" refId="ref220709" refString="Price, D. L., and F. Feer. 2012. Are there pitfalls to pitfalls? Dung beetle sampling in French Guiana. Organisms Diversity &amp; Evolution 12 (3): 325 - 331." type="journal article" year="2012">Price and Feer 2012: 327</bibRefCitation>
; MHCN.
<emphasis id="B90A9218FF97FFB1FB3CFBB7343AFF01" bold="true" box="[1204,1350,217,241]" pageId="92" pageNumber="91">“Guianas”:</emphasis>
<bibRefCitation id="EFEF33FBFF97FFB1FAD9FBB7335EFEE2" author="Pulido Herrera and Zunino" firstAuthor="Pulido Herrera" pageId="92" pageNumber="91" pagination="93 - 129" refId="ref220786" refString="Pulido Herrera, L. A., and M. Zunino. 2007. Catalogo preliminar de los Onthophagini de America (Coleoptera: Scarabaeine [sic]). Monografias 3 ercer Milenio 7: 93 - 129." type="journal article" year="2007">Pulido Herrera and Zunino 2007: 99</bibRefCitation>
.
<emphasis id="B90A9218FF97FFB1FDA4FB9733FBFEE1" bold="true" box="[556,647,249,273]" pageId="92" pageNumber="91">
<collectingCountry id="F3690E9AFF97FFB1FDA4FB9733FEFEE1" box="[556,642,249,273]" name="Brazil" pageId="92" pageNumber="91">Brazil</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF97FFB1FD05FB973538FEE2" author="Pulido Herrera and Zunino" box="[653,1092,249,274]" firstAuthor="Pulido Herrera" pageId="92" pageNumber="91" pagination="93 - 129" refId="ref220786" refString="Pulido Herrera, L. A., and M. Zunino. 2007. Catalogo preliminar de los Onthophagini de America (Coleoptera: Scarabaeine [sic]). Monografias 3 ercer Milenio 7: 93 - 129." type="journal article" year="2007">Pulido Herrera and Zunino 2007: 99</bibRefCitation>
.
<emphasis id="B90A9218FF97FFB1FBC5FB97359FFEE1" bold="true" box="[1101,1251,249,273]" pageId="92" pageNumber="91">
<collectingCountry id="F3690E9AFF97FFB1FBC5FB9735A1FEE1" box="[1101,1245,249,273]" name="Venezuela" pageId="92" pageNumber="91">Venezuela</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF97FFB1FB60FB9730C1FEC2" author="Pulido Herrera and Zunino" firstAuthor="Pulido Herrera" pageId="92" pageNumber="91" pagination="93 - 129" refId="ref220786" refString="Pulido Herrera, L. A., and M. Zunino. 2007. Catalogo preliminar de los Onthophagini de America (Coleoptera: Scarabaeine [sic]). Monografias 3 ercer Milenio 7: 93 - 129." type="journal article" year="2007">Pulido Herrera and Zunino 2007: 99</bibRefCitation>
.
<emphasis id="B90A9218FF97FFB1FE40FA77335DFEC1" bold="true" box="[456,545,281,305]" pageId="92" pageNumber="91">Other:</emphasis>
<bibRefCitation id="EFEF33FBFF97FFB1FDAFFA773258FEC2" author="Blanchard, C. E." bookContentInfo="222 p." box="[551,804,281,306]" editor="C. E. Blanchard &amp; G. A. Brulle" journalOrPublisher="Execute pendant les annees 1826" pageId="92" pageNumber="91" part="1830" refId="ref194002" refString="Blanchard, C. E. 1846. Tribu des lamellicornes. p. 155 - 194. In: C. E. Blanchard and G. A. Brulle. Voyage dans l'Amerique meridionale (…). Execute pendant les annees 1826, 1827, 1828, 1829, 1830, 1831, 1832 et 1833 par Alcide d'Orbigny. Tome sixieme, 2. e Partie: Insectes. Bertrand; Paris. 222 p." title="Tribu des lamellicornes" type="book" volumeTitle="Voyage dans l'Amerique meridionale" year="1846">Blanchard 1846: 182</bibRefCitation>
(Bolivia);
<bibRefCitation id="EFEF33FBFF97FFB1FC2AFA773455FEC1" author="Gillet, J. J. E. &amp; A. Boucomont" bookContentInfo="264 p." box="[930,1321,281,306]" editor="S. Schenkling" journalOrPublisher="Junk; Berlin" pageId="92" pageNumber="91" pagination="103 - 264" refId="ref204662" refString="Gillet, J. J. E., and A. Boucomont. 1927. Scarabaeidae: Coprinae II, Termitotroginae. p. 103 - 264. In: S. Schenkling. Coleopterorum Catalogus Pars 90. W. Junk; Berlin. 264 p." title="Scarabaeidae: Coprinae II, Termitotroginae" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 90. W" year="1927">Gillet and Boucomont 1927: 205</bibRefCitation>
(Bolivia);
<bibRefCitation id="EFEF33FBFF97FFB1FF48FA5430B1FEA2" author="Boucomont, A." box="[192,461,314,338]" journalOrPublisher="Annales de la Societe entomologique de France" pageId="92" pageNumber="91" pagination="293 - 332" part="101" refId="ref195096" refString="Boucomont, A. 1932. Synopsis des Onthophagus d'Amerique du Sud [Col. Scarab.]. Annales de la Societe entomologique de France 101: 293 - 332." title="Synopsis des Onthophagus d'Amerique du Sud [Col. Scarab.]" type="journal article" year="1932">Boucomont 1932: 322</bibRefCitation>
(Bolivia, Colombia, Ecuador, Peru);
<bibRefCitation id="EFEF33FBFF97FFB1FC16FA5735D4FEA1" author="Balthasar, V." bookContentInfo="358 p." box="[926,1192,313,338]" editor="E. Titschack" journalOrPublisher="Behre; Hamburg, Germany" pageId="92" pageNumber="91" pagination="337 - 358" refId="ref192769" refString="Balthasar, V. 1941 c. Scarabaeidae laparostictae (Col.). 68. Beitrag zur Kenntnis der Scarabaeiden Col. p. 337 - 358. In: E. Titschack. Beitrage zur Fauna Perus: nach der Ausbeute der Hamburger Sudperu-Expedition 1936, anderer Sammlungen, wie auch auf Grund von Literaturangaben. Bd I. Behre; Hamburg, Germany. 358 p. [Note. Reprinted 1951 by Gustav Fischer; Jena, Germany. 344 p.]" title="Scarabaeidae laparostictae (Col.). 68. Beitrag zur Kenntnis der Scarabaeiden Col" type="book chapter" volumeTitle="Beitrage zur Fauna Perus: nach der Ausbeute der Hamburger Sudperu-Expedition 1936, anderer Sammlungen, wie auch auf Grund von Literaturangaben. Bd I" year="1941">Balthasar 1941c: 352</bibRefCitation>
(Bolivia, Colombia, Ecuador, Peru);
<bibRefCitation id="EFEF33FBFF97FFB1FE00FA3733E2FE82" author="Blackwelder, R. E." box="[392,670,345,370]" journalOrPublisher="United States National Museum Bulletin" pageId="92" pageNumber="91" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 211</bibRefCitation>
(Bolivia);
<bibRefCitation id="EFEF33FBFF97FFB1FC95FA373541FE82" author="Medina" box="[797,1085,345,370]" etAl="et al." firstAuthor="Medina" pageId="92" pageNumber="91" pagination="131 - 144" refId="ref215451" refString="Medina, C. A., A. Lopera-Toro, A. Vitolo, and B. D. Gill. 2001. Escarabajos coprofagos (Coleoptera: Scarabaeidae: Scarabaeinae) de Colombia. Biota Colombiana 2 (2): 131 - 144." type="journal article" year="2001">Medina et al. 2001: 139</bibRefCitation>
(Colombia - as
<taxonomicName id="4C7E3589FF97FFB1FB74FA3434DCFE81" box="[1276,1440,345,369]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="91" phylum="Arthropoda" rank="species" species="clypeataus">
<emphasis id="B90A9218FF97FFB1FB74FA3434DCFE81" box="[1276,1440,345,369]" italics="true" pageId="92" pageNumber="91">O. clypeataus</emphasis>
</taxonomicName>
[sic]);
<bibRefCitation id="EFEF33FBFF97FFB1FE81FA1733B8FE62" author="Pulido Herrera and Zunino" box="[265,708,377,402]" firstAuthor="Pulido Herrera" pageId="92" pageNumber="91" pagination="93 - 129" refId="ref220786" refString="Pulido Herrera, L. A., and M. Zunino. 2007. Catalogo preliminar de los Onthophagini de America (Coleoptera: Scarabaeine [sic]). Monografias 3 ercer Milenio 7: 93 - 129." type="journal article" year="2007">Pulido Herrera and Zunino 2007: 99</bibRefCitation>
(Bolivia, Colombia, Ecuador).
</paragraph>
<paragraph id="8BC14E0AFF97FFB1FF48FACF351CFD4A" blockId="92.[192,1440,417,698]" pageId="92" pageNumber="91">
<emphasis id="B90A9218FF97FFB1FF48FACF3074FE49" bold="true" box="[192,264,417,441]" pageId="92" pageNumber="91">Note.</emphasis>
<taxonomicName id="4C7E3589FF97FFB1FE87FACF3357FE49" authorityName="Blanchard" authorityYear="1846" box="[271,555,417,441]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="91" phylum="Arthropoda" rank="species" species="clypeatus">
<emphasis id="B90A9218FF97FFB1FE87FACF3357FE49" box="[271,555,417,441]" italics="true" pageId="92" pageNumber="91">Onthophagus clypeatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7E3589FF97FFB1FDE0FACC3256FE49" authorityName="H. W. Bates" authorityYear="1887" box="[616,810,417,441]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="91" phylum="Arthropoda" rank="species" species="xanthomerus">
<emphasis id="B90A9218FF97FFB1FDE0FACC3256FE49" box="[616,810,417,441]" italics="true" pageId="92" pageNumber="91">O. xanthomerus</emphasis>
</taxonomicName>
are suspected to be different forms of the same species in
<bibRefCitation id="EFEF33FBFF97FFB1FE9EFAAC3353FE2A" author="Boucomont, A." box="[278,559,450,474]" journalOrPublisher="Annales de la Societe entomologique de France" pageId="92" pageNumber="91" pagination="293 - 332" part="101" refId="ref195096" refString="Boucomont, A. 1932. Synopsis des Onthophagus d'Amerique du Sud [Col. Scarab.]. Annales de la Societe entomologique de France 101: 293 - 332." title="Synopsis des Onthophagus d'Amerique du Sud [Col. Scarab.]" type="journal article" year="1932">Boucomont (1932: 299)</bibRefCitation>
(see the note at the bottom of that page) and in
<bibRefCitation id="EFEF33FBFF97FFB1FB0AFAAF34E0FE29" author="Balthasar, V." bookContentInfo="358 p." box="[1154,1436,449,474]" editor="E. Titschack" journalOrPublisher="Behre; Hamburg, Germany" pageId="92" pageNumber="91" pagination="337 - 358" refId="ref192769" refString="Balthasar, V. 1941 c. Scarabaeidae laparostictae (Col.). 68. Beitrag zur Kenntnis der Scarabaeiden Col. p. 337 - 358. In: E. Titschack. Beitrage zur Fauna Perus: nach der Ausbeute der Hamburger Sudperu-Expedition 1936, anderer Sammlungen, wie auch auf Grund von Literaturangaben. Bd I. Behre; Hamburg, Germany. 358 p. [Note. Reprinted 1951 by Gustav Fischer; Jena, Germany. 344 p.]" title="Scarabaeidae laparostictae (Col.). 68. Beitrag zur Kenntnis der Scarabaeiden Col" type="book chapter" volumeTitle="Beitrage zur Fauna Perus: nach der Ausbeute der Hamburger Sudperu-Expedition 1936, anderer Sammlungen, wie auch auf Grund von Literaturangaben. Bd I" year="1941">Balthasar (1941c: 352)</bibRefCitation>
. In
<bibRefCitation id="EFEF33FBFF97FFB1FF6AFA8F337EFE0A" author="Blackwelder, R. E." box="[226,514,481,506]" journalOrPublisher="United States National Museum Bulletin" pageId="92" pageNumber="91" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder (1944: 211</bibRefCitation>
, 212) these forms were treated as separate species. According to
<bibRefCitation id="EFEF33FBFF97FFB1FA9FFA8C3036FDEA" author="Boucomont, A." journalOrPublisher="Annales de la Societe entomologique de France" pageId="92" pageNumber="91" pagination="293 - 332" part="101" refId="ref195096" refString="Boucomont, A. 1932. Synopsis des Onthophagus d'Amerique du Sud [Col. Scarab.]. Annales de la Societe entomologique de France 101: 293 - 332." title="Synopsis des Onthophagus d'Amerique du Sud [Col. Scarab.]" type="journal article" year="1932">Boucomont (1932: 299)</bibRefCitation>
, the forms are distinguished by the pronotum being matte (
<taxonomicName id="4C7E3589FF97FFB1FBA3F96C35BCFDE9" authorityName="Blanchard" authorityYear="1846" box="[1067,1216,513,537]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="91" phylum="Arthropoda" rank="species" species="clypeatus">
<emphasis id="B90A9218FF97FFB1FBA3F96C35BCFDE9" box="[1067,1216,513,537]" italics="true" pageId="92" pageNumber="91">O. clypeatus</emphasis>
</taxonomicName>
) or shiny (
<taxonomicName id="4C7E3589FF97FFB1FACEF96C304EFDC9" authorityName="H. W. Bates" authorityYear="1887" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="91" phylum="Arthropoda" rank="species" species="xanthomerus">
<emphasis id="B90A9218FF97FFB1FACEF96C304EFDC9" italics="true" pageId="92" pageNumber="91">O. xanthomerus</emphasis>
</taxonomicName>
). We have collected a male Surinamese specimen in which the posterior half of the pronotum is matte and the anterior half shiny, which has led us to adopt the view of
<bibRefCitation id="EFEF33FBFF97FFB1FBC4F92C3450FDAA" author="Boucomont, A." box="[1100,1324,578,602]" journalOrPublisher="Annales de la Societe entomologique de France" pageId="92" pageNumber="91" pagination="293 - 332" part="101" refId="ref195096" refString="Boucomont, A. 1932. Synopsis des Onthophagus d'Amerique du Sud [Col. Scarab.]. Annales de la Societe entomologique de France 101: 293 - 332." title="Synopsis des Onthophagus d'Amerique du Sud [Col. Scarab.]" type="journal article" year="1932">Boucomont (1932)</bibRefCitation>
. We have thus placed all our specimens under the senior name
<taxonomicName id="4C7E3589FF97FFB1FCD6F90C3288FD89" authorityName="Blanchard" authorityYear="1846" box="[862,1012,609,633]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="91" phylum="Arthropoda" rank="species" species="clypeatus">
<emphasis id="B90A9218FF97FFB1FCD6F90C3288FD89" box="[862,1012,609,633]" italics="true" pageId="92" pageNumber="91">O. clypeatus</emphasis>
</taxonomicName>
. Otherwise, we have entered both names separately on this checklist because they are distinct species according to
<bibRefCitation id="EFEF33FBFF97FFB1FB14F9EF340AFD6A" author="Zunino" box="[1180,1398,641,666]" firstAuthor="Zunino" pageId="92" pageNumber="91" pagination="75 - 86" refId="ref230219" refString="Zunino, M. 1981. Note su alcuni Onthophagus americani e descrizione di nuove specie (Coleoptera, Scarabaeidae). Bollettino del Museo di Zoologia dell'Universita di Torino 6: 75 - 86." type="journal article" year="1981">Zunino (1981: 81)</bibRefCitation>
. To our knowledge the matter has not been given attention since
<bibRefCitation id="EFEF33FBFF97FFB1FC26F9CF3520FD4A" author="Zunino" box="[942,1116,673,698]" firstAuthor="Zunino" pageId="92" pageNumber="91" pagination="75 - 86" refId="ref230219" refString="Zunino, M. 1981. Note su alcuni Onthophagus americani e descrizione di nuove specie (Coleoptera, Scarabaeidae). Bollettino del Museo di Zoologia dell'Universita di Torino 6: 75 - 86." type="journal article" year="1981">Zunino (1981)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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