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<mods:title id="E80FE8F1F6706613D4DA258B7150D9A3">An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas</mods:title>
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<taxonomicName id="4C7E3589FF71FF57FF48F8813389FBF8" authority="Burmeister, 1847" authorityName="Burmeister" authorityYear="1847" box="[192,757,1007,1032]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="186" pageNumber="185" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B90A9218FF71FF57FF48F8813389FBF8" bold="true" box="[192,757,1007,1032]" pageId="186" pageNumber="185">
<emphasis id="B90A9218FF71FF57FF48F8813378FBF7" bold="true" box="[192,516,1007,1031]" italics="true" pageId="186" pageNumber="185">Cyclocephala variabilis</emphasis>
<bibRefCitation id="EFEF33FBFF71FF57FD84F8813389FBF8" author="Burmeister, H. C. C." bookContentInfo="584 p." box="[524,757,1007,1032]" journalOrPublisher="Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin" pageId="186" pageNumber="185" refId="ref195707" refString="Burmeister, H. C. C. 1847. Handbuch der Entomologie. Funfter Band. Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin. 584 p." title="Handbuch der Entomologie. Funfter Band" type="book" year="1847">Burmeister, 1847</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
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<paragraph id="8BC14E0AFF71FF57FF60FF78324CFBDC" blockId="186.[232,816,1046,1068]" box="[232,816,1046,1068]" pageId="186" pageNumber="185">
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<taxonomicName id="4C7E3589FF71FF57FF60FF78324CFBDC" authority="Burmeister 1847: 44" authorityName="Burmeister" authorityPageNumber="44" authorityYear="1847" box="[232,816,1046,1068]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="186" pageNumber="185" phylum="Arthropoda" rank="species" species="variabilis">
<treatmentCitation id="0ADF681BFF71FF57FF60FF7833AFFBDB" author="Burmeister, H. C. C." box="[232,723,1046,1068]" page="44" pageId="186" pageNumber="185" year="1847">
<emphasis id="B90A9218FF71FF57FF60FF783097FBDB" box="[232,491,1046,1067]" italics="true" pageId="186" pageNumber="185">Cyclocephala variabilis</emphasis>
<bibRefCitation id="EFEF33FBFF71FF57FE7AFF7833AFFBDB" author="Burmeister, H. C. C." bookContentInfo="584 p." box="[498,723,1046,1068]" journalOrPublisher="Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin" pageId="186" pageNumber="185" refId="ref195707" refString="Burmeister, H. C. C. 1847. Handbuch der Entomologie. Funfter Band. Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin. 584 p." title="Handbuch der Entomologie. Funfter Band" type="book" year="1847">Burmeister 1847: 44</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F3690E9AFF71FF57FD6AFF783255FBDC" box="[738,809,1046,1068]" name="Brazil" pageId="186" pageNumber="185">Brazil</collectingCountry>
)
</taxonomicName>
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</paragraph>
</subSubSection>
<subSubSection id="C3641D81FF71FF57FF48FF52351AFA8C" pageId="186" pageNumber="185" type="distribution">
<paragraph id="8BC14E0AFF71FF57FF48FF5234DCFAE5" blockId="186.[192,1440,1084,1301]" pageId="186" pageNumber="185">
<emphasis id="B90A9218FF71FF57FF48FF523091FBA4" bold="true" box="[192,493,1084,1108]" pageId="186" pageNumber="185">
Distribution.
<collectingCountry id="F3690E9AFF71FF57FEF2FF52309BFBA4" box="[378,487,1084,1108]" name="Guyana" pageId="186" pageNumber="185">Guyana</collectingCountry>
:
</emphasis>
RMNH -
<emphasis id="B90A9218FF71FF57FDEBFF5332B6FBA4" bold="true" box="[611,970,1084,1109]" pageId="186" pageNumber="185">NEW COUNTRY RECORD</emphasis>
(Demerara, Georgetown [= Guyana].).
<emphasis id="B90A9218FF71FF57FF48FF3330EDFB84" bold="true" box="[192,401,1116,1141]" pageId="186" pageNumber="185">
<collectingCountry id="F3690E9AFF71FF57FF48FF3330F0FB84" box="[192,396,1116,1141]" name="French Guiana" pageId="186" pageNumber="185">French Guiana</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF71FF57FE1FFF3333FCFB85" author="Endrodi, S." box="[407,640,1117,1141]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="186" pageNumber="185" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi 1966a: 326</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF71FF57FD02FF333214FB85" author="Duranton, M." box="[650,872,1117,1141]" journalOrPublisher="Coleopteres, numero special" pageId="186" pageNumber="185" pagination="1 - 31" part="1" refId="ref200092" refString="Duranton, M. 2011. Chorologie et ecologie des dynastides de Guyane francaise. Coleopteres, numero special 1: 1 - 31." title="Chorologie et ecologie des dynastides de Guyane francaise" type="journal article" year="2011">Duranton 2011: 16</bibRefCitation>
(?);
<bibRefCitation id="EFEF33FBFF71FF57FC1EFF33351CFB85" author="Ponchel" box="[918,1120,1117,1141]" firstAuthor="Ponchel" pageId="186" pageNumber="185" pagination="60 - 61" refId="ref220051" refString="Ponchel, Y. 2011. Liste actualisee des Dynastinae de Guyane (Coleoptera, Scarabaeidae). Contribution a l'etude des Coleopteres de Guyane 4: 60 - 61. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." type="journal article" year="2011">Ponchel 2011: 60</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF71FF57FBE2FF333436FB85" author="Dupuis, F." box="[1130,1354,1117,1141]" journalOrPublisher="Contribution a l'etude des coleopteres de Guyane" pageId="186" pageNumber="185" pagination="101 - 111" part="10" refId="ref199700" refString="Dupuis, F. 2016 b. Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes. Contribution a l'etude des coleopteres de Guyane 10: 101 - 111. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." title="Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes" type="journal article" year="2016">Dupuis 2016b: 109</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF71FF57FADCFF3330F7FB65" author="Moore" etAl="et al." firstAuthor="Moore" pageId="186" pageNumber="185" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. 2018b: 258</bibRefCitation>
.
<emphasis id="B90A9218FF71FF57FE1DFF12334BFB64" bold="true" box="[405,567,1148,1172]" pageId="186" pageNumber="185">
<collectingCountry id="F3690E9AFF71FF57FE1DFF123095FB64" box="[405,489,1148,1172]" name="Brazil" pageId="186" pageNumber="185">Brazil</collectingCountry>
(NA):
</emphasis>
<bibRefCitation id="EFEF33FBFF71FF57FDB5FF133245FB65" author="Burmeister, H. C. C." bookContentInfo="584 p." box="[573,825,1149,1173]" journalOrPublisher="Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin" pageId="186" pageNumber="185" refId="ref195707" refString="Burmeister, H. C. C. 1847. Handbuch der Entomologie. Funfter Band. Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Xylophila et Pectinicornia. Theod. Chr. Fr. Enslin; Berlin. 584 p." title="Handbuch der Entomologie. Funfter Band" type="book" year="1847">Burmeister 1847: 44</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF71FF57FCCCFF133573FB65" author="Arrow, G. J." bookContentInfo="124 p." box="[836,1039,1149,1173]" editor="S. Schenkling" journalOrPublisher="Junk; The Hague, Netherlands" pageId="186" pageNumber="185" pagination="3 - 124" refId="ref192146" refString="Arrow, G. J. 1937 b. Scarabaeidae: Dynastinae. p. 3 - 124. In: S. Schenkling. Coleopterorum Catalogus Pars 156. W. Junk; The Hague, Netherlands. 124 p." title="Scarabaeidae: Dynastinae" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 156. W" year="1937">Arrow 1937b: 16</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF71FF57FB93FF13344DFB65" author="Blackwelder, R. E." box="[1051,1329,1149,1173]" journalOrPublisher="United States National Museum Bulletin" pageId="186" pageNumber="185" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 253</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF71FF57FAB5FF13303AFB45" author="Endrodi, S." journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="186" pageNumber="185" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi 1966a: 326</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF71FF57FED8FFF333A5FB45" author="Andreazze, R. &amp; C. R. V. da Fonseca" box="[336,729,1181,1205]" journalOrPublisher="Acta Amazonica" pageId="186" pageNumber="185" pagination="59 - 66" part="28" refId="ref190842" refString="Andreazze, R., and C. R. V. da Fonseca. 1998. Dinastineos (Coleoptera, Scarabaeoidea, Melolonthidae) em uma area de terra firme na Amazonia central, Brasil. Acta Amazonica 28 (1): 59 - 66." title="Dinastineos (Coleoptera, Scarabaeoidea, Melolonthidae) em uma area de terra firme na Amazonia central, Brasil" type="journal article" year="1998">Andreazze and Fonseca 1998: 61</bibRefCitation>
(NA);
<bibRefCitation id="EFEF33FBFF71FF57FCAFFFF3353DFB45" author="Moore" box="[807,1089,1181,1205]" etAl="et al." firstAuthor="Moore" pageId="186" pageNumber="185" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. 2018b: 258</bibRefCitation>
.
<emphasis id="B90A9218FF71FF57FBC3FFF3359DFB45" bold="true" box="[1099,1249,1181,1205]" pageId="186" pageNumber="185">
<collectingCountry id="F3690E9AFF71FF57FBC3FFF335A7FB45" box="[1099,1243,1181,1205]" name="Venezuela" pageId="186" pageNumber="185">Venezuela</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF71FF57FB6EFFF33193FB25" author="Endrodi, S." journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="186" pageNumber="185" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi 1966a: 326</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF71FF57FF71FFD3336DFB25" author="Moore" box="[249,529,1213,1237]" etAl="et al." firstAuthor="Moore" pageId="186" pageNumber="185" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. 2018b: 258</bibRefCitation>
.
<emphasis id="B90A9218FF71FF57FD91FFD2330DFB24" bold="true" box="[537,625,1212,1236]" pageId="186" pageNumber="185">Other:</emphasis>
<bibRefCitation id="EFEF33FBFF71FF57FDFEFFD332FAFB25" author="Blackwelder, R. E." box="[630,902,1213,1237]" journalOrPublisher="United States National Museum Bulletin" pageId="186" pageNumber="185" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 253</bibRefCitation>
(Argentina);
<bibRefCitation id="EFEF33FBFF71FF57FBADFFD33473FB25" author="Endrodi, S." box="[1061,1295,1213,1237]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="186" pageNumber="185" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi 1966a: 326</bibRefCitation>
(Argentina, Bolivia, Colombia, Ecuador, Mexico, Panama), 1985a: 54, 133 (ArgentinaMexico);
<bibRefCitation id="EFEF33FBFF71FF57FB3AFFB334DCFB05" author="Lachaume" box="[1202,1440,1245,1269]" firstAuthor="Lachaume" pageId="186" pageNumber="185" pagination="7 - 56" refId="ref211183" refString="Lachaume, G. 1992. Dynastidae americains Cyclocephalini - Agaocephalini - Pentodontini - Oryctini - Phileurini. Les Coleopteres du Monde 14: 7 - 56." type="journal article" year="1992">Lachaume 1992: 12</bibRefCitation>
(ArgentinaMexico);
<bibRefCitation id="EFEF33FBFF71FF57FE34FF9333B1FAE5" author="Moore" box="[444,717,1277,1301]" etAl="et al." firstAuthor="Moore" pageId="186" pageNumber="185" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. 2018b: 258</bibRefCitation>
(Argentina, Bolivia, Colombia, Ecuador, Panama, Uruguay).
</paragraph>
<paragraph id="8BC14E0AFF71FF57FF48FE4B351AFA8C" blockId="186.[192,1440,1316,1405]" pageId="186" pageNumber="185">
<emphasis id="B90A9218FF71FF57FF48FE4B307BFACD" bold="true" box="[192,263,1317,1341]" pageId="186" pageNumber="185">Note.</emphasis>
According to
<bibRefCitation id="EFEF33FBFF71FF57FE2CFE4B33ACFACD" author="Ratcliffe" box="[420,720,1317,1341]" etAl="et al." firstAuthor="Ratcliffe" pageId="186" pageNumber="185" pagination="1 - 661" refId="ref222393" refString="Ratcliffe, B. C., R. D. Cave, and E. B. Cano. 2013. The dynastine scarab beetles of Mexico, Guatemala, and Belize (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 27: 1 - 661." type="journal article" year="2013">Ratcliffe et al. (2013: 598)</bibRefCitation>
, the record of
<taxonomicName id="4C7E3589FF71FF57FCFAFE4A35F9FACC" authorityName="Burmeister" authorityYear="1847" box="[882,1157,1316,1340]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="186" pageNumber="185" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B90A9218FF71FF57FCFAFE4A35F9FACC" box="[882,1157,1316,1340]" italics="true" pageId="186" pageNumber="185">Cyclocephala variabilis</emphasis>
</taxonomicName>
from
<collectingCountry id="F3690E9AFF71FF57FB40FE4B3460FACD" box="[1224,1308,1317,1341]" name="Mexico" pageId="186" pageNumber="185">Mexico</collectingCountry>
in
<bibRefCitation id="EFEF33FBFF71FF57FAB7FE4B302BFAAD" author="Endrodi, S." journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="186" pageNumber="185" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi (1966a: 326)</bibRefCitation>
, and by extension the copies of this record in
<bibRefCitation id="EFEF33FBFF71FF57FCE7FE2B352BFAAD" author="Endrodi, S." bookContentInfo="800 p." box="[879,1111,1349,1373]" journalOrPublisher="Junk; The Hague, Netherlands and Akademiai Kiado; Budapest" pageId="186" pageNumber="185" refId="ref201383" refString="Endrodi, S. 1985 a. The Dynastinae of the world. Series Entomologica 28. Dr W. Junk; The Hague, Netherlands and Akademiai Kiado; Budapest. 800 p." title="The Dynastinae of the world. Series Entomologica 28. Dr W" type="book" year="1985">Endrödi (1985a: 54</bibRefCitation>
, 133) and
<bibRefCitation id="EFEF33FBFF71FF57FB47FE2B3194FA8D" author="Lachaume" firstAuthor="Lachaume" pageId="186" pageNumber="185" pagination="7 - 56" refId="ref211183" refString="Lachaume, G. 1992. Dynastidae americains Cyclocephalini - Agaocephalini - Pentodontini - Oryctini - Phileurini. Les Coleopteres du Monde 14: 7 - 56." type="journal article" year="1992">Lachaume (1992: 12)</bibRefCitation>
, are incorrect, as this species only occurs in South America and
<collectingCountry id="F3690E9AFF71FF57FC71FE0B351EFA8C" box="[1017,1122,1381,1404]" name="Panama" pageId="186" pageNumber="185">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>